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<document id="595E89ED0DE81CD4DA1D9F8C5218514A" ID-DOI="10.3853/j.0067-1975.62.2010.1554" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239086" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629574225023" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2010" docId="03F987C5FFA2F76801F416E84EA1C9A6" docLanguage="en" docName="RecAustMus.62.2.217-284.pdf" docOrigin="Records of the Australian Museum 62 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada kimberleyana Köhler 2010, n.sp." docType="treatment" docUuid="11955295-7360-46AA-B23A-8E22AEEA3EFC" docUuidSource="ZooBank" docVersion="3" lastPageNumber="269" masterDocId="FFC0FFBDFF91F75C027510454D15CE74" masterDocTitle="Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)" masterLastPageNumber="284" masterPageNumber="217" pageNumber="268" updateTime="1699078283413" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="7F0747A742A30F39F37E5F8A57F543D9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BED13CC5C0FE790764ABB33A793D5B5D">
<mods:title id="5B1D442F1818C0800314CFBE3EB4523B">Uncovering Local Endemism in the Kimberley, Western Australia: Description of New Species of the Genus Amplirhagada Iredale, 1933 (Pulmonata: Camaenidae)</mods:title>
</mods:titleInfo>
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<mods:role id="79FDF2ACC80F9CD386832FE399A84410">
<mods:roleTerm id="6D8290C3D1DE53DF44925D78F8740F3D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8854F0D1710911EA49C2B72F52145790">Köhler, Frank</mods:namePart>
</mods:name>
<mods:typeOfResource id="AFAE488EE5CEC544CCCFD7D925736A31">text</mods:typeOfResource>
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<mods:title id="609231D925D81E78EA56F6860CD9BB8C">Records of the Australian Museum</mods:title>
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<mods:part id="1DFB95E7309BDF3BD524D16842AC45BD">
<mods:date id="3CF21E5922C58A8A60EF4EA4E4F09C49">2010</mods:date>
<mods:detail id="41E70FA9BF7B236B51EE2A6FDA9621ED" type="pubDate">
<mods:number id="202BFDBDA7B409E93275E9F20E04C430">2010-11-24</mods:number>
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<mods:detail id="66DE4D4AB43B2042D7D65897ACAF2FC2" type="volume">
<mods:number id="B14FEDEE07A7695EEFFB06F366A8A648">62</mods:number>
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<mods:number id="A01B939F9E35EDCA2CCE2C1054331074">2</mods:number>
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<mods:start id="332A3307DADFE6BB211B142AC40FC8C1">217</mods:start>
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<mods:url id="8F7622A6674D3BF0A5E1003C89BF588C">http://dx.doi.org/10.3853/j.0067-1975.62.2010.1554</mods:url>
</mods:location>
<mods:classification id="F6A1C47642A39618EAA078B747FDB334">journal article</mods:classification>
<mods:identifier id="88979AE3D0FEF85D0FF6CF8F19D0DB30" type="DOI">10.3853/j.0067-1975.62.2010.1554</mods:identifier>
<mods:identifier id="D7E08438F789E5ED978EA2B84DE89339" type="ISSN">2201-4349</mods:identifier>
<mods:identifier id="0F46CE734C7EA5AF9DA163661C6C78EF" type="Zenodo-Dep">5239086</mods:identifier>
<mods:identifier id="125175203B8D87212A4EDA2B1BF4CE56" type="ZooBank">749A271A-6942-4D4C-B0DF-B968462BFF0C</mods:identifier>
</mods:mods>
<treatment id="03F987C5FFA2F76801F416E84EA1C9A6" ID-DOI="http://doi.org/10.5281/zenodo.5239055" ID-Zenodo-Dep="5239055" LSID="urn:lsid:zoobank.org:act:11955295-7360-46AA-B23A-8E22AEEA3EFC" httpUri="http://treatment.plazi.org/id/03F987C5FFA2F76801F416E84EA1C9A6" lastPageId="52" lastPageNumber="269" pageId="51" pageNumber="268">
<subSubSection id="C34A6558FFA2F76F01F416E84833C8B3" box="[897,1318,1709,1735]" pageId="51" pageNumber="268" type="nomenclature">
