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<mods:title id="8E09EC66F445DF215DD001EB0719982E">UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification</mods:title>
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<mods:namePart id="F06D35AA09A406E4F4DC3B2BDC183D62">Camacho, G. P.</mods:namePart>
<mods:affiliation id="F1659543700D33F834F5C12B203CD17F">Programa de Pós-Graduação em Entomologia, Departamento de Zoologia, Universidade Federal do Paraná, 82590 - 300, Curitiba, Brazil,</mods:affiliation>
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<mods:namePart id="43D95F6B64BD0F61D9B8759794525661">Franco, W.</mods:namePart>
<mods:affiliation id="E54EA2B2478E76570D0BA37E8144A96B">Programa de Pós-Graduação em Entomologia, Departamento de Zoologia, Universidade Federal do Paraná, 82590 - 300, Curitiba, Brazil,</mods:affiliation>
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<mods:namePart id="A48AFE359605BF84567B75C38FC12E99">Branstetter, M. G.</mods:namePart>
<mods:affiliation id="A388B46092083CCE9EB306C128B1EAC2">U. S. Department of Agriculture, Agricultural Research Service (USDA-ARS), Pollinating Insects Research Unit, Utah State University, 84322, Logan, UT, USA,</mods:affiliation>
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<mods:namePart id="01B1A1DF24F5558FCB695A788D7F203F">Pie, M. R.</mods:namePart>
<mods:affiliation id="E076E3C6FF763E213A9689FBA4E04BE1">Programa de Pós-Graduação em Zoologia, Departamento de Zoologia, Universidade Federal do Paraná, 82590 - 300, Curitiba, Brazil,</mods:affiliation>
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<mods:namePart id="F7C20E4099CA02F1BBD83DACD1D94DFE">Longino, J. T.</mods:namePart>
<mods:affiliation id="61DF9C384C806F4827541DB42E806D1A">Department of Biology, University of Utah, 84112, Salt Lake City, UT, USA, and</mods:affiliation>
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<mods:title id="264B55813D332B28BE91A15F67D0C192">Insect Systematics and Diversity</mods:title>
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<mods:date id="F368BBE642AFDAC0390C73C3B072E945">2022</mods:date>
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<subSubSection id="C34F6537FFE7FFE4FF60F8BAFDB3CB98" box="[146,576,391,411]" pageId="12" pageNumber="13" type="nomenclature">
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<emphasis id="B921EAAEFFE7FFE4FF60F8BAFDB3CB98" bold="true" box="[146,576,391,411]" pageId="12" pageNumber="13">
Tribe
<taxonomicName id="4C554D3FFFE7FFE4FF3BF8BAFE68CB98" authority="Bolton" authorityName="Bolton" authorityYear="2022" baseAuthorityName="Camacho &amp; Franco &amp; Branstetter &amp; Pie &amp; Longino" baseAuthorityYear="2022" box="[201,411,391,411]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="tribe" status="new combination" tribe="Heteroponerini">Heteroponerini Bolton</taxonomicName>
<taxonomicNameLabel id="A21257D5FFE7FFE4FE52F8BAFDB3CB98" box="[416,576,391,411]" pageId="12" pageNumber="13" rank="species">new combination</taxonomicNameLabel>
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<paragraph id="8BEA36BCFFE7FFE4FF60F8F8FDC7C918" blockId="12.[145,771,452,795]" pageId="12" pageNumber="13">
<emphasis id="B921EAAEFFE7FFE4FF60F8F8FF00CBD4" box="[146,243,453,471]" italics="true" pageId="12" pageNumber="13">Diagnosis:</emphasis>
Ectatommine ants of small to medium size (head width
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, head length
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); cephalic dorsum with a longitudinal carina extending from anterior margin of clypeus to posterior margin of head (
<bibRefCitation id="EFC44B4DFFE7FFE4FE61FB21FDF9C833" author="Bolton, B." box="[403,522,540,560]" pageId="12" pageNumber="13" pagination="1 - 370" refId="ref17416" refString="Bolton, B. 2003. Synopsis and classification of Formicidae. Mem. Am. Entomol. Inst. 71: 1 - 370." type="journal article" year="2003">Bolton 2003</bibRefCitation>
); antennal scrobe present (
<bibRefCitation id="EFC44B4DFFE7FFE4FF6BFB04FEF9C84E" author="Bolton, B." box="[153,266,569,589]" pageId="12" pageNumber="13" pagination="1 - 370" refId="ref17416" refString="Bolton, B. 2003. Synopsis and classification of Formicidae. Mem. Am. Entomol. Inst. 71: 1 - 370." type="journal article" year="2003">Bolton 2003</bibRefCitation>
); eye present; acetabulum of antennal socket apparatus hemispherical (
<bibRefCitation id="EFC44B4DFFE7FFE4FEEDFB6AFE7FC868" author="Keller, R. A." box="[287,396,599,619]" pageId="12" pageNumber="13" pagination="1 - 90" refId="ref19678" refString="Keller, R. A. 2011. A phylogenetic analysis of ant morphology (Hymenoptera: Formicidae) with special reference to the Poneromorph subfamilies. Bull. Am. Mus. Nat. 355: 1 - 90." type="journal article" year="2011">Keller 2011</bibRefCitation>
); accessory chamber of antennal socket absent (
<bibRefCitation id="EFC44B4DFFE7FFE4FF2DFB49FEA3C88B" author="Keller, R. A." box="[223,336,628,648]" pageId="12" pageNumber="13" pagination="1 - 90" refId="ref19678" refString="Keller, R. A. 2011. A phylogenetic analysis of ant morphology (Hymenoptera: Formicidae) with special reference to the Poneromorph subfamilies. Bull. Am. Mus. Nat. 355: 1 - 90." type="journal article" year="2011">Keller 2011</bibRefCitation>
); labial palp with three or four palpomeres (
<bibRefCitation id="EFC44B4DFFE7FFE4FF6BFBACFEF8C8A6" author="Bolton, B." box="[153,267,657,677]" pageId="12" pageNumber="13" pagination="1 - 370" refId="ref17416" refString="Bolton, B. 2003. Synopsis and classification of Formicidae. Mem. Am. Entomol. Inst. 71: 1 - 370." type="journal article" year="2003">Bolton 2003</bibRefCitation>
,
<bibRefCitation id="EFC44B4DFFE7FFE4FEE5FBACFE71C8A6" author="Keller, R. A." box="[279,386,657,677]" pageId="12" pageNumber="13" pagination="1 - 90" refId="ref19678" refString="Keller, R. A. 2011. A phylogenetic analysis of ant morphology (Hymenoptera: Formicidae) with special reference to the Poneromorph subfamilies. Bull. Am. Mus. Nat. 355: 1 - 90." type="journal article" year="2011">Keller 2011</bibRefCitation>
); promesonotal suture complete and flexible (
<bibRefCitation id="EFC44B4DFFE7FFE4FF30FB92FECBC8C0" author="Bolton, B." box="[194,312,687,707]" pageId="12" pageNumber="13" pagination="1 - 370" refId="ref17416" refString="Bolton, B. 2003. Synopsis and classification of Formicidae. Mem. Am. Entomol. Inst. 71: 1 - 370." type="journal article" year="2003">Bolton 2003</bibRefCitation>
); ventral flap on metapleural gland opening absent (
<bibRefCitation id="EFC44B4DFFE7FFE4FF37FBF1FEC1C8E3" author="Keller, R. A." box="[197,306,716,736]" pageId="12" pageNumber="13" pagination="1 - 90" refId="ref19678" refString="Keller, R. A. 2011. A phylogenetic analysis of ant morphology (Hymenoptera: Formicidae) with special reference to the Poneromorph subfamilies. Bull. Am. Mus. Nat. 355: 1 - 90." type="journal article" year="2011">Keller 2011</bibRefCitation>
); metacoxal cavity closed (
<bibRefCitation id="EFC44B4DFFE7FFE4FDC0FBF1FD55C8E3" author="Bolton, B." box="[562,678,716,736]" pageId="12" pageNumber="13" pagination="1 - 370" refId="ref17416" refString="Bolton, B. 2003. Synopsis and classification of Formicidae. Mem. Am. Entomol. Inst. 71: 1 - 370." type="journal article" year="2003">Bolton 2003</bibRefCitation>
); petiolar sternite articulated with tergite over its entire length (
<bibRefCitation id="EFC44B4DFFE7FFE4FD7AFBD4FD0FC8FE" author="Bolton, B." box="[648,764,745,765]" pageId="12" pageNumber="13" pagination="1 - 370" refId="ref17416" refString="Bolton, B. 2003. Synopsis and classification of Formicidae. Mem. Am. Entomol. Inst. 71: 1 - 370." type="journal article" year="2003">Bolton 2003</bibRefCitation>
,
<bibRefCitation id="EFC44B4DFFE7FFE4FF60FA3AFF0EC918" author="Keller, R. A." box="[146,253,775,795]" pageId="12" pageNumber="13" pagination="1 - 90" refId="ref19678" refString="Keller, R. A. 2011. A phylogenetic analysis of ant morphology (Hymenoptera: Formicidae) with special reference to the Poneromorph subfamilies. Bull. Am. Mus. Nat. 355: 1 - 90." type="journal article" year="2011">Keller 2011</bibRefCitation>
); laterotergites of petiole present.
</paragraph>
<paragraph id="8BEA36BCFFE7FFE4FCC0F9A8FC44CAC6" blockId="12.[818,1443,148,197]" pageId="12" pageNumber="13">
<emphasis id="B921EAAEFFE7FFE4FCC0F9A8FBE2CAAB" box="[818,1041,148,168]" italics="true" pageId="12" pageNumber="13">
Genera:
<taxonomicName id="4C554D3FFFE7FFE4FC76F9A9FBE2CAAB" authorityName="Mayr" authorityYear="1862" box="[900,1041,148,168]" class="Insecta" family="Formicidae" genus="Acanthoponera" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">Acanthoponera</taxonomicName>
</emphasis>
,
<taxonomicName id="4C554D3FFFE7FFE4FBC3F9A9FB6CCAAB" box="[1073,1183,148,168]" class="Insecta" family="Formicidae" genus="Aulacopone" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAAEFFE7FFE4FBC3F9A9FB6CCAAB" box="[1073,1183,148,168]" italics="true" pageId="12" pageNumber="13">Aulacopone</emphasis>
</taxonomicName>
,
<taxonomicName id="4C554D3FFFE7FFE4FB4DF9A9FAFDCAAB" authority="Camacho &amp; Franco &amp; Branstetter &amp; Pie &amp; Longino, 2022" authorityName="Camacho &amp; Franco &amp; Branstetter &amp; Pie &amp; Longino" authorityYear="2022" box="[1215,1294,148,168]" class="Magnoliopsida" family="Asteraceae" genus="Boltonia" kingdom="Plantae" order="Asterales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="genus" status="new genus">
<emphasis id="B921EAAEFFE7FFE4FB4DF9A9FAFDCAAB" box="[1215,1294,148,168]" italics="true" pageId="12" pageNumber="13">Boltonia</emphasis>
</taxonomicName>
<emphasis id="B921EAAEFFE7FFE4FADAF9A9FA6FCAAB" bold="true" box="[1320,1436,148,168]" pageId="12" pageNumber="13">
<taxonomicNameLabel id="A21257D5FFE7FFE4FADAF9A9FA6FCAAB" box="[1320,1436,148,168]" pageId="12" pageNumber="13" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C554D3FFFE7FFE4FCC0F98FFC42CAC7" authorityName="Mayr" authorityYear="1887" box="[818,945,178,196]" class="Insecta" family="Formicidae" genus="Heteroponera" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B921EAAEFFE7FFE4FCC0F98FFC42CAC7" box="[818,945,178,196]" italics="true" pageId="12" pageNumber="13">Heteroponera</emphasis>
</taxonomicName>
.
</paragraph>
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