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<document id="8205FBA04AA2681F8D40DBBDED5DC76D" ID-DOI="10.5281/zenodo.5352660" ID-ISSN="1942-1354" ID-Zenodo-Dep="5352660" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630208527114" checkinUser="felipe" docAuthor="Dunford, James C." docDate="2009" docId="03FC87C67B18FF86FF6CF9D5FC30D9CD" docLanguage="en" docName="0090.pdf" docOrigin="Insecta Mundi 2009 (90)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Speyeria hesperis subsp. schellbachi Garth 1949" docType="treatment" docVersion="3" lastPageNumber="42" masterDocId="FFC5FFBE7B32FFADFFACFFD0FFFED816" masterDocTitle="Taxonomic overview of the greater fritillary genus Speyeria Scudder and the atlantis - hesperis species complexes, with species accounts, type images, and relevant literature (Lepidoptera: Nymphalidae)" masterLastPageNumber="74" masterPageNumber="1" pageNumber="41" updateTime="1699186102529" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="7553E03EC892FB1E44738E1FAA3FC264">Taxonomic overview of the greater fritillary genus Speyeria Scudder and the atlantis - hesperis species complexes, with species accounts, type images, and relevant literature (Lepidoptera: Nymphalidae)</mods:title>
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<mods:namePart id="EED33273C5F7AB0E28AFB06ABD1F316A">Dunford, James C.</mods:namePart>
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<mods:title id="BDE3ABBA5227122C9658A53A0938D8B4">Insecta Mundi</mods:title>
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<mods:date id="CA99D3995F247DDC49B4B5899DDED2D2">2009</mods:date>
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<mods:number id="BCD661818B1E2B28F9CFEA9E93787C29">2009-09-26</mods:number>
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<mods:number id="2770B0D37E543CE711A1F25F54111E43">2009</mods:number>
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<mods:number id="853F6CB71320D88D507E2450CD1CAFD4">90</mods:number>
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<treatment id="03FC87C67B18FF86FF6CF9D5FC30D9CD" LSID="urn:lsid:plazi:treatment:03FC87C67B18FF86FF6CF9D5FC30D9CD" httpUri="http://treatment.plazi.org/id/03FC87C67B18FF86FF6CF9D5FC30D9CD" lastPageId="43" lastPageNumber="42" pageId="42" pageNumber="41">
<subSubSection id="C34F655B7B18FF87FF6CF9D5FD0EDE0D" box="[192,752,1539,1564]" pageId="42" pageNumber="41" type="nomenclature">
<paragraph id="8BEA36D07B18FF87FF6CF9D5FD0EDE0D" blockId="42.[192,752,1539,1595]" box="[192,752,1539,1564]" pageId="42" pageNumber="41">
<heading id="D0A281BC7B18FF87FF6CF9D5FD0EDE0D" bold="true" box="[192,752,1539,1564]" fontSize="10" level="2" pageId="42" pageNumber="41" reason="3">
<taxonomicName id="4C554D537B18FF87FF6CF9D5FD0EDE0D" authority="Garth, 1949" authorityName="Garth" authorityYear="1949" box="[192,752,1539,1564]" class="Insecta" family="Nymphalidae" genus="Speyeria" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="41" phylum="Arthropoda" rank="subSpecies" species="hesperis" subSpecies="schellbachi">
<emphasis id="B921EAC27B18FF87FF6CF9D5FD0EDE0D" bold="true" box="[192,752,1539,1564]" pageId="42" pageNumber="41">
<emphasis id="B921EAC27B18FF87FF6CF9D5FDB1DE0A" bold="true" box="[192,591,1541,1564]" italics="true" pageId="42" pageNumber="41">Speyeria hesperis schellbachi</emphasis>
<bibRefCitation id="EFC44B217B18FF87FDF8F9D3FD0EDE0D" author="Garth, J. S." box="[596,752,1539,1563]" pageId="42" pageNumber="41" pagination="1" refId="ref39029" refString="Garth, J. S. 1949. Studies in Arizona Lepidoptera. I. A new subspecies of Speyeria atlantis (Edwards) from the Kaibab Plateau, Grand Canyon National Park. Bulletin of the Southern California Academy of Sciences 48: 1." type="journal article" year="1949">Garth, 1949</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C34F655B7B18FF87FF6CF9F3FEBADE2D" box="[192,324,1571,1595]" pageId="42" pageNumber="41" type="description">
<paragraph id="8BEA36D07B18FF87FF6CF9F3FEBADE2D" blockId="42.[192,752,1539,1595]" box="[192,324,1571,1595]" pageId="42" pageNumber="41">
(
<figureCitation id="136E2A557B18FF87FF64F9F3FEC2DE2D" box="[200,316,1571,1595]" captionStart="Figure 41-45" captionStartId="73.[192,276,1645,1667]" captionTargetBox="[192,1443,192,1614]" captionTargetId="figure-15@73.[192,1444,192,1614]" captionTargetPageId="73" captionText="Figure 41-45. Type images for Speyeria spp.; dorsal, ventral, and label images. 41) Speyeria hesperis electa lectotype, described as Argynnis Electa, male, CMNH. 42) Speyeria hesperis nikias lectotype, described as Argynnis nikias, male, CMNH. Now synonymized under Speyeria hesperis electa (Scott et al. 1998). 43) Speyeria hesperis schellbachi holotype, described as Speyeria atlantis schellbachi, male, LACM. 44) Speyeria hesperis nausicaa lectotype, described as Argynnis Nausicaa, male, CMNH. 45) Speyeria hesperis dorothea holotype, described as Speyeria atlantis dorothea, male, AMNH. Images 41-42 and 44-45 by James C. Dunford; 43 by Weiping Xie." figureDoi="http://doi.org/10.5281/zenodo.5352686" httpUri="https://zenodo.org/record/5352686/files/figure.png" pageId="42" pageNumber="41">Figure 43</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34F655B7B18FF87FF6CF9B4FCA8DE8D" pageId="42" pageNumber="41" type="reference_group">
<paragraph id="8BEA36D07B18FF87FF6CF9B4FD22DE6D" blockId="42.[192,854,1635,1691]" box="[192,732,1635,1659]" pageId="42" pageNumber="41">
<treatmentCitationGroup id="AB4511FE7B18FF87FF6CF9B4FD22DE6D" box="[192,732,1635,1659]" pageId="42" pageNumber="41">
<treatmentCitation id="0AF410C17B18FF87FF6CF9B4FD28DE6D" author="Garth, J. S." box="[192,726,1635,1659]" page="1" pageId="42" pageNumber="41" year="1949">
<taxonomicName id="4C554D537B18FF87FF6CF9B4FD28DE6D" ID-CoL="5L8QB" authority="Garth, 1949: 1" authorityName="Garth" authorityPageNumber="1" authorityYear="1949" box="[192,726,1635,1659]" class="Insecta" family="Nymphalidae" genus="Speyeria" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="41" phylum="Arthropoda" rank="subSpecies" species="atlantis" subSpecies="schellbachi">
<emphasis id="B921EAC27B18FF87FF6CF9B4FDDEDE6D" box="[192,544,1635,1659]" italics="true" pageId="42" pageNumber="41">Speyeria atlantis schellbachi</emphasis>
<bibRefCitation id="EFC44B217B18FF87FD89F9B3FD28DE6D" author="Garth, J. S." box="[549,726,1635,1659]" pageId="42" pageNumber="41" pagination="1" refId="ref39029" refString="Garth, J. S. 1949. Studies in Arizona Lepidoptera. I. A new subspecies of Speyeria atlantis (Edwards) from the Kaibab Plateau, Grand Canyon National Park. Bulletin of the Southern California Academy of Sciences 48: 1." type="journal article" year="1949">Garth, 1949: 1</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEA36D07B18FF87FF6CF954FCA8DE8D" blockId="42.[192,854,1635,1691]" box="[192,854,1667,1691]" pageId="42" pageNumber="41">
<taxonomicName id="4C554D537B18FF87FF6CF954FD8ADE8D" authority="Garth" authorityName="Garth" authorityYear="1949" box="[192,628,1667,1691]" class="Insecta" family="Nymphalidae" genus="Speyeria" kingdom="Animalia" order="Lepidoptera" pageId="42" pageNumber="41" phylum="Arthropoda" rank="subSpecies" species="hesperis" subSpecies="schellbachi">
<emphasis id="B921EAC27B18FF87FF6CF954FDDBDE8D" box="[192,549,1667,1691]" italics="true" pageId="42" pageNumber="41">Speyeria hesperis schellbachi</emphasis>
Garth
</taxonomicName>
[
<bibRefCitation id="EFC44B217B18FF87FD2DF953FCB5DE8D" author="Scott, J. A. &amp; N. G. Kondla &amp; S. M. Spomer" box="[641,843,1667,1691]" pageId="42" pageNumber="41" pagination="1 - 31" refId="ref45136" refString="Scott, J. A., N. G. Kondla, and S. M. Spomer. 1998. Speyeria hesperis and Speyeria atlantis are distinct species. Papilio 8: 1 - 31." type="journal article" year="1998">Scott et al. 1998</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="C34F655B7B18FF87FF6CF913FD54DECD" box="[192,682,1731,1755]" pageId="42" pageNumber="41" type="description">
<paragraph id="8BEA36D07B18FF87FF6CF913FD54DECD" blockId="42.[192,682,1731,1755]" box="[192,682,1731,1755]" pageId="42" pageNumber="41">
<emphasis id="B921EAC27B18FF87FF6CF913FE6BDECD" bold="true" box="[192,405,1731,1755]" pageId="42" pageNumber="41">Common name.</emphasis>
Schellbachs fritillary.
</paragraph>
</subSubSection>
<subSubSection id="C34F655B7B18FF86FF6CF8D3FC30D9CD" lastPageId="43" lastPageNumber="42" pageId="42" pageNumber="41" type="materials_examined">
<paragraph id="8BEA36D07B18FF87FF6CF8D3FAF2DF0D" blockId="42.[192,1292,1795,1819]" box="[192,1292,1795,1819]" pageId="42" pageNumber="41">
<emphasis id="B921EAC27B18FF87FF6CF8D3FE66DF0D" bold="true" box="[192,408,1795,1819]" pageId="42" pageNumber="41">Type deposited.</emphasis>
<materialsCitation id="3B3D3C8D7B18FF87FE32F8D3FAF2DF0D" box="[414,1292,1795,1819]" county="Natural History Museum of Los Angeles County" location="Natural History Museum of Los Angeles County" pageId="42" pageNumber="41" specimenCount="1" typeStatus="holotype">
<typeStatus id="54EE88727B18FF87FE32F8D3FDF3DF0D" box="[414,525,1795,1819]" pageId="42" pageNumber="41" type="holotype">Holotype</typeStatus>
at
<collectingCounty id="628B4E5C7B18FF87FD9CF8D3FB86DF0D" box="[560,1144,1795,1819]" pageId="42" pageNumber="41">Natural History Museum of Los Angeles County</collectingCounty>
(
<figureCitation id="136E2A557B18FF87FB2AF8D3FB01DF0D" box="[1158,1279,1795,1819]" captionStart="Figure 41-45" captionStartId="73.[192,276,1645,1667]" captionTargetBox="[192,1443,192,1614]" captionTargetId="figure-15@73.[192,1444,192,1614]" captionTargetPageId="73" captionText="Figure 41-45. Type images for Speyeria spp.; dorsal, ventral, and label images. 41) Speyeria hesperis electa lectotype, described as Argynnis Electa, male, CMNH. 42) Speyeria hesperis nikias lectotype, described as Argynnis nikias, male, CMNH. Now synonymized under Speyeria hesperis electa (Scott et al. 1998). 43) Speyeria hesperis schellbachi holotype, described as Speyeria atlantis schellbachi, male, LACM. 44) Speyeria hesperis nausicaa lectotype, described as Argynnis Nausicaa, male, CMNH. 45) Speyeria hesperis dorothea holotype, described as Speyeria atlantis dorothea, male, AMNH. Images 41-42 and 44-45 by James C. Dunford; 43 by Weiping Xie." figureDoi="http://doi.org/10.5281/zenodo.5352686" httpUri="https://zenodo.org/record/5352686/files/figure.png" pageId="42" pageNumber="41">Figure 43</figureCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="8BEA36D07B18FF86FF6CF893FB44D8CD" blockId="42.[192,1217,1859,1883]" lastBlockId="43.[192,1210,195,219]" lastPageId="43" lastPageNumber="42" pageId="42" pageNumber="41">
<emphasis id="B921EAC27B18FF87FF6CF893FE84DF4D" bold="true" box="[192,378,1859,1883]" pageId="42" pageNumber="41">
<typeStatus id="54EE88727B18FF87FF6CF893FEFBDF4D" box="[192,261,1859,1883]" pageId="42" pageNumber="41">Type</typeStatus>
locality.
</emphasis>
Neal Spring, north rim of Grand Canyon, Coconino County,
<collectingRegion id="4991F8327B18FF87FBFBF893FB42DF4D" box="[1111,1212,1859,1883]" country="United States of America" name="Arizona" pageId="42" pageNumber="41">Arizona</collectingRegion>
.
<emphasis id="B921EAC27B19FF86FF6CFF13FE64D8CD" bold="true" box="[192,410,195,219]" pageId="43" pageNumber="42">
<typeStatus id="54EE88727B19FF86FF6CFF13FEFBD8CD" box="[192,261,195,219]" pageId="43" pageNumber="42">Type</typeStatus>
label data.
</emphasis>
I have not examined the label data associated with this specimen.
</paragraph>
<paragraph id="8BEA36D07B19FF86FF6CFED3FD84D94D" blockId="43.[192,1440,259,347]" pageId="43" pageNumber="42">
<emphasis id="B921EAC27B19FF86FF6CFED3FD21D90D" bold="true" box="[192,735,259,283]" pageId="43" pageNumber="42">Identification, taxonomy, and variation.</emphasis>
Adults are bright orange-yellow dorsally and dark basally. Adult wingspan ranges from
<quantity id="4CAD9B357B19FF86FDB2FEF3FD64D92D" box="[542,666,291,315]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="43" pageNumber="42" unit="mm" value="55.0" valueMax="60.0" valueMin="50.0">50-60 mm</quantity>
. This subspecies somewhat resembles
<emphasis id="B921EAC27B19FF86FBC5FEF4FB03D92D" box="[1129,1277,291,315]" italics="true" pageId="43" pageNumber="42">
S. h.
<taxonomicName id="4C554D537B19FF86FB09FEF3FB03D92D" authorityName="Edwards" authorityYear="1879" box="[1189,1277,291,315]" class="Insecta" family="Nymphalidae" genus="Argynnis" kingdom="Animalia" order="Lepidoptera" pageId="43" pageNumber="42" phylum="Arthropoda" rank="species" species="chitone">chitone</taxonomicName>
</emphasis>
but is always silver on the ventral hindwing spots.
</paragraph>
<paragraph id="8BEA36D07B19FF86FF6CFE53FC7FD98D" blockId="43.[192,897,387,411]" box="[192,897,387,411]" pageId="43" pageNumber="42">
<emphasis id="B921EAC27B19FF86FF6CFE53FEE1D98D" bold="true" box="[192,287,387,411]" pageId="43" pageNumber="42">Range.</emphasis>
Kaibab Plateau, near the Grand Canyon,
<collectingRegion id="4991F8327B19FF86FCB4FE53FC82D98D" box="[792,892,387,411]" country="United States of America" name="Arizona" pageId="43" pageNumber="42">Arizona</collectingRegion>
.
</paragraph>
<paragraph id="8BEA36D07B19FF86FF6CFE13FC30D9CD" blockId="43.[192,974,451,475]" box="[192,974,451,475]" pageId="43" pageNumber="42">
<emphasis id="B921EAC27B19FF86FF6CFE13FE96D9CD" bold="true" box="[192,360,451,475]" pageId="43" pageNumber="42">Life history.</emphasis>
Adults are active in secluded draws along springs.
</paragraph>
</subSubSection>
</treatment>
</document>