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<document id="32D9A430394AA924639403861B48C108" ID-CLB-Dataset="6279" ID-DOI="10.5852/ejt.2021.751.1381" ID-GBIF-Dataset="4e3af683-d8a6-40de-991b-c4584d66b8cc" ID-ISSN="2118-9773" ID-Zenodo-Dep="4906007" ID-ZooBank="1D27EA41-2396-463A-B1B2-0CCF79C9F3D2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623068023717" checkinUser="felipe" docAuthor="Zamani, Alireza, Chatzaki, Maria, Esyunin, Sergei L. &amp; Marusik, Yuri M." docDate="2021" docId="03F98785036FEF731C5D7A3EFC1D6CF7" docLanguage="en" docName="ejt-751_68-114.pdf" docOrigin="European Journal of Taxonomy 751 (1)" docSource="http://dx.doi.org/10.5852/ejt.2021.751.1381" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Gnaphosa qamsarica Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik 2021, sp. nov." docType="treatment" docUuid="571ED6FD-935D-4527-8581-FEBD31C996E5" docUuidSource="ZooBank" docVersion="8" lastPageNumber="86" masterDocId="FFC0FFFD037FEF611E347F73FFC16D41" masterDocTitle="One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations" masterLastPageNumber="114" masterPageNumber="68" pageNumber="84" updateTime="1699000119853" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D2232F517430B1F68FD08102A84EE22A">One new genus and nineteen new species of ground spiders (Araneae: Gnaphosidae) from Iran, with other taxonomic considerations</mods:title>
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<mods:namePart id="0F3C60369CCED1374BF4BD0246D7445D">Zamani, Alireza</mods:namePart>
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<mods:affiliation id="2A3839A97BCB63E876FB622770E9B8A6">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland. Department of Molecular Biology and Genetics, Democritus University of Thrace, Dragana, 68100 Alexandroupolis, Greece. Perm State University, Bukireva Street 15, Perm, 614600, Russia. Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, 685000, Russia. Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa.</mods:affiliation>
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<mods:namePart id="D736E13CC01A467A74F825DBD5CE6D7E">Chatzaki, Maria</mods:namePart>
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<mods:namePart id="557FA724FC567A82F6604C455B91A1D7">Esyunin, Sergei L.</mods:namePart>
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<mods:affiliation id="A45D9DB9D8EB84C410AD43CD8B629E44">Sergei.</mods:affiliation>
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<mods:namePart id="5B8586FE3EEEB7A45222ECA9ED30766A">Marusik, Yuri M.</mods:namePart>
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<mods:date id="85B44501333525FF348E0AC3AA235CF4">2021</mods:date>
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<treatment id="03F98785036FEF731C5D7A3EFC1D6CF7" ID-DOI="http://doi.org/10.5281/zenodo.4906473" ID-GBIF-Taxon="182597840" ID-Zenodo-Dep="4906473" LSID="urn:lsid:zoobank.org:act:571ED6FD-935D-4527-8581-FEBD31C996E5" httpUri="http://treatment.plazi.org/id/03F98785036FEF731C5D7A3EFC1D6CF7" lastPageId="18" lastPageNumber="86" pageId="16" pageNumber="84">
<subSubSection id="C34A6518036FEF711C5D7A3EFC0A6829" box="[617,971,1357,1384]" pageId="16" pageNumber="84" type="nomenclature">
<paragraph id="8BEF3693036FEF711C5D7A3EFC0A6829" blockId="16.[368,1219,1357,1458]" box="[617,971,1357,1384]" pageId="16" pageNumber="84">
<heading id="D0A781FF036FEF711C5D7A3EFC0A6829" box="[617,971,1357,1384]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="84" reason="2">
<taxonomicName id="4C504D10036FEF711C5D7A3EFCA86826" ID-CoL="98YVJ" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[617,873,1357,1384]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="84" phylum="Arthropoda" rank="species" species="qamsarica" status="sp. nov.">
<emphasis id="B924EA81036FEF711C5D7A3EFCA86826" bold="true" box="[617,873,1357,1384]" italics="true" pageId="16" pageNumber="84">Gnaphosa qamsarica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757FA036FEF711D457A3DFC0A6829" box="[881,971,1358,1384]" pageId="16" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
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<uri id="FFC13A91036FEF711F447A00FB0268CF" box="[368,1219,1395,1422]" pageId="16" pageNumber="84">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
<paragraph id="8BEF3693036FEF711CC47AEBFC8268F3" blockId="16.[368,1219,1357,1458]" box="[752,835,1432,1458]" pageId="16" pageNumber="84">
<figureCitation id="136B2A16036FEF711CC47AEBFC8268F3" box="[752,835,1432,1458]" captionStart="Fig" captionStartId="17.[189,232,1928,1954]" captionTargetBox="[195,1391,1020,1890]" captionTargetId="figure-223@17.[195,1391,1020,1890]" captionTargetPageId="17" captionText="Fig. 10. Gnaphosa qamsarica sp. nov., holotype, ♀ (MHNG). A, C, E. Epigyne, ventral view. B, D, F. Epigyne, dorsal view. G. Chelicerae, posterior view. H. Habitus, dorsal view. Abbreviation: Lm = lateral margin. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906033" httpUri="https://zenodo.org/record/4906033/files/figure.png" pageId="16" pageNumber="84">Fig. 10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34A6518036FEF711E897AA8FCCF6A34" pageId="16" pageNumber="84" type="diagnosis">
<paragraph id="8BEF3693036FEF711E897AA8FEF768B4" blockId="16.[189,310,1499,1525]" box="[189,310,1499,1525]" pageId="16" pageNumber="84">
<heading id="D0A781FF036FEF711E897AA8FEF768B4" bold="true" box="[189,310,1499,1525]" fontSize="11" level="3" pageId="16" pageNumber="84" reason="3">
<emphasis id="B924EA81036FEF711E897AA8FEF768B4" bold="true" box="[189,310,1499,1525]" pageId="16" pageNumber="84">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF3693036FEF711E89797FFCCF6A34" blockId="16.[187,1399,1548,1909]" pageId="16" pageNumber="84">
The new species differs from all Palearctic congeners by the widely separated lateral margins not converging posteriorly, and widely separated receptacles leaving an open atrium with parallel rims (vs receptacles almost touching and lateral margins converging posteriorly). Furthermore, it differs by the presence of short copulatory ducts, almost vertical and in line with the receptacles (vs bent copulatory ducts parallel to the receptacles). The species most similar to
<taxonomicName id="4C504D10036FEF711CEF79D2FCBA6BFB" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[731,891,1697,1723]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="84" phylum="Arthropoda" rank="species" species="qamsarica" status="sp. nov.">
<emphasis id="B924EA81036FEF711CEF79D2FD316BFA" box="[731,752,1697,1723]" italics="true" pageId="16" pageNumber="84">G</emphasis>
.
<emphasis id="B924EA81036FEF711CCB79D2FCBA6BFB" box="[767,891,1697,1722]" italics="true" pageId="16" pageNumber="84">qamsarica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757FA036FEF711DB479D2FC186BFA" box="[896,985,1697,1723]" pageId="16" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
are:
<taxonomicName id="4C504D10036FEF711A3B79D2FAB16BFA" authority="Ponomarev, 1981" authorityName="Ponomarev" authorityYear="1981" box="[1039,1392,1697,1723]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="84" phylum="Arthropoda" rank="species" species="cumensis">
<emphasis id="B924EA81036FEF711A3B79D2FB5D6BFB" box="[1039,1180,1697,1723]" italics="true" pageId="16" pageNumber="84">G. cumensis</emphasis>
Ponomarev, 1981
</taxonomicName>
,
<taxonomicName id="4C504D10036FEF711E8879B5FD416BA1" authority="Esyunin &amp; Efimik, 1997" authorityName="Esyunin &amp; Efimik" authorityYear="1997" box="[188,640,1734,1760]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="84" phylum="Arthropoda" rank="species" species="halophila">
<emphasis id="B924EA81036FEF711E8879B5FE906BA1" box="[188,337,1734,1760]" italics="true" pageId="16" pageNumber="84">G. halophila</emphasis>
Esyunin &amp; Efimik, 1997
</taxonomicName>
,
<taxonomicName id="4C504D10036FEF711CBA79B5FC7C6BA1" authority="Thorell, 1875" authorityName="Thorell" authorityYear="1875" box="[654,957,1733,1760]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="84" phylum="Arthropoda" rank="species" species="jucunda">
<emphasis id="B924EA81036FEF711CBA79B5FCD36BA1" box="[654,786,1734,1760]" italics="true" pageId="16" pageNumber="84">G. jucunda</emphasis>
Thorell, 1875
</taxonomicName>
,
<taxonomicName id="4C504D10036FEF711DF879B5FA966BA1" authority="Ovtsharenko, Platnick" authorityName="Ovtsharenko, Platnick" box="[972,1367,1734,1760]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="84" phylum="Arthropoda" rank="species" species="saurica">
<emphasis id="B924EA81036FEF711DF879B5FB886B9E" box="[972,1097,1734,1760]" italics="true" pageId="16" pageNumber="84">G. saurica</emphasis>
Ovtsharenko, Platnick
</taxonomicName>
&amp; Song, 1992,
<taxonomicName id="4C504D10036FEF711F7E7998FD1E6A44" authority="Ovtsharenko, Platnick" authorityName="Ovtsharenko, Platnick" box="[330,735,1771,1797]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="84" phylum="Arthropoda" rank="species" species="ukrainica">
<emphasis id="B924EA81036FEF711F7E7998FE1A6A44" box="[330,475,1771,1797]" italics="true" pageId="16" pageNumber="84">G. ukrainica</emphasis>
Ovtsharenko, Platnick
</taxonomicName>
&amp; Song, 1992 and
<taxonomicName id="4C504D10036FEF711D827998FB256A44" authority="Pavesi, 1880" authorityName="Pavesi" authorityYear="1880" box="[950,1252,1771,1797]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="84" phylum="Arthropoda" rank="species" species="zeugitana">
<emphasis id="B924EA81036FEF711D827998FB886A44" box="[950,1097,1771,1797]" italics="true" pageId="16" pageNumber="84">G. zeugitana</emphasis>
Pavesi, 1880
</taxonomicName>
. The last one is the most similar in that the copulatory ducts are almost in line with the receptacles and the receptacles are widely separated. However,
<taxonomicName id="4C504D10036FEF711C0F7845FD1D6A0E" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[571,732,1846,1872]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="84" phylum="Arthropoda" rank="species" species="qamsarica" status="sp. nov.">
<emphasis id="B924EA81036FEF711C0F7845FD1D6A0E" box="[571,732,1846,1872]" italics="true" pageId="16" pageNumber="84">G. qamsarica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757FA036FEF711CD07845FCFE6A11" box="[740,831,1846,1872]" pageId="16" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="4C504D10036FEF711DEF7845FBB36A0E" authorityName="Pavesi" authorityYear="1880" box="[987,1138,1846,1872]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="84" phylum="Arthropoda" rank="species" species="zeugitana">
<emphasis id="B924EA81036FEF711DEF7845FBB36A0E" box="[987,1138,1846,1872]" italics="true" pageId="16" pageNumber="84">G. zeugitana</emphasis>
</taxonomicName>
by the parallel lateral margins and the rectangular atrium (vs triangular).
</paragraph>
</subSubSection>
<subSubSection id="C34A6518036FEF711E8978EDFB566AA9" box="[189,1175,1950,2025]" pageId="16" pageNumber="84" type="etymology">
<paragraph id="8BEF3693036FEF711E8978EDFE856AF9" blockId="16.[189,324,1950,1976]" box="[189,324,1950,1976]" pageId="16" pageNumber="84">
<heading id="D0A781FF036FEF711E8978EDFE856AF9" bold="true" box="[189,324,1950,1976]" fontSize="11" level="3" pageId="16" pageNumber="84" reason="3">
<emphasis id="B924EA81036FEF711E8978EDFE856AF9" bold="true" box="[189,324,1950,1976]" pageId="16" pageNumber="84">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF3693036FEF711E8978BDFB566AA9" blockId="16.[189,1175,1998,2025]" box="[189,1175,1998,2025]" pageId="16" pageNumber="84">
The specific epithet is an adjective and refers to the
<typeStatus id="54EB8831036FEF711D2A78BCFC906AA8" box="[798,849,1999,2025]" pageId="16" pageNumber="84">type</typeStatus>
locality of the new species.
</paragraph>
</subSubSection>
<subSubSection id="C34A6518036EEF701E897E7AFE986CFA" pageId="17" pageNumber="85" type="materials_examined">
<paragraph id="8BEF3693036EEF701E897E7AFEAD6C63" blockId="17.[189,364,264,291]" box="[189,364,264,291]" pageId="17" pageNumber="85">
<heading id="D0A781FF036EEF701E897E7AFEAD6C63" bold="true" box="[189,364,264,291]" fontSize="11" level="3" pageId="17" pageNumber="85" reason="3">
<emphasis id="B924EA81036EEF701E897E7AFEAD6C63" bold="true" box="[189,364,264,291]" pageId="17" pageNumber="85">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF3693036EEF701EDE7E44FE986CFA" blockId="17.[189,1399,311,443]" pageId="17" pageNumber="85">
<materialsCitation id="3B383CCE036EEF701EDE7E44FE986CFA" ID-GBIF-Occurrence="3127434333" collectingDate="2016-05-19" collectionCode="MHNG" collectorName="P. Ponel" country="Iran" county="Qamsar" elevation="2710" latitude="33.616665" location="Barzok Protected Area" longLatPrecision="1204" longitude="51.316666" municipality="Barzok Protected Area" pageId="17" pageNumber="85" specimenCount="1" specimenCount-female="1" stateProvince="Isfahan Province" typeStatus="holotype">
<emphasis id="B924EA81036EEF701EDE7E44FE9D6C10" bold="true" box="[234,348,311,337]" pageId="17" pageNumber="85">
<typeStatus id="54EB8831036EEF701EDE7E44FE9D6C10" box="[234,348,311,337]" pageId="17" pageNumber="85" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F3477603036EEF701E897E29FEC96C35" box="[189,264,346,372]" name="Iran" pageId="17" pageNumber="85">IRAN</collectingCountry>
•
<specimenCount id="9D56FD1A036EEF701F127E28FEFD6C34" box="[294,316,347,373]" pageId="17" pageNumber="85" type="female">♀</specimenCount>
<collectingRegion id="4994F871036EEF701F717E29FDD06C35" box="[325,529,346,372]" country="Iran" name="Esfahan" pageId="17" pageNumber="85">Isfahan Province</collectingRegion>
,
<collectingCounty id="628E4E1F036EEF701C2A7E29FDBB6C35" box="[542,634,346,372]" pageId="17" pageNumber="85">Qamsar</collectingCounty>
&amp;
<collectingMunicipality id="6B8BACE9036EEF701C977E29FC756C35" box="[675,948,346,372]" pageId="17" pageNumber="85">Barzok Protected Area</collectingMunicipality>
,
<locationDeviation id="9A5D2FB1036EEF701DF67E29FB0A6C35" box="[962,1227,346,373]" location="Qamsar" pageId="17" pageNumber="85">
<quantity id="4CA89B76036EEF701DF67E29FBCF6C35" box="[962,1038,346,373]" metricMagnitude="4" metricUnit="m" metricValue="5.5" pageId="17" pageNumber="85" unit="km" value="55.0">55 km</quantity>
SW of Qamsar
</locationDeviation>
,
<locationDeviation id="9A5D2FB1036EEF701AEC7E29FED76CD9" location="Kamoo" pageId="17" pageNumber="85">
<quantity id="4CA89B76036EEF701AEC7E29FAE46C35" box="[1240,1317,346,372]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="17" pageNumber="85" unit="km" value="14.0">14 km</quantity>
NE of Kamoo
</locationDeviation>
,
<locationDeviation id="9A5D2FB1036EEF701F157E0DFD146CD6" box="[289,725,381,408]" pageId="17" pageNumber="85">near the road of Gargash observatory</locationDeviation>
;
<geoCoordinate id="EE645054036EEF701CD47E0FFC8A6CD9" box="[736,843,380,408]" degrees="33" direction="north" minutes="37" orientation="latitude" pageId="17" pageNumber="85" precision="925" value="33.616665">33°37′ N</geoCoordinate>
,
<geoCoordinate id="EE645054036EEF701D627E0FFC7E6CD9" box="[854,959,380,408]" degrees="51" direction="east" minutes="19" orientation="longitude" pageId="17" pageNumber="85" precision="925" value="51.316666">51°19′ E</geoCoordinate>
;
<elevation id="007DD1A0036EEF701DFF7E0DFBA36CD6" box="[971,1122,381,408]" metricMagnitude="3" metricUnit="m" metricValue="2.71" pageId="17" pageNumber="85" unit="m" value="2710.0">
<quantity id="4CA89B76036EEF701DFF7E0DFBE26CD9" box="[971,1059,382,408]" metricMagnitude="3" metricUnit="m" metricValue="2.71" pageId="17" pageNumber="85" unit="m" value="2710.0">2710 m</quantity>
a.s.l.
</elevation>
;
<date id="FFEE1053036EEF701A5B7E0DFACC6CD6" box="[1135,1293,381,408]" pageId="17" pageNumber="85" value="2016-05-19">
<collectingDate id="EFAAE9BB036EEF701A5B7E0DFACC6CD6" box="[1135,1293,381,408]" pageId="17" pageNumber="85" value="2016-05-19">19 May 2016</collectingDate>
</date>
;
<collectorName id="26A55345036EEF701B2D7E0DFAB66CD6" box="[1305,1399,381,408]" pageId="17" pageNumber="85">P. Ponel</collectorName>
leg.;
<collectionCode id="ED41AE56036EEF701ECC7ED2FE976CFA" box="[248,342,417,443]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="17" pageNumber="85" type="Museum">MHNG</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34A6518036EEF731E897E91FEF86C07" lastPageId="18" lastPageNumber="86" pageId="17" pageNumber="85" type="description">
<paragraph id="8BEF3693036EEF701E897E91FE8F6CBD" blockId="17.[189,334,482,508]" box="[189,334,482,508]" pageId="17" pageNumber="85">
<heading id="D0A781FF036EEF701E897E91FE8F6CBD" bold="true" box="[189,334,482,508]" fontSize="11" level="3" pageId="17" pageNumber="85" reason="3">
<emphasis id="B924EA81036EEF701E897E91FE8F6CBD" bold="true" box="[189,334,482,508]" pageId="17" pageNumber="85">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF3693036EEF701EDE7D62FE846F6A" blockId="17.[189,1400,529,836]" box="[234,325,529,555]" pageId="17" pageNumber="85">
<emphasis id="B924EA81036EEF701EDE7D62FE846F6A" bold="true" box="[234,325,529,555]" pageId="17" pageNumber="85">Female</emphasis>
</paragraph>
<paragraph id="8BEF3693036EEF701E897D47FE096E05" blockId="17.[189,1400,529,836]" pageId="17" pageNumber="85">
Habitus as in
<figureCitation id="136B2A16036EEF701F697D47FE096F0F" box="[349,456,564,590]" captionStart="Fig" captionStartId="17.[189,232,1928,1954]" captionTargetBox="[195,1391,1020,1890]" captionTargetId="figure-223@17.[195,1391,1020,1890]" captionTargetPageId="17" captionText="Fig. 10. Gnaphosa qamsarica sp. nov., holotype, ♀ (MHNG). A, C, E. Epigyne, ventral view. B, D, F. Epigyne, dorsal view. G. Chelicerae, posterior view. H. Habitus, dorsal view. Abbreviation: Lm = lateral margin. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906033" httpUri="https://zenodo.org/record/4906033/files/figure.png" pageId="17" pageNumber="85">Fig. 10H</figureCitation>
. Total length 4.78. Carapace 1.79 long, 1.45 wide. Eye sizes and interdistance of PMEs: AME = 0.07, ALE = 0.09, PME = 0.09, PLE = 0.08, PME–PME = 0.04. Carapace, chelicerae, sternum, maxillae and labium dark brown; carapace with a distinct V-shaped mark, separating pars cephalica from pars thoracica. Chelicerae (
<figureCitation id="136B2A16036EEF701C8C7DEEFCE56FF9" box="[696,804,669,696]" captionStart="Fig" captionStartId="17.[189,232,1928,1954]" captionTargetBox="[195,1391,1020,1890]" captionTargetId="figure-223@17.[195,1391,1020,1890]" captionTargetPageId="17" captionText="Fig. 10. Gnaphosa qamsarica sp. nov., holotype, ♀ (MHNG). A, C, E. Epigyne, ventral view. B, D, F. Epigyne, dorsal view. G. Chelicerae, posterior view. H. Habitus, dorsal view. Abbreviation: Lm = lateral margin. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906033" httpUri="https://zenodo.org/record/4906033/files/figure.png" pageId="17" pageNumber="85">Fig. 10G</figureCitation>
) with 2 promarginal teeth and a serrated bifurcate retromarginal keel. Legs lighter than carapace, with few spines. Abdomen light brown, without any markings. Spinnerets light brown and uniform in color. Measurements of legs: I: 4.29 (1.15, 0.73, 0.96, 0.74, 0.71), II: 4.13 (1.12, 0.72, 0.85, 0.73, 0.71), III: 3.60 (0.98, 0.57, 0.65, 0.78, 0.62), IV: 5.40 (1.39, 0.79, 1.08, 1.26, 0.88).
</paragraph>
<paragraph id="8BEF3693036EEF701E897C1FFBA26E8D" blockId="17.[189,1399,876,973]" pageId="17" pageNumber="85">
Epigyne as in
<figureCitation id="136B2A16036EEF701F547C1FFE266EC7" box="[352,487,876,902]" captionStart="Fig" captionStartId="17.[189,232,1928,1954]" captionTargetBox="[195,1391,1020,1890]" captionTargetId="figure-223@17.[195,1391,1020,1890]" captionTargetPageId="17" captionText="Fig. 10. Gnaphosa qamsarica sp. nov., holotype, ♀ (MHNG). A, C, E. Epigyne, ventral view. B, D, F. Epigyne, dorsal view. G. Chelicerae, posterior view. H. Habitus, dorsal view. Abbreviation: Lm = lateral margin. Scale bars: 0.1 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.4906033" httpUri="https://zenodo.org/record/4906033/files/figure.png" pageId="17" pageNumber="85">Fig. 10A–F</figureCitation>
. Atrium rectangular; lateral margins (
<emphasis id="B924EA81036EEF701DAD7C1FFC7F6EC4" box="[921,958,876,901]" italics="true" pageId="17" pageNumber="85">Lm</emphasis>
) parallel, as long as scape; copulatory ducts almost as broad as receptacles, parallel to each other (i.e., almost vertical); receptacles oval and in line with copulatory ducts, separated by one receptacle width; glands indistinct.
</paragraph>
<caption id="DF2F661B036EEF701E8978FBFCB76AA8" ID-DOI="http://doi.org/10.5281/zenodo.4906033" ID-Zenodo-Dep="4906033" httpUri="https://zenodo.org/record/4906033/files/figure.png" pageId="17" pageNumber="85" startId="17.[189,232,1928,1954]" targetBox="[195,1391,1020,1890]" targetPageId="17">
<paragraph id="8BEF3693036EEF701E8978FBFCB76AA8" blockId="17.[189,1400,1928,2025]" pageId="17" pageNumber="85">
<emphasis id="B924EA81036EEF701E8978FBFEDD6AE3" bold="true" box="[189,284,1928,1954]" pageId="17" pageNumber="85">Fig. 10.</emphasis>
<taxonomicName id="4C504D10036EEF701F1278FBFDE76AE3" authority="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik, 2021" authorityName="Zamani &amp; Chatzaki &amp; Esyunin &amp; Marusik" authorityYear="2021" box="[294,550,1928,1954]" class="Arachnida" family="Gnaphosidae" genus="Gnaphosa" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="85" phylum="Arthropoda" rank="species" species="qamsarica" status="sp. nov.">
<emphasis id="B924EA81036EEF701F1278FBFDE76AE3" box="[294,550,1928,1954]" italics="true" pageId="17" pageNumber="85">Gnaphosa qamsarica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757FA036EEF701C1B78FAFD4C6AE2" box="[559,653,1929,1955]" pageId="17" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, ♀ (MHNG).
<emphasis id="B924EA81036EEF701D8478FBFBCE6AE3" bold="true" box="[944,1039,1928,1954]" pageId="17" pageNumber="85">A, C, E</emphasis>
. Epigyne, ventral view.
<emphasis id="B924EA81036EEF701B0378FBFF116A87" bold="true" pageId="17" pageNumber="85">B, D, F</emphasis>
. Epigyne, dorsal view.
<emphasis id="B924EA81036EEF701FDA78D8FDC26A84" bold="true" box="[494,515,1963,1989]" pageId="17" pageNumber="85">G</emphasis>
. Chelicerae, posterior view.
<emphasis id="B924EA81036EEF701D6B78DFFCB56A87" bold="true" box="[863,884,1964,1990]" pageId="17" pageNumber="85">H</emphasis>
. Habitus, dorsal view. Abbreviation:
<emphasis id="B924EA81036EEF701B0D78DFFA9E6A84" box="[1337,1375,1964,1989]" italics="true" pageId="17" pageNumber="85">Lm</emphasis>
= lateral margin. Scale bars: 0.1 mm, unless stated otherwise.
</paragraph>
</caption>
<paragraph id="8BEF3693036DEF731EDE7E7AFEEB6C62" blockId="18.[189,313,265,326]" box="[234,298,265,291]" pageId="18" pageNumber="86">
<emphasis id="B924EA81036DEF731EDE7E7AFEEB6C62" bold="true" box="[234,298,265,291]" pageId="18" pageNumber="86">Male</emphasis>
</paragraph>
<paragraph id="8BEF3693036DEF731E897E5FFEF86C07" blockId="18.[189,313,265,326]" box="[189,313,300,326]" pageId="18" pageNumber="86">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C34A6518036DEF731E897E1EFC1D6CF7" box="[189,988,365,438]" pageId="18" pageNumber="86" type="distribution">
<paragraph id="8BEF3693036DEF731E897E1EFE976CC6" blockId="18.[189,342,365,391]" box="[189,342,365,391]" pageId="18" pageNumber="86">
<heading id="D0A781FF036DEF731E897E1EFE976CC6" bold="true" box="[189,342,365,391]" fontSize="11" level="3" pageId="18" pageNumber="86" reason="3">
<emphasis id="B924EA81036DEF731E897E1EFE976CC6" bold="true" box="[189,342,365,391]" pageId="18" pageNumber="86">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF3693036DEF731E897EEFFC1D6CF7" blockId="18.[189,988,412,438]" box="[189,988,412,438]" pageId="18" pageNumber="86">
Known only from the
<typeStatus id="54EB8831036DEF731FF67EEFFE346CF7" box="[450,501,412,438]" pageId="18" pageNumber="86">type</typeStatus>
locality in
<collectingRegion id="4994F871036DEF731C4D7EEFFC836CF7" box="[633,834,412,438]" country="Iran" name="Esfahan" pageId="18" pageNumber="86">Isfahan Province</collectingRegion>
, central
<collectingCountry id="F3477603036DEF731D917EEFFC196CF7" box="[933,984,412,438]" name="Iran" pageId="18" pageNumber="86">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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