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<document id="42F8B9A948AF9512754BCB417CAF4B24" ID-CLB-Dataset="22649" ID-DOI="10.11646/zootaxa.4470.1.1" ID-GBIF-Dataset="615d5c4a-3b8d-4ba6-a05d-f37e432a862b" ID-ISSN="1175-5326" ID-Zenodo-Dep="3770039" ID-ZooBank="60766083-99E8-48E9-86CE-161A9EEC066B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588000738706" checkinUser="plazi" docAuthor="Garrison, Rosser W. &amp; Ellenrieder, Natalia Von" docDate="2018" docId="03AE645779369F624A9AFE9FFDCD19BD" docLanguage="en" docName="zootaxa.4470.1.1.pdf" docOrigin="Zootaxa 4470 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Argia jocosa Hagen in Selys 1865" docType="treatment" docVersion="7" lastPageNumber="18" masterDocId="FF971C2F79279F734A0DFFD3FF981C75" masterDocTitle="Damselflies of the genus Argia (Odonata: Coenagrionidae) from Ecuador with descriptions of five new species" masterLastPageNumber="69" masterPageNumber="1" pageNumber="18" updateTime="1698837067205" updateUser="ExternalLinkService">
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<mods:title id="06020F31CFAA3215323BA4AE76805015">Damselflies of the genus Argia (Odonata: Coenagrionidae) from Ecuador with descriptions of five new species</mods:title>
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<mods:name id="2BB1305B5361EE904FC76EFD62D49CCD" type="personal">
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<mods:namePart id="5606C54C957B985CE8E9C79B3F681E8F">Garrison, Rosser W.</mods:namePart>
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<mods:namePart id="A9C82E66AEA4E7DC15E4BF82AAB542E5">Ellenrieder, Natalia Von</mods:namePart>
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<mods:title id="B32A16675A3F9064788BAB89C3BC61CE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="61A183FC572F6DCB073FFB7FA66B7D86">
<mods:date id="E03484DB174BED7B7F8E2609755A7651">2018</mods:date>
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<mods:number id="98B50D20B184CCDAD0375988EA62335E">2018-09-03</mods:number>
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<mods:detail id="F5D8E268670F758218846229CF6A4A4D" type="volume">
<mods:number id="E5BF2F33C8FC07291EAF6DE5AA72EF54">4470</mods:number>
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<mods:classification id="6FC34B58B3DF766CB7C960EFCC33921E">journal article</mods:classification>
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<mods:identifier id="6521941AF0AA4883DBA8A1538253D7B0" type="DOI">10.11646/zootaxa.4470.1.1</mods:identifier>
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<mods:identifier id="F042FC4F47A1B2F78197FA01A35D0397" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03AE645779369F624A9AFE9FFDCD19BD" ID-DOI="http://doi.org/10.5281/zenodo.3799133" ID-GBIF-Taxon="163754275" ID-Zenodo-Dep="3799133" LSID="urn:lsid:plazi:treatment:03AE645779369F624A9AFE9FFDCD19BD" httpUri="http://treatment.plazi.org/id/03AE645779369F624A9AFE9FFDCD19BD" lastPageNumber="18" pageId="17" pageNumber="18">
<subSubSection id="C31D86CA79369F624A9AFE9FFDA01D13" box="[151,568,332,358]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8BB8D54179369F624A9AFE9FFDA01D13" blockId="17.[151,1436,332,427]" box="[151,568,332,358]" pageId="17" pageNumber="18">
<heading id="D0F0622D79369F624A9AFE9FFDA01D13" bold="true" box="[151,568,332,358]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<taxonomicName id="4C07AEC279369F624A9AFE9FFDA01D13" ID-CoL="GGPG" authority="Hagen in Selys, 1865" authorityName="Hagen in Selys" authorityYear="1865" box="[151,568,332,358]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="jocosa">
<emphasis id="B973095379369F624A9AFE9FFDA01D13" bold="true" box="[151,568,332,358]" pageId="17" pageNumber="18">
<emphasis id="B973095379369F624A9AFE9FFEB71D13" bold="true" box="[151,303,332,358]" italics="true" pageId="17" pageNumber="18">Argia jocosa</emphasis>
Hagen
<emphasis id="B973095379369F624B82FE9FFE3F1D13" bold="true" box="[399,423,332,358]" italics="true" pageId="17" pageNumber="18">in</emphasis>
<bibRefCitation id="EF96A8B079369F624BA2FE9FFDA01D13" author="Selys, E. M." box="[431,568,332,358]" pageId="17" pageNumber="18" pagination="375 - 417" refId="ref52356" refString="Selys, E. M. (1865) Synopsis des Agrionines, 5 me legion: Agrion. Bulletin de l'Academie Royale des Sciences, des Lettres et des Beaux Arts de Belgique, Series 2, 20, 375 - 417 (1 - 45 separate)." type="journal article" year="1865">Selys, 1865</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
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<paragraph id="8BB8D54179369F624A9AFEBDFA851DDE" blockId="17.[151,1436,332,427]" pageId="17" pageNumber="18">
<figureCitation id="133CC9C479369F624A9AFEBDFF7C1DF2" box="[151,228,366,391]" captionStart="FIGURES 1–8" captionStartId="39.[151,264,1966,1988]" captionTargetBox="[186,1424,239,1889]" captionTargetId="figure@39.[151,1437,193,1945]" captionTargetPageId="39" captionText="FIGURES 1–8. Color pattern of head. Head, anterior view (1); head posterior view (2–5); labrum, anterior view (6–8)." figureDoi="http://doi.org/10.5281/zenodo.3770041" httpUri="https://zenodo.org/record/3770041/files/figure.png" pageId="17" pageNumber="18">Figs. 3</figureCitation>
(rear of head
<specimenCount id="9D011EC879369F624B89FEBCFE0F1DF2" box="[388,407,367,391]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
), 7 (labrum
<specimenCount id="9D011EC879369F62482AFEBCFDA31DF2" box="[551,571,367,391]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
); 17 (head, thorax, S1–
<specimenCount id="9D011EC879369F62494BFEBCFCEC1DF2" box="[838,884,367,391]" pageId="17" pageNumber="18" type="male">4 ♂</specimenCount>
); 28 (thorax
<specimenCount id="9D011EC879369F624E0AFEBCFB831DF2" box="[1031,1051,367,391]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
); 50–52 (S1–
<specimenCount id="9D011EC879369F624EBBFEBCFB6A1DF2" box="[1206,1266,367,391]" pageId="17" pageNumber="18" type="male">10 ♂</specimenCount>
), 62 (S1–
<specimenCount id="9D011EC879369F624F6CFEBDFA141DF3" box="[1377,1420,366,391]" pageId="17" pageNumber="18" type="female">5 ♀</specimenCount>
), 77–79 (mesostigmal plates
<specimenCount id="9D011EC879369F624BC7FE40FE461DDE" box="[458,478,403,427]" pageId="17" pageNumber="18" type="female">♀</specimenCount>
), 95 (genital ligula
<specimenCount id="9D011EC879369F6248B0FE40FD4C1DDE" box="[701,724,403,427]" pageId="17" pageNumber="18" type="male">♂</specimenCount>
), 111 (appendages
<specimenCount id="9D011EC879369F6249A6FE40FC5A1DDE" box="[939,962,403,427]" pageId="17" pageNumber="18" type="male">♂</specimenCount>
), 128 (
<typeStatus id="54BC6BE379369F624E1DFE40FBDA1DDE" box="[1040,1090,403,427]" pageId="17" pageNumber="18">type</typeStatus>
labels), 139 (map).
</paragraph>
</subSubSection>
<subSubSection id="C31D86CA79369F624A9AFE08FE561E8D" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="8BB8D54179369F624A9AFE08FED91E1E" blockId="17.[151,1437,474,619]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="AB17F26F79369F624A9AFE08FED91E1E" pageId="17" pageNumber="18">
<treatmentCitation id="0AA6F35079369F624A9AFE08FDA91D85" author="Selys, E. M." box="[151,561,474,496]" page="408" pageId="17" pageNumber="18" year="1865">
<taxonomicName id="4C07AEC279369F624A9AFE08FDA91D85" ID-CoL="GGPG" authority="Hagen in Selys 1865: 408" authorityName="Hagen in Selys" authorityPageNumber="408" authorityYear="1865" box="[151,561,474,496]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="jocosa">
<emphasis id="B973095379369F624A9AFE08FE821D85" box="[151,282,475,496]" italics="true" pageId="17" pageNumber="18">Argia jocosa</emphasis>
Hagen in
<bibRefCitation id="EF96A8B079369F624B84FE09FDA91D85" author="Selys, E. M." box="[393,561,474,496]" pageId="17" pageNumber="18" pagination="375 - 417" refId="ref52356" refString="Selys, E. M. (1865) Synopsis des Agrionines, 5 me legion: Agrion. Bulletin de l'Academie Royale des Sciences, des Lettres et des Beaux Arts de Belgique, Series 2, 20, 375 - 417 (1 - 45 separate)." type="journal article" year="1865">Selys 1865: 408</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(36 separate), (description of
<specimenCount id="9D011EC879369F624961FE08FC191D85" box="[876,897,475,496]" pageId="17" pageNumber="18" type="male">♂</specimenCount>
, &quot;Santa-Fe de
<collectingRegion id="49C31BA379369F624E16FE08FBFF1D85" box="[1051,1127,475,496]" country="Colombia" name="Bogota" pageId="17" pageNumber="18">Bogota</collectingRegion>
, par Lindig. (Coll. Hagen.)&quot;);
<bibRefCitation id="EF96A8B079369F624ACAFE2AFECA1E7A" author="Calvert, P. P." box="[199,338,505,527]" pageId="17" pageNumber="18" pagination="17 - 72" refId="ref50307" refString="Calvert, P. P. (1901) Odonata. In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia Centrali Americana-Insecta Neuroptera. R. H. Porter &amp; Dulau Co., London, pp. 17 - 72." type="book chapter" year="1901">Calvert 1901</bibRefCitation>
; 71 (key); Calvert
<emphasis id="B973095379369F624829FE29FDA01E7A" box="[548,568,506,527]" italics="true" pageId="17" pageNumber="18">in</emphasis>
Calvert &amp; Hagen 1902: 111 (catalog);
<treatmentCitation id="0AA6F35079369F6249EAFE2AFB321E7A" author="Calvert, P. P." box="[999,1194,505,527]" page="120" pageId="17" pageNumber="18" year="1909">
<bibRefCitation id="EF96A8B079369F6249EAFE2AFB321E7A" author="Calvert, P. P." box="[999,1194,505,527]" pageId="17" pageNumber="18" pagination="73 - 280" refId="ref50529" refString="Calvert, P. P. (1909) Contributions to a knowledge of the Odonata of the Neotropical region, exclusive of Mexico and Central America. Annals of the Carnegie Museum, 6 (3), 73 - 280." type="journal article" year="1909">Calvert 1909: 120</bibRefCitation>
</treatmentCitation>
, 143 (key,
<collectingCountry id="F31095D179369F624F25FE2AFA091E7A" box="[1320,1425,505,527]" name="Colombia" pageId="17" pageNumber="18">Colombia</collectingCountry>
);
<treatmentCitation id="0AA6F35079369F624ACAFDCBFE301E5B" author="Heckman, C. W." box="[199,424,536,558]" page="361" pageId="17" pageNumber="18" year="2008">
<bibRefCitation id="EF96A8B079369F624ACAFDCBFE301E5B" author="Heckman, C. W." box="[199,424,536,558]" pageId="17" pageNumber="18" refId="ref51495" refString="Heckman, C. W. (2008) Encyclopedia of South American Aquatic Insects: Odonata-Zygoptera. Springer, Dordrecht, viii + 687 pp." type="book" year="2008">Heckman 2008: 361</bibRefCitation>
</treatmentCitation>
(key, listed erroneously from
<collectingCountry id="F31095D179369F624906FDCBFCFC1E5B" box="[779,868,536,558]" name="Ecuador" pageId="17" pageNumber="18">Ecuador</collectingCountry>
);
<treatmentCitation id="0AA6F35079369F624972FDCBFA031E5B" author="Perez-Gutierrez, L. &amp; Palacino-Rodriguez, F." box="[895,1435,536,558]" page="212" pageId="17" pageNumber="18" year="2011">
<bibRefCitation id="EF96A8B079369F624972FDCBFA031E5B" author="Perez-Gutierrez, L. &amp; Palacino-Rodriguez, F." box="[895,1435,536,558]" pageId="17" pageNumber="18" pagination="203 - 225" refId="ref52121" refString="Perez-Gutierrez, L. &amp; Palacino-Rodriguez, F. (2011) Updated checklist of the Odonata known from Colombia. Odonatologica, 40 (3), 203 - 225." type="journal article" year="2011">Pérez-Gutiérrez &amp; Palacino-Rodriguez 2011: 212</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F31095D179369F624AC3FDE5FEA11E39" box="[206,313,566,588]" name="Colombia" pageId="17" pageNumber="18">Colombia</collectingCountry>
):
<treatmentCitation id="0AA6F35079369F624B41FDE4FDA71E39" author="De Marmels, J." box="[332,575,566,588]" page="253" pageId="17" pageNumber="18" year="2012">
<bibRefCitation id="EF96A8B079369F624B41FDE4FDA71E39" author="De Marmels, J." box="[332,575,566,588]" pageId="17" pageNumber="18" pagination="249 - 262" refId="ref50706" refString="De Marmels, J. (2012) Review of the &quot; metallic group &quot; of species of Argia Rambur known from Venezuela, with description of the larva of Argia jocosa Hagen in Selys, 1865 (Odonata: Coenagrionidae). International Journal of Odonatology, 15 (3), 249 - 262. https: // doi. org / 10.1080 / 13887890.2012.717906" type="journal article" year="2012">De Marmels 2012: 253</bibRefCitation>
</treatmentCitation>
(distribution in
<collectingCountry id="F31095D179369F6248E1FDE5FCCF1E39" box="[748,855,566,588]" name="Venezuela" pageId="17" pageNumber="18">Venezuela</collectingCountry>
, illustrations and comparison with
<taxonomicName id="4C07AEC279369F624ECAFDE4FAD71E38" ID-CoL="GGN2" box="[1223,1359,567,589]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cupraurea">
<emphasis id="B973095379369F624ECAFDE4FAD71E38" box="[1223,1359,567,589]" italics="true" pageId="17" pageNumber="18">A. cupraurea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C07AEC279369F624F8BFDE4FEAB1E1E" ID-CoL="682T8" authorityName="Hagen in Selys" authorityYear="1865" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="orichalcea">
<emphasis id="B973095379369F624F8BFDE4FEAB1E1E" italics="true" pageId="17" pageNumber="18">A. orichalcea</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BB8D54179369F624A9AFD44FE561E8D" blockId="17.[151,1437,663,1480]" pageId="17" pageNumber="18">
<emphasis id="B973095379369F624A9AFD44FF441EC5" bold="true" box="[151,220,663,688]" pageId="17" pageNumber="18">Types</emphasis>
.
<materialsCitation id="3B6FDF1C79369F624AE6FD4BFB571EA1" ID-GBIF-Occurrence="2608963152" collectorName="C. H. Kennedy" location="St. Fe" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Bogota" typeStatus="lectotype">
