458 lines
56 KiB
XML
458 lines
56 KiB
XML
<document id="7C27636867ADF97E1C7D055B199652E8" ID-CLB-Dataset="157773" ID-DOI="10.11646/zootaxa.5194.1.2" ID-GBIF-Dataset="9c76b088-9845-48f6-9561-d4c6e91c4343" ID-ISSN="1175-5326" ID-Zenodo-Dep="7141848" ID-ZooBank="7C18727C-3AF0-4BE6-AFBC-EA1AC2F2B926" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1664862422389" checkinUser="plazi" docAuthor="Colloff, Matthew J." docDate="2022" docId="03EE7A4DC905FD09FF5EFA85FDCFFA8A" docLanguage="en" docName="zootaxa.5194.1.2.pdf" docOrigin="Zootaxa 5194 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tumerozetes roughleyi Colloff 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="40" masterDocId="FFD70235C901FD0EFFC9FFD6FF9AFFDF" masterDocTitle="First records of Tumerozetidae and Nodocepheidae from Australia, with descriptions of new taxa and a re-assessment of the Polypterozetoidea (Oribatida, Brachypylina)" masterLastPageNumber="57" masterPageNumber="33" pageNumber="37" updateTime="1699387054636" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="909F053FE40751909E43BD3DA6A6635B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="03E5ADC6852F173156BF8CAEFE632A41">
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<mods:title id="FE9D845F2B97940A8839C337C56986E1">First records of Tumerozetidae and Nodocepheidae from Australia, with descriptions of new taxa and a re-assessment of the Polypterozetoidea (Oribatida, Brachypylina)</mods:title>
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</mods:titleInfo>
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<mods:name id="6FB01C03685A833B7C78E366104A08D2" type="personal">
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<mods:role id="1DDC3CC674007585DBB050B4AE348549">
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<mods:roleTerm id="BA312856D8AF24605ABE323F0B9ACC44">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0CBF6681BFF517A8EC070F184C0C31D5">Colloff, Matthew J.</mods:namePart>
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<mods:typeOfResource id="7AF4F02404620CCBE8AEC6162EA11F17">text</mods:typeOfResource>
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<mods:titleInfo id="91646B1BCDA22031E4A76BD4C70EDF61">
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<mods:title id="4F6CE052C68A4CABBC057CCC4112DC82">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="D8F9DE9C3FADC06368E2FA031F60FF4B">
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<mods:date id="8A76FA6BFE327219E7D78304D9AF2E68">2022</mods:date>
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<mods:detail id="77844A86A4990554186C94EBD54AB43A" type="pubDate">
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<mods:number id="7479F2A78D5A1E9D7B18A3702952399C">2022-10-04</mods:number>
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</mods:detail>
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<mods:detail id="969ACC48F4F54598C58A7A45E22609B0" type="volume">
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<mods:number id="AADCDF4C6E053109C742D81295AAD368">5194</mods:number>
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</mods:detail>
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<mods:detail id="992E354A9A1DF44EB27C6181DE672CE3" type="issue">
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<mods:number id="A88853A2EA8C48D17FE4A00F5FA9D3E9">1</mods:number>
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<mods:extent id="1E4651B0BC591988CDA9BDA749140603" unit="page">
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<mods:start id="C19111BA7AFD89864CDA1338EC63520C">33</mods:start>
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<mods:end id="61D69C6C95C2EA228A286DC4DC3FF433">57</mods:end>
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<mods:classification id="9D8A000AFBC2784BA17022A1376624C6">journal article</mods:classification>
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<mods:identifier id="CCC45DA282E9440835B5A8EA3788C142" type="CLB-Dataset">157773</mods:identifier>
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<mods:identifier id="216FFFE116865835D9627351D79E5E81" type="DOI">10.11646/zootaxa.5194.1.2</mods:identifier>
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<mods:identifier id="C5087066CA3C13FDED0332D0699D2206" type="GBIF-Dataset">9c76b088-9845-48f6-9561-d4c6e91c4343</mods:identifier>
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<mods:identifier id="FA4281A7F665AAAADCC580013FBCB7B4" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0EF72CC8533CE9C88F7725F721C7587B" type="Zenodo-Dep">7141848</mods:identifier>
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<mods:identifier id="3941F3FAA05CB8ABCBD2EEF7231E1DBD" type="ZooBank">7C18727C-3AF0-4BE6-AFBC-EA1AC2F2B926</mods:identifier>
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</mods:mods>
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<treatment id="03EE7A4DC905FD09FF5EFA85FDCFFA8A" ID-DOI="http://doi.org/10.5281/zenodo.7141870" ID-GBIF-Taxon="202134816" ID-Zenodo-Dep="7141870" LSID="urn:lsid:plazi:treatment:03EE7A4DC905FD09FF5EFA85FDCFFA8A" httpUri="http://treatment.plazi.org/id/03EE7A4DC905FD09FF5EFA85FDCFFA8A" lastPageId="7" lastPageNumber="40" pageId="4" pageNumber="37">
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<subSubSection id="C35D98D0C905FD0AFF5EFA85FD95FAB2" box="[151,527,1362,1389]" pageId="4" pageNumber="37" type="nomenclature">
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<paragraph id="8BF8CB5BC905FD0AFF5EFA85FD95FAB2" blockId="4.[151,527,1362,1425]" box="[151,527,1362,1389]" pageId="4" pageNumber="37">
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<heading id="D0B07C37C905FD0AFF5EFA85FD95FAB2" bold="true" box="[151,527,1362,1389]" fontSize="11" level="1" pageId="4" pageNumber="37" reason="1">
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<emphasis id="B9331749C905FD0AFF5EFA85FD95FAB2" bold="true" box="[151,527,1362,1389]" pageId="4" pageNumber="37">
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<taxonomicName id="4C47B0D8C905FD0AFF5EFA85FE31FAB2" authority="Colloff, 2022" authorityName="Colloff" authorityYear="2022" box="[151,427,1362,1389]" class="Arachnida" family="Tumerozetidae" genus="Tumerozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="37" phylum="Arthropoda" rank="species" species="roughleyi" status="sp. nov.">
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<emphasis id="B9331749C905FD0AFF5EFA85FE31FAB2" bold="true" box="[151,427,1362,1389]" italics="true" pageId="4" pageNumber="37">Tumerozetes roughleyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A200AA32C905FD0AFE7BFA85FD95FAB2" box="[434,527,1363,1389]" pageId="4" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35D98D0C905FD09FF5EFAA1FD7DFE9E" lastPageId="7" lastPageNumber="40" pageId="4" pageNumber="37" type="description">
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<paragraph id="8BF8CB5BC905FD0AFF5EFAA1FEB5FA4E" blockId="4.[151,527,1362,1425]" box="[151,303,1399,1425]" pageId="4" pageNumber="37">
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||
(
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<figureCitation id="137CD7DEC905FD0AFF56FAA1FE90FA4E" box="[159,266,1399,1425]" captionStart="FIGURE 1" captionStartId="5.[152,255,1981,2005]" captionTargetBox="[253,1331,720,1949]" captionTargetId="figure-256@5.[247,1341,702,1955]" captionTargetPageId="5" captionText="FIGURE 1. Tumerozetes roughleyi sp. nov. (a) dorsal aspect; (b) ventral aspect); (c) lateral aspect; (d) detail of seta le. Scale bar = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.7141852" httpUri="https://zenodo.org/record/7141852/files/figure.png" pageId="4" pageNumber="37">Figures 1</figureCitation>
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,
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<figureCitation id="137CD7DEC905FD0AFED1FAA1FEBDFA4E" box="[280,295,1399,1425]" captionStart="FIGURE 2" captionStartId="6.[152,255,1565,1589]" captionTargetBox="[290,1300,190,1533]" captionTargetId="figure-22@6.[283,1305,181,1541]" captionTargetPageId="6" captionText="FIGURE 2. Tumerozetes roughleyi sp. nov. (a) anterior aspect; (b) bothridia and interlamellar plates; (c) waxy tegument on rostrum; (d) chelicera; (e) palp; (f) rutellum; (g) anal and genital plates; (h) bothridium. Scale bars: a–c, g, h = 50 μm; e, f = 15 μm" figureDoi="http://doi.org/10.5281/zenodo.7141854" httpUri="https://zenodo.org/record/7141854/files/figure.png" pageId="4" pageNumber="37">2</figureCitation>
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)
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</paragraph>
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<paragraph id="8BF8CB5BC905FD0AFF5EFA68FAF6F9FE" blockId="4.[151,1437,1470,2037]" pageId="4" pageNumber="37">
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<emphasis id="B9331749C905FD0AFF5EFA68FEBBFA07" bold="true" box="[151,289,1470,1496]" pageId="4" pageNumber="37">Dimensions</emphasis>
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.
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<typeStatus id="54FC75F9C905FD0AFEE6FA69FE0DFA06" box="[303,407,1471,1497]" pageId="4" pageNumber="37" type="holotype">Holotype</typeStatus>
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female length 309 μm, breadth 205 μm.
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<typeStatus id="54FC75F9C905FD0AFC97FA69FC5AFA06" box="[862,960,1471,1497]" pageId="4" pageNumber="37" type="paratype">Paratype</typeStatus>
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males: mean length (
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<emphasis id="B9331749C905FD0AFB79FA69FB24FA07" box="[1200,1214,1471,1496]" italics="true" pageId="4" pageNumber="37">n</emphasis>
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= 4) 256 μm (range 239–283 μm); mean breadth 165 μm (range 154–183 μm).
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<typeStatus id="54FC75F9C905FD0AFCFBFA35FC0EFA22" box="[818,916,1507,1533]" pageId="4" pageNumber="37" type="paratype">Paratype</typeStatus>
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females: mean length (
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<emphasis id="B9331749C905FD0AFB54FA35FB31FA23" box="[1181,1195,1507,1532]" italics="true" pageId="4" pageNumber="37">n</emphasis>
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= 10) 307 μm (range 296–321 μm); mean breadth 200 μm (range 194–211 μm). Ratio of prodorsum to total length: 0.47 (
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<typeStatus id="54FC75F9C905FD0AFB31F9D1FAFBF9FE" box="[1272,1377,1543,1569]" pageId="4" pageNumber="37" type="holotype">holotype</typeStatus>
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).
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</paragraph>
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<paragraph id="8BF8CB5BC905FD0AFF0EF9FCFAC6F82A" blockId="4.[151,1437,1470,2037]" pageId="4" pageNumber="37">
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<emphasis id="B9331749C905FD0AFF0EF9FCFE89F99B" bold="true" box="[199,275,1578,1604]" pageId="4" pageNumber="37">Adult.</emphasis>
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<emphasis id="B9331749C905FD0AFED3F9FDFE0DF99A" box="[282,407,1579,1605]" italics="true" pageId="4" pageNumber="37">Prodorsum</emphasis>
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: rostrum very broad, rounded, with margin covered by blocks of cerotegument (
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<figureCitation id="137CD7DEC905FD0AFAD6F9FDFA15F99A" box="[1311,1423,1579,1605]" captionStart="FIGURE 2" captionStartId="6.[152,255,1565,1589]" captionTargetBox="[290,1300,190,1533]" captionTargetId="figure-22@6.[283,1305,181,1541]" captionTargetPageId="6" captionText="FIGURE 2. Tumerozetes roughleyi sp. nov. (a) anterior aspect; (b) bothridia and interlamellar plates; (c) waxy tegument on rostrum; (d) chelicera; (e) palp; (f) rutellum; (g) anal and genital plates; (h) bothridium. Scale bars: a–c, g, h = 50 μm; e, f = 15 μm" figureDoi="http://doi.org/10.5281/zenodo.7141854" httpUri="https://zenodo.org/record/7141854/files/figure.png" pageId="4" pageNumber="37">Figure 2c</figureCitation>
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). Rostral setae (
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<emphasis id="B9331749C905FD0AFEF0F999FECBF9B7" box="[313,337,1615,1640]" italics="true" pageId="4" pageNumber="37">ro</emphasis>
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) short, smooth; barely visible in dorsal view. Lamellae (
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<emphasis id="B9331749C905FD0AFC18F999FC7DF9B6" box="[977,999,1615,1641]" italics="true" pageId="4" pageNumber="37">la</emphasis>
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) long, thin, narrow; lamellar setae (
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<emphasis id="B9331749C905FD0AFAB6F999FA09F9B6" box="[1407,1427,1615,1641]" italics="true" pageId="4" pageNumber="37">le</emphasis>
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) well-developed, barbed, on parallel lamellar cusps extending anteriorly as far as apex of rostrum (
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<figureCitation id="137CD7DEC905FD0AFB08F9A5FAB6F952" box="[1217,1324,1651,1677]" captionStart="FIGURE 1" captionStartId="5.[152,255,1981,2005]" captionTargetBox="[253,1331,720,1949]" captionTargetId="figure-256@5.[247,1341,702,1955]" captionTargetPageId="5" captionText="FIGURE 1. Tumerozetes roughleyi sp. nov. (a) dorsal aspect; (b) ventral aspect); (c) lateral aspect; (d) detail of seta le. Scale bar = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.7141852" httpUri="https://zenodo.org/record/7141852/files/figure.png" pageId="4" pageNumber="37">Figure 1a</figureCitation>
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). Lamella (
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<emphasis id="B9331749C905FD0AFF68F941FF2DF96E" box="[161,183,1687,1713]" italics="true" pageId="4" pageNumber="37">la</emphasis>
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) connecting anteriorly with tutorium (
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<emphasis id="B9331749C905FD0AFDA6F941FD1FF96F" box="[623,645,1687,1712]" italics="true" pageId="4" pageNumber="37">tu</emphasis>
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) in anterior view (
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<figureCitation id="137CD7DEC905FD0AFC95F941FC57F96E" box="[860,973,1687,1713]" captionStart="FIGURE 2" captionStartId="6.[152,255,1565,1589]" captionTargetBox="[290,1300,190,1533]" captionTargetId="figure-22@6.[283,1305,181,1541]" captionTargetPageId="6" captionText="FIGURE 2. Tumerozetes roughleyi sp. nov. (a) anterior aspect; (b) bothridia and interlamellar plates; (c) waxy tegument on rostrum; (d) chelicera; (e) palp; (f) rutellum; (g) anal and genital plates; (h) bothridium. Scale bars: a–c, g, h = 50 μm; e, f = 15 μm" figureDoi="http://doi.org/10.5281/zenodo.7141854" httpUri="https://zenodo.org/record/7141854/files/figure.png" pageId="4" pageNumber="37">Figure 2a</figureCitation>
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). Prodorsum inflated, with broad, lateral ridge-like interlamellar plates (
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<emphasis id="B9331749C905FD0AFE32F96DFD83F90A" box="[507,537,1723,1749]" italics="true" pageId="4" pageNumber="37">ilp</emphasis>
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) extending from base of lamellae almost to apex of rostrum, bearing smooth, curved interlamellar setae (
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<emphasis id="B9331749C905FD0AFE1DF909FE70F927" box="[468,490,1759,1784]" italics="true" pageId="4" pageNumber="37">in</emphasis>
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). Setae
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<emphasis id="B9331749C905FD0AFD84F909FDF9F927" box="[589,611,1759,1784]" italics="true" pageId="4" pageNumber="37">in</emphasis>
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projecting vertically from interlamellar plates in anterior view (
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<figureCitation id="137CD7DEC905FD0AFA98F909FF2FF8C2" captionStart="FIGURE 2" captionStartId="6.[152,255,1565,1589]" captionTargetBox="[290,1300,190,1533]" captionTargetId="figure-22@6.[283,1305,181,1541]" captionTargetPageId="6" captionText="FIGURE 2. Tumerozetes roughleyi sp. nov. (a) anterior aspect; (b) bothridia and interlamellar plates; (c) waxy tegument on rostrum; (d) chelicera; (e) palp; (f) rutellum; (g) anal and genital plates; (h) bothridium. Scale bars: a–c, g, h = 50 μm; e, f = 15 μm" figureDoi="http://doi.org/10.5281/zenodo.7141854" httpUri="https://zenodo.org/record/7141854/files/figure.png" pageId="4" pageNumber="37">Figure 2a</figureCitation>
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). Interlamellar plates sub-rectangular, slightly longer than broad, fused posteriomedially (
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<figureCitation id="137CD7DEC905FD0AFB63F8D5FA86F8C2" box="[1194,1308,1795,1821]" captionStart="FIGURE 2" captionStartId="6.[152,255,1565,1589]" captionTargetBox="[290,1300,190,1533]" captionTargetId="figure-22@6.[283,1305,181,1541]" captionTargetPageId="6" captionText="FIGURE 2. Tumerozetes roughleyi sp. nov. (a) anterior aspect; (b) bothridia and interlamellar plates; (c) waxy tegument on rostrum; (d) chelicera; (e) palp; (f) rutellum; (g) anal and genital plates; (h) bothridium. Scale bars: a–c, g, h = 50 μm; e, f = 15 μm" figureDoi="http://doi.org/10.5281/zenodo.7141854" httpUri="https://zenodo.org/record/7141854/files/figure.png" pageId="4" pageNumber="37">Figure 2b</figureCitation>
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). Apices of interlamellar plates not fused anteriorly in dorsal orientation. Heart-shaped anteriomedian projection of notogastral plate (
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<emphasis id="B9331749C905FD0AFF2BF89DFF64F8BB" box="[226,254,1867,1892]" italics="true" pageId="4" pageNumber="37">ap</emphasis>
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) with fine alveolate ornamentation and median ridges covering central part of prodorsum, positioned below and not fused with interlamellar plates, extending from anterior of
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<emphasis id="B9331749C905FD0AFC76F8B9FC4FF857" box="[959,981,1903,1928]" italics="true" pageId="4" pageNumber="37">in</emphasis>
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, converging toward dorsosejugal furrow. Anteriomedian projection of notogastral plate strongly convex in lateral view. Bothridia prominent, projecting laterally, with well-developed rim and internal circular ridges (
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<figureCitation id="137CD7DEC905FD0AFC92F861FC57F80E" box="[859,973,1975,2001]" captionStart="FIGURE 2" captionStartId="6.[152,255,1565,1589]" captionTargetBox="[290,1300,190,1533]" captionTargetId="figure-22@6.[283,1305,181,1541]" captionTargetPageId="6" captionText="FIGURE 2. Tumerozetes roughleyi sp. nov. (a) anterior aspect; (b) bothridia and interlamellar plates; (c) waxy tegument on rostrum; (d) chelicera; (e) palp; (f) rutellum; (g) anal and genital plates; (h) bothridium. Scale bars: a–c, g, h = 50 μm; e, f = 15 μm" figureDoi="http://doi.org/10.5281/zenodo.7141854" httpUri="https://zenodo.org/record/7141854/files/figure.png" pageId="4" pageNumber="37">Figure 2h</figureCitation>
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). Bothridial seta long (65 μm;
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<typeStatus id="54FC75F9C905FD0AFAE3F861FA0AF80E" box="[1322,1424,1975,2001]" pageId="4" pageNumber="37" type="holotype">holotype</typeStatus>
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), setiform, slightly inflated apically and with fine, long spinules. Short, smooth exobothridial seta (
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<emphasis id="B9331749C905FD0AFB13F80DFB69F82B" box="[1242,1267,2011,2036]" italics="true" pageId="4" pageNumber="37">ex</emphasis>
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) present.
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</paragraph>
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<paragraph id="8BF8CB5BC904FD0BFF0EFF41FC3DFE56" blockId="5.[151,1437,151,681]" pageId="5" pageNumber="38">
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<emphasis id="B9331749C904FD0BFF0EFF41FED9FF6F" box="[199,323,151,176]" italics="true" pageId="5" pageNumber="38">Notogaster</emphasis>
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: margins of relatively small humeral processes (h) convex laterally, curving ventrally and with median protuberance; humeral processes not extending as far as bothridia (
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<figureCitation id="137CD7DEC904FD0BFC01FF6DFBAEFF0A" box="[968,1076,187,213]" captionStart="FIGURE 1" captionStartId="5.[152,255,1981,2005]" captionTargetBox="[253,1331,720,1949]" captionTargetId="figure-256@5.[247,1341,702,1955]" captionTargetPageId="5" captionText="FIGURE 1. Tumerozetes roughleyi sp. nov. (a) dorsal aspect; (b) ventral aspect); (c) lateral aspect; (d) detail of seta le. Scale bar = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.7141852" httpUri="https://zenodo.org/record/7141852/files/figure.png" pageId="5" pageNumber="38">Figure 1a</figureCitation>
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). Seta
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<emphasis id="B9331749C904FD0BFBB4FF6DFB13FF0B" box="[1149,1161,187,212]" italics="true" pageId="5" pageNumber="38">c</emphasis>
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<subScript id="17C3C91EC904FD0BFB43FF1BFB08FF04" attach="left" box="[1162,1170,205,219]" fontSize="6" pageId="5" pageNumber="38">2</subScript>
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positioned immediately posterior of humeral process on small, rounded projection. Dorsosejugal furrow (
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<emphasis id="B9331749C904FD0BFBDAFF09FBB6FF26" box="[1043,1068,223,249]" italics="true" pageId="5" pageNumber="38">ds</emphasis>
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) straight, transverse, with paired, curved, longitudinal median notogastral ridges (
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<emphasis id="B9331749C904FD0BFD7CFED5FD78FEC3" box="[693,738,259,284]" italics="true" pageId="5" pageNumber="38">mnr</emphasis>
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) forming ovoid shape, extending posteriorly as far as level of lyrifissure
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<emphasis id="B9331749C904FD0BFEDAFEF1FEADFE9E" box="[275,311,295,321]" italics="true" pageId="5" pageNumber="38">iad</emphasis>
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, fused anteriorly, connecting at dorsosejugal furrow with anteriomedian projection of notogastral plate. Notogaster smooth, broader (215 μm;
|
||
<typeStatus id="54FC75F9C904FD0BFD4FFE9DFD70FEBA" box="[646,746,331,357]" pageId="5" pageNumber="38" type="holotype">holotype</typeStatus>
|
||
) than long, with 9 pairs of marginal setae (
|
||
<emphasis id="B9331749C904FD0BFB01FE9DFB4EFEBB" box="[1224,1236,331,356]" italics="true" pageId="5" pageNumber="38">c</emphasis>
|
||
<subScript id="17C3C91EC904FD0BFB1DFE8BFB46FEB4" attach="left" box="[1236,1244,349,363]" fontSize="6" pageId="5" pageNumber="38">1</subScript>
|
||
plus
|
||
<emphasis id="B9331749C904FD0BFAD1FE9DFAB4FEBA" box="[1304,1326,331,357]" italics="true" pageId="5" pageNumber="38">la</emphasis>
|
||
,
|
||
<emphasis id="B9331749C904FD0BFAF5FE9DFAC8FEBA" box="[1340,1362,331,357]" italics="true" pageId="5" pageNumber="38">lp</emphasis>
|
||
,
|
||
<emphasis id="B9331749C904FD0BFA96FE9DFAF7FEBA" box="[1375,1389,331,357]" italics="true" pageId="5" pageNumber="38">h</emphasis>
|
||
and
|
||
<emphasis id="B9331749C904FD0BFF5EFEB9FF3FFE57" box="[151,165,367,392]" italics="true" pageId="5" pageNumber="38">p</emphasis>
|
||
series);
|
||
<emphasis id="B9331749C904FD0BFECDFEB9FE80FE56" box="[260,282,367,393]" italics="true" pageId="5" pageNumber="38">la</emphasis>
|
||
,
|
||
<emphasis id="B9331749C904FD0BFEE1FEB9FEA4FE56" box="[296,318,367,393]" italics="true" pageId="5" pageNumber="38">lp</emphasis>
|
||
and
|
||
<emphasis id="B9331749C904FD0BFEBDFEB9FE18FE56" box="[372,386,367,393]" italics="true" pageId="5" pageNumber="38">h</emphasis>
|
||
<subScript id="17C3C91EC904FD0BFE4BFE57FE10FE50" attach="left" box="[386,394,385,399]" fontSize="6" pageId="5" pageNumber="38">3</subScript>
|
||
with fine spines basally. Lyrifissurae
|
||
<emphasis id="B9331749C904FD0BFCFBFEB9FCD2FE57" box="[818,840,367,392]" italics="true" pageId="5" pageNumber="38">ia</emphasis>
|
||
present.
|
||
</paragraph>
|
||
<paragraph id="8BF8CB5BC904FD0BFF0EFE45FE2EFD76" blockId="5.[151,1437,151,681]" pageId="5" pageNumber="38">
|
||
<emphasis id="B9331749C904FD0BFF0EFE45FEF2FE73" box="[199,360,403,429]" italics="true" pageId="5" pageNumber="38">Ventral aspect</emphasis>
|
||
: subcapitulum rounded apically, with tectum derived from genae, bearing a median infrabuccal cleft (
|
||
<emphasis id="B9331749C904FD0BFF13FE61FF66FE0E" box="[218,252,439,465]" italics="true" pageId="5" pageNumber="38">ibc</emphasis>
|
||
) on its ventral surface (
|
||
<figureCitation id="137CD7DEC904FD0BFDC3FE61FDE0FE0E" box="[522,634,439,465]" captionStart="FIGURE 7" captionStartId="17.[152,255,1109,1133]" captionTargetBox="[441,1148,184,1079]" captionTargetId="figure-15@17.[435,1152,181,1085]" captionTargetPageId="17" captionText="FIGURE 7. Morphology of the subcapitulum of polypterozetoid taxa included in the character analysis. (a) Polypterozetes cherubin Berlese 1916; (b) Podopterotegaeus altimonticola Piffl, 1972; (c) Nemacepheus dentatusAoki, 1968; (d) Nodocepheus laterodentatus Piffl, 1972; (e) Tumerozetes roughleyi sp. nov.; (f) Sacculella yarra gen. et sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7141872" httpUri="https://zenodo.org/record/7141872/files/figure.png" pageId="5" pageNumber="38">Figure 7e</figureCitation>
|
||
); setae
|
||
<emphasis id="B9331749C904FD0BFD19FE61FD44FE0F" box="[720,734,439,464]" italics="true" pageId="5" pageNumber="38">a</emphasis>
|
||
and
|
||
<emphasis id="B9331749C904FD0BFCDFFE61FCB0FE0F" box="[790,810,439,464]" italics="true" pageId="5" pageNumber="38">m</emphasis>
|
||
with bases not hidden by tectum in ventral orientation; setae
|
||
<emphasis id="B9331749C904FD0BFF1DFE0DFF78FE2A" box="[212,226,475,501]" italics="true" pageId="5" pageNumber="38">h</emphasis>
|
||
positioned posteriomedially on subcapitulum. Labiogenal articulation absent. Rutellum Y-shaped (
|
||
<figureCitation id="137CD7DEC904FD0BFAEDFE0DFA0AFE2A" box="[1316,1424,475,501]" captionStart="FIGURE 2" captionStartId="6.[152,255,1565,1589]" captionTargetBox="[290,1300,190,1533]" captionTargetId="figure-22@6.[283,1305,181,1541]" captionTargetPageId="6" captionText="FIGURE 2. Tumerozetes roughleyi sp. nov. (a) anterior aspect; (b) bothridia and interlamellar plates; (c) waxy tegument on rostrum; (d) chelicera; (e) palp; (f) rutellum; (g) anal and genital plates; (h) bothridium. Scale bars: a–c, g, h = 50 μm; e, f = 15 μm" figureDoi="http://doi.org/10.5281/zenodo.7141854" httpUri="https://zenodo.org/record/7141854/files/figure.png" pageId="5" pageNumber="38">Figure 2f</figureCitation>
|
||
). Chelicerae short, with digitus fixus reduced to flat, smooth, lobe-like structure; digitus mobilis curved, projecting beyond digitus fixus, bearing a comb-like fringe of several thin, straight teeth (
|
||
<figureCitation id="137CD7DEC904FD0BFBDBFDF5FB1EFDE2" box="[1042,1156,547,573]" captionStart="FIGURE 2" captionStartId="6.[152,255,1565,1589]" captionTargetBox="[290,1300,190,1533]" captionTargetId="figure-22@6.[283,1305,181,1541]" captionTargetPageId="6" captionText="FIGURE 2. Tumerozetes roughleyi sp. nov. (a) anterior aspect; (b) bothridia and interlamellar plates; (c) waxy tegument on rostrum; (d) chelicera; (e) palp; (f) rutellum; (g) anal and genital plates; (h) bothridium. Scale bars: a–c, g, h = 50 μm; e, f = 15 μm" figureDoi="http://doi.org/10.5281/zenodo.7141854" httpUri="https://zenodo.org/record/7141854/files/figure.png" pageId="5" pageNumber="38">Figure 2d</figureCitation>
|
||
). Palp setal formula 0-2- 1-3-9; setae on palpfemur, genu and seta
|
||
<emphasis id="B9331749C904FD0BFDB8FD91FDE5FDBE" box="[625,639,583,609]" italics="true" pageId="5" pageNumber="38">d</emphasis>
|
||
on tibia with sparingly spinose ornamentation, others smooth (
|
||
<figureCitation id="137CD7DEC904FD0BFA98FD91FF2FFD5A" captionStart="FIGURE 2" captionStartId="6.[152,255,1565,1589]" captionTargetBox="[290,1300,190,1533]" captionTargetId="figure-22@6.[283,1305,181,1541]" captionTargetPageId="6" captionText="FIGURE 2. Tumerozetes roughleyi sp. nov. (a) anterior aspect; (b) bothridia and interlamellar plates; (c) waxy tegument on rostrum; (d) chelicera; (e) palp; (f) rutellum; (g) anal and genital plates; (h) bothridium. Scale bars: a–c, g, h = 50 μm; e, f = 15 μm" figureDoi="http://doi.org/10.5281/zenodo.7141854" httpUri="https://zenodo.org/record/7141854/files/figure.png" pageId="5" pageNumber="38">Figure 2e</figureCitation>
|
||
). Eupathidia not on prominent tubercles; eupathidium
|
||
<emphasis id="B9331749C904FD0BFCE5FDBDFCC1FD5B" box="[812,859,619,644]" italics="true" pageId="5" pageNumber="38">acm</emphasis>
|
||
not fused with solendidum ω which is positioned posteriorly on palptarsus.
|
||
</paragraph>
|
||
<caption id="DF389BD3C904FD0BFF51F86BFEBAF825" ID-DOI="http://doi.org/10.5281/zenodo.7141852" ID-Zenodo-Dep="7141852" httpUri="https://zenodo.org/record/7141852/files/figure.png" pageId="5" pageNumber="38" startId="5.[152,255,1981,2005]" targetBox="[253,1331,720,1949]" targetPageId="5">
|
||
<paragraph id="8BF8CB5BC904FD0BFF51F86BFEBAF825" blockId="5.[151,1436,1981,2042]" pageId="5" pageNumber="38">
|
||
<emphasis id="B9331749C904FD0BFF51F86BFE83F80A" bold="true" box="[152,281,1981,2005]" pageId="5" pageNumber="38">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C47B0D8C904FD0BFEE9F868FD9EF80A" authority="Colloff, 2022" authorityName="Colloff" authorityYear="2022" box="[288,516,1981,2005]" class="Arachnida" family="Tumerozetidae" genus="Tumerozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="38" phylum="Arthropoda" rank="species" species="roughleyi" status="sp. nov.">
|
||
<emphasis id="B9331749C904FD0BFEE9F868FD9EF80A" box="[288,516,1981,2005]" italics="true" pageId="5" pageNumber="38">Tumerozetes roughleyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A200AA32C904FD0BFDC2F86BFDC4F80A" box="[523,606,1981,2005]" pageId="5" pageNumber="38" rank="species">
|
||
<emphasis id="B9331749C904FD0BFDC2F86BFDC2F80A" bold="true" box="[523,600,1981,2005]" pageId="5" pageNumber="38">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(a) dorsal aspect; (b) ventral aspect); (c) lateral aspect; (d) detail of seta
|
||
<emphasis id="B9331749C904FD0BFA8FF86BFAC2F80A" box="[1350,1368,1981,2005]" italics="true" pageId="5" pageNumber="38">le</emphasis>
|
||
. Scale bar = 100 μm
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF389BD3C907FD08FF51F9CBFA06F985" ID-DOI="http://doi.org/10.5281/zenodo.7141854" ID-Zenodo-Dep="7141854" httpUri="https://zenodo.org/record/7141854/files/figure.png" pageId="6" pageNumber="39" startId="6.[152,255,1565,1589]" targetBox="[290,1300,190,1533]" targetPageId="6">
|
||
<paragraph id="8BF8CB5BC907FD08FF51F9CBFA06F985" blockId="6.[151,1436,1565,1626]" pageId="6" pageNumber="39">
|
||
<emphasis id="B9331749C907FD08FF51F9CBFE80F9EA" bold="true" box="[152,282,1565,1589]" pageId="6" pageNumber="39">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C47B0D8C907FD08FEEBF9C8FD9DF9EA" authority="Colloff, 2022" authorityName="Colloff" authorityYear="2022" box="[290,519,1565,1589]" class="Arachnida" family="Tumerozetidae" genus="Tumerozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="roughleyi" status="sp. nov.">
|
||
<emphasis id="B9331749C907FD08FEEBF9C8FD9DF9EA" box="[290,519,1565,1589]" italics="true" pageId="6" pageNumber="39">Tumerozetes roughleyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9331749C907FD08FDC6F9CBFDFBF9EA" bold="true" box="[527,609,1565,1589]" pageId="6" pageNumber="39">
|
||
<taxonomicNameLabel id="A200AA32C907FD08FDC6F9CBFDFBF9EA" box="[527,609,1565,1589]" pageId="6" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(a) anterior aspect; (b) bothridia and interlamellar plates; (c) waxy tegument on rostrum; (d) chelicera; (e) palp; (f) rutellum; (g) anal and genital plates; (h) bothridium. Scale bars: a–c, g, h = 50 μm; e, f = 15 μm
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF8CB5BC907FD08FF0EF97DFE2EF81E" blockId="6.[151,1437,1707,2022]" pageId="6" pageNumber="39">
|
||
Epimeral plates discrete, all broadly separated in midline, sub-rectangular (I-II) or sub-triangular (III-IV); epimeral setation 3-1-3-3 (
|
||
<figureCitation id="137CD7DEC907FD08FE0FF919FDADF936" box="[454,567,1743,1769]" captionStart="FIGURE 1" captionStartId="5.[152,255,1981,2005]" captionTargetBox="[253,1331,720,1949]" captionTargetId="figure-256@5.[247,1341,702,1955]" captionTargetPageId="5" captionText="FIGURE 1. Tumerozetes roughleyi sp. nov. (a) dorsal aspect; (b) ventral aspect); (c) lateral aspect; (d) detail of seta le. Scale bar = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.7141852" httpUri="https://zenodo.org/record/7141852/files/figure.png" pageId="6" pageNumber="39">Figure 1b</figureCitation>
|
||
). Pedotectum I (pd I) and II (pd II) and patronium (
|
||
<emphasis id="B9331749C907FD08FB46F906FB31F936" box="[1167,1195,1744,1769]" italics="true" pageId="6" pageNumber="39">pa</emphasis>
|
||
) strongly developed; humeral processes appearing lobe-like in ventral orientation; discidium (
|
||
<emphasis id="B9331749C907FD08FC2AF925FC63F8D2" box="[995,1017,1779,1805]" italics="true" pageId="6" pageNumber="39">di</emphasis>
|
||
) with prominent, sharp point; welldeveloped circumpedal carina (
|
||
<emphasis id="B9331749C907FD08FE34F8CEFD8CF8EE" box="[509,534,1816,1841]" italics="true" pageId="6" pageNumber="39">cc</emphasis>
|
||
) present (
|
||
<figureCitation id="137CD7DEC907FD08FD40F8C1FD60F8EE" box="[649,762,1815,1841]" captionStart="FIGURE 1" captionStartId="5.[152,255,1981,2005]" captionTargetBox="[253,1331,720,1949]" captionTargetId="figure-256@5.[247,1341,702,1955]" captionTargetPageId="5" captionText="FIGURE 1. Tumerozetes roughleyi sp. nov. (a) dorsal aspect; (b) ventral aspect); (c) lateral aspect; (d) detail of seta le. Scale bar = 100 μm" figureDoi="http://doi.org/10.5281/zenodo.7141852" httpUri="https://zenodo.org/record/7141852/files/figure.png" pageId="6" pageNumber="39">Figure 1b</figureCitation>
|
||
). Ventral plate sub-circular, broader than long. Genital and anal plates each surrounded by ring of sclerotized cuticle; genital plates 46 µm long (
|
||
<typeStatus id="54FC75F9C907FD08FBF7F8EDFB3CF88A" box="[1086,1190,1851,1877]" pageId="6" pageNumber="39" type="holotype">holotype</typeStatus>
|
||
), with 5 pairs of setae; 1 pair of aggenital setae, 2 pairs of adanal setae and 2 pairs of anal setae; lyrifissure
|
||
<emphasis id="B9331749C907FD08FBFFF889FBC0F8A6" box="[1078,1114,1887,1913]" italics="true" pageId="6" pageNumber="39">iad</emphasis>
|
||
in para-anal position (
|
||
<figureCitation id="137CD7DEC907FD08FA98F889FF2DF842" captionStart="FIGURE 2" captionStartId="6.[152,255,1565,1589]" captionTargetBox="[290,1300,190,1533]" captionTargetId="figure-22@6.[283,1305,181,1541]" captionTargetPageId="6" captionText="FIGURE 2. Tumerozetes roughleyi sp. nov. (a) anterior aspect; (b) bothridia and interlamellar plates; (c) waxy tegument on rostrum; (d) chelicera; (e) palp; (f) rutellum; (g) anal and genital plates; (h) bothridium. Scale bars: a–c, g, h = 50 μm; e, f = 15 μm" figureDoi="http://doi.org/10.5281/zenodo.7141854" httpUri="https://zenodo.org/record/7141854/files/figure.png" pageId="6" pageNumber="39">
|
||
Figure
|
||
<quantity id="4CBF66BEC907FD08FF5EF855FF2DF842" box="[151,183,1923,1949]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="6" pageNumber="39" unit="g" value="2.0">2g</quantity>
|
||
</figureCitation>
|
||
). Anal plates lozenge-shaped, 55 µm long (
|
||
<typeStatus id="54FC75F9C907FD08FD56F855FC9DF842" box="[671,775,1923,1949]" pageId="6" pageNumber="39" type="holotype">holotype</typeStatus>
|
||
); only slightly separated from genital plates. Preanal organ (
|
||
<emphasis id="B9331749C907FD08FF68F87EFF27F81E" box="[161,189,1960,1985]" italics="true" pageId="6" pageNumber="39">po</emphasis>
|
||
) broad, flat T-shaped.
|
||
</paragraph>
|
||
<paragraph id="8BF8CB5BC907FD09FF0EF81DFA90FEC2" blockId="6.[151,1437,1707,2022]" lastBlockId="7.[151,1437,151,1365]" lastPageId="7" lastPageNumber="40" pageId="6" pageNumber="39">
|
||
<emphasis id="B9331749C907FD08FF0EF81DFEF3F83A" box="[199,361,1995,2021]" italics="true" pageId="6" pageNumber="39">Lateral aspect</emphasis>
|
||
: anterior prodorsum strongly concave. Tutorium (tu) pointed apically, similar shape as lamella, but without cusp; pedoctectum I (pd I) with sharp, well-developed cusp; pedotectum II (pd II) well-developed with blunt apex; discidium (
|
||
<emphasis id="B9331749C906FD09FE10FF6DFE75FF0A" box="[473,495,187,213]" italics="true" pageId="7" pageNumber="40">di</emphasis>
|
||
) prominent, pointed. Interlamellar plates convex, strongly inflated; seta
|
||
<emphasis id="B9331749C906FD09FAE3FF6DFADAFF0B" box="[1322,1344,187,212]" italics="true" pageId="7" pageNumber="40">in</emphasis>
|
||
pointed vertically. Anteriomedian projection of notogastral plate convex, strongly inflated. Notogastral setae positioned on faint horizontal lath of more heavily sclerotized cuticle that connects anteriorly with humeral process.
|
||
</paragraph>
|
||
<paragraph id="8BF8CB5BC906FD09FF0EFEF1FD7DFE9E" blockId="7.[151,1437,151,1365]" box="[199,743,295,321]" pageId="7" pageNumber="40">
|
||
<emphasis id="B9331749C906FD09FF0EFEF1FF66FE9F" box="[199,252,295,320]" italics="true" pageId="7" pageNumber="40">Legs</emphasis>
|
||
: monodactylous; claw lacking dorsal tooth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35D98D0C906FD09FF0EFE9DFC14FD2E" pageId="7" pageNumber="40" type="materials_examined">
|
||
<paragraph id="8BF8CB5BC906FD09FF0EFE9DFC14FD2E" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="40">
|
||
<emphasis id="B9331749C906FD09FF0EFE9DFCF0FEBA" bold="true" box="[199,874,330,357]" pageId="7" pageNumber="40">Type designation, material examined and locality data.</emphasis>
|
||
<materialsCitation id="3B2FC106C906FD09FCBCFE9DFE09FE72" ID-GBIF-Occurrence="3920727306" collectingDate="1996-03-12" collectorName="R. E. Roughley" latitude="-37.55" location="wet moss" longLatPrecision="1180" longitude="145.88333" pageId="7" pageNumber="40" specimenCount="1" stateProvince="Victoria" typeStatus="holotype">
|
||
<typeStatus id="54FC75F9C906FD09FCBCFE9DFC7AFEBA" box="[885,992,331,357]" pageId="7" pageNumber="40" type="holotype">Holotype</typeStatus>
|
||
,
|
||
<location id="8E989D80C906FD09FC26FE9DFBC4FEBA" LSID="urn:lsid:plazi:treatment:03EE7A4DC905FD09FF5EFA85FDCFFA8A:8E989D80C906FD09FC26FE9DFBC4FEBA" box="[1007,1118,331,357]" latitude="-37.55" longLatPrecision="1180" longitude="145.88333" name="wet moss" pageId="7" pageNumber="40" stateProvince="Victoria">wet moss</location>
|
||
,
|
||
<location id="8E989D80C906FD09FBA4FE9DFA03FEBA" LSID="urn:lsid:plazi:treatment:03EE7A4DC905FD09FF5EFA85FDCFFA8A:8E989D80C906FD09FBA4FE9DFA03FEBA" box="[1133,1433,331,357]" latitude="-37.55" longLatPrecision="1180" longitude="145.88333" name="Cumberland Creek Valley" pageId="7" pageNumber="40" stateProvince="Victoria">Cumberland Creek Valley</location>
|
||
,
|
||
<location id="8E989D80C906FD09FF5EFEB9FE77FE56" LSID="urn:lsid:plazi:treatment:03EE7A4DC905FD09FF5EFA85FDCFFA8A:8E989D80C906FD09FF5EFEB9FE77FE56" box="[151,493,367,393]" latitude="-37.55" longLatPrecision="1180" longitude="145.88333" name="Marysville-Woods Point Road" pageId="7" pageNumber="40" stateProvince="Victoria">Marysville-Woods Point Road</location>
|
||
,
|
||
<locationDeviation id="9A4AD279C906FD09FE3EFEB9FD32FE56" box="[503,680,367,393]" pageId="7" pageNumber="40">nr. Cambarville</locationDeviation>
|
||
,
|
||
<location id="8E989D80C906FD09FD78FEB9FC71FE56" LSID="urn:lsid:plazi:treatment:03EE7A4DC905FD09FF5EFA85FDCFFA8A:8E989D80C906FD09FD78FEB9FC71FE56" box="[689,1003,367,393]" latitude="-37.55" longLatPrecision="1180" longitude="145.88333" name="Yarra Ranges National Park" pageId="7" pageNumber="40" stateProvince="Victoria">Yarra Ranges National Park</location>
|
||
,
|
||
<collectingRegion id="498305B9C906FD09FC3CFEB9FBC8FE56" box="[1013,1106,367,393]" country="Australia" name="Victoria" pageId="7" pageNumber="40">Victoria</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE73AD9CC906FD09FB95FEB9FB2CFE56" box="[1116,1206,367,393]" degrees="37" direction="south" minutes="33" orientation="latitude" pageId="7" pageNumber="40" precision="925" value="-37.55">37º33’S</geoCoordinate>
|
||
<geoCoordinate id="EE73AD9CC906FD09FB75FEB8FAB2FE56" box="[1212,1320,366,393]" degrees="145" direction="east" minutes="53" orientation="longitude" pageId="7" pageNumber="40" precision="925" value="145.88333">145º53’E</geoCoordinate>
|
||
, coll.
|
||
<collectorName id="26B2AE8DC906FD09FAA3FEB9FE9FFE72" pageId="7" pageNumber="40">R.E. Roughley</collectorName>
|
||
,
|
||
<date id="FFF9ED9BC906FD09FEDBFE45FE15FE72" box="[274,399,403,429]" pageId="7" pageNumber="40" value="1996-03-12">
|
||
<collectingDate id="EFBD1473C906FD09FEDBFE45FE15FE72" box="[274,399,403,429]" pageId="7" pageNumber="40" value="1996-03-12">12.iii.1996</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
<typeStatus id="54FC75F9C906FD09FE52FE45FD96FE72" box="[411,524,403,429]" pageId="7" pageNumber="40" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D4100D2C906FD09FDD1FE45FD30FE72" box="[536,682,403,429]" pageId="7" pageNumber="40" type="female">four females</specimenCount>
|
||
,
|
||
<specimenCount id="9D4100D2C906FD09FD7FFE45FCBAFE72" box="[694,800,403,429]" pageId="7" pageNumber="40" type="male">one male</specimenCount>
|
||
, ANIC 297, Moss and litter, rainforest with
|
||
<taxonomicName id="4C47B0D8C906FD09FAD2FE45FDC1FE0E" authority=", Cumberland Creek Valley" authorityName="Cumberland Creek Valley" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="40" phylum="Tracheophyta" rank="species" species="cunninghami">
|
||
<emphasis id="B9331749C906FD09FAD2FE45FEB3FE0E" italics="true" pageId="7" pageNumber="40">Nothofagus cunninghami</emphasis>
|
||
, Cumberland Creek Valley
|
||
</taxonomicName>
|
||
, near picnic area [Cumberland Memorial Reserve], Yarra Ranges National Park,
|
||
<collectingRegion id="498305B9C906FD09FF13FE0DFEA2FE2A" box="[218,312,475,501]" country="Australia" name="Victoria" pageId="7" pageNumber="40">Victoria</collectingRegion>
|
||
.
|
||
<geoCoordinate id="EE73AD9CC906FD09FE8DFE0DFE3AFE2A" box="[324,416,475,501]" degrees="37" direction="south" minutes="34" orientation="latitude" pageId="7" pageNumber="40" precision="925" value="-37.566666">37°34’S</geoCoordinate>
|
||
<geoCoordinate id="EE73AD9CC906FD09FE60FE0CFD8DFE2A" box="[425,535,474,501]" degrees="145" direction="east" minutes="52" orientation="longitude" pageId="7" pageNumber="40" precision="925" value="145.86667">145°52’E</geoCoordinate>
|
||
,
|
||
<specimenCount id="9D4100D2C906FD09FDEDFE0DFDECFE2A" box="[548,630,475,501]" pageId="7" pageNumber="40" type="generic">
|
||
<quantity id="4CBF66BEC906FD09FDEDFE0DFDE8FE2A" box="[548,626,475,501]" metricMagnitude="2" metricUnit="m" metricValue="9.2" pageId="7" pageNumber="40" unit="m" value="920.0">920 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
, coll.
|
||
<collectionCode id="ED56539EC906FD09FD77FE0DFD48FE2A" box="[702,722,475,501]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="40">R</collectionCode>
|
||
.W. Taylor and
|
||
<collectionCode id="ED56539EC906FD09FC4BFE0DFC03FE2A" box="[898,921,475,501]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="7" pageNumber="40">R</collectionCode>
|
||
.J. Bartell,
|
||
<date id="FFF9ED9BC906FD09FBDBFE0DFBE5FE2A" box="[1042,1151,475,501]" pageId="7" pageNumber="40" value="1970-11-04">4.xi.1970</date>
|
||
.
|
||
<typeStatus id="54FC75F9C906FD09FB45FE0DFB67FE2A" box="[1164,1277,475,501]" pageId="7" pageNumber="40" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D4100D2C906FD09FAC3FE0DFA02FE2A" box="[1290,1432,475,501]" pageId="7" pageNumber="40" type="female">five females</specimenCount>
|
||
, moss (
|
||
<taxonomicName id="4C47B0D8C906FD09FF29FE29FE6CFDC6" authorityName="Paris" authorityYear="1904" box="[224,502,511,537]" class="Bryopsida" family="Dicranaceae" genus="Dicranoloma" kingdom="Plantae" order="Dicranales" pageId="7" pageNumber="40" phylum="Bryophyta" rank="species" species="billiardieri">
|
||
<emphasis id="B9331749C906FD09FF29FE29FE6CFDC6" box="[224,502,511,537]" italics="true" pageId="7" pageNumber="40">Dicranoloma billiardieri</emphasis>
|
||
</taxonomicName>
|
||
),
|
||
<taxonomicName id="4C47B0D8C906FD09FDC5FE29FCBCFDC6" box="[524,806,511,537]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="40" phylum="Tracheophyta" rank="species" species="cunninghami">
|
||
<emphasis id="B9331749C906FD09FDC5FE29FCBCFDC6" box="[524,806,511,537]" italics="true" pageId="7" pageNumber="40">Nothofagus cunninghami</emphasis>
|
||
</taxonomicName>
|
||
forest, The Beeches, Lady Talbot Drive near Warburton, Yarra Ranges National Park,
|
||
<collectingRegion id="498305B9C906FD09FE20FDF5FDDCFDE2" box="[489,582,547,573]" country="Australia" name="Victoria" pageId="7" pageNumber="40">Victoria</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE73AD9CC906FD09FD9DFDF5FD29FDE2" box="[596,691,547,573]" degrees="37" direction="south" minutes="29" orientation="latitude" pageId="7" pageNumber="40" precision="925" value="-37.483334">37°29’S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE73AD9CC906FD09FD08FDF4FCB5FDE2" box="[705,815,546,573]" degrees="145" direction="east" minutes="50" orientation="longitude" pageId="7" pageNumber="40" precision="925" value="145.83333">145°50’E</geoCoordinate>
|
||
,
|
||
<specimenCount id="9D4100D2C906FD09FCF7FDF5FC0BFDE2" box="[830,913,547,573]" pageId="7" pageNumber="40" type="generic">
|
||
<quantity id="4CBF66BEC906FD09FCF7FDF5FC17FDE2" box="[830,909,547,573]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="40" unit="m" value="800.0">800 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
, coll. G. Perdomo, May, 2009.
|
||
<typeStatus id="54FC75F9C906FD09FAC8FDF5FAE8FDE2" box="[1281,1394,547,573]" pageId="7" pageNumber="40" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D4100D2C906FD09FA49FDF5FF41FDBE" pageId="7" pageNumber="40" type="male">22 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D4100D2C906FD09FF21FD91FEFCFDBE" box="[232,358,583,609]" pageId="7" pageNumber="40" type="female">49 females</specimenCount>
|
||
, moss on Sassafras (
|
||
<taxonomicName id="4C47B0D8C906FD09FD98FD91FCECFDBE" box="[593,886,583,609]" class="Magnoliopsida" family="Atherospermataceae" genus="Atherosperma" kingdom="Plantae" order="Laurales" pageId="7" pageNumber="40" phylum="Tracheophyta" rank="species" species="moschatum">
|
||
<emphasis id="B9331749C906FD09FD98FD91FCECFDBE" box="[593,886,583,609]" italics="true" pageId="7" pageNumber="40">Atherosperma moschatum</emphasis>
|
||
</taxonomicName>
|
||
), cool temperate rainforest,
|
||
<quantity id="4CBF66BEC906FD09FB7EFD91FA88FDBE" box="[1207,1298,583,609]" metricMagnitude="3" metricUnit="m" metricValue="1.077" pageId="7" pageNumber="40" unit="m" value="1077.0">1077 m</quantity>
|
||
., Errinundra Saddle, Errinundra National Park,
|
||
<collectingRegion id="498305B9C906FD09FDE9FDBDFDE7FD5A" box="[544,637,619,645]" country="Australia" name="Victoria" pageId="7" pageNumber="40">Victoria</collectingRegion>
|
||
,
|
||
<geoCoordinate id="EE73AD9CC906FD09FD43FDBDFD7CFD5A" box="[650,742,619,645]" degrees="37" direction="south" minutes="19" orientation="latitude" pageId="7" pageNumber="40" precision="925" value="-37.316666">37°19’S</geoCoordinate>
|
||
<geoCoordinate id="EE73AD9CC906FD09FD26FDBCFCC7FD5A" box="[751,861,618,645]" degrees="148" direction="east" minutes="51" orientation="longitude" pageId="7" pageNumber="40" precision="925" value="148.85">148°51’E</geoCoordinate>
|
||
,
|
||
<quantity id="4CBF66BEC906FD09FCA3FDBDFC5CFD5A" box="[874,966,619,645]" metricMagnitude="3" metricUnit="m" metricValue="1.03" pageId="7" pageNumber="40" unit="m" value="1030.0">1030 m</quantity>
|
||
., coll. M.J. Colloff,
|
||
<date id="FFF9ED9BC906FD09FB62FDBDFABCFD5A" box="[1195,1318,619,645]" pageId="7" pageNumber="40" value="2009-09-29">29.ix.2009</date>
|
||
.
|
||
<typeStatus id="54FC75F9C906FD09FAFBFDBDFA02FD5A" box="[1330,1432,619,645]" pageId="7" pageNumber="40" type="paratype">Paratype</typeStatus>
|
||
: male, moss,
|
||
<taxonomicName id="4C47B0D8C906FD09FEEEFD59FDD9FD76" box="[295,579,655,681]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="40" phylum="Tracheophyta" rank="species" species="cunninghami">
|
||
<emphasis id="B9331749C906FD09FEEEFD59FDD9FD76" box="[295,579,655,681]" italics="true" pageId="7" pageNumber="40">Nothofagus cunninghami</emphasis>
|
||
</taxonomicName>
|
||
rainforest, Melba Gully, Otway National Park,
|
||
<collectingRegion id="498305B9C906FD09FBAFFD59FB59FD76" box="[1126,1219,655,681]" country="Australia" name="Victoria" pageId="7" pageNumber="40">Victoria</collectingRegion>
|
||
, coll. M.J. Colloff,
|
||
<date id="FFF9ED9BC906FD09FF5EFD65FE83FD12" box="[151,281,691,717]" pageId="7" pageNumber="40" value="2011-07-20">20.vii.2011</date>
|
||
.
|
||
<typeStatus id="54FC75F9C906FD09FEEEFD65FE02FD12" box="[295,408,691,717]" pageId="7" pageNumber="40" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D4100D2C906FD09FE6FFD65FD8BFD12" box="[422,529,691,717]" pageId="7" pageNumber="40" type="male">12 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D4100D2C906FD09FDE9FD65FD3AFD12" box="[544,672,691,717]" pageId="7" pageNumber="40" type="female">24 females</specimenCount>
|
||
, moss,
|
||
<taxonomicName id="4C47B0D8C906FD09FD33FD65FB8DFD12" box="[762,1047,691,717]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="7" pageNumber="40" phylum="Tracheophyta" rank="species" species="cunninghami">
|
||
<emphasis id="B9331749C906FD09FD33FD65FB8DFD12" box="[762,1047,691,717]" italics="true" pageId="7" pageNumber="40">Nothofagus cunninghami</emphasis>
|
||
</taxonomicName>
|
||
rainforest, rainforest walk, Maits Rest, Otway National Park,
|
||
<collectingRegion id="498305B9C906FD09FE18FD01FDB4FD2E" box="[465,558,727,753]" country="Australia" name="Victoria" pageId="7" pageNumber="40">Victoria</collectingRegion>
|
||
, coll. M.J. Colloff,
|
||
<date id="FFF9ED9BC906FD09FCC1FD01FC10FD2E" box="[776,906,727,753]" pageId="7" pageNumber="40" value="2011-07-19">19.vii.2011</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35D98D0C906FD09FF0EFD2DFE90FCE6" pageId="7" pageNumber="40" type="etymology">
|
||
<paragraph id="8BF8CB5BC906FD09FF0EFD2DFE90FCE6" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="40">
|
||
<emphasis id="B9331749C906FD09FF0EFD2DFED4FCCA" bold="true" box="[199,334,763,789]" pageId="7" pageNumber="40">Etymology.</emphasis>
|
||
This species is named in honour and memory of the collector of the
|
||
<typeStatus id="54FC75F9C906FD09FB99FD2DFB28FCCA" box="[1104,1202,763,789]" pageId="7" pageNumber="40" type="holotype">holotype</typeStatus>
|
||
specimen, Robert E. Roughley.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35D98D0C906FD09FF0EFC94FB42FBCE" pageId="7" pageNumber="40" type="diagnosis">
|
||
<paragraph id="8BF8CB5BC906FD09FF0EFC94FB42FBCE" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="40">
|
||
<emphasis id="B9331749C906FD09FF0EFC94FED8FC83" bold="true" box="[199,322,834,860]" pageId="7" pageNumber="40">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C47B0D8C906FD09FE85FC95FDD1FC82" authority="Colloff, 2022" authorityName="Colloff" authorityYear="2022" box="[332,587,835,861]" class="Arachnida" family="Tumerozetidae" genus="Tumerozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="roughleyi" status="sp. nov.">
|
||
<emphasis id="B9331749C906FD09FE85FC95FDD1FC82" box="[332,587,835,861]" italics="true" pageId="7" pageNumber="40">Tumerozetes roughleyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9331749C906FD09FD9FFC95FD29FC82" bold="true" box="[598,691,835,861]" pageId="7" pageNumber="40">
|
||
<taxonomicNameLabel id="A200AA32C906FD09FD9FFC95FD29FC82" box="[598,691,835,861]" pageId="7" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from all other species in the genus by the following combination of characters: (1) the paired, curved, longitudinal median notogastral ridges forming an ovoid structure and extending posteriorly as far as level of lyrifissure
|
||
<emphasis id="B9331749C906FD09FCA8FC5DFC1FFC7A" box="[865,901,907,933]" italics="true" pageId="7" pageNumber="40">iad</emphasis>
|
||
: (2) setae
|
||
<emphasis id="B9331749C906FD09FC35FC5DFB88FC7A" box="[1020,1042,907,933]" italics="true" pageId="7" pageNumber="40">la</emphasis>
|
||
,
|
||
<emphasis id="B9331749C906FD09FBE9FC5DFBACFC7A" box="[1056,1078,907,933]" italics="true" pageId="7" pageNumber="40">lp</emphasis>
|
||
and
|
||
<emphasis id="B9331749C906FD09FBA7FC5DFBE6FC7A" box="[1134,1148,907,933]" italics="true" pageId="7" pageNumber="40">h</emphasis>
|
||
<subScript id="17C3C91EC906FD09FBB5FC4BFB1EFC74" attach="left" box="[1148,1156,925,939]" fontSize="6" pageId="7" pageNumber="40">3</subScript>
|
||
with fine spines basally; (3) with heart-shaped anteriomedian projection of the notogastral plate; (4) the presence of a patronium; (5) the dorsosejugal furrow coinciding with the junction between the longitudinal notogastral ridges and the anteriomedian projection of the notogastral plate and positioned markedly posterior of the level of the bothridia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35D98D0C906FD09FF0EFBCDFDCFFA8A" pageId="7" pageNumber="40" type="discussion">
|
||
<paragraph id="8BF8CB5BC906FD09FF0EFBCDFDCFFA8A" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="40">
|
||
<emphasis id="B9331749C906FD09FF0EFBCDFEA1FBEA" bold="true" box="[199,315,1051,1077]" pageId="7" pageNumber="40">Remarks.</emphasis>
|
||
<taxonomicName id="4C47B0D8C906FD09FE8CFBCDFDD9FBEA" authority="Colloff, 2022" authorityName="Colloff" authorityYear="2022" box="[325,579,1051,1077]" class="Arachnida" family="Tumerozetidae" genus="Tumerozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="roughleyi" status="sp. nov.">
|
||
<emphasis id="B9331749C906FD09FE8CFBCDFDD9FBEA" box="[325,579,1051,1077]" italics="true" pageId="7" pageNumber="40">Tumerozetes roughleyi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9331749C906FD09FD84FBCDFD30FBEA" bold="true" box="[589,682,1051,1077]" pageId="7" pageNumber="40">
|
||
<taxonomicNameLabel id="A200AA32C906FD09FD84FBCDFD30FBEA" box="[589,682,1051,1077]" pageId="7" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is morphologically most similar to
|
||
<taxonomicName id="4C47B0D8C906FD09FB83FBCDFB20FBEA" authorityName="Hammer" authorityYear="1966" box="[1098,1210,1051,1077]" class="Arachnida" family="Tumerozetidae" genus="Tumerozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="pumilis">
|
||
<emphasis id="B9331749C906FD09FB83FBCDFB20FBEA" box="[1098,1210,1051,1077]" italics="true" pageId="7" pageNumber="40">T. pumilis</emphasis>
|
||
</taxonomicName>
|
||
in the shape of the median notogastral ridges and to
|
||
<taxonomicName id="4C47B0D8C906FD09FDC5FBE9FD03FB86" authorityName="Hammer" authorityYear="1966" box="[524,665,1087,1113]" class="Arachnida" family="Tumerozetidae" genus="Tumerozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="bifurcatus">
|
||
<emphasis id="B9331749C906FD09FDC5FBE9FD03FB86" box="[524,665,1087,1113]" italics="true" pageId="7" pageNumber="40">T. bifurcatus</emphasis>
|
||
</taxonomicName>
|
||
in the shape of the anteriomedian projection of the notogastral plate. However, in
|
||
<taxonomicName id="4C47B0D8C906FD09FEE0FBB5FE0DFBA2" authorityName="Hammer" authorityYear="1966" box="[297,407,1123,1149]" class="Arachnida" family="Tumerozetidae" genus="Tumerozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="pumilis">
|
||
<emphasis id="B9331749C906FD09FEE0FBB5FE0DFBA2" box="[297,407,1123,1149]" italics="true" pageId="7" pageNumber="40">T. pumilis</emphasis>
|
||
</taxonomicName>
|
||
each notogastral ridge is considerably narrower and they converge posteriorly at a position level with setae
|
||
<emphasis id="B9331749C906FD09FE82FB51FEC3FB7E" box="[331,345,1159,1185]" italics="true" pageId="7" pageNumber="40">h</emphasis>
|
||
<subScript id="17C3C91EC906FD09FE90FB4FFEFBFB78" attach="left" box="[345,353,1177,1191]" fontSize="6" pageId="7" pageNumber="40">3</subScript>
|
||
.
|
||
<taxonomicName id="4C47B0D8C906FD09FEA6FB51FC84FB7E" authority="Hammer, 1966" authorityName="Hammer" authorityYear="1966" box="[367,798,1159,1185]" class="Arachnida" family="Tumerozetidae" genus="Tumerozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="bifurcatus">
|
||
<emphasis id="B9331749C906FD09FEA6FB51FDE8FB7E" box="[367,626,1159,1185]" italics="true" pageId="7" pageNumber="40">Tumerozetes bifurcatus</emphasis>
|
||
<bibRefCitation id="EFD6B6AAC906FD09FDB1FB51FC84FB7E" author="Hammer, M." box="[632,798,1159,1185]" pageId="7" pageNumber="40" pagination="1 - 108" refId="ref14939" refString="Hammer, M. (1966) Investigations on the Oribatid fauna of New Zealand. Part I. Biologiske Skrifter det Kongelige Dansk Videnskabernes Selskab, 15 (2), 1 - 108." type="journal article" year="1966">Hammer, 1966</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C47B0D8C906FD09FC9DFB51FB12FB7E" authority="Hammer, 1966" authorityName="Hammer" authorityYear="1966" box="[852,1160,1159,1185]" class="Arachnida" family="Tumerozetidae" genus="Tumerozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="circularis">
|
||
<emphasis id="B9331749C906FD09FC9DFB51FC41FB7E" box="[852,987,1159,1185]" italics="true" pageId="7" pageNumber="40">T. circularis</emphasis>
|
||
<bibRefCitation id="EFD6B6AAC906FD09FC2BFB51FB12FB7E" author="Hammer, M." box="[994,1160,1159,1185]" pageId="7" pageNumber="40" pagination="1 - 108" refId="ref14939" refString="Hammer, M. (1966) Investigations on the Oribatid fauna of New Zealand. Part I. Biologiske Skrifter det Kongelige Dansk Videnskabernes Selskab, 15 (2), 1 - 108." type="journal article" year="1966">Hammer, 1966</bibRefCitation>
|
||
</taxonomicName>
|
||
both have short, parallel longitudinal dorsal ridges set some distance apart and extending posteriorly only as far as a position level with setae
|
||
<emphasis id="B9331749C906FD09FF5EFB19FF37FB36" box="[151,173,1231,1257]" italics="true" pageId="7" pageNumber="40">la</emphasis>
|
||
. Also,
|
||
<taxonomicName id="4C47B0D8C906FD09FECBFB19FE0BFB36" authorityName="Hammer" authorityYear="1966" box="[258,401,1231,1257]" class="Arachnida" family="Tumerozetidae" genus="Tumerozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="bifurcatus">
|
||
<emphasis id="B9331749C906FD09FECBFB19FE0BFB36" box="[258,401,1231,1257]" italics="true" pageId="7" pageNumber="40">T. bifurcatus</emphasis>
|
||
</taxonomicName>
|
||
has bifurcate notogastral setae.
|
||
<taxonomicName id="4C47B0D8C906FD09FCCEFB19FB4AFB36" authority="Hammer, 1966" authorityName="Hammer" authorityYear="1966" box="[775,1232,1231,1257]" class="Arachnida" family="Tumerozetidae" genus="Tumerozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="indistinctus">
|
||
<emphasis id="B9331749C906FD09FCCEFB19FB86FB36" box="[775,1052,1231,1257]" italics="true" pageId="7" pageNumber="40">Tumerozetes indistinctus</emphasis>
|
||
<bibRefCitation id="EFD6B6AAC906FD09FBEFFB19FB4AFB36" author="Hammer, M." box="[1062,1232,1231,1257]" pageId="7" pageNumber="40" pagination="1 - 108" refId="ref14939" refString="Hammer, M. (1966) Investigations on the Oribatid fauna of New Zealand. Part I. Biologiske Skrifter det Kongelige Dansk Videnskabernes Selskab, 15 (2), 1 - 108." type="journal article" year="1966">Hammer, 1966</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C47B0D8C906FD09FAC5FB19FEA7FAD2" authority="Hammer, 1966" authorityName="Hammer" authorityYear="1966" class="Arachnida" family="Tumerozetidae" genus="Tumerozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="parallelus">
|
||
<emphasis id="B9331749C906FD09FAC5FB19FA07FB36" box="[1292,1437,1231,1257]" italics="true" pageId="7" pageNumber="40">T. parallelus</emphasis>
|
||
<bibRefCitation id="EFD6B6AAC906FD09FF5EFB25FEA7FAD2" author="Hammer, M." box="[151,317,1267,1293]" pageId="7" pageNumber="40" pagination="1 - 108" refId="ref14939" refString="Hammer, M. (1966) Investigations on the Oribatid fauna of New Zealand. Part I. Biologiske Skrifter det Kongelige Dansk Videnskabernes Selskab, 15 (2), 1 - 108." type="journal article" year="1966">Hammer, 1966</bibRefCitation>
|
||
</taxonomicName>
|
||
both have very long, parallel longitudinal dorsal ridges, almost contiguous and extending posteriorly as far as a position level with setae
|
||
<emphasis id="B9331749C906FD09FD8DFAC1FDC8FAEE" box="[580,594,1303,1329]" italics="true" pageId="7" pageNumber="40">h</emphasis>
|
||
<subScript id="17C3C91EC906FD09FD9BFAFFFDC0FAE8" attach="left" box="[594,602,1321,1335]" fontSize="6" pageId="7" pageNumber="40">3</subScript>
|
||
. Other than dimensions, there were no obvious external morphological differences between males and females.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |