treatments-xml/data/03/FC/87/03FC87CAFFD6C971FF5BF8C82D34FEEE.xml

<document id="DBB6EE9F1184E8A4236F3D91E0268B5B" ID-DOI="10.5281/zenodo.293430" ID-GBIF-Dataset="3052ba2b-81b1-4ef4-9c21-4ea496f4d1d5" ID-ISSN="1175-5326" ID-Zenodo-Dep="293430" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460479550326" checkinUser="plazi" docAuthor="Szederjesi, Tímea &amp; Csuzdi, Csaba" docDate="2012" docId="03FC87CAFFD6C971FF5BF8C82D34FEEE" docLanguage="en" docName="zt03304p042.pdf" docOrigin="Zootaxa 3304" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Dendrobaena retrosella Szederjesi &amp; Csuzdi, 2012, sp. nov." docType="treatment" docVersion="12" lastPageNumber="34" masterDocId="FFC5FFB2FFDEC97BFFCCFFD1284DFFF2" masterDocTitle="New and little known earthworm species from Greece (Oligochaeta: Lumbricidae, Acanthodrilidae)" masterLastPageNumber="42" masterPageNumber="25" pageNumber="33" updateTime="1698307382221" updateUser="plazi">
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<mods:title id="C430CC54A0329BE39E4309F16DD047E0">New and little known earthworm species from Greece (Oligochaeta: Lumbricidae, Acanthodrilidae)</mods:title>
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<mods:namePart id="5E76AB2F3D22CADCF27D012495EEEFF4">Csuzdi, Csaba</mods:namePart>
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<emphasis id="B921EACEFFD6C973FF5BF8C82A54F8C1" bold="true" box="[151,537,1817,1843]" pageId="8" pageNumber="33">
<taxonomicName id="4C554D5FFFD6C973FF5BF8C829F8F8C1" box="[151,437,1817,1843]" class="Clitellata" family="Lumbricidae" genus="Dendrobaena" kingdom="Animalia" order="Crassiclitellata" pageId="8" pageNumber="33" phylum="Annelida" rank="species" species="retrosella" status="sp. nov.">
<emphasis id="B921EACEFFD6C973FF5BF8C829F8F8C1" bold="true" box="[151,437,1817,1843]" italics="true" pageId="8" pageNumber="33">Dendrobaena retrosella</emphasis>
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<taxonomicNameLabel id="A21257B5FFD6C973FE70F8C82A54F8C1" box="[444,537,1817,1843]" pageId="8" pageNumber="33" rank="species">sp. nov.</taxonomicNameLabel>
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.
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/15914 adult. 
<emphasis id="B921EACEFFD6C973FDC8F8B02A1DF88A" box="[516,592,1888,1912]" italics="true" pageId="8" pageNumber="33">No. 15</emphasis>
, Trikala County, Lakmos Mts., Chaliki, open torrent and brook, and rocky subalpine grassland N of the village, 
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, 
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, leg. J. Kontschán, D. Murányi, T. Szederjesi, Zs. Ujvári, 0 9.0 5.2011.
</paragraph>
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.
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/15915 one adult ex., locality and date same as that of the 
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.
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<subSubSection id="C34F6557FFD7C972FF05FF462931FF26" pageId="9" pageNumber="34" type="etymology">
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<emphasis id="B921EACEFFD7C972FF05FF4629EAFF42" bold="true" box="[201,423,151,176]" pageId="9" pageNumber="34">Derivatio nominis.</emphasis>
From Latin (retro = backward, sella = saddle), referring to the backward displaced position of the clitellum.
</paragraph>
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<subSubSection id="C34F6557FFD7C972FF05FF0E2BADFD02" pageId="9" pageNumber="34" type="diagnosis">
<paragraph id="8BEA36DCFFD7C972FF05FF0E2C34FE96" blockId="9.[151,1436,151,932]" pageId="9" pageNumber="34">
<emphasis id="B921EACEFFD7C972FF05FF0E290EFF0A" bold="true" box="[201,323,223,248]" pageId="9" pageNumber="34">Diagnosis.</emphasis>
Length 
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, diameter 
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, setae distant. Pigmentation red-violet. First dorsal pores 5/6. Clitellum on 29–36, tubercles 31–1/n34. Male pore on 15, large. Nephridial pores regularly alternate between 
<emphasis id="B921EACEFFD7C972FA93FED52D20FEEE" box="[1375,1389,260,284]" italics="true" pageId="9" pageNumber="34">b</emphasis>
and above 
<emphasis id="B921EACEFFD7C972FF2FFEF928BCFEB2" box="[227,241,296,320]" italics="true" pageId="9" pageNumber="34">d</emphasis>
. Three pairs of vesicles in 9, 11, 12; spermathecae 
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9/10–10/ 
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</date>
in 
<emphasis id="B921EACEFFD7C972FC24FEF92BBBFEB2" box="[1000,1014,296,320]" italics="true" pageId="9" pageNumber="34">d</emphasis>
. Calciferous glands with well-developed diverticula in 11, 12. Hearts in segments 7–10, nephridial bladders sausage-shaped.
