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<document id="453A835DEB69B789738BFCFA8EDDA43D" ID-CLB-Dataset="31106" ID-DOI="10.11646/zootaxa.4365.1.1" ID-GBIF-Dataset="8e2d27e3-dec0-4ebb-bcd3-dd16e56c8092" ID-ISSN="1175-5326" ID-Zenodo-Dep="1125219" ID-ZooBank="5F132840-6E2F-42E6-8DF2-9EF191DE243C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1513931221510" checkinUser="plazi" docAuthor="Shavrin, Alexey V. &amp; Smetana, Aleš" docDate="2017" docId="03FC87DCFF989D02FF0D3CDAFDB7FBED" docLanguage="en" docName="zootaxa.4365.1.1.pdf.imf" docOrigin="Zootaxa 4365 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anthobium ivani Shavrin &amp; Smetana 2017, sp.n." docType="treatment" docVersion="8" lastPageNumber="37" masterDocId="FFC5FFA4FFBB9D26FF9A3F56FFB8FFFD" masterDocTitle="A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). I. Gracilipalpe, morchella and nigrum groups" masterLastPageNumber="39" masterPageNumber="1" pageNumber="36" updateTime="1698511183827" updateUser="ExternalLinkService">
<mods:mods id="B39CA2314F944FD56AE121CE3B50D960" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="355F7473EE198C792192A81558398F7D">A revision of Eastern Palaearctic Anthobium Leach, 1819 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). I. Gracilipalpe, morchella and nigrum groups</mods:title>
</mods:titleInfo>
<mods:name id="FBE4D660D901AF5712AF040B8738DD63" type="personal">
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<mods:namePart id="880079089A19399C10248C42F1B5EBA6">Shavrin, Alexey V.</mods:namePart>
</mods:name>
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<mods:namePart id="32B2C9C2928BA4266FE0E80DFBA1CF6F">Smetana, Aleš</mods:namePart>
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<mods:title id="7E17F72FD11F46E293FFEE22982FB0D2">Zootaxa</mods:title>
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<mods:part id="9D5D225D8B7AE61633CAB18261582128">
<mods:date id="70671BC977E211BE9E30AAA1A8E453B9">2017</mods:date>
<mods:detail id="5F3D5C66778600104CFCBCB78D01C6FD" type="pubDate">
<mods:number id="4BEFA345DFC3902DDE2608DAAFD01762">2017-12-15</mods:number>
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<mods:detail id="E8514D7D41939D85C296A21E86AC50C2" type="volume">
<mods:number id="6C2B2572A014050A1DB9657FD738C2BF">4365</mods:number>
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<mods:end id="1FFEDA0DD8BDF93BC326CB0917DB9DBF">39</mods:end>
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<mods:classification id="A195D2CA9A7D84FCC50FB0FE6F5D645A">journal article</mods:classification>
<mods:identifier id="4A975B009BAB04C059C5720B3D6FB899" type="CLB-Dataset">31106</mods:identifier>
<mods:identifier id="6B584C4751A847394EB9302BAB979ADC" type="DOI">10.11646/zootaxa.4365.1.1</mods:identifier>
<mods:identifier id="D31F86866FC78216E4075CF4143F0EC9" type="GBIF-Dataset">8e2d27e3-dec0-4ebb-bcd3-dd16e56c8092</mods:identifier>
<mods:identifier id="34FF43447BABDF65A563688CCE0D445F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="651B924A9A7056B49EB4AFAB0C98D63D" type="Zenodo-Dep">1125219</mods:identifier>
<mods:identifier id="3CC51ECB641CEAD9F783580DC1A79317" type="ZooBank">5F132840-6E2F-42E6-8DF2-9EF191DE243C</mods:identifier>
</mods:mods>
<treatment id="03FC87DCFF989D02FF0D3CDAFDB7FBED" ID-DOI="http://doi.org/10.5281/zenodo.6017664" ID-GBIF-Taxon="139444209" ID-Zenodo-Dep="6017664" LSID="urn:lsid:plazi:treatment:03FC87DCFF989D02FF0D3CDAFDB7FBED" httpUri="http://treatment.plazi.org/id/03FC87DCFF989D02FF0D3CDAFDB7FBED" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<subSubSection id="C34F6541FF989D05FF0D3CDAFDBDFC5B" box="[151,517,908,934]" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="8BEA36CAFF989D05FF0D3CDAFDBDFC5B" blockId="35.[151,517,908,967]" box="[151,517,908,934]" pageId="35" pageNumber="36">
<heading id="D0A281A6FF989D05FF0D3CDAFDBDFC5B" bold="true" box="[151,517,908,934]" fontSize="11" level="1" pageId="35" pageNumber="36" reason="1">
<emphasis id="B921EAD8FF989D05FF0D3CDAFDBDFC5B" bold="true" box="[151,517,908,934]" pageId="35" pageNumber="36">
3.3.2.5.
<taxonomicName id="4C554D49FF989D05FF6C3CDAFE7AFC5B" ID-CoL="5VCDH" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[246,450,908,934]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="ivani" status="sp. nov.">
<emphasis id="B921EAD8FF989D05FF6C3CDAFE7AFC5B" bold="true" box="[246,450,908,934]" italics="true" pageId="35" pageNumber="36">Anthobium ivani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257A3FF989D05FE503CDAFDBDFC5B" box="[458,517,908,934]" pageId="35" pageNumber="36" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F6541FF989D05FF0D3CF8FE3AFC3A" box="[151,386,942,967]" pageId="35" pageNumber="36" type="description">
<paragraph id="8BEA36CAFF989D05FF0D3CF8FE3AFC3A" blockId="35.[151,517,908,967]" box="[151,386,942,967]" pageId="35" pageNumber="36">
(
<figureCitation id="136E2A4FFF989D05FF053CF8FF44FC3A" box="[159,252,942,967]" captionStart="FIGURES 65–69" captionStartId="24.[151,264,1459,1482]" captionTargetBox="[182,1395,210,1407]" captionTargetId="figure@24.[151,1436,193,1438]" captionTargetPageId="24" captionText="FIGURES 65–69. Habitus of Anthobium: 65—A. nigrum (INDIA: Uttarkashi), 66—A. anishchenkoi (paratype), 67—A. conjunctum (holotype), 68—A. densepunctatum (holotype), 69—A. ivani (paratype).Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1125247/files/figure.png" pageId="35" pageNumber="36">Figs. 69</figureCitation>
,
<figureCitation id="136E2A4FFF989D05FE923CF9FE9EFC3A" box="[264,294,943,967]" captionStart="FIGURE 74" captionStartId="28.[151,250,982,1004]" captionTargetBox="[179,1408,193,961]" captionTargetId="figure@28.[179,1408,193,961]" captionTargetPageId="28" captionText="FIGURE 74.Distribution of Anthobium nigrum (black circles) and A. ivani (white circles) in the Himalayan Region." httpUri="https://zenodo.org/record/1125251/files/figure.png" pageId="35" pageNumber="36">74</figureCitation>
,
<figureCitation id="136E2A4FFF989D05FEA83CF9FEC1FC3A" box="[306,377,943,967]" captionStart="FIGURES 80–83" captionStartId="33.[151,264,1971,1994]" captionTargetBox="[274,1291,206,1932]" captionTargetId="figure@33.[251,1339,193,1957]" captionTargetPageId="33" captionText="FIGURES 80–83. Aedeagus of Anthobium densepunctatum (80–81—paratype), A. ivani (82–83—paratype): 80, 82— parameral view, 81, 83—ateral view. Scale bars:0.1 mm." httpUri="https://zenodo.org/record/1125261/files/figure.png" pageId="35" pageNumber="36">82–83</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34F6541FF989D05FF0D3CA0FED1F9FA" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="8BEA36CAFF989D05FF0D3CA0FC3AFB86" blockId="35.[151,1437,1014,2011]" pageId="35" pageNumber="36">
<emphasis id="B921EAD8FF989D05FF0D3CA0FE78FBF2" bold="true" box="[151,448,1014,1039]" pageId="35" pageNumber="36">
<typeStatus id="54EE8868FF989D05FF0D3CA0FF6AFBF2" box="[151,210,1014,1039]" pageId="35" pageNumber="36">Type</typeStatus>
material examined:
</emphasis>
<materialsCitation id="3B3D3C97FF989D05FE5C3CA0FC3AFB86" ID-GBIF-Occurrence="1799096745" collectingDate="1981-04-09" collectionCode="MHNG" collectorName="Löbl &amp; Smetana’" country="Nepal" elevation="0" location="below Thare Pati" municipality="below Thare Pati" pageId="35" pageNumber="36" specimenCount="1" specimenCount-male="1" stateProvince="Bagmati" typeStatus="holotype">
<typeStatus id="54EE8868FF989D05FE5C3CA0FD96FBF2" box="[454,558,1014,1039]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
<specimenCount id="9D53FD43FF989D05FDAF3CA1FDF2FBF3" box="[565,586,1015,1038]" pageId="35" pageNumber="36" type="male">♂</specimenCount>
[plastic plate with aedeagus, abdominal tergite VIII, sternite VIII and apical segment in Canadian balsam was pinned under the plate with beetle]: ‘
<collectingCountry id="F342765AFF989D05FC323B4DFBBDFBCE" box="[936,1029,1051,1075]" name="Nepal" pageId="35" pageNumber="36">NEPAL</collectingCountry>
(Prov.
<collectingRegion id="4991F828FF989D05FBCC3B4CFB0EFBCE" box="[1110,1206,1050,1075]" country="Nepal" name="Bagmati" pageId="35" pageNumber="36">Bagmati</collectingRegion>
) |
<collectingMunicipality id="6B8EACB0FF989D05FB4B3B4CFA28FBCE" box="[1233,1424,1050,1075]" pageId="35" pageNumber="36">below Thare Pati</collectingMunicipality>
|
<quantity id="4CAD9B2FFF989D05FF0D3B69FF56FBAB" box="[151,238,1087,1111]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="35" pageNumber="36" unit="m" value="3300.0">
<elevation id="0078D1F9FF989D05FF0D3B69FF77FBAA" box="[151,207,1087,1111]" pageId="35" pageNumber="36">3300</elevation>
m
</quantity>
,
<collectingDate id="EFAFE9E2FF989D05FF613B69FEF5FBAA" box="[251,333,1087,1111]" pageId="35" pageNumber="36" value="1981-04-09">
<date id="FFEB100AFF989D05FF613B69FEF5FBAA" box="[251,333,1087,1111]" pageId="35" pageNumber="36" value="1981-04-09">9.IV.81</date>
</collectingDate>
|
<collectorName id="26A0531CFF989D05FEF83B68FE2FFBAA" box="[354,407,1086,1111]" pageId="35" pageNumber="36">Löbl</collectorName>
&amp;
<collectorName id="26A0531CFF989D05FE263B69FD9EFBAA" box="[444,550,1087,1111]" pageId="35" pageNumber="36">Smetana’</collectorName>
&lt;rectangular label, printed&gt;, ‘HOLOTYPE |
<taxonomicName id="4C554D49FF989D05FBB23B68FB1BFBAB" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[1064,1187,1086,1110]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B921EAD8FF989D05FBB23B68FB1BFBAB" box="[1064,1187,1086,1110]" italics="true" pageId="35" pageNumber="36">Anthobium</emphasis>
</taxonomicName>
|
<taxonomicName id="4C554D49FF989D05FB223B69FB48FBAB" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[1208,1264,1087,1110]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="ivani" status="sp. nov.">
<emphasis id="B921EAD8FF989D05FB223B69FB48FBAB" box="[1208,1264,1087,1110]" italics="true" pageId="35" pageNumber="36">ivani</emphasis>
</taxonomicName>
<emphasis id="B921EAD8FF989D05FB623B68FA97FBAA" bold="true" box="[1272,1327,1086,1111]" pageId="35" pageNumber="36">
<taxonomicNameLabel id="A21257A3FF989D05FB623B68FA97FBAA" box="[1272,1327,1086,1111]" pageId="35" pageNumber="36" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
| Shavrin A. &amp; Smetana A. 2017’ &lt;red rectangular label, printed&gt; (
<collectionCode id="ED44AE0FFF989D05FC803B35FCCEFB86" box="[794,886,1123,1147]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="35" pageNumber="36">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BEA36CAFF989D05FF5D3BD1FED1F9FA" blockId="35.[151,1437,1014,2011]" pageId="35" pageNumber="36">
Paratypes (8 specimens):
<emphasis id="B921EAD8FF989D05FE763BD0FDEBFB62" bold="true" box="[492,595,1158,1183]" pageId="35" pageNumber="36">NEPAL:</emphasis>
1 ♂: same data as the holotype (MHNG); 1 ♀ [2 preparations of genital segment and abdominal tergite VIII, sternite VIII in Canadian balsam were pinned under the plate with beetle]: ‘NEPAL (Prov. Bagmati) | below Thare Pati |
<quantity id="4CAD9B2FFF989D05FD0E3B99FD52FB1B" box="[660,746,1231,1255]" metricMagnitude="3" metricUnit="m" metricValue="3.3" pageId="35" pageNumber="36" unit="m" value="3300.0">3300 m</quantity>
,
<date id="FFEB100AFF989D05FD6F3B99FCEDFB1A" box="[757,853,1231,1255]" pageId="35" pageNumber="36" value="1981-04-10">10.IV.81</date>
| Löbl &amp; Smetana 18B’ &lt;rectangular label, printed&gt; (NSMT); 1 ♂, 1 ♀: ‘Nepal-Himalaya | Khumbu Himal | lg. O. Jäger 1995’ &lt;rectangular label, printed&gt;, ‘Tal d. Dudh Kosi | Panyali Drangka | b. Phakdinoma | 2700mNN, 27.11.’ &lt;rectangular label, printed&gt; (SNSD);
<emphasis id="B921EAD8FF989D05FF0D3A6CFEAEFAAE" bold="true" box="[151,278,1338,1363]" pageId="35" pageNumber="36">BHUTAN:</emphasis>
3 ♀♀: ‘Bhutan | Thimphu [handwritten] |
<date id="FFEB100AFF989D05FD6D3A6DFCF6FAAE" box="[759,846,1339,1363]" pageId="35" pageNumber="36" value="1980-10">X.1980</date>
. | de Rougemont’ &lt;rectangular label, printed&gt;, &lt;one specimen with additional rectangular printed label ‘
<taxonomicName id="4C554D49FF989D05FC833A08FC08FA8B" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[793,944,1374,1398]" class="Insecta" family="Staphylinidae" genus="Lathrimaeum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B921EAD8FF989D05FC833A08FC08FA8B" box="[793,944,1374,1398]" italics="true" pageId="35" pageNumber="36">Lathrimaeum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257A3FF989D05FC583A09FC4FFA8B" box="[962,1015,1375,1398]" pageId="35" pageNumber="36" rank="species">n.sp.</taxonomicNameLabel>
[handwritten] | det. 198 | G. de Rougemont’&gt; (CRG); 1 ♀: ‘Bhutan | Chasilakha [handwritten] |
<date id="FFEB100AFF989D05FC0C3AD5FC51FA66" box="[918,1001,1411,1435]" pageId="35" pageNumber="36" value="1980-10">X.1980</date>
| de Rougemont’ &lt;rectangular label, printed&gt;, ‘
<taxonomicName id="4C554D49FF989D05FE943AF0FE1DFA43" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[270,421,1446,1470]" class="Insecta" family="Staphylinidae" genus="Lathrimaeum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAD8FF989D05FE943AF0FE1DFA43" box="[270,421,1446,1470]" italics="true" pageId="35" pageNumber="36">Lathrimaeum</emphasis>
</taxonomicName>
[handwritten] | n.sp. cf
<taxonomicName id="4C554D49FF989D05FD293AF1FCBBFA43" authorityName=": Smetana" authorityYear="2004" box="[691,771,1447,1470]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="nigrum">
<emphasis id="B921EAD8FF989D05FD293AF1FCBBFA43" box="[691,771,1447,1470]" italics="true" pageId="35" pageNumber="36">nigrum</emphasis>
</taxonomicName>
C. [handwritten] | det. 198 | G. de Rougemont’ (CRG). All paratypes with additional red rectangular printed label: ‘PARATYPE |
<taxonomicName id="4C554D49FF989D05FC4A3A9DFBF3FA1E" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[976,1099,1483,1507]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B921EAD8FF989D05FC4A3A9DFBF3FA1E" box="[976,1099,1483,1507]" italics="true" pageId="35" pageNumber="36">Anthobium</emphasis>
</taxonomicName>
|
<taxonomicName id="4C554D49FF989D05FBFD3A9DFB27FA1F" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[1127,1183,1483,1506]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="ivani" status="sp. nov.">
<emphasis id="B921EAD8FF989D05FBFD3A9DFB27FA1F" box="[1127,1183,1483,1506]" italics="true" pageId="35" pageNumber="36">ivani</emphasis>
</taxonomicName>
<emphasis id="B921EAD8FF989D05FB303A9CFB59FA1E" bold="true" box="[1194,1249,1482,1507]" pageId="35" pageNumber="36">
<taxonomicNameLabel id="A21257A3FF989D05FB303A9CFB59FA1E" box="[1194,1249,1482,1507]" pageId="35" pageNumber="36" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
| Shavrin A. &amp; Smetana A. 2017’.
</paragraph>
</subSubSection>
<subSubSection id="C34F6541FF989D02FF5D3944FB2FFD78" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="description">
<paragraph id="8BEA36CAFF989D05FF5D3944FBC3F98E" blockId="35.[151,1437,1014,2011]" pageId="35" pageNumber="36">
<emphasis id="B921EAD8FF989D05FF5D3944FEE9F9D6" bold="true" box="[199,337,1554,1579]" pageId="35" pageNumber="36">Description</emphasis>
. Measurements (n=9): HW: 0.82–0.90, HL: 0.52–0.56; AL (
<typeStatus id="54EE8868FF989D05FB9B3944FBD2F9D6" box="[1025,1130,1554,1579]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
): 1.63; OL: 0.25–0.28; PL: 0.62–0.69; PW: 1.25–1.32; ESL: 1.87–2.00; EW: 1.85–2.01; AW: 1.37–1.55; MTbL (
<typeStatus id="54EE8868FF989D05FB0B3960FB42F9B2" box="[1169,1274,1590,1615]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
): 0.97; MTrL (
<typeStatus id="54EE8868FF989D05FF05390CFEB0F98E" box="[159,264,1626,1651]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
): 0.42 (MTrL 1–4: 0.27; MTrL 5: 0.15); AedL: 0.67; TL: 4.00(
<typeStatus id="54EE8868FF989D05FC53390CFB97F98E" box="[969,1071,1626,1651]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
)–4.71.
</paragraph>
<paragraph id="8BEA36CAFF989D05FF5D3928FDB3F84A" blockId="35.[151,1437,1014,2011]" pageId="35" pageNumber="36">
Reddish-brown to black; antennomeres 3–11 brown; lateral and basal margins of pronotum reddish-brown; mouthparts and legs yellow to reddish-brown; apical part of apical segment of maxillary palpi and antennomeres 1 and 2 yellow. Head with irregular sparse punctation, somewhat deeper and denser on posterior part of infraorbital ridges, sometimes middle part of head between grooves impunctated, only with several small punctures irregularly scattered in middle; middle part of neck with irregular small punctation; pronotum with punctation slightly larger and deeper than that on head, sometimes sparser and more irregular on posterio-median portion (some
<typeStatus id="54EE8868FF989D05FAAB3865FA24F8B6" box="[1329,1436,1843,1867]" pageId="35" pageNumber="36" type="paratype">paratypes</typeStatus>
with extremely sparse and irregular punctation on middle elevation of pronotum, with several smoothed punctures on anterio-median portion and wide impunctated area in front of medio-basal impression and on anterior portion of pronotum). Habitus as in
<figureCitation id="136E2A4FFF989D05FE2E38C8FDBFF84A" box="[436,519,1950,1975]" captionStart="FIGURES 65–69" captionStartId="24.[151,264,1459,1482]" captionTargetBox="[182,1395,210,1407]" captionTargetId="figure@24.[151,1436,193,1438]" captionTargetPageId="24" captionText="FIGURES 65–69. Habitus of Anthobium: 65—A. nigrum (INDIA: Uttarkashi), 66—A. anishchenkoi (paratype), 67—A. conjunctum (holotype), 68—A. densepunctatum (holotype), 69—A. ivani (paratype).Scale bars: 1.0 mm." httpUri="https://zenodo.org/record/1125247/files/figure.png" pageId="35" pageNumber="36">Fig. 69</figureCitation>
.
</paragraph>
<paragraph id="8BEA36CAFF989D02FF5D3894FF5EFEBD" blockId="35.[151,1437,1014,2011]" lastBlockId="36.[151,1436,151,1040]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
Head 1.5–1.6 times as wide as long; middle part of head and posterio-lateral parts of infraorbital ridges slightly elevated or distinctly flattened; middle part of head between grooves without or with small semicircular impression; grooves in front of ocelli reaching anterior third of length of eye. Apical segment of maxillary palp with moderately narrow basal part slightly widened medially. Measurements of antennomeres (
<typeStatus id="54EE8868FF9F9D02FB5D3F89FA83FF05" box="[1223,1339,223,248]" pageId="36" pageNumber="37" type="holotype">holotypes</typeStatus>
): 1: 0.20 × 0.07; 2: 0.12 × 0.05; 3: 0.17 × 0.05; 4–6: 0.13 × 0.06; 7–8: 0.14 × 0.06; 9: 0.14 × 0.07; 10: 0.13 × 0.07; 11: 0.20 × 0.07.
</paragraph>
<paragraph id="8BEA36CAFF9F9D02FF5D3E1AFD4AFE75" blockId="36.[151,1436,151,1040]" pageId="36" pageNumber="37">Middle elevation of pronotum markedly convex, with deep semicircular impression in medio-basal third and sometimes with very convex parts behind impression.</paragraph>
<paragraph id="8BEA36CAFF9F9D02FF5D3EC2FE3FFE2D" blockId="36.[151,1436,151,1040]" pageId="36" pageNumber="37">Elytra approximately as wide as long, reaching basal margin of abdominal tergite V; lateral portions narrow, distinctly explanated.</paragraph>
<paragraph id="8BEA36CAFF9F9D02FF5D3E8AFDE4FD9D" blockId="36.[151,1436,151,1040]" pageId="36" pageNumber="37">
Male. Apical margins of abdominal tergite VIII and sternite VIII slightly sinuate. Aedeagus (
<figureCitation id="136E2A4FFF9F9D02FA993E8AFAE0FE09" box="[1283,1368,476,501]" captionStart="FIGURES 80–83" captionStartId="33.[151,264,1971,1994]" captionTargetBox="[274,1291,206,1932]" captionTargetId="figure@33.[251,1339,193,1957]" captionTargetPageId="33" captionText="FIGURES 80–83. Aedeagus of Anthobium densepunctatum (80–81—paratype), A. ivani (82–83—paratype): 80, 82— parameral view, 81, 83—ateral view. Scale bars:0.1 mm." httpUri="https://zenodo.org/record/1125261/files/figure.png" pageId="36" pageNumber="37">Fig. 82</figureCitation>
) very narrow, with more or less rounded apical part of median lobe; parameres wide and asymmetrical, with very wide left setiferous lobe, both lobes significantly curved mediad, with long apical and preapical setae; internal sac narrow. Aedeagus laterally as in
<figureCitation id="136E2A4FFF9F9D02FD9C3D11FDE0FD9D" box="[518,600,583,608]" captionStart="FIGURES 80–83" captionStartId="33.[151,264,1971,1994]" captionTargetBox="[274,1291,206,1932]" captionTargetId="figure@33.[251,1339,193,1957]" captionTargetPageId="33" captionText="FIGURES 80–83. Aedeagus of Anthobium densepunctatum (80–81—paratype), A. ivani (82–83—paratype): 80, 82— parameral view, 81, 83—ateral view. Scale bars:0.1 mm." httpUri="https://zenodo.org/record/1125261/files/figure.png" pageId="36" pageNumber="37">Fig. 83</figureCitation>
.
</paragraph>
<paragraph id="8BEA36CAFF9F9D02FF5D3D3AFB2FFD78" blockId="36.[151,1436,151,1040]" box="[199,1175,620,645]" pageId="36" pageNumber="37">Female. Apical margins of abdominal tergite VIII and sternite VIII rounded or straight.</paragraph>
</subSubSection>
<subSubSection id="C34F6541FF9F9D02FF5D3DD9FE40FCE9" pageId="36" pageNumber="37" type="diagnosis">
<paragraph id="8BEA36CAFF9F9D02FF5D3DD9FE40FCE9" blockId="36.[151,1436,151,1040]" pageId="36" pageNumber="37">
<emphasis id="B921EAD8FF9F9D02FF5D3DD9FE00FD55" bold="true" box="[199,440,655,680]" pageId="36" pageNumber="37">Comparative notes.</emphasis>
Based on the length of the body, shape of the pronotum with deep medio-basal impression,
<taxonomicName id="4C554D49FF9F9D02FE853DE2FECEFD31" authority="Shavrin &amp; Smetana, 2017" authorityName="Shavrin &amp; Smetana" authorityYear="2017" box="[287,374,692,716]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="ivani" status="sp. nov.">
<emphasis id="B921EAD8FF9F9D02FE853DE2FECEFD31" box="[287,374,692,716]" italics="true" pageId="36" pageNumber="37">A. ivani</emphasis>
</taxonomicName>
<emphasis id="B921EAD8FF9F9D02FEE63DE2FE0CFD30" bold="true" box="[380,436,692,717]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="A21257A3FF9F9D02FEE63DE2FE0CFD30" box="[380,436,692,717]" pageId="36" pageNumber="37" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="4C554D49FF9F9D02FDDF3DE2FD0DFD31" authorityName=": Smetana" authorityYear="2004" box="[581,693,692,716]" class="Insecta" family="Staphylinidae" genus="Anthobium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="nigrum">
<emphasis id="B921EAD8FF9F9D02FDDF3DE2FD0DFD31" box="[581,693,692,716]" italics="true" pageId="36" pageNumber="37">A. nigrum</emphasis>
</taxonomicName>
, from which it differs by shorter elytra, narrower apical part of the median lobe of the aedeagus, wider setiferous lobes of parameres significantly curved mediad, and by the details of the structure of the internal sac.
</paragraph>
</subSubSection>
<subSubSection id="C34F6541FF9F9D02FF5D3C49FEC6FCA0" pageId="36" pageNumber="37" type="distribution">
<paragraph id="8BEA36CAFF9F9D02FF5D3C49FEC6FCA0" blockId="36.[151,1436,151,1040]" pageId="36" pageNumber="37">
<emphasis id="B921EAD8FF9F9D02FF5D3C49FED8FCC5" bold="true" box="[199,352,799,824]" pageId="36" pageNumber="37">Distribution.</emphasis>
The new species is known from several localities (
<figureCitation id="136E2A4FFF9F9D02FC003C49FC53FCC5" box="[922,1003,799,824]" captionStart="FIGURE 74" captionStartId="28.[151,250,982,1004]" captionTargetBox="[179,1408,193,961]" captionTargetId="figure@28.[179,1408,193,961]" captionTargetPageId="28" captionText="FIGURE 74.Distribution of Anthobium nigrum (black circles) and A. ivani (white circles) in the Himalayan Region." httpUri="https://zenodo.org/record/1125251/files/figure.png" pageId="36" pageNumber="37">Fig. 74</figureCitation>
) in central and northeastern
<collectingCountry id="F342765AFF9F9D02FAB03C49FAD5FCC5" box="[1322,1389,799,824]" name="Nepal" pageId="36" pageNumber="37">Nepal</collectingCountry>
and
<collectingCountry id="F342765AFF9F9D02FF0D3C12FEC3FCA0" box="[151,379,836,861]" name="Bhutan" pageId="36" pageNumber="37">Kingdom of Bhutan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F6541FF9F9D02FF5D3C31FE49FC35" pageId="36" pageNumber="37" type="biology_ecology">
<paragraph id="8BEA36CAFF9F9D02FF5D3C31FE49FC35" blockId="36.[151,1436,151,1040]" pageId="36" pageNumber="37">
<emphasis id="B921EAD8FF9F9D02FF5D3C31FEF2FC7D" bold="true" box="[199,330,871,896]" pageId="36" pageNumber="37">Bionomics.</emphasis>
Specimens from
<collectingCountry id="F342765AFF9F9D02FD8A3C31FDEBFC7D" box="[528,595,871,896]" name="Nepal" pageId="36" pageNumber="37">Nepal</collectingCountry>
were collected at elevations
<quantity id="4CAD9B2FFF9F9D02FC003C3EFBF4FC82" box="[922,1100,872,896]" metricMagnitude="3" metricUnit="m" metricValue="3.0" metricValueMax="3.3" metricValueMin="2.7" pageId="36" pageNumber="37" unit="m" value="3000.0" valueMax="3300.0" valueMin="2700.0">2700 to 3300 m</quantity>
a.s.l. by splashing water over gravel and stony banks of a creek, mostly in shaded areas, together with
<taxonomicName id="4C554D49FF9F9D02FB913CDAFB18FC59" box="[1035,1184,908,932]" class="Insecta" family="Staphylinidae" genus="Ochthephilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAD8FF9F9D02FB913CDAFB18FC59" box="[1035,1184,908,932]" italics="true" pageId="36" pageNumber="37">Ochthephilus</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="4C554D49FF9F9D02FB7B3CDAFAD8FC58" authorityName="Fleming" authorityYear="1821" box="[1249,1376,908,933]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="subFamily" subFamily="Oxytelinae">Oxytelinae</taxonomicName>
) and
<taxonomicName id="4C554D49FF9F9D02FF0D3CE6FEBFFC3A" authorityName="Thomson" authorityYear="1859" box="[151,263,944,967]" class="Insecta" family="Hydrophilidae" genus="Anacaena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAD8FF9F9D02FF0D3CE6FEBFFC3A" box="[151,263,944,967]" italics="true" pageId="36" pageNumber="37">Anacaena</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="4C554D49FF9F9D02FEA63CF9FE5DFC35" authorityName="Latreille" authorityYear="1802" box="[316,485,943,968]" class="Insecta" family="Hydrophilidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="family">Hydrophilidae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C34F6541FF9F9D02FF5D3C82FDB7FBED" pageId="36" pageNumber="37" type="etymology">
<paragraph id="8BEA36CAFF9F9D02FF5D3C82FDB7FBED" blockId="36.[151,1436,151,1040]" pageId="36" pageNumber="37">
<emphasis id="B921EAD8FF9F9D02FF5D3C82FEF6FC10" bold="true" box="[199,334,980,1005]" pageId="36" pageNumber="37">Etymology.</emphasis>
Patronymic, the species is named to honour our friend and colleague, the eminent coleopterist Ivan Löbl (
<collectingRegion id="4991F828FF9F9D02FE893CAEFED5FBED" box="[275,365,1016,1040]" country="Switzerland" name="Geneve" pageId="36" pageNumber="37">Genève</collectingRegion>
,
<collectingCountry id="F342765AFF9F9D02FEE23CA1FDBBFBED" box="[376,515,1015,1040]" name="Switzerland" pageId="36" pageNumber="37">Switzerland</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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