244 lines
37 KiB
XML
244 lines
37 KiB
XML
<document id="930029AC26F8469EA61AA3012401B01C" ID-DOI="10.5852/ejt.2021.755.1397" ID-GBIF-Dataset="8721d89e-82a8-425a-8707-f53293bbef22" ID-ISSN="2118-9773" ID-Zenodo-Dep="4983284" ID-ZooBank="CEB00D62-0890-403C-8C54-968DC52C8290" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="julia" IM.tables_approvedBy="julia" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624019007013" checkinUser="felipe" docAuthor="Moritz, Leif & Wesener, Thomas" docDate="2021" docId="03FC2A12FF85FFAD1A0EFBE0FB53FA8C" docLanguage="en" docName="ejt-755_22-46.pdf" docOrigin="European Journal of Taxonomy 755 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.755.1397" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Kachincambala muelleri Moritz & Wesener 2021, gen. et sp. nov." docType="treatment" docUuid="F9CC6697-1E02-4A41-AABA-1E5EE46D7AFA" docUuidSource="ZooBank" docVersion="10" lastPageNumber="35" masterDocId="FFC5526AFF8EFFA01827FF91FFE9FFB3" masterDocTitle="Electrocambalidae fam. nov., a new family of Cambalidea from Cretaceous Burmese amber (Diplopoda, Spirostreptida)" masterLastPageNumber="46" masterPageNumber="22" pageNumber="33" updateTime="1701997971992" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C9ABF439FC960954E4CFB5DB92D752B4">Electrocambalidae fam. nov., a new family of Cambalidea from Cretaceous Burmese amber (Diplopoda, Spirostreptida)</mods:title>
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<mods:name id="480DB0A6A7EFDA48E822A6C9F92DFFDC" type="personal">
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<mods:roleTerm id="54A981E1A82978D5D5742F67803B1036">Author</mods:roleTerm>
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<mods:namePart id="CAA2C9F3A17D41156C54F6B6FAF7EE63">Moritz, Leif</mods:namePart>
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<mods:affiliation id="FAAB9D124B27FB6E8EAA91DD22223C06">Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Adenauerallee 160, D- 53113, Bonn, Germany. & Institute of Evolutionary Biology and Ecology, University of Bonn, An der Immenburg 1, 53121 Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier id="0E5EC14EC7E5E7D08F4573A544221A2D" type="email">moritz.leif@gmail.com</mods:nameIdentifier>
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<mods:namePart id="3A3AF8FBD32AC3573B5F20AEEBB961C6">Wesener, Thomas</mods:namePart>
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<mods:affiliation id="C921D27E94C92A7AC7B1ABADB7B34ECE">Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Adenauerallee 160, D- 53113, Bonn, Germany.</mods:affiliation>
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<mods:nameIdentifier id="39C3675550D8AF6FCA9AA8EA84E55637" type="email">t.wesener@leibniz-zfmk.de</mods:nameIdentifier>
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<mods:title id="32EADC67818D9ECBFA495A51F8671E57">European Journal of Taxonomy</mods:title>
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<mods:part id="D514A822A621D1CE6D3FAA60F2AB9FAF">
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<mods:date id="9FA696678CFC947944FFF997B801AC90">2021</mods:date>
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<mods:number id="6BA399CFDA6CD176F81E6355278FA89A">2021-06-17</mods:number>
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<treatment id="03FC2A12FF85FFAD1A0EFBE0FB53FA8C" ID-DOI="http://doi.org/10.5281/zenodo.4984693" ID-GBIF-Taxon="182958556" ID-Zenodo-Dep="4984693" LSID="urn:lsid:zoobank.org:act:F9CC6697-1E02-4A41-AABA-1E5EE46D7AFA" httpUri="http://treatment.plazi.org/id/03FC2A12FF85FFAD1A0EFBE0FB53FA8C" lastPageId="13" lastPageNumber="35" pageId="11" pageNumber="33">
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<subSubSection id="C34FC88FFF85FFAB1A0EFBE0FBE2FB3F" box="[553,1035,1137,1164]" pageId="11" pageNumber="33" type="nomenclature">
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<paragraph id="8BEA9B04FF85FFAB1A0EFBE0FBE2FB3F" blockId="11.[356,1232,1137,1232]" box="[553,1035,1137,1164]" pageId="11" pageNumber="33">
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<heading id="D0A22C68FF85FFAB1A0EFBE0FBE2FB3F" box="[553,1035,1137,1164]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="33" reason="2">
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<taxonomicName id="4C55E087FF85FFAB1A0EFBE0FCBDFB3F" authority="Moritz & Wesener, 2021" authorityName="Moritz & Wesener" authorityYear="2021" box="[553,852,1137,1164]" class="Diplopoda" family="Pseudonannolenidae" genus="Kachincambala" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="33" phylum="Arthropoda" rank="species" species="muelleri" status="gen. et sp. nov.">
