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<mods:title id="4132B4C3A31CEADC84745D3C8B2C5EE8">A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). I. Gracilipalpe, morchella and nigrum groups</mods:title>
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<mods:namePart id="A976A54F9154E8CB43C1416FE803E5BE">Shavrin, Alexey V.</mods:namePart>
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<treatment id="03FC87DCFFAE9D30FF0D3831FA2EFE12" ID-DOI="http://doi.org/10.5281/zenodo.6017654" ID-GBIF-Taxon="139444195" ID-Zenodo-Dep="6017654" LSID="urn:lsid:plazi:treatment:03FC87DCFFAE9D30FF0D3831FA2EFE12" httpUri="http://treatment.plazi.org/id/03FC87DCFFAE9D30FF0D3831FA2EFE12" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<subSubSection id="C34F6541FFAE9D33FF0D3831FE5DF87C" box="[151,485,1895,1921]" pageId="21" pageNumber="22" type="nomenclature">
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<heading id="D0A281A6FFAE9D33FF0D3831FE5DF87C" bold="true" box="[151,485,1895,1921]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<emphasis id="B921EAD8FFAE9D33FF0D3831FE5DF87C" bold="true" box="[151,485,1895,1921]" pageId="21" pageNumber="22">
3.3.1. Key to
<taxonomicName id="4C554D49FFAE9D33FEA03831FE2CF87C" ID-CoL="3SJ7G" baseAuthorityName="Cameron" baseAuthorityYear="1924" box="[314,404,1895,1921]" class="Insecta" family="Staphylinidae" genus="Lathrimaeum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="nigrum">
<emphasis id="B921EAD8FFAE9D33FEA03831FE2CF87C" bold="true" box="[314,404,1895,1921]" italics="true" pageId="21" pageNumber="22">nigrum</emphasis>
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group
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<subSubSection id="C34F6541FFAE9D30FF0D38FBFA2EFE12" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="key">
<key id="FFC461FBFFAE9D30FF0D38FBFA2EFE12" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<keyStep id="30A12E6FFFAE9D30FF0D38FBFA24FF50" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<paragraph id="8BEA36CAFFAE9D33FF0D38FBFA24F81D" blockId="21.[151,1436,1965,2016]" pageId="21" pageNumber="22">
<keyLead id="30A495FFFFAE9D33FF0D38FBFA24F81D" pageId="21" pageNumber="22">
1 Anterior angles of pronotum strongly protruded anteriad. Elytra markedly wider than long. Aedeagus as in
<figureCitation id="136E2A4FFFAE9D33FB7C38FBFA95F83E" box="[1254,1325,1965,1987]" captionStart="FIGURES 7778" captionStartId="30.[151,264,1431,1454]" captionTargetBox="[194,1401,190,1412]" captionTargetId="figure@30.[151,1435,173,1432]" captionTargetPageId="30" captionText="FIGURES 7778. Aedeagusof Anthobium conjunctum (paratype):77—parameral view, 78—lateral view. Scale bar: 0.1mm." httpUri="https://zenodo.org/record/1125257/files/figure.png" pageId="21" pageNumber="22">Fig. 77</figureCitation>
. Habitus as in
<figureCitation id="136E2A4FFFAE9D33FF63389CFEF8F81D" box="[249,320,1994,2016]" captionStart="FIGURES 6569" captionStartId="24.[151,264,1459,1482]" captionTargetBox="[182,1395,210,1407]" captionTargetId="figure@24.[151,1436,193,1438]" captionTargetPageId="24" captionText="FIGURES 6569. Habitus of Anthobium: 65—A. nigrum (INDIA: Uttarkashi), 66—A. anishchenkoi (paratype), 67—A. conjunctum (holotype), 68—A. densepunctatum (holotype), 69—A. ivani (paratype).Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1125247/files/figure.png" pageId="21" pageNumber="22">Fig. 67</figureCitation>
. Body length:
<quantity id="4CAD9B2FFFAE9D33FE56389CFDE3F81D" box="[460,603,1994,2016]" metricMagnitude="-3" metricUnit="m" metricValue="4.05" metricValueMax="4.2" metricValueMin="3.9" pageId="21" pageNumber="22" unit="mm" value="4.05" valueMax="4.2" valueMin="3.9">3.904.20 mm</quantity>
.
<collectingCountry id="F342765AFFAE9D33FDF9389CFD18F81D" box="[611,672,1994,2016]" name="China" pageId="21" pageNumber="22">China</collectingCountry>
:
<collectingRegion id="4991F828FFAE9D33FD33389DFD4AF81D" box="[681,754,1995,2016]" country="China" name="Yunnan" pageId="21" pageNumber="22">Yunnan</collectingRegion>
........................................
<taxonomicName id="4C554D49FFAE9D33FB40389DFADDF81D" ID-CoL="67CD9" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[1242,1381,1995,2016]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="conjunctum" status="sp. nov.">
<emphasis id="B921EAD8FFAE9D33FB40389DFADDF81D" box="[1242,1381,1995,2016]" italics="true" pageId="21" pageNumber="22">A. conjunctum</emphasis>
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<emphasis id="B921EAD8FFAE9D33FAF6389CFA24F81D" bold="true" box="[1388,1436,1994,2016]" pageId="21" pageNumber="22">
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<paragraph id="8BEA36CAFFAD9D30FF0D3FC1FA24FF50" blockId="22.[151,1437,151,495]" box="[151,1436,151,173]" pageId="22" pageNumber="23">
<keyLead id="30A495FFFFAD9D30FF0D3FC1FA24FF50" box="[151,1436,151,173]" pageId="22" pageNumber="23">- Anterior angles of pronotum slightly protruded anteriad. Elytra as wide as long or slightly longer than wide............. 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E6FFFAD9D30FF0D3FE2FA23FEDF" pageId="22" pageNumber="23">
<paragraph id="8BEA36CAFFAD9D30FF0D3FE2FA24FF15" blockId="22.[151,1437,151,495]" pageId="22" pageNumber="23">
<keyLead id="30A495FFFFAD9D30FF0D3FE2FA24FF15" pageId="22" pageNumber="23">
2 Punctation of pronotum very dense. Aedeagus very wide basally, markedly, evenly narrowed toward apex of median lobe (
<figureCitation id="136E2A4FFFAD9D30FAEF3FE2FF44FF1A" captionStart="FIGURES 8083" captionStartId="33.[151,264,1971,1994]" captionTargetBox="[274,1291,206,1932]" captionTargetId="figure@33.[251,1339,193,1957]" captionTargetPageId="33" captionText="FIGURES 8083. Aedeagus of Anthobium densepunctatum (8081—paratype), A. ivani (8283—paratype): 80, 82— parameral view, 81, 83—ateral view. Scale bars:0.1 mm." httpUri="https://zenodo.org/record/1125261/files/figure.png" pageId="22" pageNumber="23">Fig. 80</figureCitation>
). Habitus as in
<figureCitation id="136E2A4FFFAD9D30FE0A3F87FE6FFF1A" box="[400,471,209,231]" captionStart="FIGURES 6569" captionStartId="24.[151,264,1459,1482]" captionTargetBox="[182,1395,210,1407]" captionTargetId="figure@24.[151,1436,193,1438]" captionTargetPageId="24" captionText="FIGURES 6569. Habitus of Anthobium: 65—A. nigrum (INDIA: Uttarkashi), 66—A. anishchenkoi (paratype), 67—A. conjunctum (holotype), 68—A. densepunctatum (holotype), 69—A. ivani (paratype).Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1125247/files/figure.png" pageId="22" pageNumber="23">Fig. 68</figureCitation>
. Body length:
<quantity id="4CAD9B2FFFAD9D30FDF83F87FD49FF1A" box="[610,753,209,231]" metricMagnitude="-3" metricUnit="m" metricValue="3.885" metricValueMax="4.27" metricValueMin="3.5" pageId="22" pageNumber="23" unit="mm" value="3.885" valueMax="4.27" valueMin="3.5">3.504.27 mm</quantity>
.
<collectingCountry id="F342765AFFAD9D30FD633F87FC8EFF1A" box="[761,822,209,231]" name="China" pageId="22" pageNumber="23">China</collectingCountry>
:
<collectingRegion id="4991F828FFAD9D30FCA53F84FC30FF1A" box="[831,904,210,231]" country="China" name="Yunnan" pageId="22" pageNumber="23">Yunnan</collectingRegion>
........................
<taxonomicName id="4C554D49FFAD9D30FB353F84FADEFF1A" ID-CoL="5VCCC" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[1199,1382,210,231]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="densepunctatum" status="sp. nov.">
<emphasis id="B921EAD8FFAD9D30FB353F84FADEFF1A" box="[1199,1382,210,231]" italics="true" pageId="22" pageNumber="23">A. densepunctatum</emphasis>
</taxonomicName>
<emphasis id="B921EAD8FFAD9D30FAF63F84FA24FF15" bold="true" box="[1388,1436,210,232]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="A21257A3FFAD9D30FAF63F84FA24FF15" box="[1388,1436,210,232]" pageId="22" pageNumber="23" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEA36CAFFAD9D30FF0D3FB9FA23FEDF" blockId="22.[151,1437,151,495]" pageId="22" pageNumber="23">
<keyLead id="30A495FFFFAD9D30FF0D3FB9FA23FEDF" pageId="22" pageNumber="23">
- Punctation of pronotum sparse. Aedeagus of different shape, not very wide basally (
<figureCitation id="136E2A4FFFAD9D30FC653FB9FBCBFEF8" box="[1023,1139,239,261]" captionStart="FIGURES 7073" captionStartId="27.[151,264,1966,1989]" captionTargetBox="[314,1276,204,1934]" captionTargetId="figure@27.[283,1304,193,1946]" captionTargetPageId="27" captionText="FIGURES 7073. Aedeagus of Anthobium nigrum (7071—INDIA: Uttarkashi) and A. anishchenkoi (7273—paratype): 70, 72—parameral view, 71, 73—lateral view.Scale bars:0.1 mm." httpUri="https://zenodo.org/record/1125249/files/figure.png" pageId="22" pageNumber="23">Figs. 70, 72</figureCitation>
), sometimes quite narrow (
<figureCitation id="136E2A4FFFAD9D30FAEE3FB9FF42FEDF" captionStart="FIGURES 8083" captionStartId="33.[151,264,1971,1994]" captionTargetBox="[274,1291,206,1932]" captionTargetId="figure@33.[251,1339,193,1957]" captionTargetPageId="33" captionText="FIGURES 8083. Aedeagus of Anthobium densepunctatum (8081—paratype), A. ivani (8283—paratype): 80, 82— parameral view, 81, 83—ateral view. Scale bars:0.1 mm." httpUri="https://zenodo.org/record/1125261/files/figure.png" pageId="22" pageNumber="23">Fig. 82</figureCitation>
)................................................................................................ 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E6FFFAD9D30FF0D3E7FFA23FE87" pageId="22" pageNumber="23">
<paragraph id="8BEA36CAFFAD9D30FF0D3E7FFA24FEA0" blockId="22.[151,1437,151,495]" pageId="22" pageNumber="23">
<keyLead id="30A495FFFFAD9D30FF0D3E7FFA24FEA0" pageId="22" pageNumber="23">
3 Pronotum slightly flattened in cross section, with indistinct medio-basal impression. Aedeagus as in
<figureCitation id="136E2A4FFFAD9D30FB393E7FFB52FEC2" box="[1187,1258,297,319]" captionStart="FIGURES 7073" captionStartId="27.[151,264,1966,1989]" captionTargetBox="[314,1276,204,1934]" captionTargetId="figure@27.[283,1304,193,1946]" captionTargetPageId="27" captionText="FIGURES 7073. Aedeagus of Anthobium nigrum (7071—INDIA: Uttarkashi) and A. anishchenkoi (7273—paratype): 70, 72—parameral view, 71, 73—lateral view.Scale bars:0.1 mm." httpUri="https://zenodo.org/record/1125249/files/figure.png" pageId="22" pageNumber="23">Fig. 72</figureCitation>
. Habitus as in
<figureCitation id="136E2A4FFFAD9D30FAEF3E7FFF44FEA0" captionStart="FIGURES 6569" captionStartId="24.[151,264,1459,1482]" captionTargetBox="[182,1395,210,1407]" captionTargetId="figure@24.[151,1436,193,1438]" captionTargetPageId="24" captionText="FIGURES 6569. Habitus of Anthobium: 65—A. nigrum (INDIA: Uttarkashi), 66—A. anishchenkoi (paratype), 67—A. conjunctum (holotype), 68—A. densepunctatum (holotype), 69—A. ivani (paratype).Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1125247/files/figure.png" pageId="22" pageNumber="23">Fig. 66</figureCitation>
. Body length:
<quantity id="4CAD9B2FFFAD9D30FE1D3E11FDADFEA1" box="[391,533,327,349]" metricMagnitude="-3" metricUnit="m" metricValue="4.26" metricValueMax="4.37" metricValueMin="4.15" pageId="22" pageNumber="23" unit="mm" value="4.26" valueMax="4.37" valueMin="4.15">4.154.37 mm</quantity>
.
<collectingCountry id="F342765AFFAD9D30FD843E11FDE3FEA0" box="[542,603,327,349]" name="China" pageId="22" pageNumber="23">China</collectingCountry>
:
<collectingRegion id="4991F828FFAD9D30FDFE3E11FD19FEA0" box="[612,673,327,349]" country="China" name="Gansu" pageId="22" pageNumber="23">Gansu</collectingRegion>
.............................................
<taxonomicName id="4C554D49FFAD9D30FB563E11FADDFEA1" ID-CoL="5VCBK" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[1228,1381,327,348]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="anishchenkoi" status="sp. nov.">
<emphasis id="B921EAD8FFAD9D30FB563E11FADDFEA1" box="[1228,1381,327,348]" italics="true" pageId="22" pageNumber="23">A. anishchenkoi</emphasis>
</taxonomicName>
<emphasis id="B921EAD8FFAD9D30FAF63E11FA24FEA0" bold="true" box="[1388,1436,327,349]" pageId="22" pageNumber="23">
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</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEA36CAFFAD9D30FF0D3E33FA23FE87" blockId="22.[151,1437,151,495]" box="[151,1435,356,378]" pageId="22" pageNumber="23">
<keyLead id="30A495FFFFAD9D30FF0D3E33FA23FE87" box="[151,1435,356,378]" pageId="22" pageNumber="23">- Pronotum very convex, with deep medio-basal impression.................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30A12E6FFFAD9D30FF0D3ED7FA2EFE12" pageId="22" pageNumber="23">
<paragraph id="8BEA36CAFFAD9D30FF0D3ED7FA24FE48" blockId="22.[151,1437,151,495]" pageId="22" pageNumber="23">
<keyLead id="30A495FFFFAD9D30FF0D3ED7FA24FE48" pageId="22" pageNumber="23">
4 Elytra as wide as long. Apical part of median lobe of aedeagus narrow, apical portions of parameres very wide (
<figureCitation id="136E2A4FFFAD9D30FA963ED7FAECFE6A" box="[1292,1364,385,407]" captionStart="FIGURES 8083" captionStartId="33.[151,264,1971,1994]" captionTargetBox="[274,1291,206,1932]" captionTargetId="figure@33.[251,1339,193,1957]" captionTargetPageId="33" captionText="FIGURES 8083. Aedeagus of Anthobium densepunctatum (8081—paratype), A. ivani (8283—paratype): 80, 82— parameral view, 81, 83—ateral view. Scale bars:0.1 mm." httpUri="https://zenodo.org/record/1125261/files/figure.png" pageId="22" pageNumber="23">Fig. 82</figureCitation>
). Habitus as in
<figureCitation id="136E2A4FFFAD9D30FEAF3EC9FEC4FE48" box="[309,380,415,437]" captionStart="FIGURES 6569" captionStartId="24.[151,264,1459,1482]" captionTargetBox="[182,1395,210,1407]" captionTargetId="figure@24.[151,1436,193,1438]" captionTargetPageId="24" captionText="FIGURES 6569. Habitus of Anthobium: 65—A. nigrum (INDIA: Uttarkashi), 66—A. anishchenkoi (paratype), 67—A. conjunctum (holotype), 68—A. densepunctatum (holotype), 69—A. ivani (paratype).Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1125247/files/figure.png" pageId="22" pageNumber="23">Fig. 69</figureCitation>
. Body length: 4.00
<quantity id="4CAD9B2FFFAD9D30FDA53EC9FD2EFE49" box="[575,662,415,437]" metricMagnitude="-3" metricUnit="m" metricValue="4.71" pageId="22" pageNumber="23" unit="mm" value="4.71">4.71 mm</quantity>
.
<collectingCountry id="F342765AFFAD9D30FD053EC9FD62FE48" box="[671,730,415,437]" name="Nepal" pageId="22" pageNumber="23">Nepal</collectingCountry>
,
<collectingCountry id="F342765AFFAD9D30FD7E3EC9FC91FE48" box="[740,809,415,437]" name="Bhutan" pageId="22" pageNumber="23">Bhutan</collectingCountry>
........................................
<taxonomicName id="4C554D49FFAD9D30FA803EC9FADDFE49" ID-CoL="5VCDH" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[1306,1381,415,436]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="ivani" status="sp. nov.">
<emphasis id="B921EAD8FFAD9D30FA803EC9FADDFE49" box="[1306,1381,415,436]" italics="true" pageId="22" pageNumber="23">A. ivani</emphasis>
</taxonomicName>
<emphasis id="B921EAD8FFAD9D30FAF63EC9FA24FE48" bold="true" box="[1388,1436,415,437]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="A21257A3FFAD9D30FAF63EC9FA24FE48" box="[1388,1436,415,437]" pageId="22" pageNumber="23" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BEA36CAFFAD9D30FF0D3EEBFA2EFE12" blockId="22.[151,1437,151,495]" pageId="22" pageNumber="23">
<keyLead id="30A495FFFFAD9D30FF0D3EEBFA2EFE12" pageId="22" pageNumber="23">
- Elytra slightly longer than wide. Apical part of median lobe of aedeagus wide, apical portions of parameres relatively narrow (
<figureCitation id="136E2A4FFFAD9D30FF723E8FFE88FE12" box="[232,304,473,495]" captionStart="FIGURES 7073" captionStartId="27.[151,264,1966,1989]" captionTargetBox="[314,1276,204,1934]" captionTargetId="figure@27.[283,1304,193,1946]" captionTargetPageId="27" captionText="FIGURES 7073. Aedeagus of Anthobium nigrum (7071—INDIA: Uttarkashi) and A. anishchenkoi (7273—paratype): 70, 72—parameral view, 71, 73—lateral view.Scale bars:0.1 mm." httpUri="https://zenodo.org/record/1125249/files/figure.png" pageId="22" pageNumber="23">Fig. 70</figureCitation>
). Habitus as in
<figureCitation id="136E2A4FFFAD9D30FE5E3E8FFDB3FE12" box="[452,523,473,495]" captionStart="FIGURES 6569" captionStartId="24.[151,264,1459,1482]" captionTargetBox="[182,1395,210,1407]" captionTargetId="figure@24.[151,1436,193,1438]" captionTargetPageId="24" captionText="FIGURES 6569. Habitus of Anthobium: 65—A. nigrum (INDIA: Uttarkashi), 66—A. anishchenkoi (paratype), 67—A. conjunctum (holotype), 68—A. densepunctatum (holotype), 69—A. ivani (paratype).Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1125247/files/figure.png" pageId="22" pageNumber="23">Fig. 65</figureCitation>
. Body length:
<quantity id="4CAD9B2FFFAD9D30FD0C3E8FFC9CFE12" box="[662,804,473,495]" metricMagnitude="-3" metricUnit="m" metricValue="4.095" metricValueMax="4.75" metricValueMin="3.44" pageId="22" pageNumber="23" unit="mm" value="4.095" valueMax="4.75" valueMin="3.44">3.444.75 mm</quantity>
. India........................
<taxonomicName id="4C554D49FFAD9D30FB163E8CFA2EFE12" ID-CoL="3SJ7G" authority="(Cameron, 1924)" baseAuthorityName="Cameron" baseAuthorityYear="1924" box="[1164,1430,473,495]" class="Insecta" family="Staphylinidae" genus="Lathrimaeum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="nigrum">
<emphasis id="B921EAD8FFAD9D30FB163E8CFB53FE12" box="[1164,1259,474,495]" italics="true" pageId="22" pageNumber="23">A. nigrum</emphasis>
(
<bibRefCitation id="EFC44B3BFFAD9D30FB623E8FFA37FE12" author="Cameron" box="[1272,1423,473,495]" pageId="22" pageNumber="23" refString="Cameron, M. (1924) New species of Staphylinidae from India. The Transactions of the Entomological Society of London, 1924, 160 - 198. https: // doi. org / 10.1111 / j. 1365 - 2311.1924. tb 03357. x" type="journal article" year="1924">Cameron, 1924</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>