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<document id="54481F66C214CD5944232B0F4CA25011" ID-DOI="10.11646/zootaxa.2260.1.31" ID-ISSN="1175-5326" ID-Zenodo-Dep="5308147" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629650827769" checkinUser="felipe" docAuthor="Lowry, J. K. &amp; Stoddart, H. E." docDate="2009" docId="03FCB50CB90A970EFF1682BF9941FD7D" docLanguage="en" docName="zt02260p597.pdf" docOrigin="Zootaxa 2260 (1)" docSource="https://www.biotaxa.org/Zootaxa/article/view/zootaxa.2260.1.31" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tryphosella Bonnier 1893" docType="treatment" docVersion="4" lastPageNumber="581" masterDocId="FFC5CD74B91E971AFF8182289D00FF8F" masterDocTitle="Lysianassidae *" masterLastPageNumber="597" masterPageNumber="561" pageNumber="581" updateTime="1699089016593" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="4FCB14C65401F1A5543237DE847ED602">Lysianassidae *</mods:title>
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<subSubSection id="C34F5791B90A970EFF1682BF9CD7FF3E" box="[151,471,151,177]" pageId="20" pageNumber="581" type="nomenclature">
<paragraph id="8BEA041AB90A970EFF1682BF9CD7FF3E" blockId="20.[151,471,151,177]" box="[151,471,151,177]" pageId="20" pageNumber="581">
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<taxonomicName id="4C557F99B90A970EFF1682BF9CD7FF3E" ID-CoL="7Q5FT" authority="Bonnier, 1893" authorityName="Bonnier" authorityYear="1893" box="[151,471,151,177]" class="Malacostraca" family="Tryphosidae" genus="Tryphosella" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="581" phylum="Arthropoda" rank="genus">
<emphasis id="B921D808B90A970EFF1682BF9CD7FF3E" bold="true" box="[151,471,151,177]" pageId="20" pageNumber="581">
<emphasis id="B921D808B90A970EFF1682BF9C25FF3E" bold="true" box="[151,293,151,177]" italics="true" pageId="20" pageNumber="581">Tryphosella</emphasis>
<bibRefCitation id="EFC479EBB90A970EFEAD82BF9CD7FF3E" author="Bonnier, J." box="[300,471,151,177]" pageId="20" pageNumber="581" pagination="161 - 207" refId="ref14117" refString="Bonnier, J. (1893) Les amphipodes du Boulonnais (1). Bulletin Scientifique de la France et de la Belgique, 24, 161 - 207, pls 5 - 8." type="journal article" year="1893">Bonnier, 1893</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<subSubSection id="C34F5791B90A970EFF1682C89941FD7D" pageId="20" pageNumber="581" type="discussion">
<paragraph id="8BEA041AB90A970EFF1682C89CE6FDBF" blockId="20.[151,1438,224,754]" pageId="20" pageNumber="581">
<emphasis id="B921D808B90A970EFF1682C89C13FF75" bold="true" box="[151,275,224,250]" pageId="20" pageNumber="581">Remarks.</emphasis>
There is considerable variation within the genus
<taxonomicName id="4C557F99B90A970EFCF782C99906FF75" authorityName="Bonnier" authorityYear="1893" box="[886,1030,225,250]" class="Malacostraca" family="Tryphosidae" genus="Tryphosella" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="581" phylum="Arthropoda" rank="genus">
<emphasis id="B921D808B90A970EFCF782C99906FF75" box="[886,1030,225,250]" italics="true" pageId="20" pageNumber="581">Tryphosella</emphasis>
</taxonomicName>
as defined by
<bibRefCitation id="EFC479EBB90A970EFB3D82C99DEAFEAD" author="Lowry, J. K. &amp; Stoddart, H. E." pageId="20" pageNumber="581" pagination="97 - 174" refId="ref14657" refString="Lowry, J. K. &amp; Stoddart, H. E. (1995) The Amphipoda (Crustacea) of Madang Lagoon: Lysianassidae, Opisidae, Uristidae, Wandinidae and Stegocephalidae. Records of the Australian Museum, Supplement, 22, 97 - 174." type="journal article" year="1995">Lowry &amp; Stoddart (1995)</bibRefCitation>
. One of the main defining characters of
<taxonomicName id="4C557F99B90A970EFD5183209E5DFEAE" authorityName="Bonnier" authorityYear="1893" box="[720,861,264,289]" class="Malacostraca" family="Tryphosidae" genus="Tryphosella" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="581" phylum="Arthropoda" rank="genus">
<emphasis id="B921D808B90A970EFD5183209E5DFEAE" box="[720,861,264,289]" italics="true" pageId="20" pageNumber="581">Tryphosella</emphasis>
</taxonomicName>
is the morphology of gnathopod
<quantity id="4CADA9FFB90A970EFB718320981EFEAD" box="[1264,1310,264,290]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="20" pageNumber="581" unit="in" value="1.0">1 in</quantity>
which the coxa is reduced (slightly shorter than coxa 2) and tapering and the ischium is short, as evidenced in the
<typeStatus id="54EEBAB8B90A970EFAE88306989BFEC7" box="[1385,1435,302,328]" pageId="20" pageNumber="581">type</typeStatus>
species
<taxonomicName id="4C557F99B90A970EFF78837D9F10FEE0" authority="Bonnier, 1893" authorityName="Bonnier" authorityYear="1893" box="[249,528,341,367]" class="Malacostraca" family="Tryphosidae" genus="Tryphosella" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="581" phylum="Arthropoda" rank="species" species="sarsi">
<emphasis id="B921D808B90A970EFF78837D9C54FEE1" box="[249,340,341,366]" italics="true" pageId="20" pageNumber="581">T. sarsi</emphasis>
<bibRefCitation id="EFC479EBB90A970EFEDC837D9F10FEE0" author="Bonnier, J." box="[349,528,341,367]" pageId="20" pageNumber="581" pagination="161 - 207" refId="ref14117" refString="Bonnier, J. (1893) Les amphipodes du Boulonnais (1). Bulletin Scientifique de la France et de la Belgique, 24, 161 - 207, pls 5 - 8." type="journal article" year="1893">Bonnier, 1893</bibRefCitation>
</taxonomicName>
. Within the genus, especially in Australasian species, there is a tendency towards shortening the coxa to a point where it becomes about as long as broad and the ischium becomes at least twice as long as broad, as evidenced in the new species
<taxonomicName id="4C557F99B90A970EFCDD838A9ED6FE34" authorityName="Lowry &amp; Stoddart" authorityYear="2009" box="[860,982,418,443]" class="Malacostraca" family="Tryphosidae" genus="Tryphosella" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="581" phylum="Arthropoda" rank="species" species="seasana">
<emphasis id="B921D808B90A970EFCDD838A9ED6FE34" box="[860,982,418,443]" italics="true" pageId="20" pageNumber="581">T. seasana</emphasis>
</taxonomicName>
. Given the two extremes there appears to be a case for establishing a second genus, but both characters are gradational so that in the case of
<taxonomicName id="4C557F99B90A970EFA0783E19C00FD86" authorityName="Lowry &amp; Stoddart" authorityYear="2009" class="Malacostraca" family="Tryphosidae" genus="Tryphosella" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="581" phylum="Arthropoda" rank="species" species="flynnana">
<emphasis id="B921D808B90A970EFA0783E19C00FD86" italics="true" pageId="20" pageNumber="581">T. flynnana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C557F99B90A970EFEBC83D89CBDFD86" authorityName="Lowry &amp; Stoddart" authorityYear="2009" box="[317,445,496,521]" class="Malacostraca" family="Tryphosidae" genus="Tryphosella" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="581" phylum="Arthropoda" rank="species" species="seasana">
<emphasis id="B921D808B90A970EFEBC83D89CBDFD86" box="[317,445,496,521]" italics="true" pageId="20" pageNumber="581">T. seasana</emphasis>
</taxonomicName>
, which are very similar,
<taxonomicName id="4C557F99B90A970EFD6E83D89E77FD86" authorityName="Lowry &amp; Stoddart" authorityYear="2009" box="[751,887,496,521]" class="Malacostraca" family="Tryphosidae" genus="Tryphosella" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="581" phylum="Arthropoda" rank="species" species="flynnana">
<emphasis id="B921D808B90A970EFD6E83D89E77FD86" box="[751,887,496,521]" italics="true" pageId="20" pageNumber="581">T. flynnana</emphasis>
</taxonomicName>
would remain in
<taxonomicName id="4C557F99B90A970EFBD083D899E0FD86" authorityName="Bonnier" authorityYear="1893" box="[1105,1248,496,521]" class="Malacostraca" family="Tryphosidae" genus="Tryphosella" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="581" phylum="Arthropoda" rank="genus">
<emphasis id="B921D808B90A970EFBD083D899E0FD86" box="[1105,1248,496,521]" italics="true" pageId="20" pageNumber="581">Tryphosella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C557F99B90A970EFA9D83D8989CFD86" authorityName="Lowry &amp; Stoddart" authorityYear="2009" box="[1308,1436,496,521]" class="Malacostraca" family="Tryphosidae" genus="Tryphosella" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="581" phylum="Arthropoda" rank="species" species="seasana">
<emphasis id="B921D808B90A970EFA9D83D8989CFD86" box="[1308,1436,496,521]" italics="true" pageId="20" pageNumber="581">T. seasana</emphasis>
</taxonomicName>
would move to a new genus.
