342 lines
38 KiB
XML
342 lines
38 KiB
XML
<document id="870C612F9821FE401D5A2E27192F1E09" ID-DOI="10.1111/j.1096-3642.2009.00526.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5443023" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630454731098" checkinUser="felipe" docAuthor="Liew, Thor-Seng, Schilthuizen, Menno & Vermeulen, Jaap Jan" docDate="2009" docId="03F9878AFF98E8140D368D7D261DFBBC" docLanguage="en" docName="j.1096-3642.2009.00526.x.pdf" docOrigin="Zoological Journal of the Linnean Society 157 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00526.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Everettia jucundior Liew & Schilthuizen & Vermeulen 2009, SP. NOV." docType="treatment" docVersion="5" lastPageNumber="538" masterDocId="FFC0FFF2FF8FE8030C238C132365FFCA" masterDocTitle="Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo" masterLastPageNumber="550" masterPageNumber="515" pageNumber="538" updateTime="1699276879878" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="ACCDC16A4F721F92A1733E55238A4D9E">Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo</mods:title>
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<mods:name id="57F3BF3B5381E86BA696E08AE0BE2FB2" type="personal">
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<mods:roleTerm id="64E4BFF5C6A6ED46F59C16A075D169A1">Author</mods:roleTerm>
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<mods:namePart id="41141A7FAA0947A60C4244B437158B6E">Liew, Thor-Seng</mods:namePart>
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<mods:roleTerm id="6B11FF77AD494F01C8F1D441CB8B412C">Author</mods:roleTerm>
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<mods:namePart id="CF07E34B8A6E6B6962C814771A5DD030">Schilthuizen, Menno</mods:namePart>
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<mods:roleTerm id="ADF1BFB01CB45AC23BCD2487059A1469">Author</mods:roleTerm>
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<mods:namePart id="309C90B2A3CB9C7843F68E9E47901F76">Vermeulen, Jaap Jan</mods:namePart>
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<mods:typeOfResource id="2385FC18B4E30F380EEFF68BB5BFE21A">text</mods:typeOfResource>
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<mods:title id="6E4AD4894A48C679996FF80160F73F66">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="F0A6E10C7FDEF4D69623484BACAD0D0F">
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<mods:date id="7E20DF568754160E985FF09EF2EC3404">2009</mods:date>
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<mods:number id="AF8BD0EB2C56430ACE2DC846ED5F6145">2009-10-26</mods:number>
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<mods:number id="E8A5525733086234E5E077C521D34BF5">157</mods:number>
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<mods:number id="9461009B1D05AAA9FE237DD93592D6FC">3</mods:number>
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<mods:start id="2C1FC6DCC20108B2C767C11A0962E4BB">515</mods:start>
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<mods:end id="DB3C654917DD9CED570579998A81BB79">550</mods:end>
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<mods:url id="286927200D8AFB21A1317757944971A5">https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00526.x</mods:url>
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<mods:identifier id="2608B086A2F0FEFD4E50D7904727EFFC" type="DOI">10.1111/j.1096-3642.2009.00526.x</mods:identifier>
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<mods:identifier id="1B6F0534628767DE211DCF71FE0D451C" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="41C8F0BF160564528F706C60BB8B0313" type="Zenodo-Dep">5443023</mods:identifier>
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<treatment id="03F9878AFF98E8140D368D7D261DFBBC" ID-DOI="http://doi.org/10.5281/zenodo.5492195" ID-Zenodo-Dep="5492195" LSID="urn:lsid:plazi:treatment:03F9878AFF98E8140D368D7D261DFBBC" httpUri="http://treatment.plazi.org/id/03F9878AFF98E8140D368D7D261DFBBC" lastPageNumber="538" pageId="23" pageNumber="538">
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<subSubSection id="C34A6517FF98E8140D368D7D21FFFE49" box="[277,666,366,387]" pageId="23" pageNumber="538" type="nomenclature">
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<paragraph id="8BEF369CFF98E8140D368D7D21FFFE49" blockId="23.[277,666,366,387]" box="[277,666,366,387]" pageId="23" pageNumber="538">
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<heading id="D0A781F0FF98E8140D368D7D21FFFE49" box="[277,666,366,387]" centered="true" fontSize="9" level="2" pageId="23" pageNumber="538" reason="2">
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<taxonomicName id="4C504D1FFF98E8140D368D7D214EFE48" ID-CoL="6HM7M" authority="Liew & Schilthuizen & Vermeulen, 2009" authorityName="Liew & Schilthuizen & Vermeulen" authorityYear="2009" box="[277,555,366,387]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="538" phylum="Mollusca" rank="species" species="jucundior" status="sp. nov.">EVERETTIA JUCUNDIOR</taxonomicName>
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<emphasis id="B924EA8EFF98E8140E168D6221FFFE49" bold="true" box="[565,666,366,387]" pageId="23" pageNumber="538">
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<taxonomicNameLabel id="A21757F5FF98E8140E168D6221FFFE49" box="[565,666,366,387]" pageId="23" pageNumber="538" rank="species">SP. NOV.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A6517FF98E8140C878D862248FE03" pageId="23" pageNumber="538" type="reference_group">
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<paragraph id="8BEF369CFF98E8140C878D862248FE03" blockId="23.[164,778,404,457]" pageId="23" pageNumber="538">
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<treatmentCitationGroup id="AB4011B2FF98E8140C878D862248FE03" pageId="23" pageNumber="538">
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<treatmentCitation id="0AF1108DFF98E8140C878D8621AAFE60" author="Laidlaw FF" box="[164,719,404,426]" page="200" pageId="23" pageNumber="538" year="1931">