</paragraph>
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<emphasis id="B921EACEFFD7C972FF05FEA129F0FE7A" bold="true" box="[201,445,368,392]" pageId="9" pageNumber="34">External characters.</emphasis>
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<typeStatus id="54EE887EFFD7C972FE09FEA12A61FE7A" box="[453,556,368,392]" pageId="9" pageNumber="34" type="holotype">Holotype</typeStatus>
</emphasis>
<quantity id="4CAD9B39FFD7C972FDFFFEBE2ACEFE75" box="[563,643,367,391]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="9" pageNumber="34" unit="mm" value="35.0">35 mm</quantity>
long and 
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wide; number of segments 107. 
<emphasis id="B921EACEFFD7C972FB69FEA02D41FE7A" box="[1189,1292,369,392]" italics="true" pageId="9" pageNumber="34">
<typeStatus id="54EE887EFFD7C972FB69FEA02D41FE7A" box="[1189,1292,369,392]" pageId="9" pageNumber="34" type="paratype">Paratype</typeStatus>
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<quantity id="4CAD9B39FFD7C972FAD8FEBE2D2EFE75" box="[1300,1379,367,391]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="9" pageNumber="34" unit="mm" value="37.0">37 mm</quantity>
long and 
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wide; number of segments 145. Colour red-violet, especially dark on dorsum. Prostomium epilobous ½ closed. First dorsal pore in intersegmental furrow 5/6. Setae distant. Setal arrangement behind clitellum: 
<emphasis id="B921EACEFFD7C972FF5BFE0C28FEFE06" box="[151,179,477,500]" italics="true" pageId="9" pageNumber="34">aa</emphasis>
: 
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: 
<emphasis id="B921EACEFFD7C972FF12FE0D28B5FE06" box="[222,248,476,500]" italics="true" pageId="9" pageNumber="34">bc</emphasis>
: 
<emphasis id="B921EACEFFD7C972FECDFE0D2956FE06" box="[257,283,476,500]" italics="true" pageId="9" pageNumber="34">cd</emphasis>
: 
<emphasis id="B921EACEFFD7C972FEEFFE0D2972FE06" box="[291,319,476,500]" italics="true" pageId="9" pageNumber="34">dd</emphasis>
= 1.3:1:1.2:1.1:2.4. Male pore large on 15, between setae 
<emphasis id="B921EACEFFD7C972FC1FFE0D2BB6FE06" box="[979,1019,476,500]" italics="true" pageId="9" pageNumber="34">b–c</emphasis>
, surrounded by a glandular crescent. Nephridial pores regularly alternate between setal line 
<emphasis id="B921EACEFFD7C972FCD9FDD12B6EFDEA" box="[789,803,512,536]" italics="true" pageId="9" pageNumber="34">b</emphasis>
and above 
<emphasis id="B921EACEFFD7C972FC63FDD12BF0FDEA" box="[943,957,512,536]" italics="true" pageId="9" pageNumber="34">d</emphasis>
. Clitellum on segments 29–36, tubercula pubertatis on segments 31–1/n 34 (
<figureCitation id="136E2A59FFD7C972FDD2FDF52A29FDCE" box="[542,612,548,572]" captionStart="FIGURE 2" captionStartId="8.[151,250,1684,1707]" captionTargetBox="[155,1432,1179,1655]" captionTargetId="figure@8.[151,1436,1143,1662]" captionTargetPageId="8" captionText="FIGURE 2. Dendrobaena retrosella sp. nov. Ventro-lateral view of the clitellar region. cl = clitellum, mp = male pore, scale bar = 10 mm." httpUri="https://zenodo.org/record/293432/files/figure.png" pageId="9" pageNumber="34">Fig. 2</figureCitation>
). Genital papillae surround setae 
<emphasis id="B921EACEFFD7C972FC17FDF42C48FDCE" box="[987,1029,548,572]" italics="true" pageId="9" pageNumber="34">a, b</emphasis>
on segments 9, 28.
</paragraph>
<paragraph id="8BEA36DCFFD7C972FF05FD992BADFD02" blockId="9.[151,1436,151,932]" pageId="9" pageNumber="34">
<emphasis id="B921EACEFFD7C972FF05FD9929FBFD92" bold="true" box="[201,438,584,608]" pageId="9" pageNumber="34">Internal characters.</emphasis>
All septa membraneous. Testes and funnels free, paired in segments 10–11. Seminal vesicles present in 9, 11 and 12, small in segment 9. Spermathecae two pairs, clearly stalked in 9/10, 10/11 with external openings in setal line 
<emphasis id="B921EACEFFD7C972FE7FFD41298CFD5A" box="[435,449,656,680]" italics="true" pageId="9" pageNumber="34">d</emphasis>
. Calciferous glands with well-developed diverticula in 11, 12. Paired hearts in segments 7–10. Nephridial bladders simple, sausage-shaped. Crop in segments 15–16, and gizzard in segments 17–18. Typhosolis simple, lamelliform. Longitudinal muscle layer of pinnate 
<typeStatus id="54EE887EFFD7C972FC64FD092B91FD02" box="[936,988,728,752]" pageId="9" pageNumber="34">type</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C34F6557FFD7C972FF05FD2D298BFC56" pageId="9" pageNumber="34" type="discussion">
<paragraph id="8BEA36DCFFD7C972FF05FD2D298BFC56" blockId="9.[151,1436,151,932]" pageId="9" pageNumber="34">
<emphasis id="B921EACEFFD7C972FF05FD2D2970FCE6" bold="true" box="[201,317,764,788]" pageId="9" pageNumber="34">Remarks.</emphasis>
The main characteristic of the new species is the backward shift of the clitellum. The same phenomenon can be observed in three other Greek 
<taxonomicName id="4C554D5FFFD7C972FDB1FCF12B5BFCCA" box="[637,790,800,824]" class="Clitellata" family="Lumbricidae" genus="Dendrobaena" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="34" phylum="Annelida" rank="genus">
<emphasis id="B921EACEFFD7C972FDB1FCF12B5BFCCA" box="[637,790,800,824]" italics="true" pageId="9" pageNumber="34">Dendrobaena</emphasis>
</taxonomicName>
species, 
<taxonomicName id="4C554D5FFFD7C972FCB2FCF02D5BFCC5" authority="Zicsi &amp; Michalis, 1993" authorityName="Zicsi &amp; Michalis" authorityYear="1993" box="[894,1302,799,824]" class="Clitellata" family="Lumbricidae" genus="Dendrobaena" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="34" phylum="Annelida" rank="species" species="epirotica">
<emphasis id="B921EACEFFD7C972FCB2FCF02C49FCCA" box="[894,1028,801,824]" italics="true" pageId="9" pageNumber="34">D. epirotica</emphasis>
<bibRefCitation id="EFC44B2DFFD7C972FBC1FCCE2D5BFCC5" author="Zicsi" box="[1037,1302,799,823]" pageId="9" pageNumber="34" refString="Zicsi, A. &amp; Michalis, K. (1993) Zwei neue Dendrobaena-Arten aus Grichenland (Oligochaeta: Lumbricidae). Acta zoologica hungarica, 39, 301 - 310." type="journal article" year="1993">Zicsi &amp; Michalis, 1993</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C554D5FFFD7C972FAEEFCF029AEFCAE" authority="Zicsi &amp; Michalis, 1993" authorityName="Zicsi &amp; Michalis" authorityYear="1993" class="Clitellata" family="Lumbricidae" genus="Dendrobaena" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="34" phylum="Annelida" rank="species" species="pindonensis">
<emphasis id="B921EACEFFD7C972FAEEFCF02882FCAE" italics="true" pageId="9" pageNumber="34">D. pindonensis</emphasis>
<bibRefCitation id="EFC44B2DFFD7C972FF14FC9529AEFCAE" author="Zicsi" box="[216,483,836,860]" pageId="9" pageNumber="34" refString="Zicsi, A. &amp; Michalis, K. (1993) Zwei neue Dendrobaena-Arten aus Grichenland (Oligochaeta: Lumbricidae). Acta zoologica hungarica, 39, 301 - 310." type="journal article" year="1993">Zicsi &amp; Michalis, 1993</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C554D5FFFD7C972FDD2FC942B79FCAE" authority="Zicsi, 1974" authorityName="Zicsi" authorityYear="1974" box="[542,820,836,860]" class="Clitellata" family="Lumbricidae" genus="Dendrobaena" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="34" phylum="Annelida" rank="species" species="mahnerti">
<emphasis id="B921EACEFFD7C972FDD2FC942AE5FCAE" box="[542,680,836,860]" italics="true" pageId="9" pageNumber="34">D. mahnerti</emphasis>
<bibRefCitation id="EFC44B2DFFD7C972FD7DFC952B79FCAE" author="Zicsi" box="[689,820,836,860]" pageId="9" pageNumber="34" refString="Zicsi, A. (1974) Ein neue Dendrobaena-Art (Oligochaeta: Lumbricidae) aus Griechenland. Acta zoologica hungarica, 20, 449 - 451." type="journal article" year="1974">Zicsi, 1974</bibRefCitation>
</taxonomicName>
, as well as in the Turkish 
<taxonomicName id="4C554D5FFFD7C972FBAAFC942882FC8D" authority="Omodeo &amp; Rota, 1989" authorityName="Omodeo &amp; Rota" authorityYear="1989" class="Clitellata" family="Lumbricidae" genus="Dendrobaena" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="34" phylum="Annelida" rank="species" species="bruna">
<emphasis id="B921EACEFFD7C972FBAAFC942C83FCAE" box="[1126,1230,836,860]" italics="true" pageId="9" pageNumber="34">D. bruna</emphasis>
<bibRefCitation id="EFC44B2DFFD7C972FB1BFC952882FC8D" author="Omodeo" pageId="9" pageNumber="34" refString="Omodeo, P. &amp; Rota, E. (1989) Earthworms of Turkey. Bolletino di Zoologia, 56, 167 - 199." type="journal article" year="1989">Omodeo &amp; Rota, 1989</bibRefCitation>
</taxonomicName>
and in 
<taxonomicName id="4C554D5FFFD7C972FEEEFCB82A7FFC8D" authority="Karaman, 1973" authorityName="Karaman" authorityYear="1973" box="[290,562,871,896]" class="Clitellata" family="Lumbricidae" genus="Dendrobaena" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="34" phylum="Annelida" rank="species" species="zicsii">
<emphasis id="B921EACEFFD7C972FEEEFCB82933FC72" box="[290,382,873,896]" italics="true" pageId="9" pageNumber="34">D. zicsii</emphasis>
<bibRefCitation id="EFC44B2DFFD7C972FE49FCB82A7FFC8D" author="Karaman" box="[389,562,871,896]" pageId="9" pageNumber="34" refString="Karaman, S. (1973) Drugi prilog poznavanju kisnih glista Srbije. Zbornik radova Prirodno - matemati c kog fakulteta, Pristina. 1, 177 - 182." type="journal article" year="1973">Karaman, 1973</bibRefCitation>
</taxonomicName>
described from 
<collectingCountry id="F342764CFFD7C972FD25FCB62B79FC8D" box="[745,820,871,895]" name="Serbia" pageId="9" pageNumber="34">Serbia</collectingCountry>
. The differences between the above-mentioned species are summarized in 
<tableCitation id="C6D70367FFD7C972FEA1FC5D298FFC56" box="[365,450,908,932]" captionStart="TABLE 2" captionStartId="9.[151,235,980,1001]" captionTargetBox="[151,1358,1029,1460]" captionTargetPageId="9" captionText="TABLE 2. Main characteristics of the Balkanic-Anatolian Dendrobaena species with clitellum shifted backward." httpUri="http://table.plazi.org/id/DF2A6654FFD7C972FF5BFC052D5EFC1B" pageId="9" pageNumber="34" tableUuid="DF2A6654FFD7C972FF5BFC052D5EFC1B">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
<caption id="DF2A6654FFD7C972FF5BFC052D5EFC1B" ID-Table-UUID="DF2A6654FFD7C972FF5BFC052D5EFC1B" box="[151,1299,979,1002]" httpUri="http://table.plazi.org/id/DF2A6654FFD7C972FF5BFC052D5EFC1B" pageId="9" pageNumber="34" targetBox="[151,1358,1029,1460]" targetIsTable="true" targetPageId="9">
<paragraph id="8BEA36DCFFD7C972FF5BFC052D5EFC1B" blockId="9.[151,1299,979,1002]" box="[151,1299,979,1002]" pageId="9" pageNumber="34">
<emphasis id="B921EACEFFD7C972FF5BFC052949FC1B" bold="true" box="[151,260,980,1001]" pageId="9" pageNumber="34">TABLE 2.</emphasis>
Main characteristics of the Balkanic-Anatolian 
<taxonomicName id="4C554D5FFFD7C972FD26FC052B39FC18" box="[746,884,980,1002]" class="Clitellata" family="Lumbricidae" genus="Dendrobaena" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="34" phylum="Annelida" rank="genus">
<emphasis id="B921EACEFFD7C972FD26FC052B39FC18" box="[746,884,980,1002]" italics="true" pageId="9" pageNumber="34">Dendrobaena</emphasis>
</taxonomicName>
species with clitellum shifted backward.
</paragraph>
</caption>
<paragraph id="8BEA36DCFFD7C972FDEBFBD42ACAFA46" pageId="9" pageNumber="34">
<table id="F955C47CFFD73684FF5BFBD42D03FA46" box="[151,1358,1029,1460]" gridcols="3" gridrows="3" pageId="9" pageNumber="34">
<tr id="3565349EFFD73684FF5BFBD42D03FB7E" box="[151,1358,1029,1164]" gridrow="0" pageId="9" pageNumber="34">
<th id="76B45DE2FFD73684FF5BFBD42AE0FB7E" box="[151,685,1029,1164]" gridcol="0" gridrow="0" pageId="9" pageNumber="34">
Clitellum 
<taxonomicName id="4C554D5FFFD7C972FF50FBE12A4AFBB6" authority="Zicsi &amp; Michalis, 1993" authorityName="Zicsi &amp; Michalis" authorityYear="1993" box="[156,519,1070,1093]" class="Clitellata" family="Lumbricidae" genus="Dendrobaena" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="34" phylum="Annelida" rank="species" species="epirotica">
<emphasis id="B921EACEFFD7C972FF50FBE12958FBB7" box="[156,277,1072,1093]" italics="true" pageId="9" pageNumber="34">D. epirotica</emphasis>
Zicsi &amp; Michalis, 1993
</taxonomicName>
31–38 
<taxonomicName id="4C554D5FFFD7C972FF50FB8829A4FB9C" authority="Zicsi &amp; Michalis" authorityName="Zicsi &amp; Michalis" box="[156,489,1112,1134]" class="Clitellata" family="Lumbricidae" genus="Dendrobaena" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="34" phylum="Annelida" rank="species" species="pindonensis">
<emphasis id="B921EACEFFD7C972FF50FB882979FB9C" box="[156,308,1112,1134]" italics="true" pageId="9" pageNumber="34">D. pindonensis</emphasis>
Zicsi &amp; Michalis
</taxonomicName>
, 32,33–40 1993
</th>
<th id="76B45DE2FFD73684FD3FFBD42CCEFB7E" box="[755,1155,1029,1164]" gridcol="1" gridrow="0" pageId="9" pageNumber="34">Tubercula pubertatis Seminal vesicles 33–35,36 9,11,12 35–37 9,11,12</th>
<th id="76B45DE2FFD73684FB0CFBD42D03FB7E" box="[1216,1358,1029,1164]" gridcol="2" gridrow="0" pageId="9" pageNumber="34">Spermathecae 9/10, 10/11 d 9/10, 10/11 d</th>
</tr>
<tr id="3565349EFFD73684FF5BFB712D03FAC3" box="[151,1358,1184,1329]" gridrow="1" pageId="9" pageNumber="34">
<th id="76B45DE2FFD73684FF5BFB712AE0FAC3" box="[151,685,1184,1329]" gridcol="0" gridrow="1" pageId="9" pageNumber="34">
<taxonomicName id="4C554D5FFFD7C972FF50FB7029C2FB47" authority="Zicsi, 1974" authorityName="Zicsi" authorityYear="1974" box="[156,399,1184,1206]" class="Clitellata" family="Lumbricidae" genus="Dendrobaena" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="34" phylum="Annelida" rank="species" species="mahnerti">
<emphasis id="B921EACEFFD7C972FF50FB70295AFB44" box="[156,279,1184,1206]" italics="true" pageId="9" pageNumber="34">D. mahnerti</emphasis>
Zicsi, 1974
</taxonomicName>
1/3 35,35–42 
<taxonomicName id="4C554D5FFFD7C972FF50FB1B29A9FB2D" authority="Omodeo &amp; Rota, 1989" authorityName="Omodeo &amp; Rota" authorityYear="1989" box="[156,484,1225,1247]" class="Clitellata" family="Lumbricidae" genus="Dendrobaena" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="34" phylum="Annelida" rank="species" species="bruna">
<emphasis id="B921EACEFFD7C972FF50FB1B28B5FB2D" box="[156,248,1225,1247]" italics="true" pageId="9" pageNumber="34">D. bruna</emphasis>
Omodeo &amp; Rota, 1989
</taxonomicName>
27–38 
<taxonomicName id="4C554D5FFFD7C972FF50FB2229DFFAFA" authority="Karaman, 1973" authorityName="Karaman" authorityYear="1973" box="[156,402,1267,1288]" class="Clitellata" family="Lumbricidae" genus="Dendrobaena" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="34" phylum="Annelida" rank="species" species="zicsii">
<emphasis id="B921EACEFFD7C972FF50FB2228A2FAFA" box="[156,239,1267,1288]" italics="true" pageId="9" pageNumber="34">D. zicsii</emphasis>
Karaman, 1973
</taxonomicName>
30–1/2 36 
<taxonomicName id="4C554D5FFFD7C972FF50FACD2958FAC3" box="[156,277,1308,1329]" class="Clitellata" family="Lumbricidae" genus="Dendrobaena" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="34" phylum="Annelida" rank="species" species="retrosella" status="sp. nov.">
<emphasis id="B921EACEFFD7C972FF50FACD2958FAC3" box="[156,277,1308,1329]" italics="true" pageId="9" pageNumber="34">D. retrosella</emphasis>
</taxonomicName>
<emphasis id="B921EACEFFD7C972FED7FACD2926FAC3" bold="true" box="[283,363,1308,1329]" pageId="9" pageNumber="34">
<taxonomicNameLabel id="A21257B5FFD7C972FED7FACD2926FAC3" box="[283,363,1308,1329]" pageId="9" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
29–36
</th>
<td id="76B45DE2FFD73684FD3FFB712CCEFAC3" box="[755,1155,1184,1329]" gridcol="1" gridrow="1" pageId="9" pageNumber="34">38–40 9,11,12 32–36 9,11,12 32–34 11,12 31–1/n 34 9,11,12</td>
<td id="76B45DE2FFD73684FB0CFB712D03FAC3" box="[1216,1358,1184,1329]" gridcol="2" gridrow="1" pageId="9" pageNumber="34">9/10, 10/11 d 9/10, 10/11 d 9/10, 10/11 d 9/10, 10/11 d</td>
</tr>
<tr id="3565349EFFD73684FF5BFA4B2D03FA46" box="[151,1358,1434,1460]" gridrow="2" pageId="9" pageNumber="34" rowspan-1="1" rowspan-2="1">
<th id="76B45DE2FFD73684FF5BFA4B2AE0FA46" box="[151,685,1434,1460]" gridcol="0" gridrow="2" pageId="9" pageNumber="34">
<taxonomicName id="4C554D5FFFD7C972FF5BFA4B2ACAFA46" authority="Rosa, 1886" authorityName="Rosa" authorityYear="1886" box="[151,647,1434,1460]" class="Clitellata" family="Lumbricidae" genus="Dendrobaena" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="34" phylum="Annelida" rank="subSpecies" species="veneta" subSpecies="veneta">
<emphasis id="B921EACEFFD7C972FF5BFA4B29ABFA46" bold="true" box="[151,486,1434,1460]" italics="true" pageId="9" pageNumber="34">Dendrobaena veneta veneta</emphasis>
(Rosa, 1886)
</taxonomicName>
</th>
</tr>
</table>
</paragraph>
<subSubSection id="C34F6557FFD7C972FF5BFA302950F9C6" pageId="9" pageNumber="34" type="reference_group">
<paragraph id="8BEA36DCFFD7C972FF5BFA302A68FA05" blockId="9.[151,1436,1505,1588]" box="[151,549,1505,1527]" pageId="9" pageNumber="34">
<treatmentCitationGroup id="AB4511F2FFD7C972FF5BFA302A68FA05" box="[151,549,1505,1527]" pageId="9" pageNumber="34">
<taxonomicName id="4C554D5FFFD7C972FF5BFA302A6CFA05" ID-CoL="85LBZ" authority="Rosa, 1886: 674" authorityName="Rosa" authorityPageNumber="674" authorityYear="1886" box="[151,545,1505,1527]" class="Clitellata" family="Lumbricidae" genus="Allolobophora" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="34" phylum="Annelida" rank="species" species="veneta">
<emphasis id="B921EACEFFD7C972FF5BFA30293EFA05" box="[151,371,1505,1527]" italics="true" pageId="9" pageNumber="34">Allolobophora veneta</emphasis>
<treatmentCitation id="0AF410CDFFD7C972FEB5FA332A6CFA05" author="Rosa" box="[377,545,1505,1527]" page="674" pageId="9" pageNumber="34" year="1886">
<bibRefCitation id="EFC44B2DFFD7C972FEB5FA3329A2FA05" author="Rosa" box="[377,495,1505,1527]" pageId="9" pageNumber="34" refString="Rosa, D. (1886) Note sui lombrici del Veneto. Atti del Reale Istituto Veneto di Scienze, 4, 673 - 687." type="journal article" year="1886">Rosa, 1886</bibRefCitation>
: 674
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEA36DCFFD7C972FF5BF9D12950F9C6" blockId="9.[151,1436,1505,1588]" pageId="9" pageNumber="34">
<treatmentCitationGroup id="AB4511F2FFD7C972FF5BF9D12950F9C6" pageId="9" pageNumber="34">
<taxonomicName id="4C554D5FFFD7C972FF5BF9D12ADFF9E4" ID-CoL="84ZKD" authority="Zicsi &amp; Michalis 1981: 258" authorityName="Zicsi &amp; Michalis" authorityPageNumber="258" authorityYear="1981" box="[151,658,1536,1558]" class="Clitellata" family="Lumbricidae" genus="Dendrobaena" kingdom="Animalia" order="Crassiclitellata" pageId="9" pageNumber="34" phylum="Annelida" rank="species" species="veneta">
<emphasis id="B921EACEFFD7C972FF5BF9D12924F9E4" box="[151,361,1536,1558]" italics="true" pageId="9" pageNumber="34">Dendrobaena veneta</emphasis>
: 
<treatmentCitation id="0AF410CDFFD7C972FEBBF9D12ADFF9E4" author="Zicsi" box="[375,658,1536,1558]" page="258" pageId="9" pageNumber="34" year="1981">
<bibRefCitation id="EFC44B2DFFD7C972FEBBF9D12A12F9E7" author="Zicsi" box="[375,607,1536,1558]" pageId="9" pageNumber="34" refString="Zicsi, A. &amp; Michalis, K. (1981) Ubersicht der Regenwurm-fauna Griechenlands (Oligochaeta: Lumbricidae). Acta zoologica hungarica, 27, 239 - 264." type="journal article" year="1981">Zicsi &amp; Michalis 1981</bibRefCitation>
: 258
</treatmentCitation>
</taxonomicName>
. 
<treatmentCitation id="0AF410CDFFD7C972FD50F9D12B29F9E4" author="Michalis" box="[668,868,1536,1558]" page="355" pageId="9" pageNumber="34" year="1982">
<bibRefCitation id="EFC44B2DFFD7C972FD50F9D12B7CF9E7" author="Michalis" box="[668,817,1536,1558]" pageId="9" pageNumber="34" refString="Michalis, K. (1982) Katalog der Oligochaetenfauna Griechenlands. Biologia Gallo-Hellenica, 9, 343 - 362." type="journal article" year="1982">Michalis 1982</bibRefCitation>
: 355
</treatmentCitation>
; 1995: 16. Csuzdi &amp; 
<treatmentCitation id="0AF410CDFFD7C972FBF7F9D12C97F9E7" author="Zicsi" box="[1083,1242,1536,1558]" page="127" pageId="9" pageNumber="34" year="2003">
<bibRefCitation id="EFC44B2DFFD7C972FBF7F9D12CE6F9E7" author="Zicsi" box="[1083,1195,1536,1558]" pageId="9" pageNumber="34" refString="Csuzdi, Cs. &amp; Zicsi, A. (2003) Earthworms of Hungary (Annelida: Oligochaeta; Lumbricidae). In: Csuzdi, Cs. &amp; Mahunka, S. (eds.): Pedozoologica Hungarica 1. Hungarian Natural History Museum, Budapest, pp. 271." type="journal article" year="2003">Zicsi 2003</bibRefCitation>
: 127
</treatmentCitation>
(for complete synonymy).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34F6557FFD7C971FF5BF9B02D34FEEE" lastPageId="10" lastPageNumber="35" pageId="9" pageNumber="34" type="materials_examined">
<paragraph id="8BEA36DCFFD7C971FF5BF9B02D34FEEE" blockId="9.[151,1436,1633,2017]" lastBlockId="10.[151,1436,151,284]" lastPageId="10" lastPageNumber="35" pageId="9" pageNumber="34">
<emphasis id="B921EACEFFD7C972FF5BF9B029CCF988" bold="true" box="[151,385,1633,1658]" pageId="9" pageNumber="34">Material examined.</emphasis>
<collectionCode id="ED44AE19FFD7C972FE44F9B02992F98B" LSID="urn:lsid:biocol.org:col:34991" box="[392,479,1633,1657]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="34">HNHM</collectionCode>
/15720 four adult ex., 
<emphasis id="B921EACEFFD7C972FD14F9B32B6EF98B" box="[728,803,1633,1657]" italics="true" pageId="9" pageNumber="34">No. 17</emphasis>
, Trikala County, Athamon Mts., Athamania, small river and mixed rocky forest S of the village, 
<quantity id="4CAD9B39FFD7C972FDAEF9542AE9F96F" box="[610,676,1669,1693]" metricMagnitude="2" metricUnit="m" metricValue="9.35" pageId="9" pageNumber="34" unit="m" value="935.0">935m</quantity>
, 
<geoCoordinate id="EE61501BFFD7C972FD7CF9542B0FF96F" box="[688,834,1669,1693]" direction="north" orientation="latitude" pageId="9" pageNumber="34" precision="1" value="39.51573">N39°30.944’</geoCoordinate>
<geoCoordinate id="EE61501BFFD7C972FC86F9542B96F96F" box="[842,987,1669,1693]" direction="east" orientation="longitude" pageId="9" pageNumber="34" precision="1" value="21.24995">E21°14.997’</geoCoordinate>
, leg. J. Kontschán, D. Murányi, T. Szederjesi, Zs. Ujvári, 0 9.0 5.2011. 
<collectionCode id="ED44AE19FFD7C972FDDEF9782A24F933" LSID="urn:lsid:biocol.org:col:34991" box="[530,617,1705,1729]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="34">HNHM</collectionCode>
/15724 five adult ex., 
<collectionCode id="ED44AE19FFD7C972FCB9F9782B81F933" LSID="urn:lsid:biocol.org:col:34991" box="[885,972,1705,1729]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="34">HNHM</collectionCode>
/15730 one adult ex., 
<emphasis id="B921EACEFFD7C972FB19F97B2D6BF933" box="[1237,1318,1705,1729]" italics="true" pageId="9" pageNumber="34">No. 14</emphasis>
, Ioannina County, Vouliasta, plane tree gallery, spring lake and upper section of Louros River in the village, 
<quantity id="4CAD9B39FFD7C972FA99F91C2DDAF917" box="[1365,1431,1741,1765]" metricMagnitude="2" metricUnit="m" metricValue="2.35" pageId="9" pageNumber="34" unit="m" value="235.0">235m</quantity>
, 
<geoCoordinate id="EE61501BFFD7C972FF5BF9202965F8FB" box="[151,296,1777,1801]" direction="north" orientation="latitude" pageId="9" pageNumber="34" precision="1" value="39.432316">N39°25.939’</geoCoordinate>
<geoCoordinate id="EE61501BFFD7C972FEFFF920298EF8FB" box="[307,451,1777,1801]" direction="east" orientation="longitude" pageId="9" pageNumber="34" precision="1" value="20.843416">E20°50.605’</geoCoordinate>
, leg. J. Kontschán, D. Murányi, T. Szederjesi, Zs. Ujvári, 0 4.0 5.2011. 
<collectionCode id="ED44AE19FFD7C972FB35F9202D1DF8FB" LSID="urn:lsid:biocol.org:col:34991" box="[1273,1360,1777,1801]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="34">HNHM</collectionCode>
/15736 one adult ex., 
<emphasis id="B921EACEFFD7C972FEF7F8C729CAF8DF" box="[315,391,1813,1837]" italics="true" pageId="9" pageNumber="34">No. 29</emphasis>
, Phthiotis County, Agios Georgios, Sperchios River and its gallery forest W of the village, 
<quantity id="4CAD9B39FFD7C972FF5BF8E82894F8A3" box="[151,217,1849,1873]" metricMagnitude="2" metricUnit="m" metricValue="3.65" pageId="9" pageNumber="34" unit="m" value="365.0">365m</quantity>
, 
<geoCoordinate id="EE61501BFFD7C972FF2BF8E82935F8A3" box="[231,376,1849,1873]" direction="north" orientation="latitude" pageId="9" pageNumber="34" precision="1" value="38.95015">N38°57.009’</geoCoordinate>
<geoCoordinate id="EE61501BFFD7C972FE4DF8E82A5FF8A3" box="[385,530,1849,1873]" direction="east" orientation="longitude" pageId="9" pageNumber="34" precision="1" value="21.9452">E21°56.712’</geoCoordinate>
, leg. J. Kontschán, D. Murányi, T. Szederjesi, Zs. Ujvári, 0 8.0 5.2011. 
<collectionCode id="ED44AE19FFD7C972FAF3F8E82DD9F8A3" LSID="urn:lsid:biocol.org:col:34991" box="[1343,1428,1849,1873]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="34">HNHM</collectionCode>
/ 15740 one adult ex., 
<emphasis id="B921EACEFFD7C972FE4AF88F299CF887" box="[390,465,1885,1909]" italics="true" pageId="9" pageNumber="34">No. 31</emphasis>
, Aetolia-Acarnania County, Kamaroula, small river and its littoral scrub N of the village, 
<quantity id="4CAD9B39FFD7C972FF1FF850295BF86B" box="[211,278,1921,1945]" metricMagnitude="2" metricUnit="m" metricValue="1.3" pageId="9" pageNumber="34" unit="m" value="130.0">130m</quantity>
, 
<geoCoordinate id="EE61501BFFD7C972FEECF85029FFF86B" box="[288,434,1921,1945]" direction="north" orientation="latitude" pageId="9" pageNumber="34" precision="1" value="38.638367">N38°38.302’</geoCoordinate>
<geoCoordinate id="EE61501BFFD7C972FE74F8502A04F86B" box="[440,585,1921,1945]" direction="east" orientation="longitude" pageId="9" pageNumber="34" precision="1" value="21.451534">E21°27.092’</geoCoordinate>
, leg. Kontschán, D. Murányi, T. Szederjesi, Zs. Ujvári, 0 7.0 5.2011. 
<collectionCode id="ED44AE19FFD7C972FAF3F8502DD9F86B" LSID="urn:lsid:biocol.org:col:34991" box="[1343,1428,1921,1945]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="34">HNHM</collectionCode>
/ 15750 one adult ex., 
<emphasis id="B921EACEFFD7C972FE4FF8772983F84F" box="[387,462,1957,1981]" italics="true" pageId="9" pageNumber="34">No. 16</emphasis>
, Trikala County, Neraida Mts., Desi, karst spring and its outlet, and bushy grassland E of the village, 
<quantity id="4CAD9B39FFD7C972FE80F81829D1F813" box="[332,412,1993,2017]" metricMagnitude="3" metricUnit="m" metricValue="1.04" pageId="9" pageNumber="34" unit="m" value="1040.0">1040m</quantity>
, 
<geoCoordinate id="EE61501BFFD7C972FE63F8182A0DF813" box="[431,576,1993,2017]" direction="north" orientation="latitude" pageId="9" pageNumber="34" precision="1" value="39.560616">N39°33.637’</geoCoordinate>
<geoCoordinate id="EE61501BFFD7C972FD82F8182A92F813" box="[590,735,1993,2017]" direction="east" orientation="longitude" pageId="9" pageNumber="34" precision="1" value="21.371916">E21°22.315’</geoCoordinate>
, leg. J. Kontschán, D. Murányi, T. Szederjesi, Zs. Ujvári, 0 9.0 5.2011. 
<collectionCode id="ED44AE19FFD4C971FEE8FF462936FF5D" LSID="urn:lsid:biocol.org:col:34991" box="[292,379,151,175]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="35">HNHM</collectionCode>
/15752 two adult ex., 
<emphasis id="B921EACEFFD4C971FDB8FF482A8DFF42" box="[628,704,152,176]" italics="true" pageId="10" pageNumber="35">No. 15</emphasis>
, Trikala County, Lakmos Mts., Chaliki, open torrent and brook, and rocky subalpine grassland N of the village, 
<quantity id="4CAD9B39FFD4C971FD7CFF6D2B4CFF26" box="[688,769,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.225" pageId="10" pageNumber="35" unit="m" value="1225.0">1225m</quantity>
, 
<geoCoordinate id="EE61501BFFD4C971FCC0FF6C2BD3FF26" box="[780,926,188,212]" direction="north" orientation="latitude" pageId="10" pageNumber="35" precision="1" value="39.698467">N39°41.908’</geoCoordinate>
<geoCoordinate id="EE61501BFFD4C971FC6AFF6D2C78FF26" box="[934,1077,188,212]" direction="east" orientation="longitude" pageId="10" pageNumber="35" precision="1" value="21.18395">E21°11.037’</geoCoordinate>
, leg. J. Kontschán, D. Murányi, T. Szederjesi, Zs. Ujvári, 0 9.0 5.2011. 
<collectionCode id="ED44AE19FFD4C971FD8FFF0E2AD7FF05" LSID="urn:lsid:biocol.org:col:34991" box="[579,666,223,247]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="10" pageNumber="35">HNHM</collectionCode>
/15857 four adult ex., 
<emphasis id="B921EACEFFD4C971FC59FF302B9FFF0A" box="[917,978,224,248]" italics="true" pageId="10" pageNumber="35">No. 6</emphasis>
, Ioannina County, Kalpaki, Vellas Monasteri, karstic spring, 
<quantity id="4CAD9B39FFD4C971FE4BFED52987FEEE" box="[391,458,260,284]" metricMagnitude="2" metricUnit="m" metricValue="4.2" pageId="10" pageNumber="35" unit="m" value="420.0">420m</quantity>
, 
<geoCoordinate id="EE61501BFFD4C971FE19FED52A3EFEEE" box="[469,627,260,284]" direction="north" orientation="latitude" pageId="10" pageNumber="35" precision="1" value="39.849167">N39°50’57.0”</geoCoordinate>
<geoCoordinate id="EE61501BFFD4C971FDB6FED52B55FEEE" box="[634,792,260,284]" direction="east" orientation="longitude" pageId="10" pageNumber="35" precision="1" value="20.623917">E20°37’26.1”</geoCoordinate>
, leg. L. Dányi, J. Kontschán, D. Murányi, 12.0 5.2006.
</paragraph>
</subSubSection>
</treatment>
</document>