<typeStatus id="54BC6BE379369F624AE6FD4BFE131EC5" box="[235,395,664,688]" pageId="17" pageNumber="18" type="lectotype">LECTOTYPE</typeStatus>
by present designation,
<specimenCount id="9D011EC879369F624893FD4BFD2E1EDA" box="[670,694,664,687]" pageId="17" pageNumber="18" type="male">♂</specimenCount>
. &quot;
<location id="8ED8839A79369F6248C6FD4BFC961EDA" LSID="urn:lsid:plazi:treatment:03AE645779369F624A9AFE9FFDCD19BD:8ED8839A79369F6248C6FD4BFC961EDA" box="[715,782,664,688]" name="St. Fe" pageId="17" pageNumber="18" stateProvince="Bogota">St. Fe</location>
de
<collectingRegion id="49C31BA379369F62493AFD4BFC101EC5" box="[823,904,664,688]" country="Colombia" name="Bogota" pageId="17" pageNumber="18">Bogota</collectingRegion>
/ Lindig 1863&quot;, &quot;
<taxonomicName id="4C07AEC279369F624E40FD4BFB6E1EC5" ID-CoL="GGPG" authority="Hag." authorityName="Hagen in Selys" authorityYear="1865" box="[1101,1270,663,688]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="jocosa">A. jocosa Hag.</taxonomicName>
/ an
<taxonomicName id="4C07AEC279369F624F22FD4BFA051EDA" ID-CoL="GGN3" authorityName=": Donnelly" authorityYear="1992" box="[1327,1437,664,688]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cuprea">A. cuprea</taxonomicName>
var?&quot;, &quot;Penis/ drawn [written by
<collectorName id="26F2B09779369F624815FD6FFD5C1EA0" box="[536,708,700,725]" pageId="17" pageNumber="18">C.H. Kennedy</collectorName>
]&quot;, &quot;Hagen [printed]&quot;. MCZ 12194 (
<figureCitation id="133CC9C479369F624E6DFD6FFB5E1EA1" box="[1120,1222,700,725]" captionStart="FIGURES 128–130" captionStartId="61.[151,264,1909,1931]" captionTargetBox="[167,1421,212,1876]" captionTargetId="figure@61.[151,1437,193,1888]" captionTargetPageId="61" captionText="FIGURES 128–130. Type labels." figureDoi="http://doi.org/10.5281/zenodo.3770084" httpUri="https://zenodo.org/record/3770084/files/figure.png" pageId="17" pageNumber="18">Fig. 128</figureCitation>
)
</materialsCitation>
.
<materialsCitation id="3B6FDF1C79369F624ED0FD6FFE521E8D" ID-GBIF-Occurrence="2608963367" collectionCode="MCZ" collectorName="C. H. Kennedy" location="St. Fe" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Bogota" typeStatus="paralectotype">
<typeStatus id="54BC6BE379369F624ED0FD6FFAED1EA0" box="[1245,1397,700,725]" pageId="17" pageNumber="18" type="paralectotype">Paralectotype</typeStatus>
<specimenCount id="9D011EC879369F624F8DFD6EFA0F1EA1" box="[1408,1431,701,724]" pageId="17" pageNumber="18" type="male">♂</specimenCount>
, same locality label. [
<collectionCode id="ED164D8479369F624B73FD33FE5B1E8D" box="[382,451,736,760]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="17" pageNumber="18" type="Museum">MCZ</collectionCode>
]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31D86CA79369F624ACAFCD7FDCD19BD" pageId="17" pageNumber="18" type="description">
<paragraph id="8BB8D54179369F624ACAFCD7FDE51FFD" blockId="17.[151,1437,663,1480]" pageId="17" pageNumber="18">
<emphasis id="B973095379369F624ACAFCD7FCF81F68" bold="true" box="[199,864,772,797]" pageId="17" pageNumber="18">
Specimens examined (only those from
<collectingCountry id="F31095D179369F6248A8FCD7FC941F68" box="[677,780,772,797]" name="Ecuador" pageId="17" pageNumber="18">Ecuador</collectingCountry>
listed)
</emphasis>
.
<materialsCitation id="3B6FDF1C79369F62497FFCD7FDE21FFD" ID-GBIF-Occurrence="2608963051" collectingDate="2014-01-11" collectionCode="RWG" collectorName="David Wagner &amp; William Haber &amp; Fred Morrison" country="Ecuador" elevation="285" latitude="-0.0058333334" location="Lago Agrio" longLatPrecision="21" longitude="-76.64361" municipality="Bellavista River" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Sucumbios">
<specimenCount id="9D011EC879369F62497FFCD7FC3B1F69" box="[882,931,772,796]" pageId="17" pageNumber="18" type="male">1 ♂</specimenCount>
,
<collectingRegion id="49C31BA379369F6249BEFCD7FBED1F69" box="[947,1141,772,797]" country="Ecuador" name="Sucumbios" pageId="17" pageNumber="18">Sucumbios Prov.</collectingRegion>
,
<collectingMunicipality id="6BDC4F3B79369F624E8BFCD7FAD91F68" box="[1158,1345,772,797]" pageId="17" pageNumber="18">Bellavista River</collectingMunicipality>
,
<quantity id="4CFF78A479369F624F5CFCD7FA041F68" box="[1361,1436,772,797]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="17" pageNumber="18" unit="km" value="32.0">32 km</quantity>
southeast of
<location id="8ED8839A79369F624B2BFCFBFE311F35" LSID="urn:lsid:plazi:treatment:03AE645779369F624A9AFE9FFDCD19BD:8ED8839A79369F624B2BFCFBFE311F35" box="[294,425,807,832]" country="Ecuador" latitude="-0.0058333334" longLatPrecision="21" longitude="-76.64361" municipality="Bellavista River" name="Lago Agrio" pageId="17" pageNumber="18" stateProvince="Sucumbios">Lago Agrio</location>
[&quot;A wide river, 3–4 meters across with good flow, bottom stony with some deeper holes, bordered by secondary forest (W-721)&quot;] (
<geoCoordinate id="EE33B38679369F62488EFC9FFD6D1F11" box="[643,757,844,868]" degrees="0" direction="south" minutes="0" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="21" value="-0.0058333334">0°0'21&quot; S</geoCoordinate>
,
<geoCoordinate id="EE33B38679369F62490AFC9FFC061F11" box="[775,926,844,869]" degrees="76" direction="west" minutes="38" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="37" value="-76.64361">76°38'37&quot; W</geoCoordinate>
,
<quantity id="4CFF78A479369F6249A3FC9FFC661F11" box="[942,1022,844,869]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="17" pageNumber="18" unit="m" value="285.0">
<elevation id="002A327279369F6249A3FC9FFC661F11" box="[942,1022,844,869]" metricMagnitude="2" metricUnit="m" metricValue="2.85" pageId="17" pageNumber="18" unit="m" value="285.0">285 m</elevation>
</quantity>
),
<date id="FFB9F38179369F624E15FC9FFB461F11" box="[1048,1246,844,869]" pageId="17" pageNumber="18" value="2014-01-11">
<collectingDate id="EFFD0A6979369F624E15FC9FFB461F11" box="[1048,1246,844,869]" pageId="17" pageNumber="18" value="2014-01-11">11 January 2014</collectingDate>
</date>
,
<collectorName id="26F2B09779369F624EE2FC9FFA001F10" box="[1263,1432,844,869]" pageId="17" pageNumber="18">David Wagner</collectorName>
,
<collectorName id="26F2B09779369F624A9AFCBCFEA61FFD" box="[151,318,879,904]" pageId="17" pageNumber="18">William Haber</collectorName>
,
<collectorName id="26F2B09779369F624B47FCBCFE741FFD" box="[330,492,879,904]" pageId="17" pageNumber="18">Fred Morrison</collectorName>
leg. [
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]
</materialsCitation>
.
</paragraph>
<paragraph id="8BB8D54179369F624ACAFC46FC1019D1" blockId="17.[151,1437,663,1480]" pageId="17" pageNumber="18">
<bibRefCitation id="EF96A8B079369F624ACAFC46FE2D1FD9" author="De Marmels, J." box="[199,437,916,941]" pageId="17" pageNumber="18" pagination="249 - 262" refId="ref50706" refString="De Marmels, J. (2012) Review of the &quot; metallic group &quot; of species of Argia Rambur known from Venezuela, with description of the larva of Argia jocosa Hagen in Selys, 1865 (Odonata: Coenagrionidae). International Journal of Odonatology, 15 (3), 249 - 262. https: // doi. org / 10.1080 / 13887890.2012.717906" type="journal article" year="2012">De Marmels (2012)</bibRefCitation>
provided a thorough review of this species within
<collectingCountry id="F31095D179369F624E36FC47FB361FD8" box="[1083,1198,916,941]" name="Venezuela" pageId="17" pageNumber="18">Venezuela</collectingCountry>
and cleared up its misidentification with
<taxonomicName id="4C07AEC279369F624B9AFC6BFDB31FA5" box="[407,555,952,976]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cupraurea">
<emphasis id="B973095379369F624B9AFC6BFDB31FA5" box="[407,555,952,976]" italics="true" pageId="17" pageNumber="18">A. cupraurea</emphasis>
</taxonomicName>
. Its male is the most variable of all of the metallic red species. The
<typeStatus id="54BC6BE379369F624F38FC64FA041FA5" box="[1333,1436,951,976]" pageId="17" pageNumber="18" type="lectotype">lectotype</typeStatus>
(Fig. 50) is characterized, as
<bibRefCitation id="EF96A8B079369F624BD7FC0FFDE61F81" author="Calvert, P. P." box="[474,638,988,1013]" pageId="17" pageNumber="18" pagination="17 - 72" refId="ref50307" refString="Calvert, P. P. (1901) Odonata. In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia Centrali Americana-Insecta Neuroptera. R. H. Porter &amp; Dulau Co., London, pp. 17 - 72." type="book chapter" year="1901">Calvert (1901)</bibRefCitation>
noted by &quot;Segments 3–7 black on dorsum, basal half of 3–6 blue, 8–10 blue on dorsum.&quot; Most males of this species (including all those known from
<collectingCountry id="F31095D179369F6249FBFC2CFBF3186D" box="[1014,1131,1023,1048]" name="Venezuela" pageId="17" pageNumber="18">Venezuela</collectingCountry>
) differ in having S3–5 blue for two-thirds or more of their length (as in Fig. 52). The male from Bellavista River,
<collectingCountry id="F31095D179369F624E75FBF7FB4D1848" box="[1144,1237,1060,1085]" name="Ecuador" pageId="17" pageNumber="18">Ecuador</collectingCountry>
(Fig. 52) has the inferior black on S3–6 thicker than on males from
<collectingCountry id="F31095D179369F6248D1FB94FCD21815" box="[732,842,1095,1120]" name="Colombia" pageId="17" pageNumber="18">Colombia</collectingCountry>
and
<collectingCountry id="F31095D179369F624989FB94FC631815" box="[900,1019,1095,1120]" name="Venezuela" pageId="17" pageNumber="18">Venezuela</collectingCountry>
. Another male from Lloraudo River,
<collectingCountry id="F31095D179369F624A9AFBBFFE9D18F0" box="[151,261,1132,1157]" name="Colombia" pageId="17" pageNumber="18">Colombia</collectingCountry>
has S3–7 predominantly black (Fig. 51) thus approaching the color pattern of
<taxonomicName id="4C07AEC279369F624E93FBBEFB7218F1" authorityName=": Donnelly" authorityYear="1992" box="[1182,1258,1133,1156]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="cuprea">
<emphasis id="B973095379369F624E93FBBEFB7218F1" box="[1182,1258,1133,1156]" italics="true" pageId="17" pageNumber="18">cuprea</emphasis>
</taxonomicName>
-like species
<taxonomicName id="4C07AEC279369F624F89FBBFFF4918DD" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="dives">
<emphasis id="B973095379369F624F89FBBFFF4918DD" italics="true" pageId="17" pageNumber="18">A. dives</emphasis>
</taxonomicName>
and
<taxonomicName id="4C07AEC279369F624B05FB43FEE818DD" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[264,368,1168,1192]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="philipi">
<emphasis id="B973095379369F624B05FB43FEE818DD" box="[264,368,1168,1192]" italics="true" pageId="17" pageNumber="18">A. philipi</emphasis>
</taxonomicName>
. The thoracic dorsum is normally entirely metallic (as in
<figureCitation id="133CC9C479369F6249F7FB5CFBD318DD" box="[1018,1099,1167,1192]" captionStart="FIGURES 19–22" captionStartId="42.[151,264,1830,1852]" captionTargetBox="[172,1401,213,1802]" captionTargetId="figure@42.[150,1436,193,1809]" captionTargetPageId="42" captionText="FIGURES 19–22. Color pattern of male head, thorax, and S1–4. Head, dorsal view, thorax and S1–4, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3770047" httpUri="https://zenodo.org/record/3770047/files/figure.png" pageId="17" pageNumber="18">Fig. 20</figureCitation>
) but the male from Bellavista River,
<collectingCountry id="F31095D179369F624AE8FB67FEDA18B8" box="[229,322,1204,1229]" name="Ecuador" pageId="17" pageNumber="18">Ecuador</collectingCountry>
(
<figureCitation id="133CC9C479369F624B58FB67FE3218B9" box="[341,426,1204,1229]" captionStart="FIGURES 14–18" captionStartId="41.[151,264,1893,1915]" captionTargetBox="[181,1407,200,1849]" captionTargetId="figure@41.[150,1436,193,1872]" captionTargetPageId="41" captionText="FIGURES 14–18. Color pattern of male head, thorax, and S1–4. Head, dorsal view, thorax and S1–4, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3770045" httpUri="https://zenodo.org/record/3770045/files/figure.png" pageId="17" pageNumber="18">Fig. 17</figureCitation>
) has a narrow blue antehumeral stripe and another male from Reserva Natural El Rey Zamuro,
<collectingCountry id="F31095D179369F624AF2FB04FEF51885" box="[255,365,1239,1264]" name="Colombia" pageId="17" pageNumber="18">Colombia</collectingCountry>
has a similar pale antehumeral stripe that is about as wide as the middorsal black stripe. The middorsal and humeral stripe, the latter of which is divided above at its upper third, show no metallic coloration although metallic reflections are present on the prothorax and head. All of these males conform to what we believe to represent
<taxonomicName id="4C07AEC279369F624B27FA97FE0F1929" authorityName="Hagen in Selys" authorityYear="1865" box="[298,407,1348,1372]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="jocosa">
<emphasis id="B973095379369F624B27FA97FE0F1929" box="[298,407,1348,1372]" italics="true" pageId="17" pageNumber="18">A. jocosa</emphasis>
</taxonomicName>
as the male appendages and genital ligula (unique for this species) match those of the
<typeStatus id="54BC6BE379369F624A9AFAB4FF6619F5" box="[151,254,1383,1408]" pageId="17" pageNumber="18" type="lectotype">lectotype</typeStatus>
(
<figureCitation id="133CC9C479369F624B00FAB4FEF919F5" box="[269,353,1383,1408]" captionStart="FIGURES 94–97" captionStartId="53.[151,264,1884,1906]" captionTargetBox="[166,1425,211,1855]" captionTargetId="figure@53.[151,1436,193,1862]" captionTargetPageId="53" captionText="FIGURES 94–97. Genital ligula. (a) lateral view; (b) ectal view; (c) entolateral view." figureDoi="http://doi.org/10.5281/zenodo.3770064" httpUri="https://zenodo.org/record/3770064/files/figure.png" pageId="17" pageNumber="18">Fig. 95</figureCitation>
). The Bellavista River pair represents a new record for
<collectingCountry id="F31095D179369F6249C0FAB4FBB519F5" box="[973,1069,1383,1408]" name="Ecuador" pageId="17" pageNumber="18">Ecuador</collectingCountry>
. We also examined an additional
<specimenCount id="9D011EC879369F624A9AFA5FFF6A19D1" box="[151,242,1420,1445]" pageId="17" pageNumber="18" type="male">296 ♂♂</specimenCount>
and
<specimenCount id="9D011EC879369F624B25FA5FFEF619D0" box="[296,366,1420,1445]" pageId="17" pageNumber="18" type="female">75♀♀</specimenCount>
from
<collectingCountry id="F31095D179369F624BBEFA5FFDB919D0" box="[435,545,1420,1445]" name="Colombia" pageId="17" pageNumber="18">Colombia</collectingCountry>
and
<collectingCountry id="F31095D179369F62485AFA5FFD5219D0" box="[599,714,1420,1445]" name="Venezuela" pageId="17" pageNumber="18">Venezuela</collectingCountry>
for this account.
</paragraph>
<paragraph id="8BB8D54179369F624ACAFA63FDCD19BD" blockId="17.[151,1437,663,1480]" box="[199,597,1455,1480]" pageId="17" pageNumber="18">
See under
<taxonomicName id="4C07AEC279369F624B30FA63FE3D19BD" authorityName="Garrison &amp; von Ellenrieder" authorityYear="2018" box="[317,421,1456,1480]" class="Insecta" family="Coenagrionidae" genus="Argia" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="philipi">
<emphasis id="B973095379369F624B30FA63FE3D19BD" box="[317,421,1456,1480]" italics="true" pageId="17" pageNumber="18">A. philipi</emphasis>
</taxonomicName>
account below.
</paragraph>
</subSubSection>
</treatment>
</document>
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