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<document id="51BA93BCE8040881B3D4B6BC94D894B6" ID-DOI="10.11646/zootaxa.3986.2.5" ID-GBIF-Dataset="0d76d10a-3a7e-495d-9489-869cef93683d" ID-ISSN="1175-5326" ID-Zenodo-Dep="243017" ID-ZooBank="3DA5305F-4D0B-4031-8240-9574842B336F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461262227298" checkinUser="plazi" docAuthor="Kamiński, Marcin Jan" docDate="2015" docId="03FC87F6D5452F3AFF6AC460FC60FE5B" docLanguage="en" docName="zt03986p226.pdf" docOrigin="Zootaxa 3986 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Quadrideres kazimierzi Kamiński, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="222" masterDocId="FFC5FF8ED5472F3FFFFDC521FFECFF8D" masterDocTitle="A taxonomic revision of the Quadrideres simplicipes species-group (Coleoptera: Tenebrionidae: Pedinini), including description of a new species from Tanzania" masterLastPageNumber="226" masterPageNumber="217" pageNumber="219" updateTime="1698609764883" updateUser="plazi">
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<mods:title id="F496C53EC6A9B2A7E21C2DB956FE2CF3">A taxonomic revision of the Quadrideres simplicipes species-group (Coleoptera: Tenebrionidae: Pedinini), including description of a new species from Tanzania</mods:title>
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<mods:namePart id="5AF50C9D5295C476FE47C573A4D9E45A">Kamiński, Marcin Jan</mods:namePart>
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<mods:date id="79398B5E4C7A0145C8A1806E63BFCED6">2015</mods:date>
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<treatment id="03FC87F6D5452F3AFF6AC460FC60FE5B" ID-DOI="http://doi.org/10.5281/zenodo.6114056" ID-GBIF-Taxon="119637306" ID-Zenodo-Dep="6114056" LSID="urn:lsid:plazi:treatment:03FC87F6D5452F3AFF6AC460FC60FE5B" httpUri="http://treatment.plazi.org/id/03FC87F6D5452F3AFF6AC460FC60FE5B" lastPageId="5" lastPageNumber="222" pageId="2" pageNumber="219">
<subSubSection id="C34F656BD5452F3DFF6AC460FDF8FED6" box="[151,532,321,347]" pageId="2" pageNumber="219" type="nomenclature">
<paragraph id="8BEA36E0D5452F3DFF6AC460FDF8FED6" blockId="2.[151,532,321,347]" box="[151,532,321,347]" pageId="2" pageNumber="219">
<heading id="D0A2818CD5452F3DFF6AC460FDF8FED6" bold="true" box="[151,532,321,347]" fontSize="11" level="1" pageId="2" pageNumber="219" reason="1">
<emphasis id="B921EAF2D5452F3DFF6AC460FDF8FED6" bold="true" box="[151,532,321,347]" pageId="2" pageNumber="219">
<taxonomicName id="4C554D63D5452F3DFF6AC460FE5CFED6" box="[151,432,321,347]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="kazimierzi" status="sp. nov.">
<emphasis id="B921EAF2D5452F3DFF6AC460FE5CFED6" bold="true" box="[151,432,321,347]" italics="true" pageId="2" pageNumber="219">Quadrideres kazimierzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2125789D5452F3DFE4AC460FDF8FED6" box="[439,532,321,347]" pageId="2" pageNumber="219" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F656BD5452F3DFF6AC4A6FD43FDF5" pageId="2" pageNumber="219" type="materials_examined">
<paragraph id="8BEA36E0D5452F3DFF6AC4A6FB58FDD8" blockId="2.[151,1437,391,1292]" pageId="2" pageNumber="219">
<emphasis id="B921EAF2D5452F3DFF6AC4A6FE55FE2D" bold="true" box="[151,441,391,416]" pageId="2" pageNumber="219">
<typeStatus id="54EE8842D5452F3DFF6AC4A6FF3EFE2D" box="[151,210,391,416]" pageId="2" pageNumber="219">Type</typeStatus>
material.
