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<document id="59932B2FD00AA7018C6C854304680CB3" ID-DOI="10.11646/zootaxa.5267.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7840783" ID-ZooBank="9CD59054-8D7D-413F-B9FD-29EAFE7E511D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1681816955386" checkinUser="plazi" docAuthor="Lehmann, Ingo, Zahiri, Reza &amp; Husemann, Martin" docDate="2023" docId="03DC6C76CC2D4C39F7DFFC89FC4CFB5C" docLanguage="en" docName="zootaxa.5267.1.1.pdf" docOrigin="Zootaxa 5267 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5267.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Lukeniana kammeri Lehmann, Zahiri &amp; Husemann 2023, sp. nov." docType="treatment" docUuid="C0AFCE0D-E1B3-484E-9AF0-14FDF9C1A2B5" docUuidSource="ZooBank" docVersion="3" lastPageNumber="30" masterDocId="FFE5140ECC314C24F748FFEFFF9EFFB5" masterDocTitle="Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa" masterLastPageNumber="106" masterPageNumber="1" pageNumber="29" updateTime="1699487052491" updateUser="plazi">
<mods:mods id="82B49ADAA264C70E3B9EBB131DF92985" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="587B0BF0B6DB8FC8676B803014DF098B">
<mods:title id="A55160580FAE379EE5E0DF44D1FF06AC">Revision of the Metarbelodes Strand, 1909 genus-group (Lepidoptera: Cossoidea: Metarbelidae) with descriptions of two new genera and 33 new species from high elevations of eastern and southern Africa</mods:title>
</mods:titleInfo>
<mods:name id="5E366031A071635BA83644738678189C" type="personal">
<mods:role id="991AAC87F15726B03676C70EFDD05EF8">
<mods:roleTerm id="D8975A5AA18679A32A1EC7BC58F42D78">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="60CF0B9C170DC31D9599386197B6D315">Lehmann, Ingo</mods:namePart>
<mods:nameIdentifier id="3F71B4FB2B4833FD60BE902E9ABDE40E" type="email">martin.husemann@uni-hamburg.de</mods:nameIdentifier>
</mods:name>
<mods:name id="2F26808AEACBF153C4B0839ED41C063B" type="personal">
<mods:role id="84EA66B0901B77BB3BBF1559B0FB1E13">
<mods:roleTerm id="E41227EDB8D734F26C2E7A34E30C4250">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8BFC4B537516101DCF0797CA14B50B20">Zahiri, Reza</mods:namePart>
<mods:nameIdentifier id="B0B6A6CA390B7C60164D9A643F2BA940" type="email">reza.zahiri@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="7613BF81E94FBCBAA0493C31CDEAB081" type="personal">
<mods:role id="D2F4021B3257334A97B3DD06F73AA885">
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</mods:role>
<mods:namePart id="99A4BC385194412226FAD26F9B1BA3E6">Husemann, Martin</mods:namePart>
<mods:nameIdentifier id="2CF3BA44E9C88C70EBB21624C888FABF" type="email">martin.husemann@uni-hamburg.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="226D550D8C90D88AA417FB68B55D0C6B">text</mods:typeOfResource>
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<mods:titleInfo id="82438A26AA663CB4D0B08B34FA48587F">
<mods:title id="ED6F63F071A9EEFC46D1A3FEF623CF68">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="498DF396CA14E1BB7A48FFB2956E9252">
<mods:date id="D4FF842947B68F2D6183A19EF1B44CF7">2023</mods:date>
<mods:detail id="94E6DAA200598CE7D75A3C8A3EC1715D" type="pubDate">
<mods:number id="A232C694032940F98D3AD2B2FE3DE681">2023-04-18</mods:number>
</mods:detail>
<mods:detail id="1B87675465D559F198541678002F342F" type="volume">
<mods:number id="32E39D7BCB0CB6303EF8674F3F5EF514">5267</mods:number>
</mods:detail>
<mods:detail id="EC5C81CF2E6CCA2E6D747BA1D4BAAF5A" type="issue">
<mods:number id="C97003F5722620A404246595B1433342">1</mods:number>
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<mods:start id="E7A0575E272C31A96DA36E0E66DFB347">1</mods:start>
<mods:end id="5B771C3F16179E88B1AD04FF8DFCC563">106</mods:end>
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<mods:url id="946EF2E92D93B97E6DA8CAD88F724B0E">http://dx.doi.org/10.11646/zootaxa.5267.1.1</mods:url>
</mods:location>
<mods:classification id="F95109F539CCE16BF1C871DE41BE8DDF">journal article</mods:classification>
<mods:identifier id="1D8F024E189F1969A5894B781E5AC104" type="DOI">10.11646/zootaxa.5267.1.1</mods:identifier>
<mods:identifier id="155B7F22C4A2642E20D2757C757BA05B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="9667AB908CAF19F821E2991BB0F58E88" type="Zenodo-Dep">7840783</mods:identifier>
<mods:identifier id="60DDB37AF79016B727DF78957F8889B8" type="ZooBank">9CD59054-8D7D-413F-B9FD-29EAFE7E511D</mods:identifier>
</mods:mods>
<treatment id="03DC6C76CC2D4C39F7DFFC89FC4CFB5C" LSID="urn:lsid:zoobank.org:act:C0AFCE0D-E1B3-484E-9AF0-14FDF9C1A2B5" httpUri="http://treatment.plazi.org/id/03DC6C76CC2D4C39F7DFFC89FC4CFB5C" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<subSubSection id="C36F8EEBCC2D4C38F7DFFC89FC1EFC34" box="[151,896,870,897]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8BCADD60CC2D4C38F7DFFC89FC1EFC34" blockId="28.[151,987,870,969]" box="[151,896,870,897]" pageId="28" pageNumber="29">
<heading id="D0826A0CCC2D4C38F7DFFC89FC1EFC34" bold="true" box="[151,896,870,897]" fontSize="11" level="2" pageId="28" pageNumber="29" reason="4">
<emphasis id="B9010172CC2D4C38F7DFFC89FC1EFC34" bold="true" box="[151,896,870,897]" pageId="28" pageNumber="29">
<taxonomicName id="4C75A6E3CC2D4C38F7DFFC89FC85FC34" ID-CoL="B8RBD" authority="Lehmann, Zahiri &amp; Husemann" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[151,795,870,897]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="kammeri" status="sp. nov.">
<emphasis id="B9010172CC2D4C38F7DFFC89FE11FC34" bold="true" box="[151,399,870,897]" italics="true" pageId="28" pageNumber="29">Lukeniana kammeri</emphasis>
Lehmann, Zahiri &amp; Husemann
</taxonomicName>
<taxonomicNameLabel id="A232BC09CC2D4C38F46AFC88FC1EFC34" box="[802,896,871,897]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC2D4C38F7DFFC64FBF4FBA4" pageId="28" pageNumber="29" type="description">
<paragraph id="8BCADD60CC2D4C38F7DFFC64FC45FC7C" blockId="28.[151,987,870,969]" pageId="28" pageNumber="29">
<figureCitation id="134EC1E5CC2D4C38F7DFFC64FF75FC10" box="[151,235,907,933]" captionStart="FIGURE 6" captionStartId="37.[151,250,1791,1815]" captionTargetBox="[151,1436,181,1767]" captionTargetId="figure-19@37.[151,1436,181,1767]" captionTargetPageId="37" captionText="FIGURE 6. a. Lukeniana kammeri, holotype, male, Kenya, County Kiambu, Thika; b. L. mbalaensis, paratype, female, Zambia, Northern Province, Mbala; c. L. mbalaensis, holotype, male, Zambia, Northern Province, Mbala; d. L. hausmanni, paratype, male, Zambia, Northern Province, 23 km southeast Mbala; e. L. lenzi, paratype, male, Zambia, Northern Province, Mutinondo Wilderness Area; f. L. robplowesi, holotype, male, Zambia, Northern Province, Mbala; g. L. utaheidenreichae, holotype, male, Mozambique, Manica Province, Barr De Chikamba Real; h. L. stueningi, holotype, male, Zambia, Muchinga Province, Nyika National Park, Manyenjere Forest." figureDoi="http://doi.org/10.5281/zenodo.7840800" httpUri="https://zenodo.org/record/7840800/files/figure.png" pageId="28" pageNumber="29">Figs 6a</figureCitation>
,
<figureCitation id="134EC1E5CC2D4C38F7BFFC64FE81FC10" box="[247,287,907,933]" captionStart="FIGURE 16" captionStartId="67.[151,250,1673,1697]" captionTargetBox="[159,1428,391,1636]" captionTargetId="figure-109@67.[151,1436,381,1649]" captionTargetPageId="67" captionText="FIGURE 16. a. Lukeniana madrandelei, paratype, male, DRC, Katanga Province, Lubumbashi; b. L. friederikebauerae, paratype, male, Zambia, Lusaka Province, Lusaka; c. L. chapmani, holotype, male, Malawi, Southern Region, Mount Mulanje, Likabula; d. L. michaelgrzimeki, holotype, male, Tanzania, Arusha Region, Mount Meru, Momella; e. L. kammeri, holotype, male, Kenya, County Kiambu, Thika; f. L. timdavenporti, holotype, male, Kenya, Rift Valley Province, Mount Elgon National Park, Chepnyalil Cave." figureDoi="http://doi.org/10.5281/zenodo.7840829" httpUri="https://zenodo.org/record/7840829/files/figure.png" pageId="28" pageNumber="29">16e</figureCitation>
<uri id="FFE4D162CC2D4C38F7DFFC41FC45FC7C" box="[151,987,942,969]" pageId="28" pageNumber="29">
urn:lsid:zoobank.org:act:
<uuid id="FFD3E7B5CC2D4C38F6E5FC41FC45FC7C" box="[429,987,942,969]" pageId="28" pageNumber="29">C0AFCE0D-E1B3-484E-9AF0-14FDF9C1A2B5</uuid>
</uri>
</paragraph>
<paragraph id="8BCADD60CC2D4C38F7DFFC18FBF4FBA4" blockId="28.[151,1130,1015,1041]" box="[151,1130,1015,1041]" pageId="28" pageNumber="29">
<materialsCitation id="3B1DD73DCC2D4C38F7DFFC18FBF8FBA4" box="[151,1126,1015,1041]" collectionCode="NMK" country="Kenya" location="National Museums" municipality="Nairobi" pageId="28" pageNumber="29" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CE63C2CC2D4C38F7DFFC18FF51FBA4" box="[151,207,1015,1041]" pageId="28" pageNumber="29">Type</typeStatus>
locality and repository:
<collectingCountry id="F3629DF0CC2D4C38F6A8FC18FDB2FBA4" box="[480,556,1015,1041]" name="Kenya" pageId="28" pageNumber="29">Kenya</collectingCountry>
, the
<location id="8EAA8BBBCC2D4C38F529FC18FCABFBA4" LSID="urn:lsid:plazi:treatment:03DC6C76CC2D4C39F7DFFC89FC4CFB5C:8EAA8BBBCC2D4C38F529FC18FCABFBA4" box="[609,821,1015,1041]" country="Kenya" municipality="Nairobi" name="National Museums" pageId="28" pageNumber="29">National Museums</location>
of
<collectingCountry id="F3629DF0CC2D4C38F412FC18FC38FBA4" box="[858,934,1015,1041]" name="Kenya" pageId="28" pageNumber="29">Kenya</collectingCountry>
,
<collectingMunicipality id="6BAE471ACC2D4C38F4F9FC18FB99FBA4" box="[945,1031,1015,1041]" pageId="28" pageNumber="29">Nairobi</collectingMunicipality>
(
<collectionCode id="ED6445A5CC2D4C38F35EFC18FBC0FBA4" box="[1046,1118,1015,1041]" country="Kenya" httpUri="http://grbio.org/cool/h6fg-emd8" name="National Museums of Kenya" pageId="28" pageNumber="29" type="Museum">NMK</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC2D4C38F7DFFBD1FF6DFB14" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8BCADD60CC2D4C38F7DFFBD1FF6DFB14" blockId="28.[151,1437,1086,2013]" pageId="28" pageNumber="29">
<emphasis id="B9010172CC2D4C38F7DFFBD1FE1AFBED" bold="true" box="[151,388,1086,1112]" pageId="28" pageNumber="29">Material examined.</emphasis>
<materialsCitation id="3B1DD73DCC2D4C38F6C6FBD0FF6DFB14" collectingDate="1950-10" collectionCode="NMK" collectorName="E. Pinhey &amp; I. Lehmann" country="Kenya" county="Colony" location="Thika District" municipality="Thika" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Kiambu" typeStatus="holotype">
<typeStatus id="54CE63C2CC2D4C38F6C6FBD0FE68FBEC" box="[398,502,1087,1113]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="F3629DF0CC2D4C38F501FBD0FD0CFBEC" box="[585,658,1087,1113]" name="Kenya" pageId="28" pageNumber="29">Kenya</collectingCountry>
<collectingCounty id="62ABA5ECCC2D4C38F5D5FBD0FD6FFBEC" box="[669,753,1087,1113]" pageId="28" pageNumber="29">Colony</collectingCounty>
, now
<collectingCountry id="F3629DF0CC2D4C38F472FBD0FC18FBEC" box="[826,902,1087,1113]" name="Kenya" pageId="28" pageNumber="29">Kenya</collectingCountry>
,
<collectingMunicipality id="6BAE471ACC2D4C38F4DCFBD0FC4BFBEC" box="[916,981,1087,1113]" pageId="28" pageNumber="29">Thika</collectingMunicipality>
(until
<locationDeviation id="9A78C442CC2D4C38F36FFBD0FAD2FBEC" box="[1063,1356,1087,1113]" location="Central Province" pageId="28" pageNumber="29">
<quantity id="4C8D7085CC2D4C38F36FFBD0FBE1FBEC" box="[1063,1151,1087,1113]" metricMagnitude="1" metricUnit="m" metricValue="5.11302" pageId="28" pageNumber="29" unit="in" value="2013.0">2013 in</quantity>
Central Province
</locationDeviation>
,
<location id="8EAA8BBBCC2D4C38F213FBD0FF70FBC8" LSID="urn:lsid:plazi:treatment:03DC6C76CC2D4C39F7DFFC89FC4CFB5C:8EAA8BBBCC2D4C38F213FBD0FF70FBC8" country="Kenya" county="Colony" municipality="Thika" name="Thika District" pageId="28" pageNumber="29" stateProvince="Kiambu">Thika District</location>
, now in
<collectingRegion id="49B11382CC2D4C38F61FFB8CFD8DFBC8" box="[343,531,1123,1149]" country="Kenya" name="Kiambu" pageId="28" pageNumber="29">Kiambu County</collectingRegion>
),
<date id="FFCBFBA0CC2D4C38F561FB8CFD56FBC8" box="[553,712,1122,1149]" pageId="28" pageNumber="29" value="1950-10">
<collectingDate id="EF8F0248CC2D4C38F561FB8CFD56FBC8" box="[553,712,1122,1149]" pageId="28" pageNumber="29" value="1950-10">October 1950</collectingDate>
</date>
,
<collectorName id="2680B8B6CC2D4C38F59FFB8CFCD9FBC8" box="[727,839,1123,1149]" pageId="28" pageNumber="29">E. Pinhey</collectorName>
leg., genitalia slide number 29/032016
<collectorName id="2680B8B6CC2D4C38F250FB8CFA02FBC8" box="[1304,1436,1123,1149]" pageId="28" pageNumber="29">I. Lehmann</collectorName>
(
<collectionCode id="ED6445A5CC2D4C38F7D7FB68FF79FB14" box="[159,231,1159,1185]" country="Kenya" httpUri="http://grbio.org/cool/h6fg-emd8" name="National Museums of Kenya" pageId="28" pageNumber="29" type="Museum">NMK</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC2D4C39F78FFB45FE3CFE18" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="description">
<paragraph id="8BCADD60CC2D4C38F78FFB45FCE7FA84" blockId="28.[151,1437,1086,2013]" pageId="28" pageNumber="29">
<emphasis id="B9010172CC2D4C38F78FFB45FEC7FB71" bold="true" box="[199,345,1194,1220]" pageId="28" pageNumber="29">Description.</emphasis>
Male.
<emphasis id="B9010172CC2D4C38F6E0FB44FE7BFB70" box="[424,485,1195,1221]" italics="true" pageId="28" pageNumber="29">Head</emphasis>
: Yellow ochre; eyes light brown with small black spots; antenna 0.44 length of forewing, bipectinate, with branches 4.5 width of shaft and covered with ivory-yellow scales laterally, shaft covered with ivory-yellow scales dorsally; antennal tips not spatulate but equally narrow with one longer acuminate scale, bending towards apex and touching each other; labial palpi ochre.
</paragraph>
<paragraph id="8BCADD60CC2D4C38F78FFAD4FE2DF92C" blockId="28.[151,1437,1086,2013]" pageId="28" pageNumber="29">
<emphasis id="B9010172CC2D4C38F78FFAD4FE86FAE0" box="[199,280,1339,1365]" italics="true" pageId="28" pageNumber="29">Thorax</emphasis>
: Patagia and tegulae with long hair-like cream-coloured scales mixed with ochre, glossy. A small crest of light cream-coloured scales on metathorax. Hindlegs ochre with fine hair-like scales, shiny; one pair of narrow tibial spurs present, outer spur
<emphasis id="B9010172CC2D4C38F6B9FA6CFD8DFA29" box="[497,531,1411,1436]" italics="true" pageId="28" pageNumber="29">ca.</emphasis>
<quantity id="4C8D7085CC2D4C38F552FA6CFDECFA28" box="[538,626,1411,1437]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="28" pageNumber="29" unit="mm" value="1.1">1.1 mm</quantity>
, inner spur
<emphasis id="B9010172CC2D4C38F5BEFA6CFC89FA29" box="[758,791,1411,1436]" italics="true" pageId="28" pageNumber="29">ca.</emphasis>
<quantity id="4C8D7085CC2D4C38F456FA6CFCE9FA28" box="[798,887,1411,1437]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="28" pageNumber="29" unit="mm" value="0.9">0.9 mm</quantity>
. Forewing length
<quantity id="4C8D7085CC2D4C38F30AFA6DFB38FA28" box="[1090,1190,1410,1437]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="28" pageNumber="29" unit="mm" value="12.5">12.5 mm</quantity>
(wingspan
<quantity id="4C8D7085CC2D4C38F260FA6DFA0EFA28" box="[1320,1424,1410,1437]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" pageId="28" pageNumber="29" unit="mm" value="26.5">26.5 mm</quantity>
). Forewing upperside ochre with few faded striae of dark ochre, glossy towards termen, upper basal part of wing dark ochre, costal margin dark ochre without striae; termen without lunules; subterminal line dark ochre, faded from near apex to end of CuA
<subScript id="17F1DF25CC2D4C38F6FBF9EEFE25F9BA" attach="left" box="[435,443,1537,1551]" fontSize="6" pageId="28" pageNumber="29">2</subScript>
; CuA
<subScript id="17F1DF25CC2D4C38F549F9EEFD97F9BA" attach="left" box="[513,521,1537,1551]" fontSize="6" pageId="28" pageNumber="29">2</subScript>
broad, ivory-yellow mixed with pure white, edged by an equally broad band of dark ochre above; all remaining veins not distinctly coloured; cilia long,
<quantity id="4C8D7085CC2D4C38F489F9FCFB86F998" box="[961,1048,1555,1581]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="28" pageNumber="29" unit="mm" value="1.3">1.3 mm</quantity>
, cream-coloured, shiny. Underside of forewing roughly scaled, largely cream-coloured, dark ochre along costa, glossy, subterminal line not visible. Hindwing upperside cream-coloured mixed with pure white, glossy, cilia concolorous with forewing, long,
<quantity id="4C8D7085CC2D4C38F208F9B4FA06F9C0" box="[1344,1432,1627,1653]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="28" pageNumber="29" unit="mm" value="1.3">1.3 mm</quantity>
; underside as in forewing.
</paragraph>
<paragraph id="8BCADD60CC2D4C39F78FF94CFD81FE3C" blockId="28.[151,1437,1086,2013]" lastBlockId="29.[151,1437,151,1257]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<emphasis id="B9010172CC2D4C38F78FF94CFEAFF908" box="[199,305,1699,1725]" italics="true" pageId="28" pageNumber="29">Abdomen</emphasis>
: Mainly ochre mixed with ivory-yellow, shiny; abdominal tuft short,
<emphasis id="B9010172CC2D4C38F321F94CFB14F909" box="[1129,1162,1699,1724]" italics="true" pageId="28" pageNumber="29">ca.</emphasis>
one-fifth of abdominal length. Genitalia (
<figureCitation id="134EC1E5CC2D4C38F62DF928FE5AF954" box="[357,452,1735,1761]" captionStart="FIGURE 16" captionStartId="67.[151,250,1673,1697]" captionTargetBox="[159,1428,391,1636]" captionTargetId="figure-109@67.[151,1436,381,1649]" captionTargetPageId="67" captionText="FIGURE 16. a. Lukeniana madrandelei, paratype, male, DRC, Katanga Province, Lubumbashi; b. L. friederikebauerae, paratype, male, Zambia, Lusaka Province, Lusaka; c. L. chapmani, holotype, male, Malawi, Southern Region, Mount Mulanje, Likabula; d. L. michaelgrzimeki, holotype, male, Tanzania, Arusha Region, Mount Meru, Momella; e. L. kammeri, holotype, male, Kenya, County Kiambu, Thika; f. L. timdavenporti, holotype, male, Kenya, Rift Valley Province, Mount Elgon National Park, Chepnyalil Cave." figureDoi="http://doi.org/10.5281/zenodo.7840829" httpUri="https://zenodo.org/record/7840829/files/figure.png" pageId="28" pageNumber="29">Fig. 16e</figureCitation>