<paragraph id="8BEF36D3FFA2F76F01F416E84833C8B3" blockId="51.[897,1318,1709,1735]" box="[897,1318,1709,1735]" pageId="51" pageNumber="268">
<heading id="D0A781BFFFA2F76F01F416E84833C8B3" box="[897,1318,1709,1735]" centered="true" fontSize="11" level="2" pageId="51" pageNumber="268" reason="2">
<emphasis id="B924EAC1FFA2F76F01F416E84833C8B3" bold="true" box="[897,1318,1709,1735]" pageId="51" pageNumber="268">
<taxonomicName id="4C504D50FFA2F76F01F416E849F7C8B3" ID-CoL="D7RF" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[897,1250,1709,1735]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="51" pageNumber="268" phylum="Mollusca" rank="species" species="kimberleyana" status="sp. nov.">
<emphasis id="B924EAC1FFA2F76F01F416E849F7C8B3" bold="true" box="[897,1250,1709,1735]" italics="true" pageId="51" pageNumber="268">Amplirhagada kimberleyana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFA2F76F069F16E84833C8B3" box="[1258,1318,1709,1735]" pageId="51" pageNumber="268" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34A6558FFA2F76F016F16B749FBC9A5" pageId="51" pageNumber="268" type="materials_examined">
<paragraph id="8BEF36D3FFA2F76F016F16B7489BC915" blockId="51.[794,1422,1778,1889]" pageId="51" pageNumber="268">
<materialsCitation id="3B383C8EFFA2F76F016F16B7489CC915" collectingDate="1988-07-09" collectionCode="V" collectorName="Kessner &amp; A. Longbottom" county="Kimberley" latitude="-15.536112" location="Camden Sound" longLatPrecision="21" longitude="124.39889" municipality="Bonaparte Archipelago" pageId="51" pageNumber="268" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="B924EAC1FFA2F76F016F16B74EBEC97E" bold="true" box="[794,939,1778,1802]" pageId="51" pageNumber="268">
<typeStatus id="54EB8871FFA2F76F016F16B74E46C97E" box="[794,851,1778,1802]" pageId="51" pageNumber="268">Type</typeStatus>
locality
</emphasis>
(
<figureCitation id="136B2A56FFA2F76F01C216B74EE0C97D" box="[951,1013,1778,1801]" captionStart="Figure 1" captionStartId="1.[121,184,1433,1454]" captionTargetBox="[121,1421,142,1408]" captionTargetId="figure-129@1.[121,1421,138,1414]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species in the Kimberley region, northwestern Australia. (1) A. euroa n.sp., Adolphus Island. (2) A. solemiana n.sp., Middle Osborn Island. (3) A. indistincta n.sp., Southwest Osborn Island, Kidney Island. (4) A. combeana, Cassini Island. (5) A. mckenziei n.sp., Oliver Island. (6) A. ponderi n.sp., Kingsmill Island. (7) A. montesquieuana n.sp., Fenelon Island. (8) A. descartesana n.sp., Descartes Island. (9) A. katerana, Katers Island. (10) A. puescheli n.sp., unnamed island near Prudhoe Island. (11) A. decora n.sp., A. kessneri n.sp., Bigge Island. (12) A. berthierana n.sp., Berthier Island. (13) A. lamarckiana n.sp., Lamarck Island.(14) A. anderdonensis n.sp., unnamed island, Anderdon Islands. (15) A. tricenaria n.sp., Prince Frederick Harbour, north of Hunter River mouth. (16) A. regia n.sp., A. boongareensis n.sp., A. gibsoni n.sp., and Amplirhagada sp., Boongaree Island. (17) A. yorkensis n.sp., Coronation Island. (18) A. buffonensis n.sp., Buffon Island. (19) A. uwinsensis n.sp., Uwins Island. (20) A. sphaeroidea n.sp., St. Andrews Island. (21) A. basilica n.sp., and A. camdenensis n.sp., Augustus Island. (22) A. kimberleyana n.sp., NW of Wilson Point. (23) A. gemina n.sp., 1.5 km SE of Hall Point. (24) A. dubitabile n.sp., Steep Island. (25) A. napierana, north Napier ranges. (26) A. percita, Napier ranges. (27) A. castra, A. mitchelliana, and A. varia, Mitchell Plateau." figureDoi="http://doi.org/10.5281/zenodo.5239088" httpUri="https://zenodo.org/record/5239088/files/figure.png" pageId="51" pageNumber="268">Fig. 1</figureCitation>
).
<collectingRegion id="4994F831FFA2F76F067C16B749DDC97D" box="[1033,1224,1778,1801]" country="Australia" name="Western Australia" pageId="51" pageNumber="268">Western Australia</collectingRegion>
,
<collectingCounty id="628E4E5FFFA2F76F06A516B74856C97D" box="[1232,1347,1778,1801]" pageId="51" pageNumber="268">Kimberley</collectingCounty>
,
<collectingMunicipality id="6B8BACA9FFA2F76F073916B74ECAC952" pageId="51" pageNumber="268">Bonaparte Archipelago</collectingMunicipality>
,
<location id="8E8F6008FFA2F76F019D174A4985C952" LSID="urn:lsid:plazi:treatment:03F987C5FFA2F76801F416E84EA1C9A6:8E8F6008FFA2F76F019D174A4985C952" box="[1000,1168,1807,1830]" county="Kimberley" latitude="-15.536112" longLatPrecision="21" longitude="124.39889" municipality="Bonaparte Archipelago" name="Camden Sound" pageId="51" pageNumber="268" stateProvince="Western Australia">Camden Sound</location>
,
<location id="8E8F6008FFA2F76F06EF174A489CC952" LSID="urn:lsid:plazi:treatment:03F987C5FFA2F76801F416E84EA1C9A6:8E8F6008FFA2F76F06EF174A489CC952" box="[1178,1417,1807,1830]" county="Kimberley" latitude="-15.536112" longLatPrecision="21" longitude="124.39889" municipality="Bonaparte Archipelago" name="northern Slate Islands" pageId="51" pageNumber="268" stateProvince="Western Australia">northern Slate Islands</location>
,
<locationDeviation id="9A5D2FF1FFA2F76F016F17684977C930" box="[794,1122,1837,1860]" location="Wilson Point" pageId="51" pageNumber="268">
<quantity id="4CA89B36FFA2F76F016F17684E7FC930" box="[794,874,1837,1860]" metricMagnitude="3" metricUnit="m" metricValue="2.4" pageId="51" pageNumber="268" unit="km" value="2.4">2.4 km</quantity>
NW of Wilson Point
</locationDeviation>
,
<geoCoordinate id="EE645014FFA2F76F0604176849FAC930" box="[1137,1263,1837,1860]" degrees="15" direction="south" minutes="32" orientation="latitude" pageId="51" pageNumber="268" precision="15" seconds="10" value="-15.536112">15°32'10&quot;S</geoCoordinate>
<geoCoordinate id="EE645014FFA2F76F068C1768489CC930" box="[1273,1417,1837,1860]" degrees="124" direction="east" minutes="23" orientation="longitude" pageId="51" pageNumber="268" precision="15" seconds="56" value="124.39889">124°23'56&quot;E</geoCoordinate>
; KC-005 (leg.
<collectionCode id="ED41AE16FFA2F76F01D9170F4ED5C915" box="[940,960,1866,1889]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="51" pageNumber="268" type="Museum">V</collectionCode>
.
<collectorName id="26A55305FFA2F76F01BC170F4935C915" box="[969,1056,1866,1889]" pageId="51" pageNumber="268">Kessner</collectorName>
&amp;
<collectorName id="26A55305FFA2F76F064A170F49FCC915" box="[1087,1257,1866,1889]" pageId="51" pageNumber="268">A. Longbottom</collectorName>
,
<date id="FFEE1013FFA2F76F0686170F4894C915" box="[1267,1409,1866,1889]" pageId="51" pageNumber="268" value="1988-07-09">
<collectingDate id="EFAAE9FBFFA2F76F0686170F4894C915" box="[1267,1409,1866,1889]" pageId="51" pageNumber="268" value="1988-07-09">09 July 1988</collectingDate>
</date>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEF36D3FFA2F76F016F17C849FBC9A5" blockId="51.[794,1421,1933,2001]" pageId="51" pageNumber="268">
<emphasis id="B924EAC1FFA2F76F016F17C84EB6C9D4" bold="true" box="[794,931,1933,1952]" pageId="51" pageNumber="268">Type material</emphasis>
.
<materialsCitation id="3B383C8EFFA2F76F01C717CB49E6C9D5" accessionNumber="S34625" box="[946,1267,1934,1953]" collectionCode="WAM" pageId="51" pageNumber="268" specimenCount="1" typeStatus="holotype">
<typeStatus id="54EB8871FFA2F76F01C717CB4913C9D5" box="[946,1030,1934,1953]" pageId="51" pageNumber="268" type="holotype">Holotype</typeStatus>
<collectionCode id="ED41AE16FFA2F76F067817CB4956C9D5" box="[1037,1091,1934,1953]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="51" pageNumber="268" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFA2F76F063E17CB4985C9D5" box="[1099,1168,1934,1953]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34625" pageId="51" pageNumber="268" type="EnaNcbi">S34625</accessionNumber>
(Pl. 2.13)
</materialsCitation>
.
<materialsCitation id="3B383C8EFFA2F76F068A17CB49FFC9A5" accessionNumber="S41480" collectionCode="WAM" pageId="51" pageNumber="268" specimenCode="FMNH 219055" specimenCount="1" typeStatus="paratype">
<typeStatus id="54EB8871FFA2F76F068A17CB4843C9D5" box="[1279,1366,1934,1953]" pageId="51" pageNumber="268" type="paratype">Paratypes</typeStatus>
AMS C463760 (5 preserved specimens),
<specimenCode id="DBF69EA8FFA2F76F060417E34817C9CD" box="[1137,1282,1958,1977]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="51" pageNumber="268" type="Museum">FMNH 219055</specimenCode>
(21 preserved specimens),
<collectionCode id="ED41AE16FFA2F76F01F317FB4EAEC9A5" box="[902,955,1982,2001]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="51" pageNumber="268" type="Museum">WAM</collectionCode>
<accessionNumber id="9403AB30FFA2F76F01B517FB4916C9A5" box="[960,1027,1982,2001]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S41480" pageId="51" pageNumber="268" type="EnaNcbi">S41480</accessionNumber>
(15 preserved specimens)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF2F665BFFA5F76801A610EA49F5CF60" ID-DOI="http://doi.org/10.5281/zenodo.5239263" ID-Zenodo-Dep="5239263" httpUri="https://zenodo.org/record/5239263/files/figure.png" pageId="52" pageNumber="269" startId="52.[979,1042,175,196]" targetBox="[839,1463,161,1422]" targetPageId="52">
<paragraph id="8BEF36D3FFA5F76801A610EA49F5CF60" blockId="52.[979,1461,175,276]" pageId="52" pageNumber="269">
Figure 81. Interior of penial chamber of
<taxonomicName id="4C504D50FFA5F76801A6108F49F3CEAB" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[979,1254,202,223]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="52" pageNumber="269" phylum="Mollusca" rank="species" species="kimberleyana" status="sp. nov.">
<emphasis id="B924EAC1FFA5F76801A6108F49F3CEAB" box="[979,1254,202,223]" italics="true" pageId="52" pageNumber="269">Amplirhagada kimberleyana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFA5F7680699108F480ECEAB" box="[1260,1307,202,223]" pageId="52" pageNumber="269" rank="species">n.sp.</taxonomicNameLabel>
, paratype AMS C463760 (9 July, scale 5 mm). Compare with Fig. 4 for labelling of structures.