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<emphasis id="B9214716FF85FFAB1A0EFBE0FCBDFB3F" bold="true" box="[553,852,1137,1164]" italics="true" pageId="11" pageNumber="33">Kachincambala muelleri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A212FA6DFF85FFAB1B7CFBE3FBE2FB3F" box="[859,1035,1138,1164]" pageId="11" pageNumber="33" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="C34FC88FFF85FFAB1943FB05FCD5FB63" pageId="11" pageNumber="33" type="description">
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<paragraph id="8BEA9B04FF85FFAB1943FB05FB39FB1C" blockId="11.[356,1232,1137,1232]" box="[356,1232,1172,1199]" pageId="11" pageNumber="33">
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<uri id="FFC49706FF85FFAB1943FB05FB39FB1C" box="[356,1232,1172,1199]" pageId="11" pageNumber="33">
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urn:lsid:zoobank.org:act:
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<uuid id="FFF3A1D1FF85FFAB1AA3FB05FB39FB1C" box="[644,1232,1172,1199]" pageId="11" pageNumber="33">F9CC6697-1E02-4A41-AABA-1E5EE46D7AFA</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BEA9B04FF85FFAB1AD0FB27FCD5FB63" blockId="11.[356,1232,1137,1232]" box="[759,828,1206,1232]" pageId="11" pageNumber="33">
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<figureCitation id="136E8781FF85FFAB1AD0FB27FCD5FB63" box="[759,828,1206,1232]" captionStart="Fig" captionStartId="12.[188,231,1787,1813]" captionTargetBox="[188,1399,529,1730]" captionTargetId="figure-117@12.[188,1399,529,1750]" captionTargetPageId="12" captionText="Fig. 4. Kachincambala muelleri gen. et sp. nov., holotype, ♂ (ZFMK-MYR10225). A. Habitus, dorsal view, photograph. B. Head and anterior body rings, ventral view, volume rendering. C. Head and anterior body rings, lateral view, volume rendering. D. head, frontal view, volume rendering. E. Head, ventral view, volume rendering. F. Gonopods, frontal view, segmentation. G. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; at = antennae; co = collum; cx = coxite; em = eumentum; gn = gnathochilarium; L = leg; la = labrum; ll = lamella limgualis; md = mandible; om = ommatidia; pe = penis; pg = posterior gonopod; pm = promentum; st = gnathochilarial stipes; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983295" httpUri="https://zenodo.org/record/4983295/files/figure.png" pageId="11" pageNumber="33">Fig. 4</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34FC88FFF85FFAB189AFB66FADAFA5F" pageId="11" pageNumber="33" type="diagnosis">
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||
<paragraph id="8BEA9B04FF85FFAB189AFB66FEDFFAA2" blockId="11.[189,310,1271,1297]" box="[189,310,1271,1297]" pageId="11" pageNumber="33">
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<heading id="D0A22C68FF85FFAB189AFB66FEDFFAA2" bold="true" box="[189,310,1271,1297]" fontSize="11" level="3" pageId="11" pageNumber="33" reason="3">
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<emphasis id="B9214716FF85FFAB189AFB66FEDFFAA2" bold="true" box="[189,310,1271,1297]" pageId="11" pageNumber="33">Diagnosis</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8BEA9B04FF85FFAB189AFAB4FADAFA5F" blockId="11.[187,1399,1317,1517]" pageId="11" pageNumber="33">
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<taxonomicName id="4C55E087FF85FFAB189AFAB4FE0DFA8C" authority="Moritz & Wesener, 2021" authorityName="Moritz & Wesener" authorityYear="2021" box="[189,484,1317,1343]" class="Diplopoda" family="Pseudonannolenidae" genus="Kachincambala" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="33" phylum="Arthropoda" rank="species" species="muelleri" status="gen. et sp. nov.">
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<emphasis id="B9214716FF85FFAB189AFAB4FE0DFA8C" box="[189,484,1317,1343]" italics="true" pageId="11" pageNumber="33">Kachincambala muelleri</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A212FA6DFF85FFAB19CAFAB7FD4AFAF3" box="[493,675,1318,1344]" pageId="11" pageNumber="33" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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differs from the only other species of the genus,
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<taxonomicName id="4C55E087FF85FFAB1CD1FAB7FA9FFA8C" authority="Moritz & Wesener, 2021" authorityName="Moritz & Wesener" authorityYear="2021" box="[1270,1398,1317,1343]" class="Diplopoda" family="Pseudonannolenidae" genus="Kachincambala" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="33" phylum="Arthropoda" rank="species" species="distorta" status="gen. et sp. nov.">
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<emphasis id="B9214716FF85FFAB1CD1FAB7FA9FFA8C" box="[1270,1398,1317,1343]" italics="true" pageId="11" pageNumber="33">K. distorta</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A212FA6DFF85FFAB189AFAD9FE86FAD1" box="[189,367,1352,1378]" pageId="11" pageNumber="33" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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, in gonopod characters. The anterior gonopod coxite is more stout in
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<taxonomicName id="4C55E087FF85FFAB1CBAFAD8FAC8FAD1" authority="Moritz & Wesener, 2021" authorityName="Moritz & Wesener" authorityYear="2021" box="[1181,1313,1352,1378]" class="Diplopoda" family="Pseudonannolenidae" genus="Kachincambala" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="33" phylum="Arthropoda" rank="species" species="muelleri" status="gen. et sp. nov.">
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<emphasis id="B9214716FF85FFAB1CBAFAD8FAC8FAD1" box="[1181,1313,1352,1378]" italics="true" pageId="11" pageNumber="33">K. muelleri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A212FA6DFF85FFAB1D0FFAD9FEF3FA36" pageId="11" pageNumber="33" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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, less than twice as long as wide, while it is more narrow in
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<taxonomicName id="4C55E087FF85FFAB1BEAFAFAFBA3FA37" authority="Moritz & Wesener, 2021" authorityName="Moritz & Wesener" authorityYear="2021" box="[973,1098,1386,1412]" class="Diplopoda" family="Pseudonannolenidae" genus="Kachincambala" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="33" phylum="Arthropoda" rank="species" species="distorta" status="gen. et sp. nov.">
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<emphasis id="B9214716FF85FFAB1BEAFAFAFBA3FA37" box="[973,1098,1386,1412]" italics="true" pageId="11" pageNumber="33">K. distorta</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A212FA6DFF85FFAB1C76FAFAFAE9FA36" box="[1105,1280,1387,1413]" pageId="11" pageNumber="33" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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The inner lobe is apically rounded, nearly triangular and bends mesally. The outer lobe of the anterior gonopod is short and broad, oriented more laterally than in
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<taxonomicName id="4C55E087FF85FFAB1AF9FA20FCB3FA79" authority="Moritz & Wesener, 2021" authorityName="Moritz & Wesener" authorityYear="2021" box="[734,858,1456,1482]" class="Diplopoda" family="Pseudonannolenidae" genus="Kachincambala" kingdom="Animalia" order="Spirostreptida" pageId="11" pageNumber="33" phylum="Arthropoda" rank="species" species="distorta" status="gen. et sp. nov.">
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<emphasis id="B9214716FF85FFAB1AF9FA20FCB3FA79" box="[734,858,1456,1482]" italics="true" pageId="11" pageNumber="33">K. distorta</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A212FA6DFF85FFAB1B78FA21FBE3FA79" box="[863,1034,1456,1482]" pageId="11" pageNumber="33" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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Telopodite of anterior gonopod is broad, shorter than coxite. Posterior gonopod simple, broad, reaching over inner lobe of coxite.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34FC88FFF85FFAB189AF985FD4DF912" pageId="11" pageNumber="33" type="etymology">
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<paragraph id="8BEA9B04FF85FFAB189AF985FEADF99D" blockId="11.[189,324,1556,1582]" box="[189,324,1556,1582]" pageId="11" pageNumber="33">
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||
<heading id="D0A22C68FF85FFAB189AF985FEADF99D" bold="true" box="[189,324,1556,1582]" fontSize="11" level="3" pageId="11" pageNumber="33" reason="3">
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<emphasis id="B9214716FF85FFAB189AF985FEADF99D" bold="true" box="[189,324,1556,1582]" pageId="11" pageNumber="33">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BEA9B04FF85FFAB189AF9D3FD4DF912" blockId="11.[189,1398,1601,1697]" pageId="11" pageNumber="33">
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In honour of Patrick Müller, who granted us access to his large collection of inclusions in Burmese amber for study, which resulted in the discovery of the specimens studied here, and who donated the
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<typeStatus id="54EE25A6FF85FFAB1CC6F9F4FAFAF9CC" box="[1249,1299,1637,1663]" pageId="11" pageNumber="33">type</typeStatus>
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material to the ZFMK. Noun in the adjective case.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34FC88FFF85FFAB189AF959FC3FF8E5" pageId="11" pageNumber="33" type="materials_examined">
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<paragraph id="8BEA9B04FF85FFAB189AF959FE85F952" blockId="11.[189,364,1735,1762]" box="[189,364,1735,1762]" pageId="11" pageNumber="33">
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<heading id="D0A22C68FF85FFAB189AF959FE85F952" bold="true" box="[189,364,1735,1762]" fontSize="11" level="3" pageId="11" pageNumber="33" reason="3">
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<emphasis id="B9214716FF85FFAB189AF959FE85F952" bold="true" box="[189,364,1735,1762]" pageId="11" pageNumber="33">Type material</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BEA9B04FF85FFAB18CDF967FC3FF8E5" blockId="11.[189,1399,1782,1878]" pageId="11" pageNumber="33">
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<materialsCitation id="3B3D9159FF85FFAB18CDF967FC3FF8E5" ID-GBIF-Occurrence="3303859309" collectionCode="ZFMK-MYR, BuB" country="Myanmar" county="Hukawng Valley" location="Hukawng Valley" municipality="Mid-Cretaceous Burmese" pageId="11" pageNumber="33" specimenCode="BuB3757, ZFMK-MYR10225" specimenCount="1" specimenCount-male="1" stateProvince="Kachin State" typeStatus="holotype">
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<emphasis id="B9214716FF85FFAB18CDF967FEB5F8A3" bold="true" box="[234,348,1782,1808]" pageId="11" pageNumber="33">
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<typeStatus id="54EE25A6FF85FFAB18CDF967FEB5F8A3" box="[234,348,1782,1808]" pageId="11" pageNumber="33" type="holotype">Holotype</typeStatus>
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</emphasis>
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<collectingCountry id="F342DB94FF85FFAB189AF889FEBFF881" box="[189,342,1816,1842]" name="Myanmar" pageId="11" pageNumber="33">MYANMAR</collectingCountry>
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•
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<specimenCount id="9D53508DFF85FFAB1957F888FE60F880" box="[368,393,1817,1843]" pageId="11" pageNumber="33" type="male">♂</specimenCount>
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;
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<collectingRegion id="499155E6FF85FFAB19B1F889FDD8F881" box="[406,561,1816,1842]" country="Myanmar" name="Kachin" pageId="11" pageNumber="33">Kachin State</collectingRegion>
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,
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<collectingCounty id="628BE388FF85FFAB1A1AF889FCEBF881" box="[573,770,1816,1842]" pageId="11" pageNumber="33">Hukawng Valley</collectingCounty>
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,
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<geologicalTimeScale id="F9751BAFFF85FFAB1B29F889FC22F881" box="[782,971,1816,1842]" pageId="11" pageNumber="33">Mid-Cretaceous</geologicalTimeScale>
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Burmese amber inclusion; formerly in the collection of Patrick Müller (
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<specimenCode id="DBF3337FFF85FFAB1A7CF8AAFD3BF8E5" box="[603,722,1851,1878]" collectionCode="BuB" pageId="11" pageNumber="33">BuB3757</specimenCode>
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);
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<specimenCode id="DBF3337FFF85FFAB1AC1F8AAFC3BF8E5" box="[742,978,1851,1878]" collectionCode="ZFMK-MYR" pageId="11" pageNumber="33">ZFMK-MYR10225</specimenCode>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
|
||
<subSubSection id="C34FC88FFF85FFAD189AF8EDFB53FA8C" lastPageId="13" lastPageNumber="35" pageId="11" pageNumber="33" type="description">
|
||
<paragraph id="8BEA9B04FF85FFAB189AF8EDFEA7F825" blockId="11.[189,334,1916,1942]" box="[189,334,1916,1942]" pageId="11" pageNumber="33">
|
||
<heading id="D0A22C68FF85FFAB189AF8EDFEA7F825" bold="true" box="[189,334,1916,1942]" fontSize="11" level="3" pageId="11" pageNumber="33" reason="3">
|
||
<emphasis id="B9214716FF85FFAB189AF8EDFEA7F825" bold="true" box="[189,334,1916,1942]" pageId="11" pageNumber="33">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEA9B04FF85FFAB189AF83BFDB7F85B" blockId="11.[189,1398,1962,2025]" pageId="11" pageNumber="33">
|
||
HABITUS.>41 body rings; cylindrical and slender, up to at least
|
||
<quantity id="4CAD36E1FF85FFAB1B96F83BFBEDF877" box="[945,1028,1962,1988]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="33" unit="mm" value="12.0">12 mm</quantity>
|
||
long and
|
||
<quantity id="4CAD36E1FF85FFAB1C5DF83BFB3CF877" box="[1146,1237,1962,1989]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="33" unit="mm" value="0.5">0.5 mm</quantity>
|
||
wide. Colour dark brown to greyish (
|
||
<figureCitation id="136E8781FF85FFAB19E8F85FFDB8F85B" box="[463,593,1998,2024]" captionStart="Fig" captionStartId="12.[188,231,1787,1813]" captionTargetBox="[188,1399,529,1730]" captionTargetId="figure-117@12.[188,1399,529,1750]" captionTargetPageId="12" captionText="Fig. 4. Kachincambala muelleri gen. et sp. nov., holotype, ♂ (ZFMK-MYR10225). A. Habitus, dorsal view, photograph. B. Head and anterior body rings, ventral view, volume rendering. C. Head and anterior body rings, lateral view, volume rendering. D. head, frontal view, volume rendering. E. Head, ventral view, volume rendering. F. Gonopods, frontal view, segmentation. G. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; at = antennae; co = collum; cx = coxite; em = eumentum; gn = gnathochilarium; L = leg; la = labrum; ll = lamella limgualis; md = mandible; om = ommatidia; pe = penis; pg = posterior gonopod; pm = promentum; st = gnathochilarial stipes; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983295" httpUri="https://zenodo.org/record/4983295/files/figure.png" pageId="11" pageNumber="33">Fig. 4A–C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEA9B04FF82FFAC189AFE98FB81FE60" blockId="12.[189,1399,265,467]" pageId="12" pageNumber="34">
|
||
HEAD. As wide as collum and body rings 6 onwards, wider than body rings 2–5. Head and labrum with setae. Labrum with three teeth (
|
||
<figureCitation id="136E8781FF82FFAC1A1AFEBDFD76FEF5" box="[573,671,300,326]" captionStart="Fig" captionStartId="12.[188,231,1787,1813]" captionTargetBox="[188,1399,529,1730]" captionTargetId="figure-117@12.[188,1399,529,1750]" captionTargetPageId="12" captionText="Fig. 4. Kachincambala muelleri gen. et sp. nov., holotype, ♂ (ZFMK-MYR10225). A. Habitus, dorsal view, photograph. B. Head and anterior body rings, ventral view, volume rendering. C. Head and anterior body rings, lateral view, volume rendering. D. head, frontal view, volume rendering. E. Head, ventral view, volume rendering. F. Gonopods, frontal view, segmentation. G. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; at = antennae; co = collum; cx = coxite; em = eumentum; gn = gnathochilarium; L = leg; la = labrum; ll = lamella limgualis; md = mandible; om = ommatidia; pe = penis; pg = posterior gonopod; pm = promentum; st = gnathochilarial stipes; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983295" httpUri="https://zenodo.org/record/4983295/files/figure.png" pageId="12" pageNumber="34">Fig. 4D</figureCitation>
|
||
).>30 ommatidia arranged in several rows. Incisura lateralis present, closed.Antennae ca
|
||
<quantity id="4CAD36E1FF82FFAC1A23FEDEFDB5FED9" box="[516,604,335,362]" metricMagnitude="-1" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="34" unit="mm" value="800.0">800 µm</quantity>
|
||
long, covered by setae, 7<1<2<4=5<3<6, with four apical sensory cones (
|
||
<figureCitation id="136E8781FF82FFAC1929FEE3FE81FE3E" box="[270,360,370,397]" captionStart="Fig" captionStartId="12.[188,231,1787,1813]" captionTargetBox="[188,1399,529,1730]" captionTargetId="figure-117@12.[188,1399,529,1750]" captionTargetPageId="12" captionText="Fig. 4. Kachincambala muelleri gen. et sp. nov., holotype, ♂ (ZFMK-MYR10225). A. Habitus, dorsal view, photograph. B. Head and anterior body rings, ventral view, volume rendering. C. Head and anterior body rings, lateral view, volume rendering. D. head, frontal view, volume rendering. E. Head, ventral view, volume rendering. F. Gonopods, frontal view, segmentation. G. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; at = antennae; co = collum; cx = coxite; em = eumentum; gn = gnathochilarium; L = leg; la = labrum; ll = lamella limgualis; md = mandible; om = ommatidia; pe = penis; pg = posterior gonopod; pm = promentum; st = gnathochilarial stipes; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983295" httpUri="https://zenodo.org/record/4983295/files/figure.png" pageId="12" pageNumber="34">Fig. 4B</figureCitation>
|
||
). Mandible base with small cardo ca ¼ of length of large stipes (
|
||
<figureCitation id="136E8781FF82FFAC1C63FEE3FB76FE3E" box="[1092,1183,370,397]" captionStart="Fig" captionStartId="12.[188,231,1787,1813]" captionTargetBox="[188,1399,529,1730]" captionTargetId="figure-117@12.[188,1399,529,1750]" captionTargetPageId="12" captionText="Fig. 4. Kachincambala muelleri gen. et sp. nov., holotype, ♂ (ZFMK-MYR10225). A. Habitus, dorsal view, photograph. B. Head and anterior body rings, ventral view, volume rendering. C. Head and anterior body rings, lateral view, volume rendering. D. head, frontal view, volume rendering. E. Head, ventral view, volume rendering. F. Gonopods, frontal view, segmentation. G. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; at = antennae; co = collum; cx = coxite; em = eumentum; gn = gnathochilarium; L = leg; la = labrum; ll = lamella limgualis; md = mandible; om = ommatidia; pe = penis; pg = posterior gonopod; pm = promentum; st = gnathochilarial stipes; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983295" httpUri="https://zenodo.org/record/4983295/files/figure.png" pageId="12" pageNumber="34">Fig. 4C</figureCitation>
|
||
). Gnathochilarium with mentum divided in eumentum and promentum, large triangular lamella lingualis, large stipites occupying entire length of gnathochilarium, with inner and outer palp (
|
||
<figureCitation id="136E8781FF82FFAC1BD8FE28FBB2FE60" box="[1023,1115,441,467]" captionStart="Fig" captionStartId="12.[188,231,1787,1813]" captionTargetBox="[188,1399,529,1730]" captionTargetId="figure-117@12.[188,1399,529,1750]" captionTargetPageId="12" captionText="Fig. 4. Kachincambala muelleri gen. et sp. nov., holotype, ♂ (ZFMK-MYR10225). A. Habitus, dorsal view, photograph. B. Head and anterior body rings, ventral view, volume rendering. C. Head and anterior body rings, lateral view, volume rendering. D. head, frontal view, volume rendering. E. Head, ventral view, volume rendering. F. Gonopods, frontal view, segmentation. G. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; at = antennae; co = collum; cx = coxite; em = eumentum; gn = gnathochilarium; L = leg; la = labrum; ll = lamella limgualis; md = mandible; om = ommatidia; pe = penis; pg = posterior gonopod; pm = promentum; st = gnathochilarial stipes; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983295" httpUri="https://zenodo.org/record/4983295/files/figure.png" pageId="12" pageNumber="34">Fig. 4E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF2ACB8CFF82FFAC189BF96AFB1EF85B" ID-DOI="http://doi.org/10.5281/zenodo.4983295" ID-Zenodo-Dep="4983295" httpUri="https://zenodo.org/record/4983295/files/figure.png" pageId="12" pageNumber="34" startId="12.[188,231,1787,1813]" targetBox="[188,1399,529,1730]" targetPageId="12">
|
||
<paragraph id="8BEA9B04FF82FFAC189BF96AFB1EF85B" blockId="12.[187,1400,1787,2025]" pageId="12" pageNumber="34">
|
||
<emphasis id="B9214716FF82FFAC189BF96AFEE2F8A6" bold="true" box="[188,267,1787,1813]" pageId="12" pageNumber="34">Fig. 4.</emphasis>
|
||
<taxonomicName id="4C55E087FF82FFAC1934F96AFDDEF8A6" authority="Moritz & Wesener, 2021" authorityName="Moritz & Wesener" authorityYear="2021" box="[275,567,1787,1813]" class="Diplopoda" family="Pseudonannolenidae" genus="Kachincambala" kingdom="Animalia" order="Spirostreptida" pageId="12" pageNumber="34" phylum="Arthropoda" rank="species" species="muelleri" status="gen. et sp. nov.">
|
||
<emphasis id="B9214716FF82FFAC1934F96AFDDEF8A6" box="[275,567,1787,1813]" italics="true" pageId="12" pageNumber="34">Kachincambala muelleri</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A212FA6DFF82FFAC1A18F96DFD1DF8A5" box="[575,756,1788,1814]" pageId="12" pageNumber="34" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂ (ZFMK-MYR10225).
|
||
<emphasis id="B9214716FF82FFAC1C87F96AFB5CF8A6" bold="true" box="[1184,1205,1787,1813]" pageId="12" pageNumber="34">A</emphasis>
|
||
. Habitus, dorsal view, photograph.
|
||
<emphasis id="B9214716FF82FFAC19B4F88FFE4EF88B" bold="true" box="[403,423,1822,1848]" pageId="12" pageNumber="34">B</emphasis>
|
||
. Head and anterior body rings, ventral view, volume rendering.
|
||
<emphasis id="B9214716FF82FFAC1CA1F88FFB72F88B" bold="true" box="[1158,1179,1822,1848]" pageId="12" pageNumber="34">C</emphasis>
|
||
. Head and anterior body rings, lateral view, volume rendering.
|
||
<emphasis id="B9214716FF82FFAC1A8EF8D3FD57F8EF" bold="true" box="[681,702,1858,1884]" pageId="12" pageNumber="34">D</emphasis>
|
||
. head, frontal view, volume rendering.
|
||
<emphasis id="B9214716FF82FFAC1C5DF8D3FB64F8EF" bold="true" box="[1146,1165,1858,1884]" pageId="12" pageNumber="34">E</emphasis>
|
||
. Head, ventral view, volume rendering.
|
||
<emphasis id="B9214716FF82FFAC19BAF8F4FE47F8CC" bold="true" box="[413,430,1893,1919]" pageId="12" pageNumber="34">F</emphasis>
|
||
. Gonopods, frontal view, segmentation.
|
||
<emphasis id="B9214716FF82FFAC1BA9F8F5FC4CF8CD" bold="true" box="[910,933,1892,1918]" pageId="12" pageNumber="34">G</emphasis>
|
||
. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; at = antennae; co = collum; cx = coxite; em = eumentum; gn = gnathochilarium; L = leg; la = labrum; ll = lamella limgualis; md = mandible; om = ommatidia; pe = penis; pg = posterior gonopod; pm = promentum; st = gnathochilarial stipes; te = telopodite.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEA9B04FF83FFAD189AFE98FCECFE07" blockId="13.[188,1399,265,437]" pageId="13" pageNumber="35">
|
||
TRUNK. Collum large, laterally with triangular extension. Body rings 2–5 narrower than collum and following body rings. Body rings totally fused from body ring 5 onwards, anterior four body rings with sternite free from pleuro-tergite. Body rings from body ring 5 onwards with distinctly arched metazonite and wider than prozonite. Metazonite ventrally with 7–9 striae. Ozopores from body ring 6 onwards positioned ventro-lateral on metazonite (
|
||
<figureCitation id="136E8781FF83FFAD1ABCFE0BFD11FE07" box="[667,760,410,436]" captionStart="Fig" captionStartId="12.[188,231,1787,1813]" captionTargetBox="[188,1399,529,1730]" captionTargetId="figure-117@12.[188,1399,529,1750]" captionTargetPageId="12" captionText="Fig. 4. Kachincambala muelleri gen. et sp. nov., holotype, ♂ (ZFMK-MYR10225). A. Habitus, dorsal view, photograph. B. Head and anterior body rings, ventral view, volume rendering. C. Head and anterior body rings, lateral view, volume rendering. D. head, frontal view, volume rendering. E. Head, ventral view, volume rendering. F. Gonopods, frontal view, segmentation. G. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; at = antennae; co = collum; cx = coxite; em = eumentum; gn = gnathochilarium; L = leg; la = labrum; ll = lamella limgualis; md = mandible; om = ommatidia; pe = penis; pg = posterior gonopod; pm = promentum; st = gnathochilarial stipes; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983295" httpUri="https://zenodo.org/record/4983295/files/figure.png" pageId="13" pageNumber="35">Fig. 4C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEA9B04FF83FFAD189AFE4CFC6EFDD7" blockId="13.[189,1399,477,612]" pageId="13" pageNumber="35">
|
||
LEGS. Body rings 3 and 4 seemingly legless, from body ring 5 onwards 2 leg pairs per body ring (
|
||
<figureCitation id="136E8781FF83FFAD1D28FE4CFA80FE44" box="[1295,1385,477,503]" captionStart="Fig" captionStartId="12.[188,231,1787,1813]" captionTargetBox="[188,1399,529,1730]" captionTargetId="figure-117@12.[188,1399,529,1750]" captionTargetPageId="12" captionText="Fig. 4. Kachincambala muelleri gen. et sp. nov., holotype, ♂ (ZFMK-MYR10225). A. Habitus, dorsal view, photograph. B. Head and anterior body rings, ventral view, volume rendering. C. Head and anterior body rings, lateral view, volume rendering. D. head, frontal view, volume rendering. E. Head, ventral view, volume rendering. F. Gonopods, frontal view, segmentation. G. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; at = antennae; co = collum; cx = coxite; em = eumentum; gn = gnathochilarium; L = leg; la = labrum; ll = lamella limgualis; md = mandible; om = ommatidia; pe = penis; pg = posterior gonopod; pm = promentum; st = gnathochilarial stipes; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983295" httpUri="https://zenodo.org/record/4983295/files/figure.png" pageId="13" pageNumber="35">Fig. 4B</figureCitation>
|
||
). Anterior three leg pairs shorter than following legs but not reduced in segmentation. Midbody legs with coxa, prefemur, femur, postfemur, tibia, tarsus. Podomeres covered by setae. Tarsus> tibia> prefemur = femur> postfemur> coxa. Accessory spine on tarsus absent.
|
||
</paragraph>
|
||
<paragraph id="8BEA9B04FF83FFAD189AFD1DFDBCFD15" blockId="13.[189,597,652,678]" box="[189,597,652,678]" pageId="13" pageNumber="35">POSTERIOR BODY RINGS. Unknown.</paragraph>
|
||
<paragraph id="8BEA9B04FF83FFAD189AFD5EFD84FCC9" blockId="13.[189,1399,719,890]" pageId="13" pageNumber="35">
|
||
MALE SEXUAL CHARACTERS. Leg pair 1 reduced in size, not in segmentation (
|
||
<figureCitation id="136E8781FF83FFAD1C12FD5EFB7DFD5A" box="[1077,1172,719,745]" captionStart="Fig" captionStartId="12.[188,231,1787,1813]" captionTargetBox="[188,1399,529,1730]" captionTargetId="figure-117@12.[188,1399,529,1750]" captionTargetPageId="12" captionText="Fig. 4. Kachincambala muelleri gen. et sp. nov., holotype, ♂ (ZFMK-MYR10225). A. Habitus, dorsal view, photograph. B. Head and anterior body rings, ventral view, volume rendering. C. Head and anterior body rings, lateral view, volume rendering. D. head, frontal view, volume rendering. E. Head, ventral view, volume rendering. F. Gonopods, frontal view, segmentation. G. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; at = antennae; co = collum; cx = coxite; em = eumentum; gn = gnathochilarium; L = leg; la = labrum; ll = lamella limgualis; md = mandible; om = ommatidia; pe = penis; pg = posterior gonopod; pm = promentum; st = gnathochilarial stipes; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983295" httpUri="https://zenodo.org/record/4983295/files/figure.png" pageId="13" pageNumber="35">Fig. 4A</figureCitation>
|
||
). Anterior gonopod plate-like, coxite with apically rounded inner lobe and shorter outer lobe, which extends laterad. Inner lobe ca two times as long as outer lobe. Telopodite as long as coxite’s inner lobe, apically rounded. Presence of flagellum unknown. Posterior gonopod as long as anterior gonopod; apically with rounded lobe extending laterally (
|
||
<figureCitation id="136E8781FF83FFAD19C5FCF1FD89FCC9" box="[482,608,864,890]" captionStart="Fig" captionStartId="12.[188,231,1787,1813]" captionTargetBox="[188,1399,529,1730]" captionTargetId="figure-117@12.[188,1399,529,1750]" captionTargetPageId="12" captionText="Fig. 4. Kachincambala muelleri gen. et sp. nov., holotype, ♂ (ZFMK-MYR10225). A. Habitus, dorsal view, photograph. B. Head and anterior body rings, ventral view, volume rendering. C. Head and anterior body rings, lateral view, volume rendering. D. head, frontal view, volume rendering. E. Head, ventral view, volume rendering. F. Gonopods, frontal view, segmentation. G. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; at = antennae; co = collum; cx = coxite; em = eumentum; gn = gnathochilarium; L = leg; la = labrum; ll = lamella limgualis; md = mandible; om = ommatidia; pe = penis; pg = posterior gonopod; pm = promentum; st = gnathochilarial stipes; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983295" httpUri="https://zenodo.org/record/4983295/files/figure.png" pageId="13" pageNumber="35">Fig. 4F–G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEA9B04FF83FFAD189AFC32FD71FC0E" blockId="13.[189,664,931,957]" box="[189,664,931,957]" pageId="13" pageNumber="35">FEMALE SEXUAL CHARACTERS. Unknown.</paragraph>
|
||
<paragraph id="8BEA9B04FF83FFAD189AFC74FDA9FC4C" blockId="13.[189,576,997,1023]" box="[189,576,997,1023]" pageId="13" pageNumber="35">
|
||
<heading id="D0A22C68FF83FFAD189AFC74FDA9FC4C" bold="true" box="[189,576,997,1023]" fontSize="11" level="3" pageId="13" pageNumber="35" reason="3">
|
||
<emphasis id="B9214716FF83FFAD189AFC74FDA9FC4C" bold="true" box="[189,576,997,1023]" pageId="13" pageNumber="35">Measurements and taphonomy</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEA9B04FF83FFAD189AFB84FCDDFB9C" blockId="13.[189,820,1045,1072]" box="[189,820,1045,1072]" pageId="13" pageNumber="35">
|
||
MEASUREMENTS. Ca
|
||
<quantity id="4CAD36E1FF83FFAD198CFB84FE17FB83" box="[427,510,1045,1072]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="35" unit="mm" value="12.0">12 mm</quantity>
|
||
long,
|
||
<quantity id="4CAD36E1FF83FFAD1A6FFB84FD4BFB83" box="[584,674,1045,1072]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="35" unit="mm" value="0.5">0.5 mm</quantity>
|
||
in diameter.
|
||
</paragraph>
|
||
<paragraph id="8BEA9B04FF83FFAD189AFBC9FE28FB25" blockId="13.[189,1398,1111,1174]" pageId="13" pageNumber="35">AMBER DESCRIPTION. Flat, triangular, 15.5 × 8.9 × 3.0 mm. Yellow transparent with brownish areas close to animal. With crack.</paragraph>
|
||
<paragraph id="8BEA9B04FF83FFAD189AFB2EFD3FFB6B" blockId="13.[189,726,1214,1241]" box="[189,726,1214,1241]" pageId="13" pageNumber="35">SYNINCLUSIONS. Insect remains, stellate hairs.</paragraph>
|
||
<paragraph id="8BEA9B04FF83FFAD189AFA90FB53FA8C" blockId="13.[189,1399,1281,1344]" pageId="13" pageNumber="35">
|
||
TAPHONOMIC FEATURES. Specimen straight. Posterior body rings missing (cut away). Mid body partially obscured by crack. Gonopods everted. Defence fluid released from ozopores (
|
||
<figureCitation id="136E8781FF83FFAD1C69FAB4FB44FA8C" box="[1102,1197,1317,1343]" captionStart="Fig" captionStartId="12.[188,231,1787,1813]" captionTargetBox="[188,1399,529,1730]" captionTargetId="figure-117@12.[188,1399,529,1750]" captionTargetPageId="12" captionText="Fig. 4. Kachincambala muelleri gen. et sp. nov., holotype, ♂ (ZFMK-MYR10225). A. Habitus, dorsal view, photograph. B. Head and anterior body rings, ventral view, volume rendering. C. Head and anterior body rings, lateral view, volume rendering. D. head, frontal view, volume rendering. E. Head, ventral view, volume rendering. F. Gonopods, frontal view, segmentation. G. Gonopods, lateral view, segmentation. Abbreviations: ag = anterior gonopod; at = antennae; co = collum; cx = coxite; em = eumentum; gn = gnathochilarium; L = leg; la = labrum; ll = lamella limgualis; md = mandible; om = ommatidia; pe = penis; pg = posterior gonopod; pm = promentum; st = gnathochilarial stipes; te = telopodite." figureDoi="http://doi.org/10.5281/zenodo.4983295" httpUri="https://zenodo.org/record/4983295/files/figure.png" pageId="13" pageNumber="35">Fig. 4A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |