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<document id="579FF2D2FD91D120D31B10FE4201094B" ID-DOI="10.5281/zenodo.282718" ID-GBIF-Dataset="23edf4a6-4cc2-4d80-a98e-d91c9ee2ff67" ID-ISSN="1175-5326" ID-Zenodo-Dep="282718" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460548329905" checkinUser="plazi" docAuthor="Shamshev, Igor &amp; Grootaert, Patrick" docDate="2012" docId="03FC87D26A66FFB9288AFD3AFD4BFB1D" docLanguage="en" docName="zt03525p050.pdf" docOrigin="Zootaxa 3525" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Platypalpus layiaphilus Shamshev &amp; Grootaert, sp. nov." docType="treatment" docVersion="8" lastPageNumber="44" masterDocId="FFC5FFAA6A6EFFB0281DFFE7FF89FFB8" masterDocTitle="Proposed change in status of the Nearctic genus Charadrodromia Melander (Diptera: Hybotidae, Tachydromiinae), with description of four new species" masterLastPageNumber="50" masterPageNumber="35" pageNumber="43" updateTime="1698317995506" updateUser="plazi">
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<mods:title id="59BB9486FAE9EAD7548D9AFEE32B5B06">Proposed change in status of the Nearctic genus Charadrodromia Melander (Diptera: Hybotidae, Tachydromiinae), with description of four new species</mods:title>
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<mods:namePart id="8FB70A5D405EEE6EC2B15429A31DD5FE">Shamshev, Igor</mods:namePart>
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<mods:namePart id="9F3417B928D237D0F9255705DDAAF5BC">Grootaert, Patrick</mods:namePart>
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<mods:date id="D7F2163172E930C7A36F8DE4ACD68A70">2012</mods:date>
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<treatment id="03FC87D26A66FFB9288AFD3AFD4BFB1D" ID-DOI="http://doi.org/10.5281/zenodo.6168210" ID-GBIF-Taxon="119529107" ID-Zenodo-Dep="6168210" LSID="urn:lsid:plazi:treatment:03FC87D26A66FFB9288AFD3AFD4BFB1D" httpUri="http://treatment.plazi.org/id/03FC87D26A66FFB9288AFD3AFD4BFB1D" lastPageId="9" lastPageNumber="44" pageId="8" pageNumber="43">
<subSubSection id="C34F654F6A66FFB8288AFD3AFEA4FCA0" pageId="8" pageNumber="43" type="nomenclature">
<paragraph id="8BEA36C46A66FFB8288AFD3AFCC9FD4F" blockId="8.[151,832,733,792]" box="[151,832,733,759]" pageId="8" pageNumber="43">
<heading id="D0A281A86A66FFB8288AFD3AFCC9FD4F" bold="true" box="[151,832,733,759]" fontSize="11" level="1" pageId="8" pageNumber="43" reason="1">
<emphasis id="B921EAD66A66FFB8288AFD3AFCC9FD4F" bold="true" box="[151,832,733,759]" pageId="8" pageNumber="43">
<taxonomicName id="4C554D476A66FFB8288AFD3AFD52FD4F" ID-CoL="9HTZX" authority="Shamshev &amp; Grootaert" authorityName="Shamshev &amp; Grootaert" box="[151,731,733,759]" class="Insecta" family="Hybotidae" genus="Platypalpus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="layiaphilus" status="sp. nov.">
<emphasis id="B921EAD66A66FFB8288AFD3AFE38FD4F" bold="true" box="[151,433,733,759]" italics="true" pageId="8" pageNumber="43">Platypalpus layiaphilus</emphasis>
Shamshev &amp; Grootaert
</taxonomicName>
<taxonomicNameLabel id="A21257AD6A66FFB82AFFFD3AFCC9FD4F" box="[738,832,733,759]" pageId="8" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEA36C46A66FFB8288AFCE7FEA4FCA0" blockId="8.[151,832,733,792]" box="[151,301,768,792]" pageId="8" pageNumber="43">
(
<figureCitation id="136E2A416A66FFB82882FCE7FEACFCA0" box="[159,293,768,792]" captionStart="FIGURES 15 17" captionStartId="8.[151,263,1683,1706]" captionTargetBox="[176,1423,1073,1657]" captionTargetId="figure@8.[153,1438,1061,1662]" captionTargetPageId="8" captionText="FIGURES 15 17. Platypalpus layiaphilus sp. nov., male. 15. right epandrial lamella, lateral view. 16. cerci and epandrium, dorsal view. 17. left epandrial lamella and left cercus, lateral view." httpUri="https://zenodo.org/record/282723/files/figure.png" pageId="8" pageNumber="43">Figs. 1517</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34F654F6A66FFB8288AFCAFFCDCF896" pageId="8" pageNumber="43" type="materials_examined">
<paragraph id="8BEA36C46A66FFB8288AFCAFFED9FC48" blockId="8.[151,1436,839,1008]" pageId="8" pageNumber="43">
<emphasis id="B921EAD66A66FFB8288AFCAFFEB4FCD8" bold="true" box="[151,317,839,864]" pageId="8" pageNumber="43">
<typeStatus id="54EE88666A66FFB8288AFCAFFF59FCD8" box="[151,208,840,864]" pageId="8" pageNumber="43">Type</typeStatus>
material
</emphasis>
:
<typeStatus id="54EE88666A66FFB82952FCA0FE64FCD8" box="[335,493,839,864]" pageId="8" pageNumber="43" type="holotype">
<emphasis id="B921EAD66A66FFB82952FCA0FE64FCD8" bold="true" box="[335,493,839,864]" pageId="8" pageNumber="43">HOLOTYPE</emphasis>
</typeStatus>
, 3 labelled:
<collectingCountry id="F34276546A66FFB82A99FCAFFD5DFCD8" box="[644,724,840,864]" name="United States of America" pageId="8" pageNumber="43">U.S.A.</collectingCountry>
: CALIFORNIA: / Mendocino County, Hop- / land, University of California Hopland / Field Station /
<date id="FFEB10046A66FFB82A54FC8BFD57FC3C" box="[585,734,876,901]" pageId="8" pageNumber="43" value="1986-05-20">20 May 1986</date>
/ Paul H. Arnaud, Jr.; Two
<taxonomicName id="4C554D476A66FFB82C0FFC8BFB02FC3C" box="[1042,1163,876,900]" class="Insecta" family="Empididae" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="family">Empididae</taxonomicName>
in one / flower of
<taxonomicName id="4C554D476A66FFB82D41FC8BFA15FC3C" box="[1372,1436,876,900]" class="Magnoliopsida" family="Asteraceae" genus="Layia" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="44" phylum="Tracheophyta" rank="genus">Layia</taxonomicName>
platy- / glosa ssp. campestris / Keck (det. B. Bartholomew) / seepage along gravel / road, midday;
<typeStatus id="54EE88666A66FFB82CFCFC77FAC0FC10" box="[1249,1353,912,936]" pageId="8" pageNumber="43" type="holotype">Holotype</typeStatus>
[pink];
<typeStatus id="54EE88666A66FFB8288AFC53FF76FC74" box="[151,255,948,972]" pageId="8" pageNumber="43" type="holotype">Holotype</typeStatus>
/
<taxonomicName id="4C554D476A66FFB8290CFC53FD93FC74" box="[273,538,948,972]" class="Insecta" family="Hybotidae" genus="Platypalpus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="layiaphilus">Platypalpus layiaphilus</taxonomicName>
/ Shamshev et Grootaert sp. nov. (
<collectionCode id="ED44AE016A66FFB82BB6FC53FC6EFC74" LSID="urn:lsid:biocol.org:col:34700" box="[939,999,948,972]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="8" pageNumber="43">CAS</collectionCode>
).
<emphasis id="B921EAD66A66FFB82BE0FC53FB1CFC75" bold="true" box="[1021,1173,948,973]" pageId="8" pageNumber="43">
<typeStatus id="54EE88666A66FFB82BE0FC53FB1CFC75" box="[1021,1173,948,973]" pageId="8" pageNumber="43" type="paratype">PARATYPE</typeStatus>
</emphasis>
. 1 3, same data as in
<typeStatus id="54EE88666A66FFB8288AFC3FFF70FC48" box="[151,249,984,1008]" pageId="8" pageNumber="43" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED44AE016A66FFB82915FC3FFECDFC48" LSID="urn:lsid:biocol.org:col:34700" box="[264,324,984,1008]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="8" pageNumber="43">CAS</collectionCode>
).
</paragraph>
<caption id="DF2A664C6A66FFB8288AF974FCBEF970" httpUri="https://zenodo.org/record/282723/files/figure.png" pageId="8" pageNumber="43" targetBox="[176,1423,1073,1657]" targetPageId="8">
<paragraph id="8BEA36C46A66FFB8288AF974FCBEF970" blockId="8.[151,1436,1683,1736]" pageId="8" pageNumber="43">
<emphasis id="B921EAD66A66FFB8288AF974FEC7F911" bold="true" box="[151,334,1683,1706]" pageId="8" pageNumber="43">FIGURES 1517</emphasis>
.
<taxonomicName id="4C554D476A66FFB82941F973FDC4F912" box="[348,589,1684,1706]" class="Insecta" family="Hybotidae" genus="Platypalpus" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="layiaphilus" status="sp. nov.">
<emphasis id="B921EAD66A66FFB82941F973FDC4F912" box="[348,589,1684,1706]" italics="true" pageId="8" pageNumber="43">Platypalpus layiaphilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257AD6A66FFB82A4BF973FD20F911" box="[598,681,1684,1705]" pageId="8" pageNumber="43" rank="species">
<emphasis id="B921EAD66A66FFB82A4BF973FD20F911" bold="true" box="[598,681,1684,1705]" pageId="8" pageNumber="43">sp. nov.</emphasis>
</taxonomicNameLabel>
, male.
<emphasis id="B921EAD66A66FFB82AEAF973FC99F911" bold="true" box="[759,784,1684,1705]" pageId="8" pageNumber="43">15</emphasis>
. right epandrial lamella, lateral view.
<emphasis id="B921EAD66A66FFB82C81F973FB3CF911" bold="true" box="[1180,1205,1684,1705]" pageId="8" pageNumber="43">16</emphasis>
. cerci and epandrium, dorsal view.
<emphasis id="B921EAD66A66FFB8290AF954FEB9F970" bold="true" box="[279,304,1715,1736]" pageId="8" pageNumber="43">17</emphasis>
. left epandrial lamella and left cercus, lateral view.
</paragraph>
</caption>
<paragraph id="8BEA36C46A66FFB828DAF915FCDCF896" blockId="8.[151,1436,1778,2019]" pageId="8" pageNumber="43">
<emphasis id="B921EAD66A66FFB828DAF915FEDFF8B3" bold="true" box="[199,342,1778,1803]" pageId="8" pageNumber="43">Recognition</emphasis>
. This species is distinguished by palpus and antenna brown; postpedicel nearly as long as wide; fore tarsi yellow; dorsocentrals arranged in 23 irregular rows.
</paragraph>
</subSubSection>
<subSubSection id="C34F654F6A66FFB928DAF8DDFE2BFC49" lastPageId="9" lastPageNumber="44" pageId="8" pageNumber="43" type="description">
<paragraph id="8BEA36C46A66FFB928DAF8DDFF66FF6C" blockId="8.[151,1436,1778,2019]" lastBlockId="9.[151,1437,151,1189]" lastPageId="9" lastPageNumber="44" pageId="8" pageNumber="43">
<emphasis id="B921EAD66A66FFB828DAF8DDFED8F8EB" bold="true" box="[199,337,1850,1875]" pageId="8" pageNumber="43">Description</emphasis>
. Body length
<quantity id="4CAD9B216A66FFB829EDF8DDFDF3F8EA" box="[496,634,1850,1874]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="8" pageNumber="43" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.31.4 mm</quantity>
; wing length
<quantity id="4CAD9B216A66FFB82B0CF8DDFCE3F8EA" box="[785,874,1850,1874]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="43" unit="mm" value="1.4">1.4 mm</quantity>
.
<emphasis id="B921EAD66A66FFB82B69F8DCFC39F8EB" bold="true" box="[884,944,1851,1875]" pageId="8" pageNumber="43">Male</emphasis>
. Head black in ground-colour. Frons broad, finely brownish grey pollinose, rather subshining, widened toward ocellar tubercle. Face narrow, concolorous with frons, denser pollinose. Ocellar tubercle with 2 long brownish proclinate anterior and 2 short lateroclinate posterior bristles. Two pairs of long brownish verticals. Occiput densely greyish pollinose, with numerous brownish yellow to pale setae of different lengths. Antenna brown; postpedicel, short, nearly as long as wide (1.3X in
<typeStatus id="54EE88666A66FFB82D34F82DFA19F85A" box="[1321,1424,1994,2018]" pageId="8" pageNumber="43" type="holotype">holotype</typeStatus>
); stylus 2.0X as long as postpedicel. Proboscis brown. Palpus small, brown, with several fine setae of different lengths.
</paragraph>
<paragraph id="8BEA36C46A67FFB928DAFF38FABDFE14" blockId="9.[151,1437,151,1189]" pageId="9" pageNumber="44">Thorax black in ground-colour, entirely finely greyish pollinose, somewhat denser on pleurae, katepisternum without shining spot; with yellowish to pale setation. Postpronotal lobe with 1 long inclinate bristle and several short setae. Mesonotum with 1 presutural supra-alar, 2 notopleurals, 1 postalar and 4 scutellar bristles (apical pair very long and cruciate, lateral pair short); additionally, notopleural depression with several setulae and similar setulae present on supra-alar space; acrostichals arranged in 2 irregular rows, short, distinctly separated from dorsocentrals by bare space; dorsocentrals arranged in 23 irregular rows, short, 2 prescutellar pairs long.</paragraph>
<paragraph id="8BEA36C46A67FFB928DAFE5FFAE6FDE7" blockId="9.[151,1437,151,1189]" pageId="9" pageNumber="44">
Legs entirely yellow to dusky yellow. Fore femur thickened, mostly covered with short setae, bearing rows of yellow anteroventral and posteroventral setae becoming longer basally. Fore tibia slightly thickened. Mid femur somewhat narrower than fore femur; with short black spinule-like anteroventral and longer posteroventral setae. Mid tibia without apical spur, with several stronger black setulae subapically.
<collectingCountry id="F34276546A67FFB92C18FDC3FBB4FD84" box="[1029,1085,548,572]" name="India" pageId="9" pageNumber="44">Hind</collectingCountry>
femur with short anteroventral setae.
<collectingCountry id="F34276546A67FFB928C0FDA0FE9CFDE7" box="[221,277,583,607]" name="India" pageId="9" pageNumber="44">Hind</collectingCountry>
tibia unmodified, bearing 1 short anterodorsal subapical seta, otherwise covered with short setulae.
</paragraph>
<paragraph id="8BEA36C46A67FFB928DAFD8BFC18FD69" blockId="9.[151,1437,151,1189]" pageId="9" pageNumber="44">Wing hyaline, with brownish yellow to pale yellow veins. One long brown costal bristle. Veins R4+5 and M straight and parallel near wing margin; CuA2 somewhat concave; cells br and bm contiguous, rather short, bm broader than br. Squama yellow, with pale setae. Halter pale yellow.</paragraph>
<paragraph id="8BEA36C46A67FFB928DAFD3AFB5CFC75" blockId="9.[151,1437,151,1189]" pageId="9" pageNumber="44">
Abdomen brown in ground-colour, finely greyish pollinose, viewed dorsally rather subshining; covered with short pale setae. Terminalia (
<figureCitation id="136E2A416A67FFB929C0FCE6FDEDFCA1" box="[477,612,769,793]" captionStart="FIGURES 15 17" captionStartId="8.[151,263,1683,1706]" captionTargetBox="[176,1423,1073,1657]" captionTargetId="figure@8.[153,1438,1061,1662]" captionTargetPageId="8" captionText="FIGURES 15 17. Platypalpus layiaphilus sp. nov., male. 15. right epandrial lamella, lateral view. 16. cerci and epandrium, dorsal view. 17. left epandrial lamella and left cercus, lateral view." httpUri="https://zenodo.org/record/282723/files/figure.png" pageId="9" pageNumber="44">Figs. 1517</figureCitation>
) rather large, concolorous with abdomen; left cercus broadened apically, with several unmodified setae of different lengths; right cercus digitiform, narrow, somewhat longer than left cercus, with several unmodified setae of different lengths; left epandrial lamella subrectangular basally, produced with finger-like projection apically and mid-ventral tubercle; covered with numerous setae of different lengths on apical part and with several long setae sitting on ventral tubercle; right epandrial lamella subconical, with numerous unmodified setae of different lengths; right surstylus undifferentiated from epandrium.
</paragraph>
<paragraph id="8BEA36C46A67FFB928DAFC3DFE2BFC49" blockId="9.[151,1437,151,1189]" box="[199,418,985,1010]" pageId="9" pageNumber="44">
<emphasis id="B921EAD66A67FFB928DAFC3DFE97FC4A" bold="true" box="[199,286,986,1010]" pageId="9" pageNumber="44">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34F654F6A67FFB928DAFC19FED3FB81" pageId="9" pageNumber="44" type="etymology">
<paragraph id="8BEA36C46A67FFB928DAFC19FED3FB81" blockId="9.[151,1437,151,1189]" pageId="9" pageNumber="44">
<emphasis id="B921EAD66A67FFB928DAFC19FECFFBAE" bold="true" box="[199,326,1022,1046]" pageId="9" pageNumber="44">Etymology</emphasis>
. The name of the new species reflects that it was collected in a flower of
<taxonomicName id="4C554D476A67FFB92CB4FC18FAE1FBAD" box="[1193,1384,1021,1046]" class="Magnoliopsida" family="Asteraceae" genus="Layia" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="44" phylum="Tracheophyta" rank="species" species="platyglosa">
<emphasis id="B921EAD66A67FFB92CB4FC18FAE1FBAD" box="[1193,1384,1021,1046]" italics="true" pageId="9" pageNumber="44">Layia platyglosa</emphasis>
</taxonomicName>
ssp.
<emphasis id="B921EAD66A67FFB9288AFBC5FE99FB81" box="[151,272,1058,1081]" italics="true" pageId="9" pageNumber="44">campestris</emphasis>
Keck.
</paragraph>
</subSubSection>
<subSubSection id="C34F654F6A67FFB928DAFBA2FDBBFBE5" box="[199,562,1093,1118]" pageId="9" pageNumber="44" type="distribution">
<paragraph id="8BEA36C46A67FFB928DAFBA2FDBBFBE5" blockId="9.[151,1437,151,1189]" box="[199,562,1093,1118]" pageId="9" pageNumber="44">
<emphasis id="B921EAD66A67FFB928DAFBA2FED0FBE6" bold="true" box="[199,345,1093,1118]" pageId="9" pageNumber="44">Distribution</emphasis>
.
<collectingCountry id="F34276546A67FFB9297AFBA2FE16FBE5" box="[359,415,1093,1117]" name="United States of America" pageId="9" pageNumber="44">USA</collectingCountry>
(California).
</paragraph>
</subSubSection>
<subSubSection id="C34F654F6A67FFB928DAFB8DFD4BFB1D" pageId="9" pageNumber="44" type="discussion">
<paragraph id="8BEA36C46A67FFB928DAFB8DFD4BFB1D" blockId="9.[151,1437,151,1189]" pageId="9" pageNumber="44">
<emphasis id="B921EAD66A67FFB928DAFB8DFEBDFB3A" bold="true" box="[199,308,1130,1154]" pageId="9" pageNumber="44">Remarks</emphasis>
.
<taxonomicName id="4C554D476A67FFB9295BFB8EFDDDFB39" box="[326,596,1129,1153]" class="Insecta" family="Hybotidae" genus="Platypalpus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="layiaphilus" status="sp. nov.">
<emphasis id="B921EAD66A67FFB9295BFB8EFDDDFB39" box="[326,596,1129,1153]" italics="true" pageId="9" pageNumber="44">Platypalpus layiaphilus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257AD6A67FFB92A42FB8DFD34FB3A" box="[607,701,1130,1154]" pageId="9" pageNumber="44" rank="species">
<emphasis id="B921EAD66A67FFB92A42FB8DFD34FB3A" bold="true" box="[607,701,1130,1154]" pageId="9" pageNumber="44">sp. nov.</emphasis>
</taxonomicNameLabel>
is most similar to
<taxonomicName id="4C554D476A67FFB92BBFFB8DFBB8FB39" box="[930,1073,1130,1153]" class="Insecta" family="Hybotidae" genus="Platypalpus" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="cummingi" status="sp. nov.">
<emphasis id="B921EAD66A67FFB92BBFFB8DFC39FB39" box="[930,944,1130,1153]" italics="true" pageId="9" pageNumber="44">P</emphasis>
.
<emphasis id="B921EAD66A67FFB92BDEFB8DFBB8FB39" box="[963,1073,1130,1153]" italics="true" pageId="9" pageNumber="44">cummingi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21257AD6A67FFB92C21FB8DFB12FB3A" box="[1084,1179,1130,1154]" pageId="9" pageNumber="44" rank="species">
<emphasis id="B921EAD66A67FFB92C21FB8DFB12FB3A" bold="true" box="[1084,1179,1130,1154]" pageId="9" pageNumber="44">sp. nov.</emphasis>
</taxonomicNameLabel>
The main differences between these species are given below in the key.
</paragraph>
</subSubSection>
</treatment>
</document>