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<taxonomicName id="4C504D1FFF98E8140C878D8621AAFE60" ID-CoL="3DHB4" authority=", Laidlaw, 1931: 196 - 200" authorityName=", Laidlaw" authorityPageNumber="196 - 200" authorityYear="1931" box="[164,719,404,426]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="538" phylum="Mollusca" rank="species" species="subconsul">
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<emphasis id="B924EA8EFF98E8140C878D8622EBFE63" box="[164,398,404,426]" italics="true" pageId="23" pageNumber="538">Everettia subconsul</emphasis>
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,
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<bibRefCitation id="EFC14B6DFF98E8140D8B8D8721AAFE60" author="Laidlaw FF" box="[424,719,404,426]" pageId="23" pageNumber="538" pagination="190 - 201" refId="ref25575" refString="Laidlaw FF. 1931. On a new sub-family Dyakiinae of the Zonitidae. Proceedings of the Malacological Society 19: 190 - 201." type="journal article" year="1931">Laidlaw, 1931: 196–200</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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, figures 3, 4, 6.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A6517FF98E8140C878DE1261DFBBC" pageId="23" pageNumber="538" type="materials_examined">
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<paragraph id="8BEF369CFF98E8140C878DE1219DFD75" blockId="23.[164,779,498,703]" pageId="23" pageNumber="538">
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<emphasis id="B924EA8EFF98E8140C878DE12228FDCD" box="[164,333,498,519]" italics="true" pageId="23" pageNumber="538">Type material:</emphasis>
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<materialsCitation id="3B383CC1FF98E8140D788DE12142FDA9" collectingDate="2001-12" country="Malaysia" latitude="4.4" location="Tawau District" longLatPrecision="1306" longitude="117.88333" municipality="State" pageId="23" pageNumber="538" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
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<collectingCountry id="F347760CFF98E8140D788DE12281FDCD" box="[347,484,498,519]" name="Malaysia" pageId="23" pageNumber="538">MALAYSIA</collectingCountry>
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:
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<collectingMunicipality id="6B8BACE6FF98E8140DD68DE12157FDCD" box="[501,562,498,519]" pageId="23" pageNumber="538">State</collectingMunicipality>
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of
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<collectingRegion id="4994F87EFF98E8140E418DE121C8FDC2" box="[610,685,498,520]" country="Malaysia" name="Sabah" pageId="23" pageNumber="538">Sabah</collectingRegion>
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:
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<location id="8E8F6047FF98E8140E9D8DE1239AFDEC" LSID="urn:lsid:plazi:treatment:03F9878AFF98E8140D368D7D261DFBBC:8E8F6047FF98E8140E9D8DE1239AFDEC" country="Malaysia" latitude="4.4" longLatPrecision="1306" longitude="117.88333" municipality="State" name="Tawau District" pageId="23" pageNumber="538" stateProvince="Sabah">Tawau District</location>
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. Tawau Hills Park (
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<geoCoordinate id="EE64505BFF98E8140DD98E03212CFDEC" box="[506,585,527,550]" degrees="4" direction="north" minutes="24" orientation="latitude" pageId="23" pageNumber="538" precision="925" value="4.4">
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4°24
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<emphasis id="B924EA8EFF98E8140E0E8E1C2156FDEC" box="[557,563,527,550]" italics="true" pageId="23" pageNumber="538">′</emphasis>
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N
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</geoCoordinate>
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,
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<geoCoordinate id="EE64505BFF98E8140E788E0321A7FDEC" box="[603,706,527,550]" degrees="117" direction="east" minutes="53" orientation="longitude" pageId="23" pageNumber="538" precision="925" value="117.88333">
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117°53
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<emphasis id="B924EA8EFF98E8140E8B8E1C21CBFDEC" box="[680,686,527,550]" italics="true" pageId="23" pageNumber="538">′</emphasis>
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E
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</geoCoordinate>
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). Collected by Jacqueline Peter King. Date:
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<date id="FFEE105CFF98E8140E4A8E3C21A3FD8F" box="[617,710,559,581]" pageId="23" pageNumber="538" value="2001-12">
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<collectingDate id="EFAAE9B4FF98E8140E4A8E3C21A3FD8F" box="[617,710,559,581]" pageId="23" pageNumber="538" value="2001-12">xii.2001</collectingDate>
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</date>
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,
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<typeStatus id="54EB883EFF98E8140EF08E3C23BDFDA9" pageId="23" pageNumber="538" type="holotype">holotype</typeStatus>
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, BOR/MOL 1375,
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<figureCitation id="136B2A19FF98E8140D908E5E2142FDA9" box="[435,551,589,611]" captionStart="Figure 2" captionStartId="7.[164,243,1411,1430]" captionTargetBox="[326,1281,196,1382]" captionTargetId="figure-180@7.[323,1283,194,1382]" captionTargetPageId="7" captionText="Figure 2. Shells. A, Everettia corrugata corrugata; B, Everettia corrugata williamsi ssp. nov.; C, Everettia interior sp. nov.; D, Everettia jucunda; E, Everettia occidentalis sp. nov.; F, Everettia jucundior sp. nov. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5443027" httpUri="https://zenodo.org/record/5443027/files/figure.png" pageId="23" pageNumber="538">Figure 2F</figureCitation>
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</materialsCitation>
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||
;
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<materialsCitation id="3B383CC1FF98E8140E158E5D2191FD75" collectingDate="2002-05-16" collectionCode="BMNH" country="Malaysia" latitude="5.5166664" location="Kinabatangan District" longLatPrecision="1305" longitude="118.21667" municipality="State" pageId="23" pageNumber="538" specimenCode="BMNH 20080629" specimenCount="1" stateProvince="Sabah" typeStatus="paratype">
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<collectingCountry id="F347760CFF98E8140E158E5D21DAFDA9" box="[566,703,590,611]" name="Malaysia" pageId="23" pageNumber="538">MALAYSIA</collectingCountry>
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:
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<collectingMunicipality id="6B8BACE6FF98E8140EED8E5D206EFDA9" box="[718,779,590,611]" pageId="23" pageNumber="538">State</collectingMunicipality>
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of
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||
<collectingRegion id="4994F87EFF98E8140CE78E7F226AFD48" box="[196,271,620,642]" country="Malaysia" name="Sabah" pageId="23" pageNumber="538">Sabah</collectingRegion>
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:
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<location id="8E8F6047FF98E8140D3E8E7F2142FD48" LSID="urn:lsid:plazi:treatment:03F9878AFF98E8140D368D7D261DFBBC:8E8F6047FF98E8140D3E8E7F2142FD48" box="[285,551,620,642]" country="Malaysia" latitude="5.5166664" longLatPrecision="1305" longitude="118.21667" municipality="State" name="Kinabatangan District" pageId="23" pageNumber="538" stateProvince="Sabah">Kinabatangan District</location>
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. Bod Tai limestone (
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<geoCoordinate id="EE64505BFF98E8140C8F8E99239EFD6A" box="[172,251,649,672]" degrees="5" direction="north" minutes="31" orientation="latitude" pageId="23" pageNumber="538" precision="925" value="5.5166664">
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||
5°31
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<emphasis id="B924EA8EFF98E8140CFC8E9A2380FD6A" box="[223,229,649,672]" italics="true" pageId="23" pageNumber="538">′</emphasis>
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N
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</geoCoordinate>
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,
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<geoCoordinate id="EE64505BFF98E8140D2A8E982215FD6A" box="[265,368,649,672]" degrees="118" direction="east" minutes="13" orientation="longitude" pageId="23" pageNumber="538" precision="925" value="118.21667">
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118°13
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<emphasis id="B924EA8EFF98E8140D758E9A2239FD6A" box="[342,348,649,672]" italics="true" pageId="23" pageNumber="538">′</emphasis>
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E
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</geoCoordinate>
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). Collected by Menno Schilthuizen. Date:
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<date id="FFEE105CFF98E8140CC88EBA223DFD75" box="[235,344,681,703]" pageId="23" pageNumber="538" value="2002-05-16">
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<collectingDate id="EFAAE9B4FF98E8140CC88EBA223DFD75" box="[235,344,681,703]" pageId="23" pageNumber="538" value="2002-05-16">16.v.2002</collectingDate>
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||
</date>
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||
,
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||
<specimenCount id="9D56FD15FF98E8140D458EB9229BFD75" box="[358,510,682,703]" pageId="23" pageNumber="538" type="generic" typeStatus="paratype">one paratype</specimenCount>
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||
in
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||
<specimenCode id="DBF69EE7FF98E8140E0B8EB92191FD75" box="[552,756,681,703]" collectionCode="BMNH" pageId="23" pageNumber="538">BMNH 20080629</specimenCode>
|
||
</materialsCitation>
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||
.
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||
</paragraph>
|
||
<paragraph id="8BEF369CFF98E8140C878EFB21D5FCF1" blockId="23.[164,779,744,827]" pageId="23" pageNumber="538">
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||
<emphasis id="B924EA8EFF98E8140C878EFB2247FD37" box="[164,290,744,765]" italics="true" pageId="23" pageNumber="538">Etymology:</emphasis>
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||
This species is named
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||
<taxonomicName id="4C504D1FFF98E8140E1A8EFB21C2FD37" authorityName="Liew & Schilthuizen & Vermeulen" authorityYear="2009" box="[569,679,744,765]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="538" phylum="Mollusca" rank="species" species="jucundior">
|
||
<emphasis id="B924EA8EFF98E8140E1A8EFB21C2FD37" box="[569,679,744,765]" italics="true" pageId="23" pageNumber="538">jucundior</emphasis>
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||
</taxonomicName>
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||
because of its shell surface sculpture: similar to (but even more jocund than) that of
|
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<taxonomicName id="4C504D1FFF98E8140DF88F3521CCFCF0" authorityName=", Schileyko, 2003:" authorityYear="1360" box="[475,681,805,827]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="538" phylum="Mollusca" rank="species" species="jucunda">
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||
<emphasis id="B924EA8EFF98E8140DF88F3521CCFCF0" box="[475,681,805,827]" italics="true" pageId="23" pageNumber="538">Everettia jucunda</emphasis>
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</taxonomicName>
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||
.
|
||
</paragraph>
|
||
<paragraph id="8BEF369CFF98E8140C878F77206EFB00" blockId="23.[164,779,868,1226]" pageId="23" pageNumber="538">
|
||
<emphasis id="B924EA8EFF98E8140C878F7722E4FCB3" box="[164,385,868,889]" italics="true" pageId="23" pageNumber="538">Material examined:</emphasis>
|
||
<materialsCitation id="3B383CC1FF98E8140DAA8F772158FC52" collectingDate="2000-07" elevation="100" location="Batu Baturong" municipality="Tawau" pageId="23" pageNumber="538" specimenCount="1" stateProvince="Sabah">
|
||
<collectingRegion id="4994F87EFF98E8140DAA8F772287FCB3" box="[393,482,868,889]" country="Malaysia" name="Sabah" pageId="23" pageNumber="538">SABAH</collectingRegion>
|
||
–
|
||
<collectingMunicipality id="6B8BACE6FF98E8140E238F772135FCB3" box="[512,592,868,889]" pageId="23" pageNumber="538">Tawau</collectingMunicipality>
|
||
: BOR/MOL 922,
|
||
<location id="8E8F6047FF98E8140C878F902237FC52" LSID="urn:lsid:plazi:treatment:03F9878AFF98E8140D368D7D261DFBBC:8E8F6047FF98E8140C878F902237FC52" box="[164,338,899,920]" municipality="Tawau" name="Batu Baturong" pageId="23" pageNumber="538" stateProvince="Sabah">Batu Baturong</location>
|
||
<quantity id="4CA89B79FF98E8140D788F9022C4FC52" box="[347,417,899,920]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="538" unit="m" value="100.0">
|
||
<elevation id="007DD1AFFF98E8140D788F9022C4FC52" box="[347,417,899,920]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="538" unit="m" value="100.0">100 m</elevation>
|
||
</quantity>
|
||
alt.,
|
||
<date id="FFEE105CFF98E8140DC28F912158FC52" box="[481,573,898,920]" pageId="23" pageNumber="538" value="2000-07">
|
||
<collectingDate id="EFAAE9B4FF98E8140DC28F912158FC52" box="[481,573,898,920]" pageId="23" pageNumber="538" value="2000-07">vii.2000</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
; BOR/MOL 1376, Tawau Hills Park,
|
||
<date id="FFEE105CFF98E8140DB68FB22283FC7D" box="[405,486,929,951]" pageId="23" pageNumber="538" value="2002-02">ii.2002</date>
|
||
.
|
||
<materialsCitation id="3B383CC1FF98E8140DDA8FB22131FC1C" collectingDate="1986-09" collectionCode="JJ" county="Lahad Datu" location="Kirk's Cave" municipality="Segama Valley" pageId="23" pageNumber="538" specimenCode="JJ 1215" specimenCount="1" stateProvince="SABAH">
|
||
<collectingCounty id="628E4E10FF98E8140DDA8FB221EAFC7D" box="[505,655,929,951]" pageId="23" pageNumber="538">Lahad Datu</collectingCounty>
|
||
:
|
||
<specimenCode id="DBF69EE7FF98E8140E818FB12063FC7D" box="[674,774,929,951]" collectionCode="JJ" pageId="23" pageNumber="538">JJ 1215</specimenCode>
|
||
,
|
||
<collectingMunicipality id="6B8BACE6FF98E8140C878FD32235FC1C" box="[164,336,960,982]" pageId="23" pageNumber="538">Segama Valley</collectingMunicipality>
|
||
,
|
||
<location id="8E8F6047FF98E8140D428FD3228BFC1F" LSID="urn:lsid:plazi:treatment:03F9878AFF98E8140D368D7D261DFBBC:8E8F6047FF98E8140D428FD3228BFC1F" box="[353,494,960,982]" county="Lahad Datu" municipality="Segama Valley" name="Kirk's Cave" pageId="23" pageNumber="538" stateProvince="SABAH">Kirk’s Cave</location>
|
||
,
|
||
<date id="FFEE105CFF98E8140DDC8FD32131FC1C" box="[511,596,960,982]" pageId="23" pageNumber="538" value="1986-09">
|
||
<collectingDate id="EFAAE9B4FF98E8140DDC8FD32131FC1C" box="[511,596,960,982]" pageId="23" pageNumber="538" value="1986-09">ix.1986</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B383CC1FF98E8140E478FD32158FC3E" collectingDate="2005-01" location="Tabin Wildlife Reserve" pageId="23" pageNumber="538" specimenCount="1" stateProvince="SABAH">
|
||
<location id="8E8F6047FF98E8140E478FD32267FC3E" LSID="urn:lsid:plazi:treatment:03F9878AFF98E8140D368D7D261DFBBC:8E8F6047FF98E8140E478FD32267FC3E" name="Tabin Wildlife Reserve" pageId="23" pageNumber="538" stateProvince="SABAH">Tabin Wildlife Reserve</location>
|
||
: BOR/MOL 5268,
|
||
<date id="FFEE105CFF98E8140DD68FCD2158FC3E" box="[501,573,990,1012]" pageId="23" pageNumber="538" value="2005-01">
|
||
<collectingDate id="EFAAE9B4FF98E8140DD68FCD2158FC3E" box="[501,573,990,1012]" pageId="23" pageNumber="538" value="2005-01">i.2005</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B383CC1FF98E8140E768FCC22A1FBD8" collectingDate="2000-05" location="Danum Valley" pageId="23" pageNumber="538" specimenCount="1" stateProvince="SABAH">
|
||
<location id="8E8F6047FF98E8140E768FCC2062FC3E" LSID="urn:lsid:plazi:treatment:03F9878AFF98E8140D368D7D261DFBBC:8E8F6047FF98E8140E768FCC2062FC3E" box="[597,775,990,1012]" name="Danum Valley" pageId="23" pageNumber="538" stateProvince="SABAH">Danum Valley</location>
|
||
: BOR/MOL 1231,
|
||
<date id="FFEE105CFF98E8140D5A8FEE22A1FBD8" box="[377,452,1021,1042]" pageId="23" pageNumber="538" value="2000-05">
|
||
<collectingDate id="EFAAE9B4FF98E8140D5A8FEE22A1FBD8" box="[377,452,1021,1042]" pageId="23" pageNumber="538" value="2000-05">v.2000</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B383CC1FF98E8140DFB8FEE211AFBFB" collectingDate="2002-04" collectorName="Batu Tomanggong Besar" location="Kinabatangan" pageId="23" pageNumber="538" specimenCount="1" stateProvince="SABAH">
|
||
<location id="8E8F6047FF98E8140DFB8FEE211AFBD9" LSID="urn:lsid:plazi:treatment:03F9878AFF98E8140D368D7D261DFBBC:8E8F6047FF98E8140DFB8FEE211AFBD9" box="[472,639,1021,1043]" name="Kinabatangan" pageId="23" pageNumber="538" stateProvince="SABAH">Kinabatangan</location>
|
||
: BOR/MOL 1387,
|
||
<collectorName id="26A5534AFF98E8140CD3880F217CFBFB" box="[240,537,1052,1073]" pageId="23" pageNumber="538">Batu Tomanggong Besar</collectorName>
|
||
,
|
||
<date id="FFEE105CFF98E8140E0E8808211AFBFB" box="[557,639,1051,1073]" pageId="23" pageNumber="538" value="2002-04">
|
||
<collectingDate id="EFAAE9B4FF98E8140E0E8808211AFBFB" box="[557,639,1051,1073]" pageId="23" pageNumber="538" value="2002-04">iv.2002</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
; BOR/MOL 5269, Batu Mawas,
|
||
<date id="FFEE105CFF98E8140DA5882922B8FB9A" box="[390,477,1082,1104]" pageId="23" pageNumber="538" value="2003-03">iii.2003</date>
|
||
; BOR/MOL 5270, Bod Tai,
|
||
<date id="FFEE105CFF98E8140C87884A238AFBA4" box="[164,239,1113,1134]" pageId="23" pageNumber="538" value="2002-05">v.2002</date>
|
||
;
|
||
<materialsCitation id="3B383CC1FF98E8140CD9884A21D8FB66" collectingDate="1965-01" collectionCode="BMNH" collectorName="Crocker Range & Keningau & Kampung Matou" location="Abai river" municipality="Abai river" pageId="23" pageNumber="538" specimenCode="BMNH 1897.5, ZMA 176863" specimenCount="1" stateProvince="SABAH">
|
||
<collectionCode id="ED41AE59FF98E8140CD9884A2229FBA4" box="[250,332,1113,1134]" pageId="23" pageNumber="538">BMNH</collectionCode>
|
||
1897.5.13.2-5,
|
||
<collectingMunicipality id="6B8BACE6FF98E8140DD5884A210FFBA5" box="[502,618,1113,1135]" pageId="23" pageNumber="538">Abai river</collectingMunicipality>
|
||
, 1897;
|
||
<specimenCode id="DBF69EE7FF98E8140E9A884A2390FB47" collectionCode="BMNH" pageId="23" pageNumber="538">BMNH 1897.5</specimenCode>
|
||
.13.1,
|
||
<location id="8E8F6047FF98E8140D1A886422D6FB47" LSID="urn:lsid:plazi:treatment:03F9878AFF98E8140D368D7D261DFBBC:8E8F6047FF98E8140D1A886422D6FB47" box="[313,435,1143,1165]" municipality="Abai river" name="Abai river" pageId="23" pageNumber="538" stateProvince="SABAH">Abai river</location>
|
||
, 1897.
|
||
<collectorName id="26A5534AFF98E8140E2C886421A4FB47" box="[527,705,1143,1165]" pageId="23" pageNumber="538">Crocker Range</collectorName>
|
||
:
|
||
<specimenCode id="DBF69EE7FF98E8140EF08864239CFB66" collectionCode="ZMA" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="23" pageNumber="538">ZMA 176863</specimenCode>
|
||
,
|
||
<collectorName id="26A5534AFF98E8140D2E8885221AFB66" box="[269,383,1174,1196]" pageId="23" pageNumber="538">Keningau</collectorName>
|
||
,
|
||
<collectorName id="26A5534AFF98E8140DB088852105FB61" box="[403,608,1174,1195]" pageId="23" pageNumber="538">Kampung Matou</collectorName>
|
||
,
|
||
<date id="FFEE105CFF98E8140E57888521D8FB66" box="[628,701,1174,1196]" pageId="23" pageNumber="538" value="1965-01">
|
||
<collectingDate id="EFAAE9B4FF98E8140E57888521D8FB66" box="[628,701,1174,1196]" pageId="23" pageNumber="538" value="1965-01">i.1965</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B383CC1FF98E8140EF288852062FB00" collectingDate="1987-01" collectionCode="JJ" collectorName="Batu Tengar" location="Segarong Hills" municipality="Semporna" pageId="23" pageNumber="538" specimenCode="JJ 1823" specimenCount="1" stateProvince="SABAH">
|
||
<collectingMunicipality id="6B8BACE6FF98E8140EF28885238CFB00" pageId="23" pageNumber="538">Semporna</collectingMunicipality>
|
||
:
|
||
<specimenCode id="DBF69EE7FF98E8140CD588A62230FB00" box="[246,341,1205,1226]" collectionCode="JJ" pageId="23" pageNumber="538">JJ 1823</specimenCode>
|
||
,
|
||
<location id="8E8F6047FF98E8140D4088A62174FB00" LSID="urn:lsid:plazi:treatment:03F9878AFF98E8140D368D7D261DFBBC:8E8F6047FF98E8140D4088A62174FB00" box="[355,529,1204,1226]" municipality="Semporna" name="Segarong Hills" pageId="23" pageNumber="538" stateProvince="SABAH">Segarong Hills</location>
|
||
,
|
||
<collectorName id="26A5534AFF98E8140E3C88A621D5FB00" box="[543,688,1205,1226]" pageId="23" pageNumber="538">Batu Tengar</collectorName>
|
||
,
|
||
<date id="FFEE105CFF98E8140E9D88A72062FB00" box="[702,775,1204,1226]" pageId="23" pageNumber="538" value="1987-01">
|
||
<collectingDate id="EFAAE9B4FF98E8140E9D88A72062FB00" box="[702,775,1204,1226]" pageId="23" pageNumber="538" value="1987-01">i.1987</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEF369CFF98E8140C8788E721DEFA68" blockId="23.[164,779,1267,1442]" pageId="23" pageNumber="538">
|
||
<emphasis id="B924EA8EFF98E8140C8788E722BCFAC2" box="[164,473,1267,1289]" italics="true" pageId="23" pageNumber="538">Diagnostic characteristics:</emphasis>
|
||
Shiny shell with dense spiral striation above shell. The species differs from
|
||
<taxonomicName id="4C504D1FFF98E8140C8789222246FA8F" authorityName=", Schileyko, 2003:" authorityYear="1360" box="[164,291,1328,1350]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="538" phylum="Mollusca" rank="species" species="jucunda">
|
||
<emphasis id="B924EA8EFF98E8140C8789222246FA8F" box="[164,291,1328,1350]" italics="true" pageId="23" pageNumber="538">E. jucunda</emphasis>
|
||
</taxonomicName>
|
||
(which has similar sculpture) in terms of the shell shape and size (
|
||
<taxonomicName id="4C504D1FFF98E8140DE28943212BFAAE" authorityName="Liew & Schilthuizen & Vermeulen" authorityYear="2009" box="[449,590,1359,1381]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="538" phylum="Mollusca" rank="species" species="jucundior">
|
||
<emphasis id="B924EA8EFF98E8140DE28943212BFAAE" box="[449,590,1359,1381]" italics="true" pageId="23" pageNumber="538">E. jucundior</emphasis>
|
||
</taxonomicName>
|
||
with five whorls and shell
|
||
<quantity id="4CA89B79FF98E8140D39897D220DFA49" box="[282,360,1390,1411]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="23" pageNumber="538" unit="mm" value="19.0">19 mm</quantity>
|
||
in width vs.
|
||
<taxonomicName id="4C504D1FFF98E8140E29897D21EFFA49" authorityName=", Schileyko, 2003:" authorityYear="1360" box="[522,650,1390,1411]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="538" phylum="Mollusca" rank="species" species="jucunda">
|
||
<emphasis id="B924EA8EFF98E8140E29897D21EFFA49" box="[522,650,1390,1411]" italics="true" pageId="23" pageNumber="538">E. jucunda</emphasis>
|
||
</taxonomicName>
|
||
– with six and a half whorls and shell
|
||
<quantity id="4CA89B79FF98E8140DD1899F2125FA68" box="[498,576,1420,1442]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="23" pageNumber="538" unit="mm" value="17.0">17 mm</quantity>
|
||
in width).
|
||
</paragraph>
|
||
<paragraph id="8BEF369CFF98E8140C8789DF20E6FDA0" blockId="23.[164,779,1483,1903]" lastBlockId="23.[826,1441,198,618]" pageId="23" pageNumber="538">
|
||
<emphasis id="B924EA8EFF98E8140C8789DF2249FA2B" box="[164,300,1484,1505]" italics="true" pageId="23" pageNumber="538">Description:</emphasis>
|
||
Shell (
|
||
<figureCitation id="136B2A19FF98E8140D8389D82293FA2A" box="[416,502,1483,1505]" captionStart="Figure 2" captionStartId="7.[164,243,1411,1430]" captionTargetBox="[326,1281,196,1382]" captionTargetId="figure-180@7.[323,1283,194,1382]" captionTargetPageId="7" captionText="Figure 2. Shells. A, Everettia corrugata corrugata; B, Everettia corrugata williamsi ssp. nov.; C, Everettia interior sp. nov.; D, Everettia jucunda; E, Everettia occidentalis sp. nov.; F, Everettia jucundior sp. nov. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5443027" httpUri="https://zenodo.org/record/5443027/files/figure.png" pageId="23" pageNumber="538">Figs 2F</figureCitation>
|
||
,
|
||
<figureCitation id="136B2A19FF98E8140E3089D82125FA2B" box="[531,576,1483,1505]" captionStart="Figure 11" captionStartId="18.[144,223,1209,1228]" captionTargetBox="[305,1263,196,1180]" captionTargetId="figure-241@18.[303,1263,196,1180]" captionTargetPageId="18" captionText="Figure 11. Shell surface sculptures (scanning electron micrographs). A, upper surface of Everettia corrugata williamsi ssp. nov.; B, underside of E. corrugata williamsi ssp. nov.; C, underside of Everettia lapidini; D, upper surface of Everettia occidentalis sp. nov.; E, corrugation at the suture of Everettia corrugata corrugata; F, upper surface of Everettia jucundior sp. nov. Scale bar = 500 Mm." figureDoi="http://doi.org/10.5281/zenodo.5443045" httpUri="https://zenodo.org/record/5443045/files/figure.png" pageId="23" pageNumber="538">11F</figureCitation>
|
||
): medium-sized, rather thin, brownish, moderately elevated, outer whorls slightly shouldered or rounded. Periphery rounded. Above the periphery, shell shiny with densely placed spiral striation,
|
||
<emphasis id="B924EA8EFF98E8140E078A552153F991" box="[548,566,1606,1627]" italics="true" pageId="23" pageNumber="538">c.</emphasis>
|
||
20–30 striae per mm. Below the periphery, shell has fine, densely placed spiral grooves. Height up to
|
||
<quantity id="4CA89B79FF98E8140E168A9021FFF952" box="[565,666,1667,1688]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="23" pageNumber="538" unit="mm" value="10.9">10.9 mm</quantity>
|
||
; width up to
|
||
<quantity id="4CA89B79FF98E8140CE28AB22240F97D" box="[193,293,1697,1719]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="23" pageNumber="538" unit="mm" value="19.5">19.5 mm</quantity>
|
||
; diameter of the first three whorls 0.9, 0.8, and
|
||
<quantity id="4CA89B79FF98E8140CFE8AD32251F91F" box="[221,308,1728,1749]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="23" pageNumber="538" unit="mm" value="1.3">1.3 mm</quantity>
|
||
, respectively; number of whorls up to five and one-fifth; height aperture up to
|
||
<quantity id="4CA89B79FF98E8140E8C8ACD2063F93E" box="[687,774,1758,1780]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="23" pageNumber="538" unit="mm" value="8.3">8.3 mm</quantity>
|
||
; width aperture up to
|
||
<quantity id="4CA89B79FF98E8140D928AEE2173F8D8" box="[433,534,1789,1810]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="23" pageNumber="538" unit="mm" value="10.1">10.1 mm</quantity>
|
||
. Genitalia (
|
||
<figureCitation id="136B2A19FF98E8140E8B8AEE219AF8D9" box="[680,767,1789,1811]" captionStart="Figure 7" captionStartId="13.[164,243,1755,1774]" captionTargetBox="[167,1442,198,1722]" captionTargetId="figure-0@13.[164,1444,194,1725]" captionTargetPageId="13" captionText="Figure 7. Genitalia. A, Everettia klemmantanica; B, Everettia lapidini sp. nov.; C, Everettia paulbasintali sp. nov.; D, Everettia subconsul; E, Everettia themis; F, Everettia layanglayang sp. nov.; G, Everettia jucundior sp. nov.; H, Everettia interior sp. nov.; I, Everettia corrugata corrugata." figureDoi="http://doi.org/10.5281/zenodo.5443037" httpUri="https://zenodo.org/record/5443037/files/figure.png" pageId="23" pageNumber="538">Fig. 7G</figureCitation>
|
||
): maximum length from genital opening to the end of dart-sac (before the visible gland tubules) up to
|
||
<quantity id="4CA89B79FF98E8140C878B4A2391F8A4" box="[164,244,1881,1902]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="23" pageNumber="538" unit="mm" value="11.0">11 mm</quantity>
|
||
. The penis, dart-sac, and vagina with thick muscular walls. P, V, GO, BC, and DS arranged around the short genital atrium, with limited space between the opening of each of the genital structures. BC short, about one-third of total DS length. Animal (
|
||
<figureCitation id="136B2A19FF98E8140F618D5320FFFE9C" box="[834,922,320,342]" captionStart="Figure 8" captionStartId="14.[144,223,1760,1779]" captionTargetBox="[147,1419,200,1728]" captionTargetId="figure-0@14.[144,1424,195,1730]" captionTargetPageId="14" captionText="Figure 8. Animal and mantle pigmentation. A, Everettia subconsul west coast lowland; B, Everettia layanglayang sp. nov.; C, Everettia monticola sp. nov.; D, Everettia safriei sp. nov.; E, Everettia interior sp. nov.; F, Everettia subconsul east coast lowland; G, Everettia corrugata corrugata; H, Everettia jucundior sp. nov.; I, Everettia klemmantanica; J, Everettia lapidini sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5443039" httpUri="https://zenodo.org/record/5443039/files/figure.png" pageId="23" pageNumber="538">Fig. 8H</figureCitation>
|
||
): black bands on either side of the tentacles, which are separated by a white band that occupies the space between the eye tentacles and extends downward to just below the eye tentacles. Grey smear on either side of the tail extending from the umbilicus region to the end of the tail. Other parts of the animal are clear from any markings. The mantle is black and covered by crowded tiny bright spots and followed by regularly spaced radial black bands in the last half whorl.
|
||
</paragraph>
|
||
<paragraph id="8BEF369CFF98E8140F198EB120C1FD3F" blockId="23.[826,1441,674,758]" pageId="23" pageNumber="538">
|
||
<emphasis id="B924EA8EFF98E8140F198EB12714FD7D" box="[826,1137,674,695]" italics="true" pageId="23" pageNumber="538">Distribution and habitat:</emphasis>
|
||
Primary forest.
|
||
<collectingRegion id="4994F87EFF98E81409708EB126FBFD72" box="[1363,1438,674,696]" country="Malaysia" name="Sabah" pageId="23" pageNumber="538">Sabah</collectingRegion>
|
||
: Kinabatangan, Tawau Hills Park, Danum valley (
|
||
<figureCitation id="136B2A19FF98E8140F618EF320F2FD3F" box="[834,919,735,758]" captionStart="Figure 5" captionStartId="10.[144,223,1342,1361]" captionTargetBox="[305,1261,197,1311]" captionTargetId="figure-168@10.[303,1263,195,1313]" captionTargetPageId="10" captionText="Figure 5. Distribution maps (Sabah). A, Everettia subconsul in the east and west coast lowland, Everettia jucunda in the west coast, and Everettia occidentalis sp. nov. in the western mountain range; B, Everettia klemmantanica in the central mountain ranges, Everettia interior sp. nov. in the interior area, Everettia paulbasintali sp. nov. in the east coast lowland, Everettia themis in the central mountain ranges; C, Everettia lapidini sp. nov. in the central mountain ranges, Everettia jucundior sp. nov. in the east coast lowland; D, Everettia monticola sp. nov. in the central mountain ranges, Everettia planispira sp. nov. in the eastern and northern parts of Sabah." figureDoi="http://doi.org/10.5281/zenodo.5443033" httpUri="https://zenodo.org/record/5443033/files/figure.png" pageId="23" pageNumber="538">Fig. 5C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEF369CFF98E8140F198F3D261DFBBC" blockId="23.[826,1441,814,1142]" pageId="23" pageNumber="538">
|
||
<emphasis id="B924EA8EFF98E8140F198F3D20C3FC89" box="[826,934,814,835]" italics="true" pageId="23" pageNumber="538">Remarks:</emphasis>
|
||
The specimens in
|
||
<bibRefCitation id="EFC14B6DFF98E81408878F3D263CFC8F" author="Laidlaw FF" box="[1188,1369,814,837]" pageId="23" pageNumber="538" pagination="190 - 201" refId="ref25575" refString="Laidlaw FF. 1931. On a new sub-family Dyakiinae of the Zonitidae. Proceedings of the Malacological Society 19: 190 - 201." type="journal article" year="1931">Laidlaw (1931)</bibRefCitation>
|
||
were collected by R. Hanitsch in
|
||
<quantity id="4CA89B79FF98E81408B68F5E2794FCA9" box="[1173,1265,845,867]" metricMagnitude="1" metricUnit="m" metricValue="4.82346" pageId="23" pageNumber="538" unit="in" value="1899.0">1899 in</quantity>
|
||
British
|
||
<collectingCountry id="F347760CFF98E814097E8F5E20EBFC4B" name="Malaysia" pageId="23" pageNumber="538">North Borneo</collectingCountry>
|
||
. According to the description by Laidlaw, these shells’ size and sculpture fit with the material we consider as
|
||
<taxonomicName id="4C504D1FFF98E8140F9C8FBA2729FC74" authorityName="Liew & Schilthuizen & Vermeulen" authorityYear="2009" box="[959,1100,937,958]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="538" phylum="Mollusca" rank="species" species="jucundior">
|
||
<emphasis id="B924EA8EFF98E8140F9C8FBA2729FC74" box="[959,1100,937,958]" italics="true" pageId="23" pageNumber="538">E. jucundior</emphasis>
|
||
</taxonomicName>
|
||
. Some material in the
|
||
<collectionCode id="ED41AE59FF98E814096C8FBA26C4FC74" box="[1359,1441,937,958]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="23" pageNumber="538" type="Museum">RMNH</collectionCode>
|
||
and JJ collections from Balikpapan (Southeast Coast of Kalimantan) and Sarawak’ Fourth Division have similar spiral striation and shell shape but we do not include them in this
|
||
<collectingRegion id="4994F87EFF98E814081E883027E0FBF3" box="[1085,1157,1059,1081]" country="Malaysia" name="Sabah" pageId="23" pageNumber="538">Sabah</collectingRegion>
|
||
<taxonomicName id="4C504D1FFF98E81408B18837279CFBF3" authorityName="Godwin-Austen" authorityYear="1891" box="[1170,1273,1060,1081]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="538" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B924EA8EFF98E81408B18837279CFBF3" box="[1170,1273,1060,1081]" italics="true" pageId="23" pageNumber="538">Everettia</emphasis>
|
||
</taxonomicName>
|
||
revision. Our genetic data also show that
|
||
<taxonomicName id="4C504D1FFF98E814085A88512662FB9D" authorityName="Liew & Schilthuizen & Vermeulen" authorityYear="2009" box="[1145,1287,1090,1111]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="538" phylum="Mollusca" rank="species" species="jucundior">
|
||
<emphasis id="B924EA8EFF98E814085A88512662FB9D" box="[1145,1287,1090,1111]" italics="true" pageId="23" pageNumber="538">E. jucundior</emphasis>
|
||
</taxonomicName>
|
||
is not closely related to other
|
||
<collectingRegion id="4994F87EFF98E8140FD988732727FBBC" box="[1018,1090,1120,1142]" country="Malaysia" name="Sabah" pageId="23" pageNumber="538">Sabah</collectingRegion>
|
||
lowland
|
||
<taxonomicName id="4C504D1FFF98E814088C8872261DFBBC" box="[1199,1400,1120,1142]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="538" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B924EA8EFF98E814088C88722673FBBC" box="[1199,1302,1121,1142]" italics="true" pageId="23" pageNumber="538">Everettia</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |