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<document id="1B46D3E3658B56A1E0B71615E56A7258" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625105984715" checkinUser="felipe" docAuthor="Shelley, Rowland M. &amp; Shear, William A." docDate="2005" docId="03F91146D877AB5BFEA1F92FC5CDE510" docLanguage="en" docName="zt01017p032.pdf" docOrigin="Zootaxa 1017 (1017)" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Stenozonium Shelley 1998" docType="treatment" docVersion="3" lastPageNumber="28" masterDocId="FFC0693ED875AB58FFA9FF9BC464E146" masterDocTitle="A new milliped of the genus Stenozonium Shelley 1998 from Washington State, U. S. A.: first record of the genus and family from North of the Columbia River (Polyzoniida: Polyzoniidae)" masterLastPageNumber="32" masterPageNumber="25" pageNumber="27" updateTime="1698980299450" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="71F5AAB54BBB20FFB7BBAAC5DEBFFCC2">A new milliped of the genus Stenozonium Shelley 1998 from Washington State, U. S. A.: first record of the genus and family from North of the Columbia River (Polyzoniida: Polyzoniidae)</mods:title>
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<paragraph id="8BEFA050D877AB5AFEA1F92FC6C0E788" blockId="2.[264,676,1716,1742]" box="[264,676,1716,1742]" pageId="2" pageNumber="27">
<heading id="D0A7173CD877AB5AFEA1F92FC6C0E788" bold="true" box="[264,676,1716,1742]" fontSize="11" level="1" pageId="2" pageNumber="27" reason="1">
<emphasis id="B9247C42D877AB5AFEA1F92FC6C0E788" bold="true" box="[264,676,1716,1742]" pageId="2" pageNumber="27">
Genus
<taxonomicName id="4C50DBD3D877AB5AFEF6F92FC6C0E788" ID-CoL="7Q2VT" authority="Shelley, 1998" authorityName="Shelley" authorityYear="1998" box="[351,676,1716,1742]" class="Diplopoda" family="Polyzoniidae" genus="Stenozonium" kingdom="Animalia" order="Polyzoniida" pageId="2" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B9247C42D877AB5AFEF6F92FC59FE788" bold="true" box="[351,507,1716,1742]" italics="true" pageId="2" pageNumber="27">Stenozonium</emphasis>
<bibRefCitation id="EFC1DDA1D877AB5AFDAAF92FC6C0E788" author="Shelley, R. M." box="[515,676,1716,1742]" pageId="2" pageNumber="27" pagination="3 - 34" refId="ref3108" refString="Shelley, R. M. (1998 [1997]) The milliped family Polyzoniidae in North America, with a classification of the global fauna (Diplopoda: Polyzoniida). Arthropoda Selecta, 6 (3 - 4), 3 - 34." type="journal article" year="1998">Shelley, 1998</bibRefCitation>
</taxonomicName>
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</paragraph>
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<paragraph id="8BEFA050D877AB5AFEA1F964C795E650" blockId="2.[264,1009,1791,1814]" box="[264,1009,1791,1814]" pageId="2" pageNumber="27">
<treatmentCitationGroup id="AB40877ED877AB5AFEA1F964C795E650" box="[264,1009,1791,1814]" pageId="2" pageNumber="27">
<treatmentCitation id="0AF18641D877AB5AFEA1F964C628E650" author="Shelley, R. M." box="[264,588,1791,1814]" page="12" pageId="2" pageNumber="27" year="1998">
<taxonomicName id="4C50DBD3D877AB5AFEA1F964C628E650" ID-CoL="7Q2VT" authority="Shelley, 1998: 12" authorityName="Shelley" authorityPageNumber="12" authorityYear="1998" box="[264,588,1791,1814]" class="Diplopoda" family="Polyzoniidae" genus="Stenozonium" kingdom="Animalia" order="Polyzoniida" pageId="2" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B9247C42D877AB5AFEA1F964C5EBE650" box="[264,399,1791,1814]" italics="true" pageId="2" pageNumber="27">Stenozonium</emphasis>
<bibRefCitation id="EFC1DDA1D877AB5AFE3FF964C628E650" author="Shelley, R. M." box="[406,588,1791,1814]" pageId="2" pageNumber="27" pagination="3 - 34" refId="ref3108" refString="Shelley, R. M. (1998 [1997]) The milliped family Polyzoniidae in North America, with a classification of the global fauna (Diplopoda: Polyzoniida). Arthropoda Selecta, 6 (3 - 4), 3 - 34." type="journal article" year="1998">Shelley, 1998:12</bibRefCitation>
</taxonomicName>
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.
<treatmentCitation id="0AF18641D877AB5AFDF7F964C741E650" author="Hoffman, R. L." box="[606,805,1791,1814]" page="32" pageId="2" pageNumber="27" year="1999">
<bibRefCitation id="EFC1DDA1D877AB5AFDF7F964C741E650" author="Hoffman, R. L." box="[606,805,1791,1814]" pageId="2" pageNumber="27" pagination="1 - 584" refId="ref2941" refString="Hoffman, R. L. (1999). Checklist of the millipeds of North and Middle America. Virginia Museum of Natural History Special Publication no. 8, 1 - 584." type="journal article" year="1999">Hoffman, 1999:32</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="0AF18641D877AB5AFC9FF964C789E650" author="Shelley, R. M." box="[822,1005,1791,1814]" page="92" pageId="2" pageNumber="27" year="2002">
<bibRefCitation id="EFC1DDA1D877AB5AFC9FF964C789E650" author="Shelley, R. M." box="[822,1005,1791,1814]" pageId="2" pageNumber="27" pagination="90 - 115" refId="ref3192" refString="Shelley, R. M. (2002) Annotated checklist of the millipeds of California (Arthropoda: Diplopoda). Monographs of the Western North American Naturalist, 1, 90 - 115." type="journal article" year="2002">Shelley, 2002:92</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<paragraph id="8BEFA050D877AB5AFEA1F8DCC02BE627" blockId="2.[264,1103,1863,1889]" box="[264,1103,1863,1889]" pageId="2" pageNumber="27">
<emphasis id="B9247C42D877AB5AFEA1F8DCC5FEE626" box="[264,410,1863,1888]" italics="true" pageId="2" pageNumber="27">
<typeStatus id="54EB1EF2D877AB5AFEA1F8DCC525E626" box="[264,321,1863,1888]" pageId="2" pageNumber="27">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="4C50DBD3D877AB5AFE18F8DCC749E627" authority="Shelley, 1998" authorityName="Shelley" authorityYear="1998" box="[433,813,1863,1889]" class="Diplopoda" family="Polyzoniidae" genus="Stenozonium" kingdom="Animalia" order="Polyzoniida" pageId="2" pageNumber="27" phylum="Arthropoda" rank="species" species="exile">
<emphasis id="B9247C42D877AB5AFE18F8DCC6E1E626" box="[433,645,1863,1888]" italics="true" pageId="2" pageNumber="27">Stenozonium exile</emphasis>
<bibRefCitation id="EFC1DDA1D877AB5AFD25F8DCC749E627" author="Shelley, R. M." box="[652,813,1863,1889]" pageId="2" pageNumber="27" pagination="3 - 34" refId="ref3108" refString="Shelley, R. M. (1998 [1997]) The milliped family Polyzoniidae in North America, with a classification of the global fauna (Diplopoda: Polyzoniida). Arthropoda Selecta, 6 (3 - 4), 3 - 34." type="journal article" year="1998">Shelley, 1998</bibRefCitation>
</taxonomicName>
, by original designation.
</paragraph>
<paragraph id="8BEFA050D876AB5BFE91FE87C758E070" blockId="3.[264,1324,284,1110]" box="[312,828,284,310]" pageId="3" pageNumber="28">
<emphasis id="B9247C42D876AB5BFE91FE87C5CBE073" box="[312,431,284,309]" italics="true" pageId="3" pageNumber="28">Diagnosis</emphasis>
. As presented by
<bibRefCitation id="EFC1DDA1D876AB5BFD21FE87C75DE070" author="Shelley, R. M." box="[648,825,284,310]" pageId="3" pageNumber="28" pagination="3 - 34" refId="ref3108" refString="Shelley, R. M. (1998 [1997]) The milliped family Polyzoniidae in North America, with a classification of the global fauna (Diplopoda: Polyzoniida). Arthropoda Selecta, 6 (3 - 4), 3 - 34." type="journal article" year="1998">Shelley (1998)</bibRefCitation>
.
</paragraph>
<paragraph id="8BEFA050D876AB5BFE91FEDFC690E0C0" blockId="3.[264,1324,284,1110]" pageId="3" pageNumber="28">
<emphasis id="B9247C42D876AB5BFE91FEDFC5AEE01B" box="[312,458,324,349]" italics="true" pageId="3" pageNumber="28">Components</emphasis>
. Three allopatric species are known; others may exist in moist coastal environments north of San Francisco Bay.
</paragraph>
<paragraph id="8BEFA050D876AB5BFE91FE0FC048E0B8" blockId="3.[264,1324,284,1110]" pageId="3" pageNumber="28">
<emphasis id="B9247C42D876AB5BFE91FE0FC5A3E0EB" box="[312,455,404,429]" italics="true" pageId="3" pageNumber="28">Distribution</emphasis>
. Occupying widely segregated areas in the Coast Range of northern
<collectingRegion id="49946EB2D876AB5BFEA1FE27C5E6E090" box="[264,386,444,470]" country="United States of America" name="California" pageId="3" pageNumber="28">California</collectingRegion>
, central
<collectingRegion id="49946EB2D876AB5BFE4CFE27C624E090" box="[485,576,444,470]" country="United States of America" name="Oregon" pageId="3" pageNumber="28">Oregon</collectingRegion>
, and the Olympic Peninsula of
<collectingRegion id="49946EB2D876AB5BFC19FE27C05BE090" box="[944,1087,444,470]" country="United States of America" name="Washington" pageId="3" pageNumber="28">Washington</collectingRegion>
; the
<collectingRegion id="49946EB2D876AB5BFBDFFE27C0AAE090" box="[1142,1230,444,470]" country="United States of America" name="Oregon" pageId="3" pageNumber="28">Oregon</collectingRegion>
species also ranges eastward onto the western slope of the Cascades (Fig. 2).
</paragraph>
<paragraph id="8BEFA050D876AB5BFE91FD97C5CDE510" blockId="3.[264,1324,284,1110]" pageId="3" pageNumber="28">
<emphasis id="B9247C42D876AB5BFE91FD97C5FAE363" box="[312,414,524,549]" italics="true" pageId="3" pageNumber="28">Remarks</emphasis>
. While additional species of
<taxonomicName id="4C50DBD3D876AB5BFCA2FD97C7C4E363" authorityName="Shelley" authorityYear="1998" box="[779,928,524,549]" class="Diplopoda" family="Polyzoniidae" genus="Stenozonium" kingdom="Animalia" order="Polyzoniida" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B9247C42D876AB5BFCA2FD97C7C4E363" box="[779,928,524,549]" italics="true" pageId="3" pageNumber="28">Stenozonium</emphasis>
</taxonomicName>
may yet be encountered and the ranges of the known ones will surely expand with additional samples, the lacunae between the three components will surely never be completely filled because of their sizes (180 &amp;
<quantity id="4CA80DB5D876AB5BFEA1FD1FC505E3D8" box="[264,353,644,670]" metricMagnitude="5" metricUnit="m" metricValue="3.7819584" pageId="3" pageNumber="28" unit="mi" value="235.0">235 mi</quantity>
(288 &amp;
<quantity id="4CA80DB5D876AB5BFE70FD1FC658E3D8" box="[473,572,644,670]" metricMagnitude="5" metricUnit="m" metricValue="3.76" pageId="3" pageNumber="28" unit="km" value="376.0">376 km</quantity>
)). We therefore suspect that
<taxonomicName id="4C50DBD3D876AB5BFC17FD1FC037E3DB" authorityName="Shelley" authorityYear="1998" box="[958,1107,644,669]" class="Diplopoda" family="Polyzoniidae" genus="Stenozonium" kingdom="Animalia" order="Polyzoniida" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B9247C42D876AB5BFC17FD1FC037E3DB" box="[958,1107,644,669]" italics="true" pageId="3" pageNumber="28">Stenozonium</emphasis>
</taxonomicName>
truly consists of allopatric species, in contrast to the continuous distributions of species and populations in the other Pacific polyzoniid genera, excepting the Monterey County,
<collectingRegion id="49946EB2D876AB5BFB07FD4FC14CE3A8" box="[1198,1320,724,750]" country="United States of America" name="California" pageId="3" pageNumber="28">California</collectingRegion>
, population of
<taxonomicName id="4C50DBD3D876AB5BFE06FD67C696E253" authorityName="Cook &amp; Loomis" authorityYear="1928" box="[431,754,764,789]" class="Diplopoda" family="Polyzoniidae" genus="Bdellozonium" kingdom="Animalia" order="Polyzoniida" pageId="3" pageNumber="28" phylum="Arthropoda" rank="species" species="cerviculatum">
<emphasis id="B9247C42D876AB5BFE06FD67C696E253" box="[431,754,764,789]" italics="true" pageId="3" pageNumber="28">Bdellozonium cerviculatum</emphasis>
</taxonomicName>
, and in contrast to the continuous pattern in the Pacific species of
<taxonomicName id="4C50DBD3D876AB5BFE4FFCBFC604E27B" authorityName="Wood" authorityYear="1864" box="[486,608,804,829]" class="Diplopoda" family="Hirudisomatidae" genus="Octoglena" kingdom="Animalia" order="Polyzoniida" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B9247C42D876AB5BFE4FFCBFC604E27B" box="[486,608,804,829]" italics="true" pageId="3" pageNumber="28">Octoglena</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFC1DDA1D876AB5BFDDAFCBFC770E278" author="Shelley, R. M." box="[627,788,804,830]" pageId="3" pageNumber="28" pagination="103 - 143" refId="ref3079" refString="Shelley, R. M. (1995) The milliped family Hirudisomatidae in the New World (Polyzoniida). Brimleyana, 23, 103 - 143." type="journal article" year="1995">Shelley 1995</bibRefCitation>
,
<bibRefCitation id="EFC1DDA1D876AB5BFC8DFCBFC707E278" author="Shelley, R. M." box="[804,867,804,830]" pageId="3" pageNumber="28" pagination="3 - 34" refId="ref3108" refString="Shelley, R. M. (1998 [1997]) The milliped family Polyzoniidae in North America, with a classification of the global fauna (Diplopoda: Polyzoniida). Arthropoda Selecta, 6 (3 - 4), 3 - 34." type="journal article" year="1998">1998</bibRefCitation>
). Allopatry patterns are indicators of age, evidence that a taxon has existed long enough for extinctions to generate anatomical and geographical discontinuities and isolate populations and species from each other. That
<taxonomicName id="4C50DBD3D876AB5BFEA1FC07C5F8E2F3" authorityName="Shelley" authorityYear="1998" box="[264,412,924,949]" class="Diplopoda" family="Polyzoniidae" genus="Stenozonium" kingdom="Animalia" order="Polyzoniida" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B9247C42D876AB5BFEA1FC07C5F8E2F3" box="[264,412,924,949]" italics="true" pageId="3" pageNumber="28">Stenozonium</emphasis>
</taxonomicName>
alone among west­Nearctic polyzoniidan genera exhibits allopatry suggests greater age and that it antedates the other genera. We cannot hypothesize time periods, but the geographic evidence suggests that
<taxonomicName id="4C50DBD3D876AB5BFD7BFC77C703E543" authorityName="Shelley" authorityYear="1998" box="[722,871,1004,1029]" class="Diplopoda" family="Polyzoniidae" genus="Stenozonium" kingdom="Animalia" order="Polyzoniida" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B9247C42D876AB5BFD7BFC77C703E543" box="[722,871,1004,1029]" italics="true" pageId="3" pageNumber="28">Stenozonium</emphasis>
</taxonomicName>
is a product of an early dichotomy in the
<taxonomicName id="4C50DBD3D876AB5BFE9CFB8FC5AAE568" authorityName="Newport" authorityYear="1844" box="[309,462,1044,1070]" class="Diplopoda" family="Polyzoniidae" kingdom="Animalia" order="Polyzoniida" pageId="3" pageNumber="28" phylum="Arthropoda" rank="family">Polyzoniidae</taxonomicName>
and that both
<taxonomicName id="4C50DBD3D876AB5BFDD7FB8FC77BE56B" authorityName="Cook &amp; Loomis" authorityYear="1928" box="[638,799,1044,1069]" class="Diplopoda" family="Polyzoniidae" genus="Bdellozonium" kingdom="Animalia" order="Polyzoniida" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B9247C42D876AB5BFDD7FB8FC77BE56B" box="[638,799,1044,1069]" italics="true" pageId="3" pageNumber="28">Bdellozonium</emphasis>
</taxonomicName>
and
<taxonomicName id="4C50DBD3D876AB5BFCF5FB8FC7B5E56B" authorityName="Cook &amp; Loomis" authorityYear="1928" box="[860,977,1044,1069]" class="Diplopoda" family="Polyzoniidae" genus="Buzonium" kingdom="Animalia" order="Polyzoniida" pageId="3" pageNumber="28" phylum="Arthropoda" rank="genus">
<emphasis id="B9247C42D876AB5BFCF5FB8FC7B5E56B" box="[860,977,1044,1069]" italics="true" pageId="3" pageNumber="28">Buzonium</emphasis>
</taxonomicName>
/
<taxonomicName id="4C50DBD3D876AB5BFC73FB8FC028E568" authorityName="Shelley" authorityYear="1998" box="[986,1100,1044,1070]" class="Diplopoda" family="Polyzoniidae" kingdom="Animalia" order="Polyzoniida" pageId="3" pageNumber="28" phylum="Arthropoda" rank="tribe" tribe="Buzoniini">Buzoniini</taxonomicName>
are comparatively &quot;young&quot; taxa.
</paragraph>
</subSubSection>
</treatment>
</document>
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