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<document id="EFD30E191C1225C6A0F658E5E95A1CE1" ID-DOI="10.5281/zenodo.178011" ID-GBIF-Dataset="b02a028f-ba14-458e-be60-cd539f28d9d1" ID-ISSN="1175-5326" ID-Zenodo-Dep="178011" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459878369919" checkinUser="plazi" docAuthor="Tian, Mingyi &amp; Deuve, Thierry" docDate="2007" docId="03EE11580173FFBBFF68FE83FCAC22C3" docLanguage="en" docName="zt01546p022.pdf" docOrigin="Zootaxa 1546" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Brachinus philippinensis Tian &amp; Deuve, new species" docType="treatment" docVersion="8" lastPageNumber="18" masterDocId="FFD769200170FFB8FFFFFFDCFFB5251F" masterDocTitle="Review of four species of the genus Brachinus Weber (Coleoptera: Caraboidea: Brachinidae) from the Philippines" masterLastPageNumber="22" masterPageNumber="15" pageNumber="18" updateTime="1698227246572" updateUser="plazi">
<mods:mods id="33C58475FB19EE3AA57B7E22E541C42C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="8E65DA1AFFB439C0118DBA3A43833AD7">Review of four species of the genus Brachinus Weber (Coleoptera: Caraboidea: Brachinidae) from the Philippines</mods:title>
</mods:titleInfo>
<mods:name id="4EAB884CD46A9F839D4F6EE42D65F554" type="personal">
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<mods:namePart id="62524C5C95D6B7F3FC9D0AC3C396BA71">Tian, Mingyi</mods:namePart>
</mods:name>
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<mods:namePart id="672B20E36846CA4B38842C4AB5505DC1">Deuve, Thierry</mods:namePart>
</mods:name>
<mods:typeOfResource id="189623C2DD2DF0AE934411A125A47304">text</mods:typeOfResource>
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<mods:title id="215A42AE54D669C0D2A50C1C5FD3DF6E">Zootaxa</mods:title>
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<mods:date id="601735D3F04F7D959D42D9B4F82A6F27">2007</mods:date>
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<mods:number id="8824EE77886EF7F78218D9912B4CEC98">1546</mods:number>
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<mods:classification id="DC8C098822710F68D222B7FBD0FB9513">journal article</mods:classification>
<mods:identifier id="42ED3A193E3213E7355C4E9ED6BD7387" type="DOI">10.5281/zenodo.178011</mods:identifier>
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<mods:identifier id="D55DD60A19D44A656C58F64CC86D7371" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F522498CD5260C7C11AE519A583E04AE" type="Zenodo-Dep">178011</mods:identifier>
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<treatment id="03EE11580173FFBBFF68FE83FCAC22C3" ID-DOI="http://doi.org/10.5281/zenodo.6244092" ID-GBIF-Taxon="119370432" ID-Zenodo-Dep="6244092" LSID="urn:lsid:plazi:treatment:03EE11580173FFBBFF68FE83FCAC22C3" httpUri="http://treatment.plazi.org/id/03EE11580173FFBBFF68FE83FCAC22C3" lastPageNumber="18" pageId="3" pageNumber="18">
<subSubSection id="C35DF3C50173FFBBFF68FE83FEA22483" pageId="3" pageNumber="18" type="nomenclature">
<paragraph id="8BF8A04E0173FFBBFF68FE83FCAE2466" blockId="3.[151,795,351,412]" box="[151,795,351,377]" pageId="3" pageNumber="18">
<heading id="D0B017220173FFBBFF68FE83FCAE2466" bold="true" box="[151,795,351,377]" fontSize="11" level="1" pageId="3" pageNumber="18" reason="1">
<emphasis id="B9337C5C0173FFBBFF68FE83FCAE2466" bold="true" box="[151,795,351,377]" pageId="3" pageNumber="18">
<taxonomicName id="4C47DBCD0173FFBBFF68FE83FDCA2466" ID-CoL="MSYV" authority="Tian &amp; Deuve" authorityName="Tian &amp; Deuve" box="[151,639,351,377]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="philippinensis" status="sp. nov.">
<emphasis id="B9337C5C0173FFBBFF68FE83FE722466" bold="true" box="[151,455,351,377]" italics="true" pageId="3" pageNumber="18">Brachinus philippinensis</emphasis>
Tian &amp; Deuve
</taxonomicName>
,
<taxonomicNameLabel id="A200C1270173FFBBFD73FE83FCAE2466" box="[652,795,351,377]" pageId="3" pageNumber="18" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF8A04E0173FFBBFF68FE5EFEA22483" blockId="3.[151,795,351,412]" box="[151,279,386,412]" pageId="3" pageNumber="18">
(
<figureCitation id="137CBCCB0173FFBBFF60FE5EFF462483" box="[159,243,386,412]" captionStart="FIGURES 1 4" captionStartId="2.[151,269,1773,1797]" captionTargetBox="[273,1303,204,1739]" captionTargetId="figure@2.[269,1317,197,1747]" captionTargetPageId="2" captionText="FIGURES 1 4. Habiti of Brachinus ssp. from the Philippines. 1. B. solidipalpis n. sp., paratype, male, 2. B. philippinensis n. sp., holotype, female, 3. B. piceus Chaudior, holotype, male, 4. B. luzonicus Chaudoir, female. Scale bar: 5.0 mm." httpUri="https://zenodo.org/record/178012/files/figure.png" pageId="3" pageNumber="18">Figs. 2</figureCitation>
,
<figureCitation id="137CBCCB0173FFBBFF00FE5EFEBB2483" box="[255,270,386,412]" captionStart="FIGURES 5 8" captionStartId="4.[151,269,1584,1608]" captionTargetBox="[195,1416,185,1554]" captionTargetId="figure@4.[151,1436,182,1560]" captionTargetPageId="4" captionText="FIGURES 5 8. Pronota and aedeagi of Brachinus ssp. from the Philippines. 5, 9 10. B. solidipalpis n. sp., holotype, 6. B. philippinensis n. sp., holotype, 7, 11 12. B. piceus Chaudior, holotype, 8, 13 14. B. luzonicus Chaudoir. Scale bar: 5.0 mm for 8, 10.0 mm for others." httpUri="https://zenodo.org/record/178013/files/figure.png" pageId="3" pageNumber="18">6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35DF3C50173FFBBFF68FE0EFB8B270B" pageId="3" pageNumber="18" type="materials_examined">
<paragraph id="8BF8A04E0173FFBBFF68FE0EFB8B270B" blockId="3.[151,1437,466,2012]" pageId="3" pageNumber="18">
<emphasis id="B9337C5C0173FFBBFF68FE0EFEA524F3" bold="true" box="[151,272,466,492]" pageId="3" pageNumber="18">
<typeStatus id="54FC1EEC0173FFBBFF68FE0EFEBE24F3" box="[151,267,466,492]" pageId="3" pageNumber="18" type="holotype">Holotype</typeStatus>
.
</emphasis>
Female, labeled “
<collectingCountry id="F350E0DE0173FFBBFE12FE0EFDC324F3" box="[493,630,466,492]" name="Philippines" pageId="3" pageNumber="18">Philippines</collectingCountry>
, Ch. Semper, Tuga,
<date id="FFF9868E0173FFBBFC8DFE0EFC6824F3" box="[882,989,466,492]" pageId="3" pageNumber="18" value="1921-04-30">21/30. 4.</date>
62”, “Museum Paris, 1953, Coll. R. Oberthür”, from Tawi-Tawi
<collectingCountry id="F350E0DE0173FFBBFE1DFE26FD98270B" box="[482,557,506,532]" name="Iceland" pageId="3" pageNumber="18">Island</collectingCountry>
, southwest
<collectingCountry id="F350E0DE0173FFBBFD4AFE26FC89270B" box="[693,828,506,532]" name="Philippines" pageId="3" pageNumber="18">Philippines</collectingCountry>
, deposited in
<collectionCode id="ED56388B0173FFBBFC22FE26FB8E270B" LSID="urn:lsid:biocol.org:col:34988" box="[989,1083,506,532]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="18">MNHN</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C35DF3C50173FFBBFF38FDFEFE3B23F3" pageId="3" pageNumber="18" type="diagnosis">
<paragraph id="8BF8A04E0173FFBBFF38FDFEFE3227AB" blockId="3.[151,1437,466,2012]" pageId="3" pageNumber="18">
<emphasis id="B9337C5C0173FFBBFF38FDFEFEFC2723" bold="true" box="[199,329,546,572]" pageId="3" pageNumber="18">Diagnosis:</emphasis>
Large sized, rust brown, legs with femur (except apex) darker, mouthparts with palpi slender, mentum without pit, but deep furrows, labial palpiger bisetose; head as wide as pronotum; lateral margin of pronotum deeply sinuate before hind angle, hind angle acute, proepisternum faintly visible from above; elytra indistinctly carinate.
</paragraph>
<paragraph id="8BF8A04E0173FFBBFF38FD1EFCF227C3" blockId="3.[151,1437,466,2012]" box="[199,839,706,732]" pageId="3" pageNumber="18">
Length: 16.0 mm; width:
<quantity id="4CBF0DAB0173FFBBFE0CFD1EFDE527C3" box="[499,592,706,732]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="18" unit="mm" value="6.5">6.5 mm</quantity>
. Habitus as in
<figureCitation id="137CBCCB0173FFBBFD04FD1EFCF627C3" box="[763,835,706,732]" captionStart="FIGURES 1 4" captionStartId="2.[151,269,1773,1797]" captionTargetBox="[273,1303,204,1739]" captionTargetId="figure@2.[269,1317,197,1747]" captionTargetPageId="2" captionText="FIGURES 1 4. Habiti of Brachinus ssp. from the Philippines. 1. B. solidipalpis n. sp., paratype, male, 2. B. philippinensis n. sp., holotype, female, 3. B. piceus Chaudior, holotype, male, 4. B. luzonicus Chaudoir, female. Scale bar: 5.0 mm." httpUri="https://zenodo.org/record/178012/files/figure.png" pageId="3" pageNumber="18">Fig. 2</figureCitation>
.
</paragraph>
<paragraph id="8BF8A04E0173FFBBFF38FD36FD2B2663" blockId="3.[151,1437,466,2012]" pageId="3" pageNumber="18">Rust brown, but frontal impressions, antennomeres 511, legs with femur (except apex), coxae, trochanters, and gula yellow, mouthparts palpi, apex of femur, antennomeres 14, elytral epipleuron and ventral surface light brown, lateral sides of abdominal ventrites IIVI and whole ventrites VII dark brown; head with a pair of large and yellow spots between eyes.</paragraph>
<paragraph id="8BF8A04E0173FFBBFF38FC56FF6C2013" blockId="3.[151,1437,466,2012]" pageId="3" pageNumber="18">Head longer than wide (HL/HW=1.12), labrum slightly rounded at front margin, sexsetose, clypeus with shorter additional setae; mandibular scrobe with several setae on upper margin, glabrous on median portion; eyes small, frons and vertex convex, frontal impressions deep and wide, convergent backwards; with several setae along sides of eyes, neck well marked, with coarse transversal wrinkles, glabrous on frons and vertex; ligula broad, pubescent at apex, adnated to paraglossae posteriorly; palpi pubescent, maxillary palpomere 4 slightly dilated, labial palpomere 3 strongly dilated; labial palpomere 2 bisetose; mentum without tooth, with two deep furrows at base, and several setae on median portion, two setae on either side, submentum with a row of long setae; labial palpiger bisetose. Antennae extending to middle of elytra, pubescent from apex of antennomere 2, antennomere 3 longer than antennomeres 1 and 2 combined, antennomere 1 with several long setae.</paragraph>
<paragraph id="8BF8A04E0173FFBBFF38FAC6FE7B20B3" blockId="3.[151,1437,466,2012]" pageId="3" pageNumber="18">
Pronotum (
<figureCitation id="137CBCCB0173FFBBFEB2FAC6FE23202B" box="[333,406,1306,1332]" captionStart="FIGURES 5 8" captionStartId="4.[151,269,1584,1608]" captionTargetBox="[195,1416,185,1554]" captionTargetId="figure@4.[151,1436,182,1560]" captionTargetPageId="4" captionText="FIGURES 5 8. Pronota and aedeagi of Brachinus ssp. from the Philippines. 5, 9 10. B. solidipalpis n. sp., holotype, 6. B. philippinensis n. sp., holotype, 7, 11 12. B. piceus Chaudior, holotype, 8, 13 14. B. luzonicus Chaudoir. Scale bar: 5.0 mm for 8, 10.0 mm for others." httpUri="https://zenodo.org/record/178013/files/figure.png" pageId="3" pageNumber="18">Fig. 6</figureCitation>
) wider than long (PW/PL=1.13), widest at about one-sixth from apex, lateral margin deeply sinuate near hind angle, with a pair of lateral setae; basal transverse impression distinct; surface rather glabrous, distinctly wrinkled and punctured at base, median line clear; proepisternum slightly tumescent and faintly visible from above.
</paragraph>
<paragraph id="8BF8A04E0173FFBBFF38FA66FC942353" blockId="3.[151,1437,466,2012]" pageId="3" pageNumber="18">Elytra elongate-ovate (EL/EW=1.48), convex, without basal border, sides broad, not paralleled each other, widest at one-third from apex, shoulders slightly prominent, apex truncate, membrane narrow, with fringe of short setae and few sparsely long setae; depressions flat, wide, pubescent (indistinct on median portion), costae slightly convex, but not carinate, glabrous.</paragraph>
<paragraph id="8BF8A04E0173FFBBFF38F986FA992383" blockId="3.[151,1437,466,2012]" pageId="3" pageNumber="18">Fore tibia with subapical spur externally located. Meso- and meta-sterna glabrous, abdominal ventrites densely setose and punctate. Proepisternum glabrous, slightly tumescent, faintly visible from above.</paragraph>
<paragraph id="8BF8A04E0173FFBBFF38F976FB8D23DB" blockId="3.[151,1437,466,2012]" box="[199,1080,1706,1732]" pageId="3" pageNumber="18">Microsculpture meshes densely isodiametric on head, pronotum and elytra.</paragraph>
<paragraph id="8BF8A04E0173FFBBFF38F90EFE3B23F3" blockId="3.[151,1437,466,2012]" box="[199,398,1746,1772]" pageId="3" pageNumber="18">Male: Unknown.</paragraph>
</subSubSection>
<subSubSection id="C35DF3C50173FFBBFF38F926FC85227B" pageId="3" pageNumber="18" type="discussion">
<paragraph id="8BF8A04E0173FFBBFF38F926FC85227B" blockId="3.[151,1437,466,2012]" pageId="3" pageNumber="18">
<emphasis id="B9337C5C0173FFBBFF38F926FEF7220B" bold="true" box="[199,322,1786,1812]" pageId="3" pageNumber="18">Remarks:</emphasis>
This new species is similar to Chinese species
<taxonomicName id="4C47DBCD0173FFBBFC98F927FBDF220B" authority="Chaudoir" authorityName="Chaudoir" box="[871,1130,1786,1812]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="B9337C5C0173FFBBFC98F927FC41220B" box="[871,1012,1787,1812]" italics="true" pageId="3" pageNumber="18">B. chinensis</emphasis>
Chaudoir
</taxonomicName>
, but elytral costae not evidently carinate, pronotum deeply sinuated near hind angle, proepisternum slightly tumescent and visible from above, and median portion of elytra without pubescence.
</paragraph>
</subSubSection>
<subSubSection id="C35DF3C50173FFBBFF38F8AEFE3F22AB" pageId="3" pageNumber="18" type="etymology">
<paragraph id="8BF8A04E0173FFBBFF38F8AEFE3F22AB" blockId="3.[151,1437,466,2012]" pageId="3" pageNumber="18">
<emphasis id="B9337C5C0173FFBBFF38F8AEFEED2293" bold="true" box="[199,344,1906,1932]" pageId="3" pageNumber="18">Etymology:</emphasis>
The specific epithet,
<taxonomicName id="4C47DBCD0173FFBBFDABF8AFFD4D2293" box="[596,760,1907,1932]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="18" phylum="Arthropoda" rank="species" species="philippinensis">
<emphasis id="B9337C5C0173FFBBFDABF8AFFD4D2293" box="[596,760,1907,1932]" italics="true" pageId="3" pageNumber="18">philippinensis</emphasis>
</taxonomicName>
, is an adjective derived from the
<typeStatus id="54FC1EEC0173FFBBFB7EF8AEFB062293" box="[1153,1203,1906,1932]" pageId="3" pageNumber="18">type</typeStatus>
locality of this species, the
<collectingCountry id="F350E0DE0173FFBBFF01F846FE3322AB" box="[254,390,1946,1972]" name="Philippines" pageId="3" pageNumber="18">Philippines</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35DF3C50173FFBBFF38F81EFCAC22C3" box="[199,793,1986,2012]" pageId="3" pageNumber="18" type="distribution">
<paragraph id="8BF8A04E0173FFBBFF38F81EFCAC22C3" blockId="3.[151,1437,466,2012]" box="[199,793,1986,2012]" pageId="3" pageNumber="18">
<emphasis id="B9337C5C0173FFBBFF38F81EFEDF22C3" bold="true" box="[199,362,1986,2012]" pageId="3" pageNumber="18">Distribution:</emphasis>
The
<collectingCountry id="F350E0DE0173FFBBFE59F81EFD9F22C3" box="[422,554,1986,2012]" name="Philippines" pageId="3" pageNumber="18">Philippines</collectingCountry>
(Tawi-Tawi
<collectingCountry id="F350E0DE0173FFBBFD3FF81EFCB822C3" box="[704,781,1986,2012]" name="Iceland" pageId="3" pageNumber="18">Island</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>