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<document id="6D142B05764F1D12CE902E59C5D95CB9" ID-DOI="10.5281/zenodo.5352660" ID-ISSN="1942-1354" ID-Zenodo-Dep="5352660" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630208527114" checkinUser="felipe" docAuthor="Dunford, James C." docDate="2009" docId="03FC87C67B2FFFB0FF6CFF15FC6ADBAD" docLanguage="en" docName="0090.pdf" docOrigin="Insecta Mundi 2009 (90)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Speyeria atlantis subsp. hollandi" docType="treatment" docVersion="3" lastPageNumber="28" masterDocId="FFC5FFBE7B32FFADFFACFFD0FFFED816" masterDocTitle="Taxonomic overview of the greater fritillary genus Speyeria Scudder and the atlantis - hesperis species complexes, with species accounts, type images, and relevant literature (Lepidoptera: Nymphalidae)" masterLastPageNumber="74" masterPageNumber="1" pageNumber="28" updateTime="1699186102529" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="07789BFE201D361D7B6BE8A00CC036D6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="21EF4B4F4E4B860B700FF2E1AF1723AF">
<mods:title id="18CFFB665A919F8A756E626B7A874987">Taxonomic overview of the greater fritillary genus Speyeria Scudder and the atlantis - hesperis species complexes, with species accounts, type images, and relevant literature (Lepidoptera: Nymphalidae)</mods:title>
</mods:titleInfo>
<mods:name id="498F67116EEE7B227F9412DD729D1F5E" type="personal">
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</mods:role>
<mods:namePart id="2818D5E2706C59FA314D7AE97433CA24">Dunford, James C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A8A776EE01389C544088B09FEA507A3F">text</mods:typeOfResource>
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<mods:title id="69D6A74F1344CAB4A9EBF9DEF7E834A2">Insecta Mundi</mods:title>
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<mods:part id="760FEA29FE2A8825A7407B797EC2C5B9">
<mods:date id="F39010DC137CA136A2392928EC17E50D">2009</mods:date>
<mods:detail id="D2250A14B4F3AA1D2F1C702FDF41F771" type="pubDate">
<mods:number id="19717ED07E7044A5401FCBAC4B53DD3F">2009-09-26</mods:number>
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<mods:number id="A4EF0C8D45B7F3E7FACF5FAF1F7B6DD6">2009</mods:number>
</mods:detail>
<mods:detail id="82018676CC596222BEEEF8C00A4C123D" type="issue">
<mods:number id="97C584AE407564DCAE823BF9A92514D2">90</mods:number>
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<mods:classification id="D6D7F48AAE4EA51436CA20F85EF6216C">journal article</mods:classification>
<mods:identifier id="CEFD1EA92D9C8C3BB9DA532C34956B97" type="DOI">10.5281/zenodo.5352660</mods:identifier>
<mods:identifier id="B878771FA661397D5D0D53FB86DA7CDA" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="BEC74B9DBC840463A38AC48405CB15BF" type="Zenodo-Dep">5352660</mods:identifier>
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<treatment id="03FC87C67B2FFFB0FF6CFF15FC6ADBAD" LSID="urn:lsid:plazi:treatment:03FC87C67B2FFFB0FF6CFF15FC6ADBAD" httpUri="http://treatment.plazi.org/id/03FC87C67B2FFFB0FF6CFF15FC6ADBAD" lastPageNumber="28" pageId="29" pageNumber="28">
<subSubSection id="C34F655B7B2FFFB0FF6CFF15FC18D8CD" box="[192,998,195,220]" pageId="29" pageNumber="28" type="nomenclature">
<paragraph id="8BEA36D07B2FFFB0FF6CFF15FC18D8CD" blockId="29.[192,998,195,251]" box="[192,998,195,220]" pageId="29" pageNumber="28">
<heading id="D0A281BC7B2FFFB0FF6CFF15FC18D8CD" bold="true" box="[192,998,195,220]" fontSize="10" level="2" pageId="29" pageNumber="28" reason="3">
<taxonomicName id="4C554D537B2FFFB0FF6CFF15FC18D8CD" ID-CoL="5L8Q5" authority="(Chermock and Chermock, 1940)" baseAuthorityName="Chermock and Chermock" baseAuthorityYear="1940" box="[192,998,195,220]" class="Insecta" family="Nymphalidae" genus="Speyeria" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="28" phylum="Arthropoda" rank="subSpecies" species="atlantis" subSpecies="hollandi">
<emphasis id="B921EAC27B2FFFB0FF6CFF15FC18D8CD" bold="true" box="[192,998,195,220]" pageId="29" pageNumber="28">
<emphasis id="B921EAC27B2FFFB0FF6CFF15FDDBD8CA" bold="true" box="[192,549,197,220]" italics="true" pageId="29" pageNumber="28">Speyeria atlantis hollandi</emphasis>
(
<bibRefCitation id="EFC44B217B2FFFB0FD9FFF13FC20D8CD" author="Chermock, F. H. &amp; R. L. Chermock" box="[563,990,195,219]" pageId="29" pageNumber="28" pagination="81 - 83" refId="ref35091" refString="Chermock, F. H., and R. L. Chermock. 1940. Some new diurnal Lepidoptera from the Riding Mountains and the Sand Ridge, Manitoba. The Canadian Entomologist 72: 81 - 83." type="journal article" year="1940">Chermock and Chermock, 1940</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F655B7B2FFFB0FF6CFF33FEB8D8ED" box="[192,326,227,251]" pageId="29" pageNumber="28" type="description">
<paragraph id="8BEA36D07B2FFFB0FF6CFF33FEB8D8ED" blockId="29.[192,998,195,251]" box="[192,326,227,251]" pageId="29" pageNumber="28">
(
<figureCitation id="136E2A557B2FFFB0FF64FF33FEC0D8ED" box="[200,318,227,251]" captionStart="Figure 21-25" captionStartId="69.[192,276,1750,1772]" captionTargetBox="[192,1426,198,1725]" captionTargetId="figure-15@69.[192,1426,190,1725]" captionTargetPageId="69" captionText="Figure 21-25. Type images for Speyeria spp.; dorsal, ventral, and label images. 21) Speyeria atlantis canadensis lectotype, described as Argynnis atlantis canadensis, male, AMNH. Now synonymized under Speyeria atlantis (Scott et al. 1998). 22) Speyeria atlantis hollandi holotype, described as Argynnis atlantis Hollandi, male, CNIC. 23) Speyeria atlantis sorocko holotype, described as Speyeria atlantis sorocko, male, AMNH. 24) Speyeria atlantis pahasapa holotype, described as Speyeria atlantis pahasapa, male, AMNH.25) Speyeria hesperis neotype, described as Argynnis Hesperis, male, CMNH. Images 21 and 23-25 by James C. Dunford; 22 by Norbert Kondla." figureDoi="http://doi.org/10.5281/zenodo.5352678" httpUri="https://zenodo.org/record/5352678/files/figure.png" pageId="29" pageNumber="28">Figure 22</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34F655B7B2FFFB0FF6CFEF4FB60D94D" pageId="29" pageNumber="28" type="reference_group">
<paragraph id="8BEA36D07B2FFFB0FF6CFEF4FC3ED92D" blockId="29.[192,1182,291,347]" box="[192,960,291,315]" pageId="29" pageNumber="28">
<treatmentCitationGroup id="AB4511FE7B2FFFB0FF6CFEF4FC3ED92D" box="[192,960,291,315]" pageId="29" pageNumber="28">
<treatmentCitation id="0AF410C17B2FFFB0FF6CFEF4FC42D92D" author="Chermock, F. H. &amp; R. L. Chermock" box="[192,956,291,315]" page="82" pageId="29" pageNumber="28" year="1940">
<taxonomicName id="4C554D537B2FFFB0FF6CFEF4FC42D92D" ID-CoL="93QQW" authority="Hollandi Chermock and Chermock, 1940: 82" authorityName="Hollandi Chermock and Chermock" authorityPageNumber="82" authorityYear="1940" box="[192,956,291,315]" class="Insecta" family="Nymphalidae" genus="Argynnis" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="atlantis">
<emphasis id="B921EAC27B2FFFB0FF6CFEF4FDF4D92D" box="[192,522,291,315]" italics="true" pageId="29" pageNumber="28">Argynnis atlantis Hollandi</emphasis>
<bibRefCitation id="EFC44B217B2FFFB0FDBCFEF3FC42D92D" author="Chermock, F. H. &amp; R. L. Chermock" box="[528,956,291,315]" pageId="29" pageNumber="28" pagination="81 - 83" refId="ref35091" refString="Chermock, F. H., and R. L. Chermock. 1940. Some new diurnal Lepidoptera from the Riding Mountains and the Sand Ridge, Manitoba. The Canadian Entomologist 72: 81 - 83." type="journal article" year="1940">Chermock and Chermock, 1940: 82</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEA36D07B2FFFB0FF6CFE94FB60D94D" blockId="29.[192,1182,291,347]" box="[192,1182,323,347]" pageId="29" pageNumber="28">
<taxonomicName id="4C554D537B2FFFB0FF6CFE94FCB6D94D" ID-CoL="5L8Q5" authority="(Chermock and Chermock)" baseAuthorityName="Chermock and Chermock" baseAuthorityYear="1940" box="[192,840,323,347]" class="Insecta" family="Nymphalidae" genus="Speyeria" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="28" phylum="Arthropoda" rank="subSpecies" species="atlantis" subSpecies="hollandi">
<emphasis id="B921EAC27B2FFFB0FF6CFE94FE02D94D" box="[192,508,323,347]" italics="true" pageId="29" pageNumber="28">Speyeria atlantis hollandi</emphasis>
(Chermock and Chermock)
</taxonomicName>
[
<bibRefCitation id="EFC44B217B2FFFB0FCF8FE93FB6CD94D" author="dos Passos C. F. &amp; L. P. Grey" box="[852,1170,323,347]" pageId="29" pageNumber="28" pagination="1 - 30" refId="ref35903" refString="dos Passos C. F., and L. P. Grey. 1947. Systematic catalogue of Speyeria (Lepidoptera, Nymphalidae) with designations of types and fixations of the type localities. American Museum Novitates 1370: 1 - 30." type="journal article" year="1947">dos Passos and Grey 1947</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="C34F655B7B2FFFB0FF6CFE53FD78D98D" box="[192,646,387,411]" pageId="29" pageNumber="28" type="description">
<paragraph id="8BEA36D07B2FFFB0FF6CFE53FD78D98D" blockId="29.[192,646,387,411]" box="[192,646,387,411]" pageId="29" pageNumber="28">
<emphasis id="B921EAC27B2FFFB0FF6CFE53FE6AD98D" bold="true" box="[192,404,387,411]" pageId="29" pageNumber="28">Common name.</emphasis>
Holland’s fritillary.
</paragraph>
</subSubSection>
<subSubSection id="C34F655B7B2FFFB0FF6CFE13FC6ADBAD" pageId="29" pageNumber="28" type="materials_examined">
<paragraph id="8BEA36D07B2FFFB0FF6CFE13FBD8D9CD" blockId="29.[192,1062,451,475]" box="[192,1062,451,475]" pageId="29" pageNumber="28">
<emphasis id="B921EAC27B2FFFB0FF6CFE13FE69D9CD" bold="true" box="[192,407,451,475]" pageId="29" pageNumber="28">Type deposited.</emphasis>
<typeStatus id="54EE88727B2FFFB0FE30FE13FDF4D9CD" box="[412,522,451,475]" pageId="29" pageNumber="28" type="holotype">Holotype</typeStatus>
at Canadian National Collection (
<figureCitation id="136E2A557B2FFFB0FC0DFE13FBE4D9CD" box="[929,1050,451,475]" captionStart="Figure 21-25" captionStartId="69.[192,276,1750,1772]" captionTargetBox="[192,1426,198,1725]" captionTargetId="figure-15@69.[192,1426,190,1725]" captionTargetPageId="69" captionText="Figure 21-25. Type images for Speyeria spp.; dorsal, ventral, and label images. 21) Speyeria atlantis canadensis lectotype, described as Argynnis atlantis canadensis, male, AMNH. Now synonymized under Speyeria atlantis (Scott et al. 1998). 22) Speyeria atlantis hollandi holotype, described as Argynnis atlantis Hollandi, male, CNIC. 23) Speyeria atlantis sorocko holotype, described as Speyeria atlantis sorocko, male, AMNH. 24) Speyeria atlantis pahasapa holotype, described as Speyeria atlantis pahasapa, male, AMNH.25) Speyeria hesperis neotype, described as Argynnis Hesperis, male, CMNH. Images 21 and 23-25 by James C. Dunford; 22 by Norbert Kondla." figureDoi="http://doi.org/10.5281/zenodo.5352678" httpUri="https://zenodo.org/record/5352678/files/figure.png" pageId="29" pageNumber="28">Figure 22</figureCitation>
).
</paragraph>
<paragraph id="8BEA36D07B2FFFB0FF6CFDD3FCAEDA0D" blockId="29.[192,848,515,539]" box="[192,848,515,539]" pageId="29" pageNumber="28">
<emphasis id="B921EAC27B2FFFB0FF6CFDD3FE87DA0D" bold="true" box="[192,377,515,539]" pageId="29" pageNumber="28">
<typeStatus id="54EE88727B2FFFB0FF6CFDD3FEFADA0D" box="[192,260,515,539]" pageId="29" pageNumber="28">Type</typeStatus>
locality.
</emphasis>
Riding Mountains,
<collectingRegion id="4991F8327B2FFFB0FDCBFDD3FD1EDA0D" box="[615,736,515,539]" country="Canada" name="Manitoba" pageId="29" pageNumber="28">Manitoba</collectingRegion>
,
<collectingCountry id="F34276407B2FFFB0FD46FDD3FCB3DA0D" box="[746,845,515,539]" name="Canada" pageId="29" pageNumber="28">Canada</collectingCountry>
.
</paragraph>
<paragraph id="8BEA36D07B2FFFB0FF6CFD93FBBCDA8D" blockId="29.[192,1440,579,667]" pageId="29" pageNumber="28">
<emphasis id="B921EAC27B2FFFB0FF6CFD93FE5DDA4D" bold="true" box="[192,419,579,603]" pageId="29" pageNumber="28">Type label data.</emphasis>
“
<taxonomicName id="4C554D537B2FFFB0FE1BFD93FD68DA4D" box="[439,662,579,603]" class="Insecta" family="Nymphalidae" genus="Speyeria" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="atlantis">ARG. ATLANTIS</taxonomicName>
.
<collectionCode id="ED44AE157B2FFFB0FD0FFD94FD44DA4D" box="[675,698,580,603]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="29" pageNumber="28">R</collectionCode>
. HOLLANDI [male]
<typeStatus id="54EE88727B2FFFB0FC6EFD93FBA0DA4D" box="[962,1118,579,603]" pageId="29" pageNumber="28" type="holotype">HOLOTYPE</typeStatus>
F. H. &amp;
<collectionCode id="ED44AE157B2FFFB0FB7EFD94FB17DA4D" box="[1234,1257,580,603]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="29" pageNumber="28">R</collectionCode>
. L. Chermock;
<typeStatus id="54EE88727B2FFFB0FF6CFDB3FEA5DA6D" box="[192,347,611,635]" pageId="29" pageNumber="28" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C554D537B2FFFB0FECDFDB3FE05DA6D" box="[353,507,611,635]" class="Insecta" family="Nymphalidae" genus="Speyeria" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="atlantis">Arg. atlantis</taxonomicName>
<collectionCode id="ED44AE157B2FFFB0FDACFDB4FDE9DA6D" box="[512,535,612,635]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="29" pageNumber="28">R</collectionCode>
.
<collectingCountry id="F34276407B2FFFB0FD8CFDB3FD7DDA6D" box="[544,643,611,635]" name="Netherlands" pageId="29" pageNumber="28">Holland</collectingCountry>
No. 4370 F. H. &amp;
<collectionCode id="ED44AE157B2FFFB0FCFBFDB4FC90DA6D" box="[855,878,612,635]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="29" pageNumber="28">R</collectionCode>
. L. Chermock; RIDING MTS MANITOBA VII- 24-34; Can. Dep. Agr. Photo. Specimen No. 4093
<date id="FFEB10107B2FFFB0FCA7FD53FC6ADA8D" box="[779,916,643,667]" pageId="29" pageNumber="28" value="1986-04-24">24-IV-1986</date>
Negative No”.
</paragraph>
<paragraph id="8BEA36D07B2FFFB0FF6CFD13FAC6DB0D" blockId="29.[192,1440,707,796]" pageId="29" pageNumber="28">
<emphasis id="B921EAC27B2FFFB0FF6CFD13FD1CDACD" bold="true" box="[192,738,707,731]" pageId="29" pageNumber="28">Identification, taxonomy, and variation.</emphasis>
The discal and basal areas located on the ventral forewing and hindwing are deep brown compared to paler
<taxonomicName id="4C554D537B2FFFB0FC88FD34FC5BDAED" box="[804,933,739,763]" class="Insecta" family="Nymphalidae" genus="Speyeria" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="atlantis">
<emphasis id="B921EAC27B2FFFB0FC88FD34FCC8DAED" box="[804,822,740,763]" italics="true" pageId="29" pageNumber="28">S</emphasis>
.
<emphasis id="B921EAC27B2FFFB0FCE9FD33FC5BDAED" box="[837,933,739,763]" italics="true" pageId="29" pageNumber="28">atlantis</emphasis>
</taxonomicName>
forms. This subspecies is considered the western terminus of the
<taxonomicName id="4C554D537B2FFFB0FE46FCD4FD97DB0D" box="[490,617,771,795]" class="Insecta" family="Nymphalidae" genus="Speyeria" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="28" phylum="Arthropoda" rank="species" species="atlantis">
<emphasis id="B921EAC27B2FFFB0FE46FCD4FE02DB0D" box="[490,508,772,795]" italics="true" pageId="29" pageNumber="28">S</emphasis>
.
<emphasis id="B921EAC27B2FFFB0FDA5FCD3FD97DB0D" box="[521,617,771,795]" italics="true" pageId="29" pageNumber="28">atlantis</emphasis>
</taxonomicName>
cline (
<bibRefCitation id="EFC44B217B2FFFB0FD1BFCD3FCBCDB0D" author="Howe, W. H." box="[695,834,771,795]" pageId="29" pageNumber="28" refId="ref41056" refString="Howe, W. H. 1975. The butterflies of North America. Doubleday and Company, Inc.; Garden City, New York. 633 p." type="book" year="1975">Howe 1975</bibRefCitation>
). Adult wingspan ranges from
<quantity id="4CAD9B357B2FFFB0FB1BFCD3FACADB0D" box="[1207,1332,771,795]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="29" pageNumber="28" unit="mm" value="55.0" valueMax="60.0" valueMin="50.0">50-60 mm</quantity>
.
</paragraph>
<paragraph id="8BEA36D07B2FFFB0FF6CFC93FEC2DB6D" blockId="29.[192,1440,835,891]" pageId="29" pageNumber="28">
<emphasis id="B921EAC27B2FFFB0FF6CFC93FEDEDB4D" bold="true" box="[192,288,835,859]" pageId="29" pageNumber="28">Range.</emphasis>
This subspecies occurs in the Riding Mountains of
<collectingRegion id="4991F8327B2FFFB0FC23FC93FBF8DB4D" box="[911,1030,835,859]" country="Canada" name="Manitoba" pageId="29" pageNumber="28">Manitoba</collectingRegion>
and Peace River region of
<collectingRegion id="4991F8327B2FFFB0FAE4FC93FEC6DB6D" country="Canada" name="British Columbia" pageId="29" pageNumber="28">British Columbia</collectingRegion>
.
</paragraph>
<paragraph id="8BEA36D07B2FFFB0FF6CFC73FC6ADBAD" blockId="29.[192,916,931,955]" box="[192,916,931,955]" pageId="29" pageNumber="28">
<emphasis id="B921EAC27B2FFFB0FF6CFC73FE96DBAD" bold="true" box="[192,360,931,955]" pageId="29" pageNumber="28">Life history.</emphasis>
Nothing could be gleaned from the literature.
</paragraph>
</subSubSection>
</treatment>
</document>
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