</paragraph>
</caption>
<caption id="DF2F665BFFA5F76802D312334C6ACCB5" ID-DOI="http://doi.org/10.5281/zenodo.5239265" ID-Zenodo-Dep="5239265" httpUri="https://zenodo.org/record/5239265/files/figure.png" pageId="52" pageNumber="269" startId="52.[166,228,630,651]" targetBox="[171,788,141,616]" targetPageId="52">
<paragraph id="8BEF36D3FFA5F76802D312334C6ACCB5" blockId="52.[166,794,630,705]" pageId="52" pageNumber="269">
Figure 80. Genitalia of
<taxonomicName id="4C504D50FFA5F768030B12324F98CCF8" authority="Köhler, 2010" authorityName="Köhler" authorityYear="2010" box="[382,653,631,652]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="52" pageNumber="269" phylum="Mollusca" rank="species" species="kimberleyana" status="sp. nov.">
<emphasis id="B924EAC1FFA5F768030B12324F98CCF8" box="[382,653,631,652]" italics="true" pageId="52" pageNumber="269">Amplirhagada kimberleyana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757BAFFA5F76800E412334FD5CCFF" box="[657,704,630,651]" pageId="52" pageNumber="269" rank="species">n.sp.</taxonomicNameLabel>
, paratype AMS C463760 (9 July, scale 10 mm). Compare with Fig. 3 for labelling of structures.
</paragraph>
</caption>
<subSubSection id="C34A6558FFA5F76802D3134C4EA1C9A6" pageId="52" pageNumber="269" type="etymology">
<paragraph id="8BEF36D3FFA5F76802D3134C4FAECD55" blockId="52.[166,699,777,801]" box="[166,699,777,801]" pageId="52" pageNumber="269">
<emphasis id="B924EAC1FFA5F76802D3134C4C34CD55" bold="true" box="[166,289,777,801]" pageId="52" pageNumber="269">Etymology</emphasis>
. In reference to the Kimberley region.
</paragraph>
<paragraph id="8BEF36D3FFA5F76802D313014CEDCA82" blockId="52.[166,795,836,1270]" pageId="52" pageNumber="269">
<emphasis id="B924EAC1FFA5F76802D313014DF5CD28" bold="true" box="[166,224,836,860]" pageId="52" pageNumber="269">Shell</emphasis>
(
<figureCitation id="136B2A56FFA5F768028113014C6ACD2F" box="[244,383,836,859]" captionStart="Figure 79" captionStartId="51.[121,184,1552,1573]" captionTargetBox="[121,1421,151,1543]" captionTargetId="figure-147@51.[121,1421,151,1543]" captionTargetPageId="51" captionText="Figure 79. SEM photographs of Amplirhagada kimberleyana n.sp. (A–D) Shell, paratype FMNH 219055: (A) apical whorl viewed from above (scale 200 µm); (B) sculpture on first three whorls viewed from above (scale 1 mm); (C) close-up showing sculpture on penultimate whorl (scale 200 µm); (D) close-up showing surface of last whorl (scale 200 µm).(E) Jaw, paratypeAMS C463760 (scale 100 µm). (F–G) Radula, paratype AMS C463760: (F) central and lateral teeth (scale 20 µm); (G) outer lateral and inner marginal teeth (scale 20 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239261" httpUri="https://zenodo.org/record/5239261/files/figure.png" pageId="52" pageNumber="269">Fig. 79A–D</figureCitation>
, Pl. 2.13). Semi-globose to broadly conical, with medium high spire; thin to solid (translucent). Periphery evenly rounded to slightly angulate; upper sector of whorl and basal sector rounded. Umbilicus forming a chink to narrowly winding opening, 60–100 percent concealed by columellar reflection. Background colour beige-brown; subsutural band absent or diffuse, brown; mid-whorl band absent or diffuse, brown, thin, visible on last whorl only; ventral colour lighter, horn or whitish; outer lip colour differs from shell, light brown to whitish; inner lip whitish. Protoconch c.
<quantity id="4CA89B36FFA5F76802D3142F4DF4CAF5" box="[166,225,1130,1153]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="52" pageNumber="269" unit="mm" value="3.0">3 mm</quantity>
in diameter, comprising 2 whorls, smooth. Teleoconch smooth except of axial growth lines. Angle of aperture 45°; outer lip rounded, moderately thick, slightly expanded, not or slightly reflected, basal and palatal node absent. Parietal wall of inner lip inconspicuous.
</paragraph>
<paragraph id="8BEF36D3FFA5F76802D3155C4F9CCB1F" blockId="52.[166,794,1305,1387]" pageId="52" pageNumber="269">
<emphasis id="B924EAC1FFA5F76802D3155C4C81CB45" bold="true" box="[166,404,1305,1329]" pageId="52" pageNumber="269">Pallial morphology</emphasis>
. Pallial cavity moderately deep, extending
<emphasis id="B924EAC1FFA5F768036015724C08CB31" bold="true" box="[277,285,1335,1349]" pageId="52" pageNumber="269">
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⁄
<emphasis id="B924EAC1FFA5F7680354157B4C3CCB38" bold="true" box="[289,297,1342,1356]" pageId="52" pageNumber="269">
<subScript id="17D43496FFA5F7680354157B4C3CCB38" attach="left" box="[289,297,1342,1356]" fontSize="6" pageId="52" pageNumber="269">4</subScript>
</emphasis>
whorl. Pigmentation on mantle mottled, black. Kidney extending about half of pallial cavity.
</paragraph>
<paragraph id="8BEF36D3FFA5F76802D315CA4C67C9A7" blockId="52.[166,794,1423,2003]" pageId="52" pageNumber="269">
<emphasis id="B924EAC1FFA5F76802D315CA4C9DCBD3" bold="true" box="[166,392,1423,1447]" pageId="52" pageNumber="269">Genital morphology</emphasis>
(
<figureCitation id="136B2A56FFA5F76803E215CA4F0CCBD2" box="[407,537,1423,1446]" captionStart-0="Figure 80" captionStart-1="Figure 81" captionStartId-0="52.[166,228,630,651]" captionStartId-1="52.[979,1042,175,196]" captionTargetBox-0="[171,788,141,616]" captionTargetBox-1="[839,1463,161,1422]" captionTargetId-0="figure-455@52.[166,793,128,621]" captionTargetId-1="figure-479@52.[839,1466,151,1423]" captionTargetPageId-0="52" captionTargetPageId-1="52" captionText-0="Figure 80. Genitalia of Amplirhagada kimberleyana n.sp., paratype AMS C463760 (9 July, scale 10 mm). Compare with Fig. 3 for labelling of structures." captionText-1="Figure 81. Interior of penial chamber of Amplirhagada kimberleyana n.sp., paratype AMS C463760 (9 July, scale 5 mm). Compare with Fig. 4 for labelling of structures." figureDoi-0="http://doi.org/10.5281/zenodo.5239265" figureDoi-1="http://doi.org/10.5281/zenodo.5239263" httpUri-0="https://zenodo.org/record/5239265/files/figure.png" httpUri-1="https://zenodo.org/record/5239263/files/figure.png" pageId="52" pageNumber="269">Figs. 80–81</figureCitation>
). Penis straight, more or less of same length as anterior part of oviduct or longer. Vas deferens coils once before entering penis. Penial retractor muscle shorter than penis. Penial verge moderately long (
<emphasis id="B924EAC1FFA5F76802DA16414DA2C866" bold="true" box="[175,183,1540,1554]" pageId="52" pageNumber="269">
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⁄
<emphasis id="B924EAC1FFA5F76802C9164E4DCCC866" bold="true" box="[188,217,1540,1563]" pageId="52" pageNumber="269">
<subScript id="17D43496FFA5F76802C9164E4DD1C86D" attach="right" box="[188,196,1547,1561]" fontSize="6" pageId="52" pageNumber="269">8</subScript>
–
<superScript id="7C259B9BFFA5F76802A416414DCCC866" attach="none" box="[209,217,1540,1554]" fontSize="6" pageId="52" pageNumber="269">1</superScript>
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⁄
<emphasis id="B924EAC1FFA5F76802AB164E4DF3C86D" bold="true" box="[222,230,1547,1561]" pageId="52" pageNumber="269">
<subScript id="17D43496FFA5F76802AB164E4DF3C86D" attach="left" box="[222,230,1547,1561]" fontSize="6" pageId="52" pageNumber="269">4</subScript>
</emphasis>
penial chamber), slender to spatulate, with pointed tip. Penial wall pustules large, arranged in rows on apical to median portion of the penial chamber. Main stimulatory pilaster undifferentiated, formed by few rows of slightly enlarged, hooked pustules. Base of penial wall smooth, supporting two smooth and narrow pilasters. Vas deferens entering penial sheath in upper third. Vagina moderately long, tubular to posteriorly slightly inflated; inner vaginal wall with smooth longitudinal pilasters. Spermathecal duct wide, internally with corrugated longitudinal pilasters. Spermathecal head globular, connected with oviduct by connective tissue, internally smooth, with thin wall. Free oviduct comprising about half of anterior part of oviduct, more or less straight. Spermoviduct of same length as anterior part of oviduct. Talon embedded in albumen gland at junction with spermoviduct.
</paragraph>
<paragraph id="8BEF36D3FFA5F768013216464899C971" blockId="52.[839,1467,1539,1797]" pageId="52" pageNumber="269">
<emphasis id="B924EAC1FFA5F768013216464929C86F" bold="true" box="[839,1084,1539,1563]" pageId="52" pageNumber="269">Radular morphology</emphasis>
(
<figureCitation id="136B2A56FFA5F768063A164649C0C86E" box="[1103,1237,1539,1562]" captionStart="Figure 79" captionStartId="51.[121,184,1552,1573]" captionTargetBox="[121,1421,151,1543]" captionTargetId="figure-147@51.[121,1421,151,1543]" captionTargetPageId="51" captionText="Figure 79. SEM photographs of Amplirhagada kimberleyana n.sp. (A–D) Shell, paratype FMNH 219055: (A) apical whorl viewed from above (scale 200 µm); (B) sculpture on first three whorls viewed from above (scale 1 mm); (C) close-up showing sculpture on penultimate whorl (scale 200 µm); (D) close-up showing surface of last whorl (scale 200 µm).(E) Jaw, paratypeAMS C463760 (scale 100 µm). (F–G) Radula, paratype AMS C463760: (F) central and lateral teeth (scale 20 µm); (G) outer lateral and inner marginal teeth (scale 20 µm)." figureDoi="http://doi.org/10.5281/zenodo.5239261" httpUri="https://zenodo.org/record/5239261/files/figure.png" pageId="52" pageNumber="269">Fig. 79F–G</figureCitation>
). Rectangular. Tooth formula C+16+3+22. With 147 rows of teeth, 31.9 rows per mm (n = 1). Central teeth with sharply pointed, triangular mesocones, shorter than base of tooth; ectocones vestigial. Lateral teeth with sharply pointed, triangular mesocones, length equal to base of tooth; ectocones small, endocones vestigial. Marginal teeth with elongate to triangular mesocones; ectocones shorter and narrower than mesocones, divided into two denticles; endocones greatly reduced.
</paragraph>
<paragraph id="8BEF36D3FFA5F7680132176D4EA1C9A6" blockId="52.[839,1467,1832,2002]" pageId="52" pageNumber="269">
<emphasis id="B924EAC1FFA5F7680132176D495FC934" bold="true" box="[839,1098,1832,1856]" pageId="52" pageNumber="269">Comparative remarks</emphasis>
. Shell broadly conical to domeshaped but smaller than species with similar shell shape. Inner penial wall with strong developed pustulation and undifferentiated main pilaster is diagnostic. Material of the present species has been labelled as “
<taxonomicName id="4C504D50FFA5F76806A517DB489BC9C1" box="[1232,1422,1950,1973]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="52" pageNumber="269" phylum="Mollusca" rank="species" species="undetermined">
<emphasis id="B924EAC1FFA5F76806A517DB487FC9C1" box="[1232,1386,1950,1973]" italics="true" pageId="52" pageNumber="269">Amplirhagada</emphasis>
sp.
</taxonomicName>
64” by Solem.
</paragraph>
</subSubSection>
</treatment>
</document>
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