treatments-xml/data/03/FC/87/03FC87E9FF8B0629FF4CCE2DF992C532.xml

<document id="A26098ED68DFE2F90E11E9BC1150D6AE" ID-CLB-Dataset="27891" ID-DOI="10.11646/zootaxa.4526.2.4" ID-GBIF-Dataset="10aed39d-f494-4164-b41f-5c2e6bcb7be4" ID-ISSN="1175-5326" ID-Zenodo-Dep="2611530" ID-ZooBank="541D8D11-DCC0-4E5C-8B23-81D92D0F1BE9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553697344389" checkinUser="plazi" docAuthor="Sobral, Rafael, Grossi, Paschoal C. &amp; De Morais, José W." docDate="2018" docId="03FC87E9FF8B0629FF4CCE2DF992C532" docLanguage="en" docName="zootaxa.4526.2.4.pdf" docOrigin="Zootaxa 4526 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Aegopsis vazdemelloi Sobral &amp; Grossi &amp; De Morais 2018, new species" docType="treatment" docVersion="8" lastPageNumber="192" masterDocId="FFC5FF91FF840638FFDBC840FB0BC106" masterDocTitle="Two new species of Aegopsis Burmeister, 1847 (Coleoptera: Scarabaeidae: Dynastinae) from the central Brazilian Cerrado" masterLastPageNumber="194" masterPageNumber="175" pageNumber="190" updateTime="1698714249358" updateUser="ExternalLinkService">
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<mods:title id="AD422155CB7D28528E5580A26D90D94E">Two new species of Aegopsis Burmeister, 1847 (Coleoptera: Scarabaeidae: Dynastinae) from the central Brazilian Cerrado</mods:title>
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<mods:namePart id="4C5DB476A10781C53A84F1F77B7B9386">Sobral, Rafael</mods:namePart>
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<mods:namePart id="2253D31F14F832C7E7C690C7B4FDD17A">Grossi, Paschoal C.</mods:namePart>
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<treatment id="03FC87E9FF8B0629FF4CCE2DF992C532" ID-DOI="http://doi.org/10.5281/zenodo.5976895" ID-GBIF-Taxon="154507625" ID-Zenodo-Dep="5976895" LSID="urn:lsid:plazi:treatment:03FC87E9FF8B0629FF4CCE2DF992C532" httpUri="http://treatment.plazi.org/id/03FC87E9FF8B0629FF4CCE2DF992C532" lastPageId="17" lastPageNumber="192" pageId="15" pageNumber="190">
<subSubSection id="C34F6574FF8B0637FF4CCE2DF803C78E" box="[151,776,1645,1672]" pageId="15" pageNumber="190" type="nomenclature">
<paragraph id="8BEA36FFFF8B0637FF4CCE2DF803C78E" blockId="15.[151,776,1645,1705]" box="[151,776,1645,1672]" pageId="15" pageNumber="190">
<heading id="D0A28193FF8B0637FF4CCE2DF803C78E" bold="true" box="[151,776,1645,1672]" fontSize="11" level="1" pageId="15" pageNumber="190" reason="1">
<emphasis id="B921EAEDFF8B0637FF4CCE2DF803C78E" bold="true" box="[151,776,1645,1672]" pageId="15" pageNumber="190">
<taxonomicName id="4C554D7CFF8B0637FF4CCE2DF960C78E" ID-CoL="65974" authority="Sobral &amp; Grossi" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[151,619,1645,1672]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="190" phylum="Arthropoda" rank="species" species="vazdemelloi" status="sp. nov.">
<emphasis id="B921EAEDFF8B0637FF4CCE2DFA9CC781" bold="true" box="[151,407,1645,1671]" italics="true" pageId="15" pageNumber="190">Aegopsis vazdemelloi</emphasis>
Sobral &amp; Grossi
</taxonomicName>
, 
<taxonomicNameLabel id="A2125796FF8B0637FDA3CE2EF803C78E" box="[632,776,1646,1672]" pageId="15" pageNumber="190" rank="species">new species</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C34F6574FF8B0637FF4CCED0FA52C7AF" box="[151,345,1680,1705]" pageId="15" pageNumber="190" type="description">
<paragraph id="8BEA36FFFF8B0637FF4CCED0FA52C7AF" blockId="15.[151,776,1645,1705]" box="[151,345,1680,1705]" pageId="15" pageNumber="190">
(
<figureCitation id="136E2A7AFF8B0637FF44CED0FA0BC7AF" box="[159,256,1680,1705]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStart-6="FIGURE 8" captionStartId-0="4.[151,250,1882,1904]" captionStartId-1="5.[151,250,1876,1898]" captionStartId-2="6.[151,250,1693,1715]" captionStartId-3="7.[151,250,1292,1314]" captionStartId-4="9.[151,250,1775,1797]" captionStartId-5="10.[151,250,1892,1914]" captionStartId-6="11.[151,250,1322,1344]" captionTargetBox-0="[200,1377,193,1853]" captionTargetBox-1="[231,1354,193,1844]" captionTargetBox-2="[151,1415,201,1637]" captionTargetBox-3="[151,1436,193,1270]" captionTargetBox-4="[152,1421,205,1743]" captionTargetBox-5="[185,1389,202,1867]" captionTargetBox-6="[151,1436,193,1283]" captionTargetId-0="figure@4.[200,1377,193,1853]" captionTargetId-1="figure@5.[231,1354,193,1844]" captionTargetId-2="figure@6.[151,1436,193,1672]" captionTargetId-3="figure@7.[151,1436,193,1270]" captionTargetId-4="figure@9.[151,1436,193,1754]" captionTargetId-5="figure@10.[181,1389,193,1871]" captionTargetId-6="figure@11.[151,1436,193,1283]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="11" captionText-0="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." captionText-1="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." captionText-2="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." captionText-3="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." captionText-4="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." captionText-5="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." captionText-6="FIGURE 8. A, Mandible of Aegopsis bolboceridus female in ventral view; B, mandible of Aegopsis diceratops female in ventral view; C, mandible of Aegopsis vazdemelloi female in ventral view; D, mentum of Aegopsis bolboceridus female in ventral view; E, mentum of Aegopsis diceratops female in ventral view; F, mentum of Aegopsis vazdemelloi female in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; arrow indicating middle part of mesal brushes in B; arrow indicating internal lobe in C. Scale bars: 0.5 mm." httpUri-0="https://zenodo.org/record/2611534/files/figure.png" httpUri-1="https://zenodo.org/record/2611536/files/figure.png" httpUri-2="https://zenodo.org/record/2611538/files/figure.png" httpUri-3="https://zenodo.org/record/2611541/files/figure.png" httpUri-4="https://zenodo.org/record/2611544/files/figure.png" httpUri-5="https://zenodo.org/record/2611546/files/figure.png" httpUri-6="https://zenodo.org/record/2611548/files/figure.png" pageId="15" pageNumber="190">Fig. 2–8</figureCitation>
, 
<figureCitation id="136E2A7AFF8B0637FED7CED1FA5AC7AF" box="[268,337,1681,1705]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStartId-0="13.[151,250,1970,1992]" captionStartId-1="14.[151,250,1333,1355]" captionTargetBox-0="[194,1312,193,1920]" captionTargetBox-1="[150,1437,193,1311]" captionTargetId-0="figure@13.[194,1388,193,1949]" captionTargetId-1="figure@14.[150,1437,193,1311]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURE 10. Habitus of Aegopsis vazdemelloi. A, Major male in dorsal view; B, minor male in dorsal view; C, female in dorsal view. Scale bars: 3 mm." captionText-1="FIGURE 11. Distribution map of the species in the Aegopsis bolboceridus species group." httpUri-0="https://zenodo.org/record/2611553/files/figure.png" httpUri-1="https://zenodo.org/record/2611555/files/figure.png" pageId="15" pageNumber="190">10–11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34F6574FF8B0637FF4CCE98FBF0C6CF" pageId="15" pageNumber="190" type="diagnosis">
<paragraph id="8BEA36FFFF8B0637FF4CCE98FBF0C6CF" blockId="15.[151,1437,1752,2029]" pageId="15" pageNumber="190">
<emphasis id="B921EAEDFF8B0637FF4CCE98FA1AC7F7" bold="true" box="[151,273,1752,1777]" pageId="15" pageNumber="190">Diagnosis.</emphasis>
Males: clypeus subrectangular, apex slightly emarginate (
<figureCitation id="136E2A7AFF8B0637FC78CE98F8F0C7F7" box="[931,1019,1752,1777]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="15" pageNumber="190">Fig. 2E</figureCitation>
); internal lobe of mandibles strongly sclerotized, mesal brush not crossing the internal lobe (
<figureCitation id="136E2A7AFF8B0637FCD0CEBCF86FC613" box="[779,868,1788,1813]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="15" pageNumber="190">Fig. 5C</figureCitation>
); mentum subtriangular (
<figureCitation id="136E2A7AFF8B0637FB5FCEBCFFD0C613" box="[1156,1243,1788,1813]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="15" pageNumber="190">Fig. 5F</figureCitation>
); labrum slightly prominent at middle, sides round (
<figureCitation id="136E2A7AFF8B0637FDC0CF60F97DC63F" box="[539,630,1824,1849]" captionStart="FIGURE 4" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[151,1415,201,1637]" captionTargetId="figure@6.[151,1436,193,1672]" captionTargetPageId="6" captionText="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611538/files/figure.png" pageId="15" pageNumber="190">Fig. 4G</figureCitation>
); small punctures on posterior angles of pronotum with micropunctures between them (
<figureCitation id="136E2A7AFF8B0637FE9FCF04FA90C65B" box="[324,411,1860,1885]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="15" pageNumber="190">Fig. 2F</figureCitation>
); parameres, in ventral view, with apex acute, ventral carina basally acute, inferior edge of parameres round (
<figureCitation id="136E2A7AFF8B0637FEBECF28FACAC687" box="[357,449,1896,1921]" captionStart="FIGURE 6" captionStartId="9.[151,250,1775,1797]" captionTargetBox="[152,1421,205,1743]" captionTargetId="figure@9.[151,1436,193,1754]" captionTargetPageId="9" captionText="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." httpUri="https://zenodo.org/record/2611544/files/figure.png" pageId="15" pageNumber="190">Fig. 6H</figureCitation>
). Females: clypeus round, punctures deep and coalescent on vertex (
<figureCitation id="136E2A7AFF8B0637FB0ACF28FE21C687" box="[1233,1322,1896,1921]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="15" pageNumber="190">Fig. 7E</figureCitation>
); distance from canthi carina to canthi apex larger than canthi thickness (
<figureCitation id="136E2A7AFF8B0637FC85CFCCF8BFC6A3" box="[862,948,1932,1957]" captionStart="FIGURE 3" captionStartId="5.[151,250,1876,1898]" captionTargetBox="[231,1354,193,1844]" captionTargetId="figure@5.[231,1354,193,1844]" captionTargetPageId="5" captionText="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611536/files/figure.png" pageId="15" pageNumber="190">Fig. 3F</figureCitation>
); sensorial area of maxillary palpus rugose (
<figureCitation id="136E2A7AFF8B0637FF44CFF0FBE5C6CF" box="[159,238,1968,1993]" captionStart="FIGURE 4" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[151,1415,201,1637]" captionTargetId="figure@6.[151,1436,193,1672]" captionTargetPageId="6" captionText="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611538/files/figure.png" pageId="15" pageNumber="190">
Fig. 
<date id="FFEB103FFF8B0637FF0FCFF1FBE5C6CF" box="[212,238,1969,1993]" pageId="15" pageNumber="190">4I</date>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34F6574FF8B0628FF1CCF94F8E1C509" lastPageId="16" lastPageNumber="191" pageId="15" pageNumber="190" type="materials_examined">
<paragraph id="8BEA36FFFF8B0628FF1CCF94FF0DC2CE" blockId="15.[151,1437,1752,2029]" lastBlockId="16.[151,1437,151,2013]" lastPageId="16" lastPageNumber="191" pageId="15" pageNumber="190">
<emphasis id="B921EAEDFF8B0637FF1CCF94F96EC6EB" bold="true" box="[199,613,2004,2029]" pageId="15" pageNumber="190">
<typeStatus id="54EE885DFF8B0637FF1CCF94FA3FC6EB" box="[199,308,2004,2029]" pageId="15" pageNumber="190" type="holotype">Holotype</typeStatus>
description, male. 
<emphasis id="B921EAEDFF8B0637FDC0CF94F957C6EB" bold="true" box="[539,604,2004,2029]" italics="true" pageId="15" pageNumber="190">Color</emphasis>
:
</emphasis>
Surface lustrous; dorsum, venter, and appendages reddish brown with a weak metallic shine (
<figureCitation id="136E2A7AFF940628FE51C8D7FAFEC1B6" box="[394,501,151,176]" captionStart="FIGURE 10" captionStartId="13.[151,250,1970,1992]" captionTargetBox="[194,1312,193,1920]" captionTargetId="figure@13.[194,1388,193,1949]" captionTargetPageId="13" captionText="FIGURE 10. Habitus of Aegopsis vazdemelloi. A, Major male in dorsal view; B, minor male in dorsal view; C, female in dorsal view. Scale bars: 3 mm." httpUri="https://zenodo.org/record/2611553/files/figure.png" pageId="16" pageNumber="191">Fig. 10A</figureCitation>
). 
<emphasis id="B921EAEDFF940628FDD1C8D7F95AC1B6" bold="true" box="[522,593,151,176]" pageId="16" pageNumber="191">
<emphasis id="B921EAEDFF940628FDD1C8D7F943C1B6" bold="true" box="[522,584,151,176]" italics="true" pageId="16" pageNumber="191">Head</emphasis>
:
</emphasis>
Frons sparsely punctate, punctures small. Vertex almost four times wider than eye width. Cephalic horns moderately punctate; punctures small, mixed with larger punctures dorsally on lateral side. Clypeus densely punctate, punctures larger and marked basally, punctures fine apically; clypeus subrectangular, apex slightly emarginate at middle (
<figureCitation id="136E2A7AFF940628FD3EC944F834C01A" box="[741,831,260,285]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="16" pageNumber="191">Fig. 2E</figureCitation>
). Ocular canthi anteriorly almost straight, posteriorly projecting with a round tip; canthi carina oblique, almost straight, continuous with canthi anteriorly (
<figureCitation id="136E2A7AFF940628FAEFC967FE84C046" box="[1332,1423,295,320]" captionStart="FIGURE 3" captionStartId="5.[151,250,1876,1898]" captionTargetBox="[231,1354,193,1844]" captionTargetId="figure@5.[231,1354,193,1844]" captionTargetPageId="5" captionText="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611536/files/figure.png" pageId="16" pageNumber="191">Fig. 3E</figureCitation>
). 
<emphasis id="B921EAEDFF940628FF4CC90BFA2CC063" bold="true" box="[151,295,331,357]" pageId="16" pageNumber="191">
<emphasis id="B921EAEDFF940628FF4CC90BFA15C062" bold="true" box="[151,286,331,356]" italics="true" pageId="16" pageNumber="191">Mouthparts</emphasis>
:
</emphasis>
Labrum setose, almost semicircular, slightly prominent at middle, sides round (
<figureCitation id="136E2A7AFF940628FB6EC90CFE1AC062" box="[1205,1297,332,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[151,1415,201,1637]" captionTargetId="figure@6.[151,1436,193,1672]" captionTargetPageId="6" captionText="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611538/files/figure.png" pageId="16" pageNumber="191">Fig. 4G</figureCitation>
). Mandibles subtriangular with two prominent teeth apically and two ventral carina; apical protuberance absent; in ventral view, internal lobe strongly sclerotized, mesal brush not crossing internal lobe (
<figureCitation id="136E2A7AFF940628FBEEC9D4FF9CC0AB" box="[1077,1175,404,429]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="16" pageNumber="191">Fig. 5C</figureCitation>
). Maxilla with galea subtriangular, strongly oblique, teeth absent, anterior margin of external edge concave; maxillary palpomere IV 3.6 times longer than wide, sensorial area sparsely striate (
<figureCitation id="136E2A7AFF940628FCCBC99CF865C0F2" box="[784,878,476,501]" captionStart="FIGURE 4" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[151,1415,201,1637]" captionTargetId="figure@6.[151,1436,193,1672]" captionTargetPageId="6" captionText="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611538/files/figure.png" pageId="16" pageNumber="191">Fig. 4H</figureCitation>
). Mentum subtriangular, carina absent (
<figureCitation id="136E2A7AFF940628FAECC99CFE9BC0F3" box="[1335,1424,476,501]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="16" pageNumber="191">Fig. 5F</figureCitation>
). 
<emphasis id="B921EAEDFF940628FF4CC9BFFA1DC31E" bold="true" box="[151,278,511,536]" pageId="16" pageNumber="191">
<emphasis id="B921EAEDFF940628FF4CC9BFFA06C31E" bold="true" box="[151,269,511,536]" italics="true" pageId="16" pageNumber="191">Prothorax</emphasis>
:
</emphasis>
Pronotum bordered, disc weakly gibbous and distinctly punctate. Thoracic horn strong, apex slightly downturned. Punctures sparse medially, becoming denser laterally; small punctures on posterior angles of pronotum with micropunctures between them (
<figureCitation id="136E2A7AFF940628FD67CA07F81DC366" box="[700,790,583,608]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="16" pageNumber="191">Fig. 2F</figureCitation>
). Anterior angles and lateral edges of horn completely wrinkled. 
<taxonomicName id="4C554D7CFF940628FED3CA2BF92CC383" authority=": Subtriangular" authorityName="Subtriangular" box="[264,551,619,645]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="16" pageNumber="191" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAEDFF940628FED3CA2BFA8EC383" bold="true" box="[264,389,619,645]" pageId="16" pageNumber="191">
<emphasis id="B921EAEDFF940628FED3CA2BFA77C382" bold="true" box="[264,380,619,644]" italics="true" pageId="16" pageNumber="191">Scutellum</emphasis>
:
</emphasis>
Subtriangular
</taxonomicName>
, apex distinctly acute; punctures sparse and weak anteriorly, absent posteriorly, no setae. 
<emphasis id="B921EAEDFF940628FEDECACFFA5DC3AE" bold="true" box="[261,342,655,680]" pageId="16" pageNumber="191">
<emphasis id="B921EAEDFF940628FEDECACFFA47C3AE" bold="true" box="[261,332,655,680]" italics="true" pageId="16" pageNumber="191">Elytra</emphasis>
:
</emphasis>
Form 1.7 times longer than wide. Surface glabrous, irregularly covered by ocellate punctures, microsculpture slightly wrinkled. Elytral punctures generally irregular, with distinct sutural striae; striae not impressed, punctures large. Humeral umbone strong, apical umbone distinct crossing elytral external edges in dorsal view. 
<emphasis id="B921EAEDFF940628FEFFCABCFA6FC213" bold="true" box="[292,356,764,789]" pageId="16" pageNumber="191">
<emphasis id="B921EAEDFF940628FEFFCABCFA51C215" bold="true" box="[292,346,764,787]" italics="true" pageId="16" pageNumber="191">Legs</emphasis>
:
</emphasis>
Protibial surface with punctures coalescent dorsally; protibia with three well-developed external teeth, posteriorly decreasing in size; punctures weak and small on external edge of teeth. 
<emphasis id="B921EAEDFF940628FBAFCB5FFFE7C23E" bold="true" box="[1140,1260,799,824]" pageId="16" pageNumber="191">
<taxonomicName id="4C554D7CFF940628FBAFCB5FFFE9C23E" box="[1140,1250,799,824]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="191" phylum="Chordata" rank="genus">
<emphasis id="B921EAEDFF940628FBAFCB5FFFE9C23E" bold="true" box="[1140,1250,799,824]" italics="true" pageId="16" pageNumber="191">Pygidium</emphasis>
</taxonomicName>
:
</emphasis>
Form 2.5 times wider than longer, strongly convex. Disc glabrous on middle and posterior edges, hirsute laterally. Disc completely wrinkled with sparse punctures medially. 
<emphasis id="B921EAEDFF940628FDB2CB27F9D6C286" bold="true" box="[617,733,871,896]" pageId="16" pageNumber="191">
<emphasis id="B921EAEDFF940628FDB2CB27F9DFC286" bold="true" box="[617,724,871,896]" italics="true" pageId="16" pageNumber="191">Genitalia</emphasis>
:
</emphasis>
Parameres, in dorsal view, as in 
<figureCitation id="136E2A7AFF940628FB97CB27FF91C286" box="[1100,1178,871,896]" captionStart="FIGURE 6" captionStartId="9.[151,250,1775,1797]" captionTargetBox="[152,1421,205,1743]" captionTargetId="figure@9.[151,1436,193,1754]" captionTargetPageId="9" captionText="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." httpUri="https://zenodo.org/record/2611544/files/figure.png" pageId="16" pageNumber="191">
Fig. 
<date id="FFEB103FFF940628FBA4CB27FF91C286" box="[1151,1178,871,896]" pageId="16" pageNumber="191">6I</date>
</figureCitation>
; in lateral view, almost with same thickness as apex of phallobase, lateral carina distinctly curved at base (
<figureCitation id="136E2A7AFF940628FB9BCBCCFF90C2A3" box="[1088,1179,908,933]" captionStart="FIGURE 6" captionStartId="9.[151,250,1775,1797]" captionTargetBox="[152,1421,205,1743]" captionTargetId="figure@9.[151,1436,193,1754]" captionTargetPageId="9" captionText="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." httpUri="https://zenodo.org/record/2611544/files/figure.png" pageId="16" pageNumber="191">Fig. 6G</figureCitation>
); in ventral view, apex acute, ventral carina basally acute, inferior edge of parameres round (
<figureCitation id="136E2A7AFF940628FC44CBEFF8F2C2CE" box="[927,1017,943,968]" captionStart="FIGURE 6" captionStartId="9.[151,250,1775,1797]" captionTargetBox="[152,1421,205,1743]" captionTargetId="figure@9.[151,1436,193,1754]" captionTargetPageId="9" captionText="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." httpUri="https://zenodo.org/record/2611544/files/figure.png" pageId="16" pageNumber="191">Fig. 6H</figureCitation>
).
</paragraph>
<paragraph id="8BEA36FFFF940628FF1CCB94F8E1C509" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="191">
<emphasis id="B921EAEDFF940628FF1CCB94F90AC2EB" bold="true" box="[199,513,980,1005]" pageId="16" pageNumber="191">
Measurement of 
<typeStatus id="54EE885DFF940628FE48CB94FAF7C2EB" box="[403,508,980,1005]" pageId="16" pageNumber="191" type="holotype">holotype</typeStatus>
.
</emphasis>
Body length: 
<quantity id="4CAD9B1AFF940628FD7FCB94F807C2EA" box="[676,780,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="3.21" pageId="16" pageNumber="191" unit="mm" value="32.1">32.1 mm</quantity>
. Cephalic horns length: 
<quantity id="4CAD9B1AFF940628FBFBCB94FF71C2EA" box="[1056,1146,980,1004]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="16" pageNumber="191" unit="mm" value="7.3">7.3 mm</quantity>
. Elytral length: 
<quantity id="4CAD9B1AFF940628FAEBCB94FE92C2EA" box="[1328,1433,980,1005]" metricMagnitude="-2" metricUnit="m" metricValue="1.76" pageId="16" pageNumber="191" unit="mm" value="17.6">17.6 mm</quantity>
. Elytral width: 10.0 mm. Pronotal width: 
<quantity id="4CAD9B1AFF940628FD84CBB8F9CDC509" box="[607,710,1016,1040]" metricMagnitude="-2" metricUnit="m" metricValue="1.42" pageId="16" pageNumber="191" unit="mm" value="14.2">14.2 mm</quantity>
. Protibial length: 
<quantity id="4CAD9B1AFF940628FC57CBB8F8EDC509" box="[908,998,1016,1040]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="16" pageNumber="191" unit="mm" value="8.9">8.9 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F6574FF940628FF1CCC5CF842C64A" pageId="16" pageNumber="191" type="description">
<paragraph id="8BEA36FFFF940628FF1CCC5CF8ACC5EE" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="191">
<emphasis id="B921EAEDFF940628FF1CCC5CF900C533" bold="true" box="[199,523,1052,1077]" pageId="16" pageNumber="191">
<typeStatus id="54EE885DFF940628FF1CCC5CFA34C533" box="[199,319,1052,1077]" pageId="16" pageNumber="191" type="paratype">Paratypes</typeStatus>
variation, males.
</emphasis>
Major males with cephalic horns 2–4 times longer than vertex; body length 
<quantity id="4CAD9B1AFF940628FA86CC5CFBF1C551" metricMagnitude="-2" metricUnit="m" metricValue="2.93" metricValueMax="3.42" metricValueMin="2.44" pageId="16" pageNumber="191" unit="mm" value="29.3" valueMax="34.2" valueMin="24.4">24.4– 34.2 mm</quantity>
(
<emphasis id="B921EAEDFF940628FED1CC01FA13C55E" box="[266,280,1089,1112]" italics="true" pageId="16" pageNumber="191">n</emphasis>
= 14) and pronotal width 
<quantity id="4CAD9B1AFF940628FDE0CC7FF9D5C551" box="[571,734,1087,1112]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" metricValueMax="1.45" metricValueMin="1.21" pageId="16" pageNumber="191" unit="mm" value="13.3" valueMax="14.5" valueMin="12.1">12.1–14.5 mm</quantity>
(
<emphasis id="B921EAEDFF940628FD36CC01F9F0C55E" box="[749,763,1089,1112]" italics="true" pageId="16" pageNumber="191">n</emphasis>
= 14), sometimes ocular canthi anteriorly rounder but never protruding. Thoracic horn 0.5 or one time longer than vertex. Minor males (
<figureCitation id="136E2A7AFF940628FC2ACC24FF5EC57A" box="[1009,1109,1124,1149]" captionStart="FIGURE 10" captionStartId="13.[151,250,1970,1992]" captionTargetBox="[194,1312,193,1920]" captionTargetId="figure@13.[194,1388,193,1949]" captionTargetPageId="13" captionText="FIGURE 10. Habitus of Aegopsis vazdemelloi. A, Major male in dorsal view; B, minor male in dorsal view; C, female in dorsal view. Scale bars: 3 mm." httpUri="https://zenodo.org/record/2611553/files/figure.png" pageId="16" pageNumber="191">Fig. 10B</figureCitation>
) with cephalic horns smaller than vertex or with same length; body length 16.2–23.0 mm (
<emphasis id="B921EAEDFF940628FCB3CCC9F87DC5A6" box="[872,886,1161,1184]" italics="true" pageId="16" pageNumber="191">n</emphasis>
= 8) and pronotal width 9.8–11.0 mm (
<emphasis id="B921EAEDFF940628FA90CCC9FE52C5A6" box="[1355,1369,1161,1184]" italics="true" pageId="16" pageNumber="191">n</emphasis>
= 8), thoracic horn short, like a tubercle, not crossing anterior margin of pronotum. Ocular canthi shorter, posteriorly more distinct; punctures of pronotum larger and denser than 
<typeStatus id="54EE885DFF940628FCE6CC8FF8A8C5EE" box="[829,931,1231,1256]" pageId="16" pageNumber="191" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BEA36FFFF940628FF1CCCB4FE7FC436" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="191">
<emphasis id="B921EAEDFF940628FF1CCCB4F957C40B" bold="true" box="[199,604,1268,1293]" pageId="16" pageNumber="191">
Measurements of male 
<typeStatus id="54EE885DFF940628FE04CCB4F95CC40B" box="[479,599,1268,1293]" pageId="16" pageNumber="191" type="paratype">paratypes</typeStatus>
.
</emphasis>
Body length: 
<quantity id="4CAD9B1AFF940628FD24CCB4F8ADC40A" box="[767,934,1268,1293]" metricMagnitude="-2" metricUnit="m" metricValue="2.5200000000000005" metricValueMax="3.4200000000000004" metricValueMin="1.6199999999999999" pageId="16" pageNumber="191" unit="mm" value="25.2" valueMax="34.2" valueMin="16.2">16.2–34.2 mm</quantity>
. Cephalic horns length: 
<quantity id="4CAD9B1AFF940628FB62CCB4FE4FC40A" box="[1209,1348,1268,1293]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="6.7" metricValueMin="1.2" pageId="16" pageNumber="191" unit="mm" value="3.95" valueMax="6.7" valueMin="1.2">1.2–6.7 mm</quantity>
. Elytral length: 
<quantity id="4CAD9B1AFF940628FF37CD57FA99C436" box="[236,402,1303,1328]" metricMagnitude="-2" metricUnit="m" metricValue="1.465" metricValueMax="1.66" metricValueMin="1.27" pageId="16" pageNumber="191" unit="mm" value="14.65" valueMax="16.6" valueMin="12.7">12.7–16.6 mm</quantity>
. Elytral width: 
<quantity id="4CAD9B1AFF940628FDE5CD58F9C3C436" box="[574,712,1304,1328]" metricMagnitude="-3" metricUnit="m" metricValue="8.65" metricValueMax="9.4" metricValueMin="7.9" pageId="16" pageNumber="191" unit="mm" value="8.65" valueMax="9.4" valueMin="7.9">7.9–9.4 mm</quantity>
. Pronotal width: 
<quantity id="4CAD9B1AFF940628FC5CCD57FF14C436" box="[903,1055,1303,1328]" metricMagnitude="-2" metricUnit="m" metricValue="1.215" metricValueMax="1.45" metricValueMin="0.98" pageId="16" pageNumber="191" unit="mm" value="12.15" valueMax="14.5" valueMin="9.8">9.8–14.5 mm</quantity>
. Protibial length: 
<quantity id="4CAD9B1AFF940628FB3DCD57FE7BC436" box="[1254,1392,1303,1328]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="8.1" metricValueMin="6.4" pageId="16" pageNumber="191" unit="mm" value="7.25" valueMax="8.1" valueMin="6.4">6.4–8.1 mm</quantity>
.
</paragraph>
<paragraph id="8BEA36FFFF940628FF1CCD7CFBD8C7E6" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="191">
<emphasis id="B921EAEDFF940628FF1CCD7CF938C453" bold="true" box="[199,563,1340,1365]" pageId="16" pageNumber="191">
<typeStatus id="54EE885DFF940628FF1CCD7CFA3FC453" box="[199,308,1340,1365]" pageId="16" pageNumber="191" type="paratype">Paratype</typeStatus>
description, females.
</emphasis>
Females as males in general aspect, except the following (
<figureCitation id="136E2A7AFF940628FB0CCD7CFE34C452" box="[1239,1343,1340,1365]" captionStart="FIGURE 10" captionStartId="13.[151,250,1970,1992]" captionTargetBox="[194,1312,193,1920]" captionTargetId="figure@13.[194,1388,193,1949]" captionTargetPageId="13" captionText="FIGURE 10. Habitus of Aegopsis vazdemelloi. A, Major male in dorsal view; B, minor male in dorsal view; C, female in dorsal view. Scale bars: 3 mm." httpUri="https://zenodo.org/record/2611553/files/figure.png" pageId="16" pageNumber="191">Fig. 10C</figureCitation>
). 
<emphasis id="B921EAEDFF940628FA8FCD7BFE97C453" bold="true" box="[1364,1436,1339,1365]" pageId="16" pageNumber="191">
<emphasis id="B921EAEDFF940628FA8FCD7BFE99C452" bold="true" box="[1364,1426,1339,1364]" italics="true" pageId="16" pageNumber="191">Head</emphasis>
:
</emphasis>
Cephalic horns absent; frons with two inconspicuous tubercles (
<figureCitation id="136E2A7AFF940628FCA5CD1FF8DDC47E" box="[894,982,1375,1400]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="16" pageNumber="191">Fig. 7F</figureCitation>
); clypeus round, slightly concave from frontoclypeal suture to apex; punctures larger and denser especially on clypeus, wrinkles in front of tubercles (
<figureCitation id="136E2A7AFF940628FAB4CDC4FBB2C4C6" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="16" pageNumber="191">Fig. 7E</figureCitation>
). Ocular canthi convex anteriorly, concave posteriorly with tip thick and round; distance from canthi carina to canthi apex longer than canthi thickness (
<figureCitation id="136E2A7AFF940628FDB7CD8CF9C8C4E2" box="[620,707,1484,1509]" captionStart="FIGURE 3" captionStartId="5.[151,250,1876,1898]" captionTargetBox="[231,1354,193,1844]" captionTargetId="figure@5.[231,1354,193,1844]" captionTargetPageId="5" captionText="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611536/files/figure.png" pageId="16" pageNumber="191">Fig. 3F</figureCitation>
). 
<emphasis id="B921EAEDFF940628FD0CCD8BF86CC4E3" bold="true" box="[727,871,1483,1509]" pageId="16" pageNumber="191">
<emphasis id="B921EAEDFF940628FD0CCD8BF855C4E2" bold="true" box="[727,862,1483,1508]" italics="true" pageId="16" pageNumber="191">Mouthparts</emphasis>
:
</emphasis>
Labrum thinner, sides acute. Mandibles and teeth shorter, mesal brush reaching internal lobe but less hirsute than apex, internal lobe less sclerotized (
<figureCitation id="136E2A7AFF940628FAE9CDAFFE84C70E" box="[1330,1423,1519,1544]" captionStart="FIGURE 8" captionStartId="11.[151,250,1322,1344]" captionTargetBox="[151,1436,193,1283]" captionTargetId="figure@11.[151,1436,193,1283]" captionTargetPageId="11" captionText="FIGURE 8. A, Mandible of Aegopsis bolboceridus female in ventral view; B, mandible of Aegopsis diceratops female in ventral view; C, mandible of Aegopsis vazdemelloi female in ventral view; D, mentum of Aegopsis bolboceridus female in ventral view; E, mentum of Aegopsis diceratops female in ventral view; F, mentum of Aegopsis vazdemelloi female in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; arrow indicating middle part of mesal brushes in B; arrow indicating internal lobe in C. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611548/files/figure.png" pageId="16" pageNumber="191">Fig. 8C</figureCitation>
). Maxilla shorter, galea apically acute, maxillary palpomere IV 2.8 times longer than wide, sensorial area shorter, rugose (
<figureCitation id="136E2A7AFF940628FF2BCE77FA35C756" box="[240,318,1591,1616]" captionStart="FIGURE 4" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[151,1415,201,1637]" captionTargetId="figure@6.[151,1436,193,1672]" captionTargetPageId="6" captionText="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611538/files/figure.png" pageId="16" pageNumber="191">
Fig. 
<date id="FFEB103FFF940628FEFFCE78FA35C756" box="[292,318,1592,1616]" pageId="16" pageNumber="191">4I</date>
</figureCitation>
). Mentum shorter and less triangular, apex longer (
<figureCitation id="136E2A7AFF940628FCADCE77F8C1C756" box="[886,970,1591,1616]" captionStart="FIGURE 8" captionStartId="11.[151,250,1322,1344]" captionTargetBox="[151,1436,193,1283]" captionTargetId="figure@11.[151,1436,193,1283]" captionTargetPageId="11" captionText="FIGURE 8. A, Mandible of Aegopsis bolboceridus female in ventral view; B, mandible of Aegopsis diceratops female in ventral view; C, mandible of Aegopsis vazdemelloi female in ventral view; D, mentum of Aegopsis bolboceridus female in ventral view; E, mentum of Aegopsis diceratops female in ventral view; F, mentum of Aegopsis vazdemelloi female in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; arrow indicating middle part of mesal brushes in B; arrow indicating internal lobe in C. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611548/files/figure.png" pageId="16" pageNumber="191">Fig. 8F</figureCitation>
). 
<emphasis id="B921EAEDFF940628FC06CE78FF50C756" bold="true" box="[989,1115,1591,1616]" pageId="16" pageNumber="191">
<emphasis id="B921EAEDFF940628FC06CE78FF59C749" bold="true" box="[989,1106,1592,1615]" italics="true" pageId="16" pageNumber="191">Pronotum</emphasis>
:
</emphasis>
Weakly convex; prothoracic horn absent; punctures larger and denser, sparser on sides, wrinkles absent. 
<taxonomicName id="4C554D7CFF940628FBC4CE1BFE13C772" authority=": Punctures" authorityName="Punctures" box="[1055,1304,1627,1653]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Corynexochida" pageId="16" pageNumber="191" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAEDFF940628FBC4CE1BFF96C773" bold="true" box="[1055,1181,1627,1653]" pageId="16" pageNumber="191">
<emphasis id="B921EAEDFF940628FBC4CE1BFF9FC772" bold="true" box="[1055,1172,1627,1652]" italics="true" pageId="16" pageNumber="191">Scutellum</emphasis>
:
</emphasis>
Punctures
</taxonomicName>
sparser on middle, larger and denser near base. 
<taxonomicName id="4C554D7CFF940628FDEFCE3FF806C79E" authority=": Convex" authorityName="Convex" box="[564,781,1663,1688]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="16" pageNumber="191" phylum="Chordata" rank="genus">
<emphasis id="B921EAEDFF940628FDEFCE3FF9A0C79E" bold="true" box="[564,683,1663,1688]" pageId="16" pageNumber="191">
<emphasis id="B921EAEDFF940628FDEFCE3FF9A9C79E" bold="true" box="[564,674,1663,1688]" italics="true" pageId="16" pageNumber="191">Pygidium</emphasis>
:
</emphasis>
Convex
</taxonomicName>
, apex sharp. Surface weakly setose laterally and medially. Coalescent punctures denser laterally, round large and dense punctures on middle, wrinkles on sides and posterior edge.
</paragraph>
<paragraph id="8BEA36FFFF940628FF1CCEACFE3AC621" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="191">
<emphasis id="B921EAEDFF940628FF1CCEACF976C603" bold="true" box="[199,637,1772,1797]" pageId="16" pageNumber="191">
Measurements of female 
<typeStatus id="54EE885DFF940628FDDBCEACF973C603" box="[512,632,1772,1797]" pageId="16" pageNumber="191" type="paratype">paratypes</typeStatus>
.
</emphasis>
Body length: 
<quantity id="4CAD9B1AFF940628FCF0CEACF8DDC602" box="[811,982,1772,1796]" metricMagnitude="-2" metricUnit="m" metricValue="2.12" metricValueMax="2.41" metricValueMin="1.83" pageId="16" pageNumber="191" unit="mm" value="21.2" valueMax="24.1" valueMin="18.3">18.3–24.1 mm</quantity>
. Elytral length: 
<quantity id="4CAD9B1AFF940628FB4CCEACFE4BC602" box="[1175,1344,1772,1797]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.61" metricValueMin="1.19" pageId="16" pageNumber="191" unit="mm" value="14.0" valueMax="16.1" valueMin="11.9">11.9–16.1 mm</quantity>
. Elytral width: 
<quantity id="4CAD9B1AFF940628FF3ECF4FFA64C621" box="[229,367,1807,1832]" metricMagnitude="-3" metricUnit="m" metricValue="8.05" metricValueMax="9.3" metricValueMin="6.8" pageId="16" pageNumber="191" unit="mm" value="8.05" valueMax="9.3" valueMin="6.8">6.8–9.3 mm</quantity>
. Head length: 
<quantity id="4CAD9B1AFF940628FDCACF4FF990C621" box="[529,667,1807,1832]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" metricValueMax="3.2" metricValueMin="2.6" pageId="16" pageNumber="191" unit="mm" value="2.9" valueMax="3.2" valueMin="2.6">2.6–3.2 mm</quantity>
. Pronotal width: 
<quantity id="4CAD9B1AFF940628FC81CF50F8D7C621" box="[858,988,1808,1832]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" metricValueMax="1.1199999999999999" metricValueMin="0.9" pageId="16" pageNumber="191" unit="mm" value="10.1" valueMax="11.2" valueMin="9.0">9–11.2 mm</quantity>
. Protibial length: 
<quantity id="4CAD9B1AFF940628FB78CF4FFE26C621" box="[1187,1325,1807,1832]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" metricValueMax="6.7" metricValueMin="4.6" pageId="16" pageNumber="191" unit="mm" value="5.65" valueMax="6.7" valueMin="4.6">4.6–6.7 mm</quantity>
.
</paragraph>
<paragraph id="8BEA36FFFF940628FF1CCF74F842C64A" blockId="16.[151,1437,151,2013]" box="[199,841,1844,1869]" pageId="16" pageNumber="191">
<emphasis id="B921EAEDFF940628FF1CCF74FAE4C64B" bold="true" box="[199,495,1844,1869]" pageId="16" pageNumber="191">Geographic distribution.</emphasis>
<collectingCountry id="F342766FFF940628FE2ECF74F935C64B" box="[501,574,1844,1869]" name="Brazil" pageId="16" pageNumber="191">Brazil</collectingCountry>
: 
<collectingRegion id="4991F81DFF940628FD92CF75F9D1C64A" box="[585,730,1844,1868]" country="Brazil" name="Mato Grosso" pageId="16" pageNumber="191">Mato Grosso</collectingRegion>
(
<figureCitation id="136E2A7AFF940628FD32CF74F830C64A" box="[745,827,1844,1869]" captionStart="FIGURE 11" captionStartId="14.[151,250,1333,1355]" captionTargetBox="[150,1437,193,1311]" captionTargetId="figure@14.[150,1437,193,1311]" captionTargetPageId="14" captionText="FIGURE 11. Distribution map of the species in the Aegopsis bolboceridus species group." httpUri="https://zenodo.org/record/2611555/files/figure.png" pageId="16" pageNumber="191">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34F6574FF940629FF1CCF17FFBBC01B" lastPageId="17" lastPageNumber="192" pageId="16" pageNumber="191" type="materials_examined">
<paragraph id="8BEA36FFFF940629FF1CCF17FFBBC01B" blockId="16.[151,1437,151,2013]" lastBlockId="17.[151,1437,151,1077]" lastPageId="17" lastPageNumber="192" pageId="16" pageNumber="191">
<emphasis id="B921EAEDFF940628FF1CCF17F92FC676" bold="true" box="[199,548,1879,1904]" pageId="16" pageNumber="191">
Material examined. 
<materialsCitation id="3B3D3CA2FF940628FE6CCF17F92FC676" ID-GBIF-Occurrence="2029162621" box="[439,548,1879,1904]" collectingDate="2010-10" country="Brazil" determinerName="R. Sobral &amp; P. C. Grossi" location="Mato Grosso" pageId="16" pageNumber="191" specimenCount="2" stateProvince="Mato Grosso" typeStatus="holotype">
<typeStatus id="54EE885DFF940628FE6CCF17F92FC676" box="[439,548,1879,1904]" pageId="16" pageNumber="191" type="holotype">Holotype</typeStatus>
</materialsCitation>
</emphasis>
male dissected, labeled: a) “ 
<collectingCountry id="F342766FFF940628FCB8CF18F8C6C676" box="[867,973,1880,1904]" name="Brazil" pageId="16" pageNumber="191">BRASIL</collectingCountry>
: 
<collectingRegion id="4991F81DFF940628FC0CCF18FF67C676" box="[983,1132,1880,1904]" country="Brazil" name="Mato Grosso" pageId="16" pageNumber="191">Mato Grosso</collectingRegion>
. / Chapada dos Guimarães. / Centro. 
<date id="FFEB103FFF940628FED8CF3CFA52C692" box="[259,345,1916,1940]" pageId="16" pageNumber="191" value="2010-10">
<collectingDate id="EFAFE9D7FF940628FED8CF3CFA52C692" box="[259,345,1916,1940]" pageId="16" pageNumber="191" value="2010-10">X-2010</collectingDate>
</date>
FZ Vaz-de- / Mello”; b) “ 
<taxonomicName id="4C554D7CFF940628FD5CCF3CF871C692" authority="Sobral &amp; Grossi &amp; De Morais, 2018" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[647,890,1916,1940]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="191" phylum="Arthropoda" rank="species" species="vazdemelloi" status="sp. nov.">
<emphasis id="B921EAEDFF940628FD5CCF3CF871C692" box="[647,890,1916,1940]" italics="true" pageId="16" pageNumber="191">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
<emphasis id="B921EAEDFF940628FC5FCF3CF8CFC693" bold="true" box="[900,964,1916,1941]" pageId="16" pageNumber="191">
<taxonomicNameLabel id="A2125796FF940628FC5FCF3CF8CFC693" box="[900,964,1916,1941]" pageId="16" pageNumber="191" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
/ male 
<typeStatus id="54EE885DFF940628FBC6CF3CFFC8C692" box="[1053,1219,1916,1940]" pageId="16" pageNumber="191" type="holotype">HOLOTYPUS</typeStatus>
/ 
<determinerName id="7E36DA3BFF940628FB07CF3DFE4CC693" box="[1244,1351,1916,1941]" pageId="16" pageNumber="191">R. Sobral</determinerName>
&amp; 
<determinerName id="7E36DA3BFF940628FAB4CF3CFBEBC6BF" pageId="16" pageNumber="191">P.C. Grossi</determinerName>
det. 2016”. 
<typeStatus id="54EE885DFF940628FEB0CFE0FAD8C6BF" box="[363,467,1952,1977]" pageId="16" pageNumber="191" type="holotype">Holotype</typeStatus>
at CERPE. 
<materialsCitation id="3B3D3CA2FF940628FD87CFE0FE9CC6BF" ID-GBIF-Occurrence="2029163007" box="[604,1431,1952,1977]" collectingDate="2010-10" collectionCode="CERPE" country="Brazil" determinerName="R. Sobral &amp; P. C. Grossi" location="Mato Grosso" pageId="16" pageNumber="191" specimenCount="13" specimenCount-female="3" specimenCount-male="10" stateProvince="Mato Grosso" typeStatus="paratype">
<emphasis id="B921EAEDFF940628FD87CFE0F9D8C6BF" bold="true" box="[604,723,1952,1977]" pageId="16" pageNumber="191">
<typeStatus id="54EE885DFF940628FD87CFE0F9D8C6BF" box="[604,723,1952,1977]" pageId="16" pageNumber="191" type="paratype">Paratypes</typeStatus>
</emphasis>
: 
<specimenCount id="9D53FD76FF940628FD39CFE0F84CC6BF" box="[738,839,1952,1977]" pageId="16" pageNumber="191" type="male">10 males</specimenCount>
and 
<specimenCount id="9D53FD76FF940628FCA5CFE0F8E0C6BF" box="[894,1003,1952,1977]" pageId="16" pageNumber="191" type="female">3 females</specimenCount>
(
<collectionCode id="ED44AE3AFF940628FC21CFE0FF5EC6BE" box="[1018,1109,1952,1976]" pageId="16" pageNumber="191">CERPE</collectionCode>
) with same data as holotype
</materialsCitation>
; 
<materialsCitation id="3B3D3CA2FF940628FF4CCF84F830C6DB" ID-GBIF-Occurrence="2029163006" box="[151,827,1988,2013]" collectingDate="2010-10" collectionCode="INPA" country="Brazil" determinerName="R. Sobral &amp; P. C. Grossi" location="Mato Grosso" pageId="16" pageNumber="191" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Mato Grosso" typeStatus="paratype">
<specimenCount id="9D53FD76FF940628FF4CCF84FBE4C6DB" box="[151,239,1988,2013]" pageId="16" pageNumber="191" type="male">3 males</specimenCount>
and 
<specimenCount id="9D53FD76FF940628FEF0CF84FA91C6DB" box="[299,410,1988,2013]" pageId="16" pageNumber="191" type="female">3 females</specimenCount>
(
<collectionCode id="ED44AE3AFF940628FE70CF84FAE5C6DA" box="[427,494,1988,2012]" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="16" pageNumber="191">INPA</collectionCode>
) with same data as holotype
</materialsCitation>
; 
<materialsCitation id="3B3D3CA2FF940629FC92CF84FBF6C1B6" ID-GBIF-Occurrence="2029161438" collectingDate="2010-10" collectionCode="CEMT" country="Brazil" determinerName="R. Sobral &amp; P. C. Grossi" lastPageId="17" lastPageNumber="192" location="Mato Grosso" pageId="16" pageNumber="191" specimenCount="14" specimenCount-female="2" specimenCount-male="12" stateProvince="Mato Grosso" typeStatus="paratype">
<specimenCount id="9D53FD76FF940628FC92CF84F8BBC6DB" box="[841,944,1988,2013]" pageId="16" pageNumber="191" type="male">12 males</specimenCount>
and 
<specimenCount id="9D53FD76FF940628FC37CF84FF51C6DB" box="[1004,1114,1988,2013]" pageId="16" pageNumber="191" type="female">2 females</specimenCount>
(
<collectionCode id="ED44AE3AFF940628FBB7CF84FFB6C6DA" box="[1132,1213,1988,2012]" pageId="16" pageNumber="191">CEMT</collectionCode>
) with same data as holotype
</materialsCitation>
; 
<materialsCitation id="3B3D3CA2FF950629FED3C8D8F9F4C1B6" ID-GBIF-Occurrence="2029162131" box="[264,767,151,176]" collectingDate="2010-10" collectionCode="EPGC" country="Brazil" determinerName="R. Sobral &amp; P. C. Grossi" location="Mato Grosso" pageId="17" pageNumber="192" specimenCount="3" specimenCount-male="3" stateProvince="Mato Grosso" typeStatus="paratype">
<specimenCount id="9D53FD76FF950629FED3C8D8FA54C1B6" box="[264,351,151,176]" pageId="17" pageNumber="192" type="male">3 males</specimenCount>
(
<collectionCode id="ED44AE3AFF950629FEB4C8D8FAB1C1B6" box="[367,442,152,176]" pageId="17" pageNumber="192">EPGC</collectionCode>
) with same data as holotype
</materialsCitation>
; 
<materialsCitation id="3B3D3CA2FF950629FCD0C8D8F834C1D3" ID-GBIF-Occurrence="2029162807" collectingDate="2008-11-15" collectionCode="EPGC" country="Brazil" location="Guimaraes." municipality="Chapada" pageId="17" pageNumber="192" specimenCount="2" specimenCount-male="2" stateProvince="Mato Grosso" typeStatus="paratype">
<specimenCount id="9D53FD76FF950629FCD0C8D8F869C1B6" box="[779,866,151,176]" pageId="17" pageNumber="192" type="male">2 males</specimenCount>
(
<collectionCode id="ED44AE3AFF950629FCAAC8D8F8B7C1B6" box="[881,956,152,176]" pageId="17" pageNumber="192">EPGC</collectionCode>
) with the following data: “ 
<collectingCountry id="F342766FFF950629FB37C8D8FE5DC1B6" box="[1260,1366,152,176]" name="Brazil" pageId="17" pageNumber="192">BRASIL</collectingCountry>
: 
<collectingRegion id="4991F81DFF950629FABAC8D8FBE1C1D2" country="Brazil" name="Mato Grosso" pageId="17" pageNumber="192">Mato Grosso</collectingRegion>
. / 
<collectingMunicipality id="6B8EAC85FF950629FED9C8FCFA6FC1D3" box="[258,356,188,213]" pageId="17" pageNumber="192">Chapada</collectingMunicipality>
dos 
<location id="8E8A6024FF950629FE42C8FCF911C1D3" LSID="urn:lsid:plazi:treatment:03FC87E9FF8B0629FF4CCE2DF992C532:8E8A6024FF950629FE42C8FCF911C1D3" box="[409,538,188,213]" country="Brazil" municipality="Chapada" name="Guimaraes." pageId="17" pageNumber="192" stateProvince="Mato Grosso">Guimarães.</location>
/ 
<date id="FFEB103FFF950629FDF5C8FCF9A6C1D3" box="[558,685,188,213]" pageId="17" pageNumber="192" value="2008-11-15">
<collectingDate id="EFAFE9D7FF950629FDF5C8FCF9A6C1D3" box="[558,685,188,213]" pageId="17" pageNumber="192" value="2008-11-15">15.XI.2008</collectingDate>
</date>
AAS Pinto”
</materialsCitation>
; 
<materialsCitation id="3B3D3CA2FF950629FC92C8FCF8C9C1FE" ID-GBIF-Occurrence="2029160377" collectingDate="2010-09-25" collectionCode="EPGC" collectorName="A. P. Pinheiro" country="Brazil" location="Cuiaba. Tijucal. Manual." pageId="17" pageNumber="192" specimenCount="1" specimenCount-female="1" stateProvince="Mato Grosso" typeStatus="paratype">
<specimenCount id="9D53FD76FF950629FC92C8FCF8A1C1D3" box="[841,938,188,213]" pageId="17" pageNumber="192" type="female">1 female</specimenCount>
(
<collectionCode id="ED44AE3AFF950629FC62C8FCFF0FC1D2" box="[953,1028,188,212]" pageId="17" pageNumber="192">EPGC</collectionCode>
) with the following data: “ 
<collectingCountry id="F342766FFF950629FAF4C8FCFE92C1D2" box="[1327,1433,188,212]" name="Brazil" pageId="17" pageNumber="192">BRASIL</collectingCountry>
: 
<collectingRegion id="4991F81DFF950629FF4CC8A0FA25C1FE" box="[151,302,224,248]" country="Brazil" name="Mato Grosso" pageId="17" pageNumber="192">Mato Grosso</collectingRegion>
. / 
<location id="8E8A6024FF950629FE97C89FF96CC1FE" LSID="urn:lsid:plazi:treatment:03FC87E9FF8B0629FF4CCE2DF992C532:8E8A6024FF950629FE97C89FF96CC1FE" box="[332,615,223,248]" country="Brazil" name="Cuiaba. Tijucal. Manual." pageId="17" pageNumber="192" stateProvince="Mato Grosso">Cuiabá. Tijucal. Manual.</location>
/ 
<date id="FFEB103FFF950629FD5AC89FF80AC1FE" box="[641,769,223,248]" pageId="17" pageNumber="192" value="2010-09-25">
<collectingDate id="EFAFE9D7FF950629FD5AC89FF80AC1FE" box="[641,769,223,248]" pageId="17" pageNumber="192" value="2010-09-25">25.IX.2010</collectingDate>
</date>
/ 
<collectorName id="26A05329FF950629FCC0C8A0F8B3C1FE" box="[795,952,223,248]" pageId="17" pageNumber="192">A.P. Pinheiro</collectorName>
”
</materialsCitation>
; 
<materialsCitation id="3B3D3CA2FF950629FC0BC89FFFA7C01B" ID-GBIF-Occurrence="2029160976" collectingDate="2006-11-26" collectionCode="EPGC" collectorName="F. Z. Vaz-de-Mello" country="Brazil" location="Guimaraes." municipality="MT. Chapada" pageId="17" pageNumber="192" specimenCount="9" specimenCount-female="3" specimenCount-male="6" typeStatus="paratype">
<specimenCount id="9D53FD76FF950629FC0BC89FFF23C1FE" box="[976,1064,223,248]" pageId="17" pageNumber="192" type="male">6 males</specimenCount>
and 
<specimenCount id="9D53FD76FF950629FBB8C8A0FFD9C1FE" box="[1123,1234,223,248]" pageId="17" pageNumber="192" type="female">3 females</specimenCount>
(
<collectionCode id="ED44AE3AFF950629FB38C8A0FE25C1FE" box="[1251,1326,224,248]" pageId="17" pageNumber="192">EPGC</collectionCode>
) with the following data: “ 
<collectingCountry id="F342766FFF950629FE89C944FAB7C01A" box="[338,444,260,284]" name="Brazil" pageId="17" pageNumber="192">BRASIL</collectingCountry>
: 
<collectingMunicipality id="6B8EAC85FF950629FE1CC945F954C01B" box="[455,607,260,285]" pageId="17" pageNumber="192">MT. Chapada</collectingMunicipality>
/ dos 
<location id="8E8A6024FF950629FD79C944F828C01B" LSID="urn:lsid:plazi:treatment:03FC87E9FF8B0629FF4CCE2DF992C532:8E8A6024FF950629FD79C944F828C01B" box="[674,803,260,285]" country="Brazil" municipality="MT. Chapada" name="Guimaraes." pageId="17" pageNumber="192">Guimarães.</location>
/ 
<date id="FFEB103FFF950629FCE3C944F8B3C01B" box="[824,952,260,285]" pageId="17" pageNumber="192" value="2006-11-26">
<collectingDate id="EFAFE9D7FF950629FCE3C944F8B3C01B" box="[824,952,260,285]" pageId="17" pageNumber="192" value="2006-11-26">26.XI.2006</collectingDate>
</date>
/ 
<collectorName id="26A05329FF950629FC15C945FFAAC01B" box="[974,1185,260,285]" pageId="17" pageNumber="192">F.Z. Vaz-de-Mello</collectorName>
”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F6574FF950629FF1CC967F935C063" pageId="17" pageNumber="192" type="etymology">
<paragraph id="8BEA36FFFF950629FF1CC967F935C063" blockId="17.[151,1437,151,1077]" pageId="17" pageNumber="192">
<emphasis id="B921EAEDFF950629FF1CC967FA45C046" bold="true" box="[199,334,295,320]" pageId="17" pageNumber="192">Etymology.</emphasis>
This name is in honor of Dr. Fernando Zagury Vaz-de-Mello (UFMT), a scarab beetle specialist who collected most of the 
<typeStatus id="54EE885DFF950629FE64C90DFAE4C063" box="[447,495,333,357]" pageId="17" pageNumber="192">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="C34F6574FF950629FF1CC92FF992C532" pageId="17" pageNumber="192" type="discussion">
<paragraph id="8BEA36FFFF950629FF1CC92FF992C532" blockId="17.[151,1437,151,1077]" pageId="17" pageNumber="192">
<emphasis id="B921EAEDFF950629FF1CC92FFA30C08E" bold="true" box="[199,315,367,392]" pageId="17" pageNumber="192">Remarks.</emphasis>
<taxonomicName id="4C554D7CFF950629FE9DC930F937C08E" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[326,572,368,392]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF950629FE9DC930F937C08E" box="[326,572,368,392]" italics="true" pageId="17" pageNumber="192">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
was found only in Chapada dos Guimarães municipality, a region with a formation of mountain ranges within the Cerrado in the south-central portion of 
<collectingRegion id="4991F81DFF950629FBFDC9D5FFB7C0AA" box="[1062,1212,404,428]" country="Brazil" name="Mato Grosso" pageId="17" pageNumber="192">Mato Grosso</collectingRegion>
. The environmental conditions of the Chapada dos Guimarães are distinct from the other Cerrado areas, where 
<taxonomicName id="4C554D7CFF950629FB6AC9F8FE43C0D6" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[1201,1352,440,464]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF950629FB6AC9F8FE43C0D6" box="[1201,1352,440,464]" italics="true" pageId="17" pageNumber="192">A. diceratops</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C554D7CFF950629FA5FC9F8FA22C0F2" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF950629FA5FC9F8FA22C0F2" italics="true" pageId="17" pageNumber="192">A. bolboceridus</emphasis>
</taxonomicName>
were found, generally being more humid and with extensive valley forests, covering the crevices and recesses of rocky walls (
<bibRefCitation id="EFC44B0EFF950629FE72C9BFF9E5C31E" author="Pinto, J. R. R. &amp; Oliveira-Filho, A. T." box="[425,750,511,536]" pageId="17" pageNumber="192" pagination="53 - 67" refId="ref9782" refString="Pinto, J. R. R. &amp; Oliveira-Filho, A. T. (1999) Perfil floristico e estrutura da comunidade arborea de uma floresta de vale no Parque Nacional da Chapada dos Guimaraes, Mato Grosso, Brasil. Revista Brasileira de Botanica, 22, 53 - 67. https: // doi. org / 10.1590 / S 0100 - 84041999000100008" type="journal article" year="1999">Pinto &amp; Oliveira-Filho 1999</bibRefCitation>
). 
<taxonomicName id="4C554D7CFF950629FCDACA40F8F8C31E" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[769,1011,512,536]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF950629FCDACA40F8F8C31E" box="[769,1011,512,536]" italics="true" pageId="17" pageNumber="192">Aegopsis vazdemelloi</emphasis>
</taxonomicName>
is very similar to 
<taxonomicName id="4C554D7CFF950629FB64CA40FE7AC31E" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[1215,1393,512,536]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF950629FB64CA40FE7AC31E" box="[1215,1393,512,536]" italics="true" pageId="17" pageNumber="192">A. bolboceridus</emphasis>
</taxonomicName>
but differs in males by having the vertex four times wider than the eye width (
<figureCitation id="136E2A7AFF950629FC0ECA64FF20C33A" box="[981,1067,548,573]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="17" pageNumber="192">Fig. 2E</figureCitation>
), the mentum subtriangular (
<figureCitation id="136E2A7AFF950629FAB4CA64FBBCC366" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="17" pageNumber="192">Fig. 5F</figureCitation>
), and the presence of small punctures on the posterior pronotal angles, with micropunctation between them (
<figureCitation id="136E2A7AFF950629FAB4CA07FBB3C382" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="17" pageNumber="192">Fig. 2F</figureCitation>
). On the contrary, in 
<taxonomicName id="4C554D7CFF950629FE73CA2CF951C382" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[424,602,620,644]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF950629FE73CA2CF951C382" box="[424,602,620,644]" italics="true" pageId="17" pageNumber="192">A. bolboceridus</emphasis>
</taxonomicName>
the vertex is 3.5 times wider than the eye width (
<figureCitation id="136E2A7AFF950629FB57CA2CFFECC382" box="[1164,1255,620,645]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="17" pageNumber="192">Fig. 2A</figureCitation>
), the mentum is subrectangular (
<figureCitation id="136E2A7AFF950629FE90CACFFAAFC3AE" box="[331,420,655,680]" captionStart="FIGURE 5" captionStartId="7.[151,250,1292,1314]" captionTargetBox="[151,1436,193,1270]" captionTargetId="figure@7.[151,1436,193,1270]" captionTargetPageId="7" captionText="FIGURE 5. A, Mandible of Aegopsis bolboceridus male, in ventral view; B, mandible of Aegopsis diceratops male, in ventral view; C, mandible of Aegopsis vazdemelloi male, in ventral view; D, mentum of Aegopsis bolboceridus male, in ventral view; E, mentum of Aegopsis diceratops male, in ventral view; F, mentum of Aegopsis vazdemelloi male, in ventral view. Arrows indicating respectively apical protuberance and protuberance on internal lobe in A; Arrows indicating the internal lobe in B, C; arrow indicating the mentum carina in E. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611541/files/figure.png" pageId="17" pageNumber="192">Fig. 5D</figureCitation>
), and the micropunctures are absent between the punctures on the posterior angles, instead with the surface smooth (
<figureCitation id="136E2A7AFF950629FE6BCAF4F90CC3CA" box="[432,519,692,717]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="17" pageNumber="192">Fig. 2B</figureCitation>
). Also, males of 
<taxonomicName id="4C554D7CFF950629FD18CAF4F86DC3CA" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[707,870,692,716]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF950629FD18CAF4F86DC3CA" box="[707,870,692,716]" italics="true" pageId="17" pageNumber="192">A. vazdemelloi</emphasis>
</taxonomicName>
differ from 
<taxonomicName id="4C554D7CFF950629FC35CAF4FF89C3CA" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[1006,1154,692,716]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF950629FC35CAF4FF89C3CA" box="[1006,1154,692,716]" italics="true" pageId="17" pageNumber="192">A. diceratops</emphasis>
</taxonomicName>
in having the apex of the clypeus emarginate (
<figureCitation id="136E2A7AFF950629FE53CA97FAE8C3F6" box="[392,483,727,752]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="17" pageNumber="192">Fig. 2E</figureCitation>
), ocular canthi anteriorly almost straight (
<figureCitation id="136E2A7AFF950629FC0DCA97FF3AC3F6" box="[982,1073,727,752]" captionStart="FIGURE 3" captionStartId="5.[151,250,1876,1898]" captionTargetBox="[231,1354,193,1844]" captionTargetId="figure@5.[231,1354,193,1844]" captionTargetPageId="5" captionText="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611536/files/figure.png" pageId="17" pageNumber="192">Fig. 3E</figureCitation>
), and the ventral carina of the parameres basally acute (
<figureCitation id="136E2A7AFF950629FE64CABCF915C213" box="[447,542,764,789]" captionStart="FIGURE 6" captionStartId="9.[151,250,1775,1797]" captionTargetBox="[152,1421,205,1743]" captionTargetId="figure@9.[151,1436,193,1754]" captionTargetPageId="9" captionText="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." httpUri="https://zenodo.org/record/2611544/files/figure.png" pageId="17" pageNumber="192">Fig. 6H</figureCitation>
). In 
<taxonomicName id="4C554D7CFF950629FD82CABCF9F9C212" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[601,754,764,788]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF950629FD82CABCF9F9C212" box="[601,754,764,788]" italics="true" pageId="17" pageNumber="192">A. diceratops</emphasis>
</taxonomicName>
the apex of the clypeus is round (
<figureCitation id="136E2A7AFF950629FB4BCABCFFE6C212" box="[1168,1261,764,789]" captionStart="FIGURE 2" captionStartId="4.[151,250,1882,1904]" captionTargetBox="[200,1377,193,1853]" captionTargetId="figure@4.[200,1377,193,1853]" captionTargetPageId="4" captionText="FIGURE 2. A, Head of Aegopsis bolboceridus male, in dorsal view; B, pronotum of Aegopsis bolboceridus male, in dorsal view; C, head of Aegopsis diceratops male, in dorsal view; D, pronotum of Aegopsis diceratops male, in dorsal view; E, head of Aegopsis vazdemelloi male, in dorsal view; F, pronotum of Aegopsis vazdemelloi male, in dorsal view. Black arrow indicating emarginate apex of clypeus in A, E; white arrow indicating micropunctures on pronotum in F. Scale bars: A, C, F, 2 mm; B, D, E, 3 mm." httpUri="https://zenodo.org/record/2611534/files/figure.png" pageId="17" pageNumber="192">Fig. 2C</figureCitation>
), ocular canthi anteriorly protruded (
<figureCitation id="136E2A7AFF950629FE50CB5FFAEEC23E" box="[395,485,799,824]" captionStart="FIGURE 3" captionStartId="5.[151,250,1876,1898]" captionTargetBox="[231,1354,193,1844]" captionTargetId="figure@5.[231,1354,193,1844]" captionTargetPageId="5" captionText="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611536/files/figure.png" pageId="17" pageNumber="192">Fig. 3C</figureCitation>
), and the ventral carina of the parameres is basally round (
<figureCitation id="136E2A7AFF950629FB55CB5FFFECC23E" box="[1166,1255,799,824]" captionStart="FIGURE 6" captionStartId="9.[151,250,1775,1797]" captionTargetBox="[152,1421,205,1743]" captionTargetId="figure@9.[151,1436,193,1754]" captionTargetPageId="9" captionText="FIGURE 6. A, Aedeagus of Aegopsis bolboceridus, in lateral view; B, parameres of Aegopsis bolboceridus, in ventral view; C, parameres of Aegopsis bolboceridus, in dorsal view; D, aedeagus of Aegopsis diceratops in lateral view; E, parameres of Aegopsis diceratops in ventral view; F, parameres of Aegopsis diceratops in dorsal view; G, aedeagus of Aegopsis vazdemelloi in lateral view; H, parameres of Aegopsis vazdemelloi in ventral view; I, parameres of Aegopsis vazdemelloi in dorsal view. White arrows in the left indicating ventral carina of parameres in B, E, H; white arrows upward indicating inferior edge of parameres in B, E, H. Scale bars: A, D, G, 1 mm; B, C, E, F, H, I, 0.5 mm." httpUri="https://zenodo.org/record/2611544/files/figure.png" pageId="17" pageNumber="192">Fig. 6E</figureCitation>
). Females of 
<taxonomicName id="4C554D7CFF950629FA5FCB60FA17C25A" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF950629FA5FCB60FA17C25A" italics="true" pageId="17" pageNumber="192">A. vazdemelloi</emphasis>
</taxonomicName>
can be distinguished by the deeper and coalescent punctures on the vertex (
<figureCitation id="136E2A7AFF950629FBB4CB04FFCEC25A" box="[1135,1221,836,861]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="17" pageNumber="192">Fig. 7E</figureCitation>
), the distance from canthus carina to canthus apex longer than the canthus thickness (
<figureCitation id="136E2A7AFF950629FC5FCB27F8D1C286" box="[900,986,871,896]" captionStart="FIGURE 3" captionStartId="5.[151,250,1876,1898]" captionTargetBox="[231,1354,193,1844]" captionTargetId="figure@5.[231,1354,193,1844]" captionTargetPageId="5" captionText="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611536/files/figure.png" pageId="17" pageNumber="192">Fig. 3F</figureCitation>
), the pronotum rounder (
<figureCitation id="136E2A7AFF950629FB2CCB27FE55C286" box="[1271,1374,871,896]" captionStart="FIGURE 10" captionStartId="13.[151,250,1970,1992]" captionTargetBox="[194,1312,193,1920]" captionTargetId="figure@13.[194,1388,193,1949]" captionTargetPageId="13" captionText="FIGURE 10. Habitus of Aegopsis vazdemelloi. A, Major male in dorsal view; B, minor male in dorsal view; C, female in dorsal view. Scale bars: 3 mm." httpUri="https://zenodo.org/record/2611553/files/figure.png" pageId="17" pageNumber="192">Fig. 10C</figureCitation>
), and the sensorial area of maxillary palpi rugose (
<figureCitation id="136E2A7AFF950629FD49CBCCF9E9C2A2" box="[658,738,908,933]" captionStart="FIGURE 4" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[151,1415,201,1637]" captionTargetId="figure@6.[151,1436,193,1672]" captionTargetPageId="6" captionText="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611538/files/figure.png" pageId="17" pageNumber="192">
Fig. 
<date id="FFEB103FFF950629FD1CCBCCF9E9C2A2" box="[711,738,908,932]" pageId="17" pageNumber="192">4I</date>
</figureCitation>
). In 
<taxonomicName id="4C554D7CFF950629FCCCCBCCF8C2C2A2" authorityName=": Dechambre &amp; Grossi" authorityYear="1990" box="[791,969,908,932]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="bolboceridus">
<emphasis id="B921EAEDFF950629FCCCCBCCF8C2C2A2" box="[791,969,908,932]" italics="true" pageId="17" pageNumber="192">A. bolboceridus</emphasis>
</taxonomicName>
, the punctures are shallower and annular on the vertex (
<figureCitation id="136E2A7AFF950629FEE2CBEFFA98C2CE" box="[313,403,943,968]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="17" pageNumber="192">Fig. 7A</figureCitation>
), the distance from canthus carina to canthus apex is subequal to the canthus thickness (
<figureCitation id="136E2A7AFF950629FAB4CBEFFBB1C2EA" captionStart="FIGURE 3" captionStartId="5.[151,250,1876,1898]" captionTargetBox="[231,1354,193,1844]" captionTargetId="figure@5.[231,1354,193,1844]" captionTargetPageId="5" captionText="FIGURE 3. A, Canthus of Aegopsis bolboceridus male, in dorsal view; B, canthus of Aegopsis bolboceridus female, in dorsal view; C, canthus of Aegopsis diceratops male, in dorsal view; D, canthus of Aegopsis diceratops female, in dorsal view; E, canthus of Aegopsis vazdemelloi male, in dorsal view; F, canthus of Aegopsis vazdemelloi female, in dorsal view. White arrow indicating apex inconspicuous of canthi carina in A; black arrows indicating canthi carina split in C; black arrow indicating apex conspicuous of canthi carina in E; black arrows indicating external edge of canthi in B, D, F. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611536/files/figure.png" pageId="17" pageNumber="192">Fig. 3B</figureCitation>
), and the sensorial area of the maxillary palpi striate (
<figureCitation id="136E2A7AFF950629FCC3CB94F879C2EA" box="[792,882,980,1005]" captionStart="FIGURE 4" captionStartId="6.[151,250,1693,1715]" captionTargetBox="[151,1415,201,1637]" captionTargetId="figure@6.[151,1436,193,1672]" captionTargetPageId="6" captionText="FIGURE 4. A, Labrum of Aegopsis bolboceridus male, in dorsal view; B, maxillary palpomere IV of Aegopsis bolboceridus male, in dorsal view; C, maxillary palpomere IV of Aegopsis bolboceridus female, in dorsal view; D, labrum of Aegopsis diceratops male, in dorsal view; E, maxillary palpomere IV of Aegopsis diceratops male, in dorsal view; F, maxillary palpomere IV of Aegopsis diceratops female, in dorsal view; G, labrum of Aegopsis vazdemelloi male, in dorsal view; H, maxillary palpomere IV of Aegopsis vazdemelloi male, in dorsal view; I, maxillary palpomere IV of Aegopsis vazdemelloi female, in dorsal view. Arrows indicating the middle of the labrum in A, G; arrows indicating sensorial area in C, I. Scale bars: 0.5 mm." httpUri="https://zenodo.org/record/2611538/files/figure.png" pageId="17" pageNumber="192">Fig. 4C</figureCitation>
). In 
<taxonomicName id="4C554D7CFF950629FC7FCB94FF31C2EA" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[932,1082,980,1004]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="diceratops">
<emphasis id="B921EAEDFF950629FC7FCB94FF31C2EA" box="[932,1082,980,1004]" italics="true" pageId="17" pageNumber="192">A. diceratops</emphasis>
</taxonomicName>
, the clypeus is trapezoidal, and punctures are absent on the vertex (
<figureCitation id="136E2A7AFF950629FDF5CBB7F983C516" box="[558,648,1015,1040]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="17" pageNumber="192">Fig. 7C</figureCitation>
), while in 
<taxonomicName id="4C554D7CFF950629FCD8CBB8F8A2C516" authorityName="Sobral &amp; Grossi &amp; De Morais" authorityYear="2018" box="[771,937,1016,1040]" class="Insecta" family="Scarabaeidae" genus="Aegopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="192" phylum="Arthropoda" rank="species" species="vazdemelloi">
<emphasis id="B921EAEDFF950629FCD8CBB8F8A2C516" box="[771,937,1016,1040]" italics="true" pageId="17" pageNumber="192">A. vazdemelloi</emphasis>
</taxonomicName>
the clypeus is round, and the punctures are deeper and coalescent on the vertex (
<figureCitation id="136E2A7AFF950629FDEECC5CF987C532" box="[565,652,1052,1077]" captionStart="FIGURE 7" captionStartId="10.[151,250,1892,1914]" captionTargetBox="[185,1389,202,1867]" captionTargetId="figure@10.[181,1389,193,1871]" captionTargetPageId="10" captionText="FIGURE 7. A, Head of Aegopsis bolboceridus female in dorsal view; B, head of Aegopsis bolboceridus female in lateral view; C, head of Aegopsis diceratops female in dorsal view; D, head of Aegopsis diceratops female in lateral view; E, head of Aegopsis vazdemelloi female in dorsal view; F, head of Aegopsis vazdemelloi female in lateral view. Black arrow indicating head punctures in A; black arrow indicating clypeus trapezoidal in C; white arrow indicating vertex in C; black arrow indicating frontal tubercles in D; white arrow indicating head punctures in E. Scale bars: 1 mm." httpUri="https://zenodo.org/record/2611546/files/figure.png" pageId="17" pageNumber="192">Fig. 7E</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>