</paragraph>
<paragraph id="8BEA041AB90A970EFF4780159941FD7D" blockId="20.[151,1438,224,754]" pageId="20" pageNumber="581">
Another set of defining characters is the prominent ridge on the coxae of pereopods 4 and 5 and an apparent photophore on the basis of pereopod 5. Several species (
<taxonomicName id="4C557F99B90A970EFC25804C9945FDF2" authority="Lowry &amp; Stoddart, 2009" authorityName="Lowry &amp; Stoddart" authorityYear="2009" box="[932,1093,612,637]" class="Malacostraca" family="Tryphosidae" genus="Tryphosella" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="581" phylum="Arthropoda" rank="species" species="charlotteae" status="sp. nov.">
<emphasis id="B921D808B90A970EFC25804C9945FDF2" box="[932,1093,612,637]" italics="true" pageId="20" pageNumber="581">T. charlotteae</emphasis>
</taxonomicName>
<emphasis id="B921D808B90A970EFBCD804B99A9FDF2" bold="true" box="[1100,1193,611,637]" pageId="20" pageNumber="581">
<taxonomicNameLabel id="A2126573B90A970EFBCD804B99A9FDF2" box="[1100,1193,611,637]" pageId="20" pageNumber="581" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C557F99B90A970EFB36804C9DE3FD2B" authority="(Stebbing, 1888)" baseAuthorityName="Stebbing" baseAuthorityYear="1888" class="Malacostraca" family="Tryphosidae" genus="Tryphosella" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="581" phylum="Arthropoda" rank="species" species="miersi">
<emphasis id="B921D808B90A970EFB36804C981CFDF2" box="[1207,1308,612,637]" italics="true" pageId="20" pageNumber="581">T. miersi</emphasis>
(
<bibRefCitation id="EFC479EBB90A970EFAAA804C9DDAFD2B" author="Stebbing, T. R. R." pageId="20" pageNumber="581" pagination="1 - 1737" refId="ref15054" refString="Stebbing, T. R. R. (1888) Report on the Amphipoda collected by H. M. S. Challenger during the years 1873 - 1876. Report on the Scientific Results of the Voyage of H. M. S. Challenger during the years 1873 - 76, Zoology 29, 1 - 1737, pls 1 - 210." type="journal article" year="1888">Stebbing, 1888</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C557F99B90A970EFF7280A29F5EFD2B" authority="(Pirlot, 1936))" authorityName="" baseAuthorityName="Pirlot" baseAuthorityYear="1936" box="[243,606,650,676]" class="Malacostraca" family="Tryphosidae" genus="Tryphosella" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="581" phylum="Arthropoda" rank="species" species="mucronata">
<emphasis id="B921D808B90A970EFF7280A29C9BFD2C" box="[243,411,650,675]" italics="true" pageId="20" pageNumber="581">T. mucronata</emphasis>
(
<bibRefCitation id="EFC479EBB90A970EFE2F80A29F4AFD2B" author="Pirlot, J. M." box="[430,586,650,676]" pageId="20" pageNumber="581" pagination="237 - 328" refId="ref14927" refString="Pirlot, J. M. (1936) Les amphipodes de l'expedition du Siboga. Deuxieme partie: Les amphipodes gammarides, II. - Les amphipodes de la mer profonde. 3: Addendum et partie generale. III. - Les amphipodes littoraux. 1: Lysianassidae, Ampeliscidae, Leucothoidae, Stenothoidae, Phliantidae, Colomastigidae, Ochlesidae, Liljeborgiidae, Oedicerotidae, Synopiidae, Eusiridae, Gammaridae. Siboga-Expeditie, Monographie, 33 e, 237 - 328." type="book chapter" year="1936">Pirlot, 1936</bibRefCitation>
))
</taxonomicName>
known from
<collectingCountry id="F342448AB90A970EFC8F80A29E83FD2B" box="[782,899,650,676]" name="Australia" pageId="20" pageNumber="581">Australia</collectingCountry>
and New
<collectingCountry id="F342448AB90A970EFB8980A29963FD2B" box="[1032,1123,650,676]" name="Guinea-Bissau" pageId="20" pageNumber="581">Guinea</collectingCountry>
share these apomorphic characters. In other respects they cannot be separated from the genus
<taxonomicName id="4C557F99B90A970EFC3E80999949FD45" authorityName="Bonnier" authorityYear="1893" box="[959,1097,689,714]" class="Malacostraca" family="Tryphosidae" genus="Tryphosella" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="581" phylum="Arthropoda" rank="genus">
<emphasis id="B921D808B90A970EFC3E80999949FD45" box="[959,1097,689,714]" italics="true" pageId="20" pageNumber="581">Tryphosella</emphasis>
</taxonomicName>
and appear to form a species complex within it. At this stage we prefer to maintain all species in
<taxonomicName id="4C557F99B90A970EFC2E80F09939FD7E" authorityName="Bonnier" authorityYear="1893" box="[943,1081,728,753]" class="Malacostraca" family="Tryphosidae" genus="Tryphosella" kingdom="Animalia" order="Amphipoda" pageId="20" pageNumber="581" phylum="Arthropoda" rank="genus">
<emphasis id="B921D808B90A970EFC2E80F09939FD7E" box="[943,1081,728,753]" italics="true" pageId="20" pageNumber="581">Tryphosella</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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