<typeStatus id="54EE8842D5452F3DFEB1C4A6FE55FE2D" box="[332,441,391,416]" pageId="2" pageNumber="219" type="holotype">Holotype</typeStatus>
</emphasis>
, male (
<collectionCode id="ED44AE25D5452F3DFDF1C4A9FD8BFE2D" box="[524,615,392,416]" pageId="2" pageNumber="219">BMNH</collectionCode>
): “Tang. Terr. / Ukerewe I. / Father Conrad.”.
<typeStatus id="54EE8842D5452F3DFB94C4A6FB0DFE2D" box="[1129,1249,391,416]" pageId="2" pageNumber="219" type="paratype">
<emphasis id="B921EAF2D5452F3DFB94C4A6FB0DFE2D" bold="true" box="[1129,1249,391,416]" pageId="2" pageNumber="219">Paratypes</emphasis>
</typeStatus>
: male (
<collectionCode id="ED44AE25D5452F3DFAC8C4A9FA7CFE2D" box="[1333,1424,392,416]" pageId="2" pageNumber="219">BMNH</collectionCode>
): “
<date id="FFEB1020D5452F3DFF5FC48DFF3AFE49" box="[162,214,428,452]" pageId="2" pageNumber="219">4.IX</date>
W/682”, “Tang. Terr. / Ukerewe I. / Father Conrad.”, “
<taxonomicName id="4C554D63D5452F3DFCBDC48DFC78FE48" box="[832,916,428,453]" class="Insecta" family="Tenebrionidae" genus="Selinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="226" phylum="Arthropoda" rank="genus">Selinus</taxonomicName>
/
<taxonomicName id="4C554D63D5452F3DFC56C48DFBFEFE48" box="[939,1042,428,453]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="femineus">femineus</taxonomicName>
/ Lucas / Det. G.E.Bryant”; female (
<collectionCode id="ED44AE25D5452F3DFF62C4F1FF16FE65" box="[159,250,464,488]" pageId="2" pageNumber="219">BMNH</collectionCode>
): “Tang. Terr. / Ukerewe I. / Father Conrad.”, “1322”, “682”;
<specimenCount id="9D53FD69D5452F3DFC46C4F1FBC4FE65" box="[955,1064,463,488]" pageId="2" pageNumber="219" type="female">2 females</specimenCount>
(
<collectionCode id="ED44AE25D5452F3DFBC4C4F1FB7DFE65" box="[1081,1169,464,488]" pageId="2" pageNumber="219">ZMHB</collectionCode>
): “Victoria Nayansa / I. Ukerewe / Conrads S.G.“; female (
<collectionCode id="ED44AE25D5452F3DFDE3C4D5FD9AFD81" box="[542,630,500,524]" pageId="2" pageNumber="219">ZMHB</collectionCode>
): “D.O. Afrika / Victoria Nyansa /
<collectingCountry id="F3427670D5452F3DFBFDC4D5FBDAFD80" box="[1024,1078,500,525]" name="Iceland" pageId="2" pageNumber="219">Insel</collectingCountry>
Ukerewe / Conrads S.G”; male (
<collectionCode id="ED44AE25D5452F3DFF62C739FF1BFDBD" box="[159,247,536,560]" pageId="2" pageNumber="219">ZMHB</collectionCode>
): “Usika / Hssukuma / Hönbrch.
<date id="FFEB1020D5452F3DFD82C739FD2EFDBD" box="[639,706,536,560]" pageId="2" pageNumber="219" value="2013-09">IX.13</date>
”, “
<taxonomicName id="4C554D63D5452F3DFD17C739FCD7FDBD" box="[746,827,536,560]" class="Insecta" family="Tenebrionidae" genus="Selinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAF2D5452F3DFD17C739FCD7FDBD" box="[746,827,536,560]" italics="true" pageId="2" pageNumber="219">Selinus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C554D63D5452F3DFCA5C739FC25FDBD" box="[856,969,536,560]" class="Insecta" family="Tenebrionidae" genus="Selinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B921EAF2D5452F3DFCA5C739FC25FDBD" box="[856,969,536,560]" italics="true" pageId="2" pageNumber="219">parallelus</emphasis>
</taxonomicName>
/ Ancey”; female (
<collectionCode id="ED44AE25D5452F3DFB53C739FAE7FDBD" LSID="urn:lsid:biocol.org:col:34988" box="[1198,1291,536,560]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="219">MNHN</collectionCode>
): “Ukerewe- Nansio / Ost-Afrika / leg. P.A. Conrads.”, “
<taxonomicName id="4C554D63D5452F3DFD81C71DFCE5FDD9" box="[636,777,572,596]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAF2D5452F3DFD81C71DFCE5FDD9" box="[636,777,572,596]" italics="true" pageId="2" pageNumber="219">Quadrideres</emphasis>
</taxonomicName>
/
<taxonomicName id="4C554D63D5452F3DFCE3C71DFCB3FDD9" box="[798,863,572,596]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="lesnei">
<emphasis id="B921EAF2D5452F3DFCE3C71DFCB3FDD9" box="[798,863,572,596]" italics="true" pageId="2" pageNumber="219">lesnei</emphasis>
</taxonomicName>
/ Koch / P. Ardoin det. 1962”.
</paragraph>
<paragraph id="8BEA36E0D5452F3DFF3AC77EFD43FDF5" blockId="2.[151,1437,391,1292]" box="[199,687,607,632]" pageId="2" pageNumber="219">
<emphasis id="B921EAF2D5452F3DFF3AC77EFE9EFDF5" bold="true" box="[199,370,607,632]" pageId="2" pageNumber="219">Locus typicus.</emphasis>
Ukerewe Island (
<collectingCountry id="F3427670D5452F3DFDC5C77EFD48FDF5" box="[568,676,607,632]" name="Tanzania" pageId="2" pageNumber="219">Tanzania</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C34F656BD5452F3DFF3AC7A5FE9CFD6A" pageId="2" pageNumber="219" type="etymology">
<paragraph id="8BEA36E0D5452F3DFF3AC7A5FE9CFD6A" blockId="2.[151,1437,391,1292]" pageId="2" pageNumber="219">
<emphasis id="B921EAF2D5452F3DFF3AC7A5FEA2FD10" bold="true" box="[199,334,644,669]" pageId="2" pageNumber="219">Etymology.</emphasis>
This new species is dedicated to the memory of my grandfather Kazimierz Józef Kamiński (born on the 13th of December, 1926, in Grabina Radziwiłłowska,
<collectingCountry id="F3427670D5452F3DFC91C78BFC51FD4E" box="[876,957,682,707]" name="Poland" pageId="2" pageNumber="219">Poland</collectingCountry>
, died on the 2nd of November, 2009, in Żyrardów,
<collectingCountry id="F3427670D5452F3DFEECC7EFFE88FD6A" box="[273,356,718,743]" name="Poland" pageId="2" pageNumber="219">Poland</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C34F656BD5452F3DFF3AC7D3FE3FFC6B" pageId="2" pageNumber="219" type="diagnosis">
<paragraph id="8BEA36E0D5452F3DFF3AC7D3FE3FFC6B" blockId="2.[151,1437,391,1292]" pageId="2" pageNumber="219">
<emphasis id="B921EAF2D5452F3DFF3AC7D3FEADFC86" bold="true" box="[199,321,754,779]" pageId="2" pageNumber="219">Diagnosis.</emphasis>
This species is similar to all of the members of the
<taxonomicName id="4C554D63D5452F3DFC7CC7D2FBCDFC86" box="[897,1057,755,779]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="simplicipes">
<emphasis id="B921EAF2D5452F3DFC7CC7D2FBCDFC86" box="[897,1057,755,779]" italics="true" pageId="2" pageNumber="219">Q. simplicipes</emphasis>
</taxonomicName>
species-group (
<taxonomicName id="4C554D63D5452F3DFB2EC7D2FA9FFC86" box="[1235,1395,755,779]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="simplicipes">
<emphasis id="B921EAF2D5452F3DFB2EC7D2FA9FFC86" box="[1235,1395,755,779]" italics="true" pageId="2" pageNumber="219">Q. simplicipes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C554D63D5452F3DFA7CC7D2FEE3FCA2" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="ruandanus">
<emphasis id="B921EAF2D5452F3DFA7CC7D2FEE3FCA2" italics="true" pageId="2" pageNumber="219">Q. ruandanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C554D63D5452F3DFEB9C636FE56FCA2" box="[324,442,791,815]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="robynsi">
<emphasis id="B921EAF2D5452F3DFEB9C636FE56FCA2" box="[324,442,791,815]" italics="true" pageId="2" pageNumber="219">Q. robynsi</emphasis>
</taxonomicName>
) having the inner contours of male anterior tibiae without strongly postmedian dilatation (cf. Kamiński 2013a) and with the secondary intervals and epipleurae of elytra with coarse punctures (Fig. 7, 8).
<taxonomicName id="4C554D63D5452F3DFA7CC61AFEE6FCF4" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="kazimierzi" status="sp. nov.">
<emphasis id="B921EAF2D5452F3DFA7CC61AFEE6FCF4" italics="true" pageId="2" pageNumber="219">Q. kazimierzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2125789D5452F3DFEECC643FE84FCF4" box="[273,360,866,889]" pageId="2" pageNumber="219" rank="species">sp. nov.</taxonomicNameLabel>
can be easily distinguished from
<taxonomicName id="4C554D63D5452F3DFD1CC643FC6EFCF4" box="[737,898,865,889]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="simplicipes">
<emphasis id="B921EAF2D5452F3DFD1CC643FC6EFCF4" box="[737,898,865,889]" italics="true" pageId="2" pageNumber="219">Q. simplicipes</emphasis>
</taxonomicName>
by having an unbordered 5th abdominal ventrite (Fig. 9, 10). From hitherto described representatives of the
<taxonomicName id="4C554D63D5452F3DFCBAC6A7FC06FC10" box="[839,1002,901,925]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="simplicipes">
<emphasis id="B921EAF2D5452F3DFCBAC6A7FC06FC10" box="[839,1002,901,925]" italics="true" pageId="2" pageNumber="219">Q. simplicipes</emphasis>
</taxonomicName>
species-group,
<taxonomicName id="4C554D63D5452F3DFB5CC6A7FAD5FC10" box="[1185,1337,901,925]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="kazimierzi" status="sp. nov.">
<emphasis id="B921EAF2D5452F3DFB5CC6A7FAD5FC10" box="[1185,1337,901,925]" italics="true" pageId="2" pageNumber="219">Q. kazimierzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2125789D5452F3DFABEC6A7FA70FC10" box="[1347,1436,902,925]" pageId="2" pageNumber="219" rank="species">sp. nov.</taxonomicNameLabel>
differs by having strong lateral indentations and interrupted emargination of pronotal base, and pronotum widest at 0.75 of its length (Fig. 57).
</paragraph>
</subSubSection>
<subSubSection id="C34F656BD5452F3AFF3AC6D0FE29FE03" lastPageId="5" lastPageNumber="222" pageId="2" pageNumber="219" type="description">
<paragraph id="8BEA36E0D5452F3DFF3AC6D0FCBCFA86" blockId="2.[151,1437,391,1292]" pageId="2" pageNumber="219">
<emphasis id="B921EAF2D5452F3DFF3AC6D0FEB5FB87" bold="true" box="[199,345,1009,1034]" pageId="2" pageNumber="219">Description.</emphasis>
Measurements. Body—length = 9.011.0 mm; width ratio elytra / pronotum = 1.01.1; length ratio elytra / middle of pronotum = 2.12.2. Clypeus— clypeal emargination width / depth ratio ca. 7.0. Antennae—length ratio of antennomere 3rd / 2nd ca. 3.0. Maxillary palp—ratio width of maxillary palp / length of 3rd antennomere ca. 1.2. Pronotum—middle of pronotum length / width ratio = 0.80.9. Elytra—elytra length / width ratio = 1.31.4. Scutellum—width ratio anterior margin of elytra / scutellum at base ca. 5.5. Metaventritelength ratio cavity of hind coxa / metaventrite between the insertions of mid and hind coxae = 2.72.8. Abdomenwidth ratio process of 1st abdominal ventrite / process of metaventrite = 1.31.5. Male legs— protibiae length / width ratio = 4.14.4; profemur length / width ratio = 2.83.0.
</paragraph>
<caption id="DF2A6668D5452F3DFF6AC299FC8DF860" httpUri="https://zenodo.org/record/243018/files/figure.png" pageId="2" pageNumber="219" targetBox="[155,1433,1351,1949]" targetPageId="2">
<paragraph id="8BEA36E0D5452F3DFF6AC299FC8DF860" blockId="2.[151,1436,1976,2029]" pageId="2" pageNumber="219">
<emphasis id="B921EAF2D5452F3DFF6AC299FEDAF840" bold="true" box="[151,310,1976,1999]" pageId="2" pageNumber="219">FIGURES 14</emphasis>
. Habitus of the representatives of the
<taxonomicName id="4C554D63D5452F3DFD34C299FCA4F843" box="[713,840,1976,1998]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAF2D5452F3DFD34C299FCA4F843" box="[713,840,1976,1998]" italics="true" pageId="2" pageNumber="219">Quadrideres</emphasis>
</taxonomicName>
s
<emphasis id="B921EAF2D5452F3DFCA6C299FC2FF843" box="[859,963,1976,1998]" italics="true" pageId="2" pageNumber="219">implicipes</emphasis>
species-group:
<taxonomicName id="4C554D63D5452F3DFB95C298FB1DF843" box="[1128,1265,1976,1998]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="kazimierzi" status="sp. nov.">
<emphasis id="B921EAF2D5452F3DFB95C298FB1DF843" box="[1128,1265,1976,1998]" italics="true" pageId="2" pageNumber="219">Q. kazimierzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2125789D5452F3DFB07C298FAA1F843" box="[1274,1357,1977,1998]" pageId="2" pageNumber="219" rank="species">
<emphasis id="B921EAF2D5452F3DFB07C298FAA1F843" bold="true" box="[1274,1357,1977,1998]" pageId="2" pageNumber="219">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<emphasis id="B921EAF2D5452F3DFAA2C299FA87F840" bold="true" box="[1375,1387,1976,1997]" pageId="2" pageNumber="219">1</emphasis>
),
<taxonomicName id="4C554D63D5452F3DFA7EC298FEE5F860" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="simplicipes">
<emphasis id="B921EAF2D5452F3DFA7EC298FEE5F860" italics="true" pageId="2" pageNumber="219">Q. simplicipes</emphasis>
</taxonomicName>
(
<emphasis id="B921EAF2D5452F3DFEEAC2F6FEC8F861" bold="true" box="[279,292,2007,2028]" pageId="2" pageNumber="219">2</emphasis>
),
<taxonomicName id="4C554D63D5452F3DFEC4C2F9FE53F860" box="[313,447,2007,2029]" class="Insecta" family="Tenebrionidae" genus="Selinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B921EAF2D5452F3DFEC4C2F9FE53F860" box="[313,447,2007,2029]" italics="true" pageId="2" pageNumber="219">Q. parallelus</emphasis>
</taxonomicName>
(
<emphasis id="B921EAF2D5452F3DFE30C2F6FE36F861" bold="true" box="[461,474,2007,2028]" pageId="2" pageNumber="219">3</emphasis>
) and
<taxonomicName id="4C554D63D5452F3DFDE9C2F9FD92F860" box="[532,638,2007,2029]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="219" phylum="Arthropoda" rank="species" species="robynsi">
<emphasis id="B921EAF2D5452F3DFDE9C2F9FD92F860" box="[532,638,2007,2029]" italics="true" pageId="2" pageNumber="219">Q. robynsi</emphasis>
</taxonomicName>
(
<emphasis id="B921EAF2D5452F3DFD71C2F6FD74F861" bold="true" box="[652,664,2007,2028]" pageId="2" pageNumber="219">4</emphasis>
). Scale bar = 1mm.
</paragraph>
</caption>
<paragraph id="8BEA36E0D5442F3CFF6AC2A5FF56F857" blockId="3.[151,1437,1924,2010]" pageId="3" pageNumber="220">
<emphasis id="B921EAF2D5442F3CFF6AC2A5FEA8F817" bold="true" box="[151,324,1924,1947]" pageId="3" pageNumber="220">FIGURES 511</emphasis>
. Diagnostic characters proposed for the representatives of the
<taxonomicName id="4C554D63D5442F3CFC00C2A5FB90F817" box="[1021,1148,1924,1946]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="220" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAF2D5442F3CFC00C2A5FB90F817" box="[1021,1148,1924,1946]" italics="true" pageId="3" pageNumber="220">Quadrideres</emphasis>
</taxonomicName>
s
<emphasis id="B921EAF2D5442F3CFB69C2A5FB10F817" box="[1172,1276,1924,1946]" italics="true" pageId="3" pageNumber="220">implicipes</emphasis>
species-group. Pronotum (
<emphasis id="B921EAF2D5442F3CFEF4C287FEC2F836" bold="true" box="[265,302,1958,1980]" pageId="3" pageNumber="220">57</emphasis>
), 5th ventrite (
<emphasis id="B921EAF2D5442F3CFE41C287FE02F836" bold="true" box="[444,494,1958,1979]" pageId="3" pageNumber="220">9, 10</emphasis>
) and male protibia (
<emphasis id="B921EAF2D5442F3CFD46C287FD3DF836" bold="true" box="[699,721,1958,1979]" pageId="3" pageNumber="220">11</emphasis>
).
<taxonomicName id="4C554D63D5442F3CFD15C286FC82F831" box="[744,878,1958,1980]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="kazimierzi" status="sp. nov.">
<emphasis id="B921EAF2D5442F3CFD15C286FC82F831" box="[744,878,1958,1980]" italics="true" pageId="3" pageNumber="220">Q. kazimierzi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2125789D5442F3CFC89C287FC28F836" box="[884,964,1958,1979]" pageId="3" pageNumber="220" rank="species">
<emphasis id="B921EAF2D5442F3CFC89C287FC28F836" bold="true" box="[884,964,1958,1979]" pageId="3" pageNumber="220">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<emphasis id="B921EAF2D5442F3CFC2EC287FBF2F836" bold="true" box="[979,1054,1958,1980]" pageId="3" pageNumber="220">57, 10</emphasis>
),
<taxonomicName id="4C554D63D5442F3CFBCEC286FB54F831" box="[1075,1208,1958,1980]" class="Insecta" family="Tenebrionidae" genus="Selinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B921EAF2D5442F3CFBCEC286FB54F831" box="[1075,1208,1958,1980]" italics="true" pageId="3" pageNumber="220">Q. parallelus</emphasis>
</taxonomicName>
(
<emphasis id="B921EAF2D5442F3CFB3BC287FB1BF836" bold="true" box="[1222,1271,1958,1979]" pageId="3" pageNumber="220">8, 11</emphasis>
),
<taxonomicName id="4C554D63D5442F3CFAF1C286FA70F831" box="[1292,1436,1958,1980]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="220" phylum="Arthropoda" rank="species" species="simplicipes">
<emphasis id="B921EAF2D5442F3CFAF1C286FA70F831" box="[1292,1436,1958,1980]" italics="true" pageId="3" pageNumber="220">Q. simplicipes</emphasis>
</taxonomicName>
(
<emphasis id="B921EAF2D5442F3CFF62C2E4FF40F857" bold="true" box="[159,172,1989,2010]" pageId="3" pageNumber="220">9</emphasis>
).
</paragraph>
<caption id="DF2A6668D5432F3BFF6AC0D6FE3CF9E7" httpUri="https://zenodo.org/record/243019/files/figure.png" pageId="4" pageNumber="221" targetBox="[173,1413,184,1505]" targetPageId="4">
<paragraph id="8BEA36E0D5432F3BFF6AC0D6FE3CF9E7" blockId="4.[151,1436,1527,1642]" pageId="4" pageNumber="221">
<emphasis id="B921EAF2D5432F3BFF6AC0D6FEA1F981" bold="true" box="[151,333,1527,1550]" pageId="4" pageNumber="221">FIGURES 1213</emphasis>
. Distribution of the
<taxonomicName id="4C554D63D5432F3BFDE1C0D6FD77F980" box="[540,667,1527,1549]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="221" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAF2D5432F3BFDE1C0D6FD77F980" box="[540,667,1527,1549]" italics="true" pageId="4" pageNumber="221">Quadrideres</emphasis>
</taxonomicName>
s
<emphasis id="B921EAF2D5432F3BFD56C0D6FCFFF980" box="[683,787,1527,1549]" italics="true" pageId="4" pageNumber="221">implicipes</emphasis>
species-group illustrated using digital elevation model (
<emphasis id="B921EAF2D5432F3BFAB2C0D6FA84F981" bold="true" box="[1359,1384,1527,1548]" pageId="4" pageNumber="221">12</emphasis>
) and Afrotropical ecoregions and the distribution of the
<taxonomicName id="4C554D63D5432F3BFD55C337FC4EF9A1" box="[680,930,1558,1580]" class="Insecta" family="Tenebrionidae" genus="Quadrideres" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="221" phylum="Arthropoda" rank="species" species="simplicipes">
<emphasis id="B921EAF2D5432F3BFD55C337FC4EF9A1" box="[680,930,1558,1580]" italics="true" pageId="4" pageNumber="221">Quadrideres simplicipes</emphasis>
</taxonomicName>
species-group (
<emphasis id="B921EAF2D5432F3BFBBAC337FB7FF9A1" bold="true" box="[1095,1171,1558,1580]" pageId="4" pageNumber="221">13A, B</emphasis>
). On Fig. 13 the areas of particular ecoregions were distinguished by colors. Their names are provided in black boxes. Fig. 13B presents a close-up of the frame marked on Fig. 13A.
</paragraph>
</caption>
<paragraph id="8BEA36E0D5432F3BFF3AC3B7FE44F922" blockId="4.[151,1436,1686,2035]" box="[199,424,1686,1711]" pageId="4" pageNumber="221">
Habitus as in
<figureCitation id="136E2A65D5432F3BFE9DC3B7FE4EF922" box="[352,418,1686,1711]" captionStart="FIGURES 1 4" captionStartId="2.[151,264,1976,1999]" captionTargetBox="[155,1433,1351,1949]" captionTargetId="figure@2.[150,1436,1347,1952]" captionTargetPageId="2" captionText="FIGURES 1 4. Habitus of the representatives of the Quadrideres s implicipes species-group: Q. kazimierzi sp. nov. (1), Q. simplicipes (2), Q. parallelus (3) and Q. robynsi (4). Scale bar = 1 mm." httpUri="https://zenodo.org/record/243018/files/figure.png" pageId="4" pageNumber="221">Fig. 1</figureCitation>
.
</paragraph>
<paragraph id="8BEA36E0D5432F3BFF3AC39BFBB6F97A" blockId="4.[151,1436,1686,2035]" pageId="4" pageNumber="221">Dorsal side of head dull with coarse punctures (0.51.0 diameters apart). Frontoclypeal suture fine. Clypeal emargination shallow. Mentum with median part narrow. Submentum with short base.</paragraph>
<paragraph id="8BEA36E0D5432F3BFF3AC223FC39F80B" blockId="4.[151,1436,1686,2035]" pageId="4" pageNumber="221">
Pronotal disc dull, with coarse punctures (1.01.5 diameters apart). Anterior pronotal angles rounded. Lateral sides of pronotal disc subparallel (pronotum widest at 0.75 of its length). Pronotal disc with apophyseal and basal depressions (cf.
<bibRefCitation id="EFC44B11D5432F3BFE9DC26AFD96F8EE" author="Ras" box="[352,634,1866,1891]" pageId="4" pageNumber="221" refString="Ras, M. &amp; Kaminski, M. J. (2013) X-ray microtomography as a tool for taxonomic investigations: prothoracic skeletal structure in some Tenebrionidae (Coleoptera). Annales Zoologici, 63 (2), 381 - 391." type="journal article" year="2013">Raś &amp; Kamiński 2013</bibRefCitation>
). Lateral emargination of pronotum complete. Basal emargination interrupted laterally (Fig. 7). Pronotal hypomeron dull, without punctures.
</paragraph>
<paragraph id="8BEA36E0D5432F3BFF3AC2B3FEE6F87E" blockId="4.[151,1436,1686,2035]" pageId="4" pageNumber="221">Elytral rows with coarse punctures (1.0 diameters apart). Intervals dull, with coarse punctures (1.01.5 diameters apart). Elytral base straight. Elytral humeri rounded, slightly protruding laterad (Fig. 7). Scutellum triangular.</paragraph>
<paragraph id="8BEA36E0D5422F3AFF3AC5BBFBE8FF5A" blockId="5.[151,1437,151,470]" pageId="5" pageNumber="222">Intercoxal process of prosternum straight with complete bordering. Metaventrite without tubercle. 5th abdominal ventrite without bordering; with punctures 2.03.0 diameters apart.</paragraph>
<paragraph id="8BEA36E0D5422F3AFF3AC5C2FE1CFF76" blockId="5.[151,1437,151,470]" box="[199,496,226,251]" pageId="5" pageNumber="222">Legs in both sexes simple.</paragraph>
<paragraph id="8BEA36E0D5422F3AFF3AC427FF3AFECE" blockId="5.[151,1437,151,470]" pageId="5" pageNumber="222">Apical part of aedeagal tegmen equal to 0.25 of the rest of tegmen. Parameres narrowing towards apex. Clavae strait.</paragraph>
<paragraph id="8BEA36E0D5422F3AFF3AC473FE29FE03" blockId="5.[151,1437,151,470]" pageId="5" pageNumber="222">Paraproct longer then coxites. First sclerite wide and short; sclerites 3rd and 4th narrow and long. Bursa copulatrix without sclerite.</paragraph>
</subSubSection>
<subSubSection id="C34F656BD5422F3AFF3AC4B8FC60FE5B" pageId="5" pageNumber="222" type="distribution">
<paragraph id="8BEA36E0D5422F3AFF3AC4B8FC60FE5B" blockId="5.[151,1437,151,470]" pageId="5" pageNumber="222">
<emphasis id="B921EAF2D5422F3AFF3AC4B8FE8CFE3F" bold="true" box="[199,352,409,434]" pageId="5" pageNumber="222">Distribution.</emphasis>
This species has been collected in the following ecoregion of the East Africa (
<collectingCountry id="F3427670D5422F3AFAD8C4B8FA7DFE3F" box="[1317,1425,409,434]" name="Tanzania" pageId="5" pageNumber="222">Tanzania</collectingCountry>
): Southern
<emphasis id="B921EAF2D5422F3AFEFEC49FFE02FE58" box="[259,494,445,469]" italics="true" pageId="5" pageNumber="222">
<taxonomicName id="4C554D63D5422F3AFEFEC49FFEBDFE58" box="[259,337,446,469]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="5" pageNumber="222" phylum="Tracheophyta" rank="genus">Acacia</taxonomicName>
-Commiphora
</emphasis>
bushlands and thickets (
<figureCitation id="136E2A65D5422F3AFCFFC49CFC93FE5B" box="[770,895,445,470]" captionStart="FIGURES 12 13" captionStartId="4.[151,264,1527,1550]" captionTargetBox="[173,1413,184,1505]" captionTargetId="figure@4.[167,1420,179,1512]" captionTargetPageId="4" captionText="FIGURES 12 13. Distribution of the Quadrideres s implicipes species-group illustrated using digital elevation model (12) and Afrotropical ecoregions and the distribution of the Quadrideres simplicipes species-group (13 A, B). On Fig. 13 the areas of particular ecoregions were distinguished by colors. Their names are provided in black boxes. Fig. 13 B presents a close-up of the frame marked on Fig. 13 A." httpUri="https://zenodo.org/record/243019/files/figure.png" pageId="5" pageNumber="222">Fig. 12, 13</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>