) with uncus lobes rounded, outer edge slightly C-shaped, emargination between lobes small, slightly narrower than one uncus tip, basal edge of uncus lobes rounded at base and strongly bent at middle towards emargination, lobes densely covered with short setae ventrally; gnathos arms broad, long (almost 1.4 basal width of valva), touching semi-transtilla and almost reaching coastal margin of valva, not bent towards uncus; valva large (almost 1.5 size of one uncus lobe) and long, ovoid, broadest at base, costa slightly bent with few very long and strongly sclerotized setae near base of weakly-sclerotized projection; sacculus ending at middle of ventral edge of valva and with few short setae; weakly-sclerotized projection without any prickly appearance, tip rounded, equal in length to thorn-like process below; latter slightly bent, hollow, with a rounded tip, no dots but tiny setae present; edge strongly S-shaped ventrally from base of thorn-like process to point where sacculus normally begins; median sector of valva with only very few scattered short setae on inner side; long, broad, ovoid emargination extending between weakly-sclerotized projection and thorn-like process,
<emphasis id="B9010172CC2C4C39F41DFF30FCE9FF4D" box="[853,887,223,248]" italics="true" pageId="29" pageNumber="30">ca.</emphasis>
50% of length of valva; ventral side of valva not bent at middle. Saccus unusually small, slightly less than half size of juxta, lunular. In front of saccus a broader part of vinculum forming narrow sclerotized plate. Juxta broad with two acuminate tips, each bearing a thorn-like process, emargination between tips deep, 90% of length of juxta. Phallus long, 1.2 length of valva, not bent, straight, bilobed with a cleft at broader end.
</paragraph>
<paragraph id="8BCADD60CC2C4C39F78FFE7CFE3CFE18" blockId="29.[151,1437,151,1257]" box="[199,418,403,429]" pageId="29" pageNumber="30">
<emphasis id="B9010172CC2C4C39F78FFE7CFEBBFE18" bold="true" box="[199,293,403,429]" pageId="29" pageNumber="30">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC2C4C39F78FFE59FAF0FCE8" pageId="29" pageNumber="30" type="diagnosis">
<paragraph id="8BCADD60CC2C4C39F78FFE59FAF0FCE8" blockId="29.[151,1437,151,1257]" pageId="29" pageNumber="30">
<emphasis id="B9010172CC2C4C39F78FFE59FEDCFE65" bold="true" box="[199,322,438,464]" pageId="29" pageNumber="30">Diagnosis.</emphasis>
<taxonomicName id="4C75A6E3CC2C4C39F602FE58FDB4FE64" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[330,554,439,465]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kammeri">
<emphasis id="B9010172CC2C4C39F602FE58FDB4FE64" box="[330,554,439,465]" italics="true" pageId="29" pageNumber="30">Lukeniana kammeri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C75A6E3CC2C4C39F52CFE58FCA2FE64" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[612,828,439,465]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="michaelgrzimeki">
<emphasis id="B9010172CC2C4C39F52CFE58FCA2FE64" box="[612,828,439,465]" italics="true" pageId="29" pageNumber="30">L. michaelgrzimeki</emphasis>
</taxonomicName>
share two features of the male genitalia: (i) the basal edge of the uncus lobes is deeply bent towards a small rounded emargination between them; and (ii) strongly Sshaped edge of the valva extends ventrally from the base of the stout thorn-like process to the point where the sacculus typically ends (viewed posteriorly). The species are easily separated by the absence of long setae on the ventral side of the uncus and on the weakly-sclerotized projection (resulting in a prickly appearance) in
<taxonomicName id="4C75A6E3CC2C4C39F25EFDA8FA0BFDD4" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1302,1429,583,609]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kammeri">
<emphasis id="B9010172CC2C4C39F25EFDA8FA0BFDD4" box="[1302,1429,583,609]" italics="true" pageId="29" pageNumber="30">L. kammeri</emphasis>
</taxonomicName>
. Additionally, the uncus lobes are more rounded, the valva shorter, the sacculus continuous and the aedeagus strongly bent near its narrower end in
<taxonomicName id="4C75A6E3CC2C4C39F6AFFD60FD21FD1C" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[487,703,655,681]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="michaelgrzimeki">
<emphasis id="B9010172CC2C4C39F6AFFD60FD21FD1C" box="[487,703,655,681]" italics="true" pageId="29" pageNumber="30">L. michaelgrzimeki</emphasis>
</taxonomicName>
, whereas in
<taxonomicName id="4C75A6E3CC2C4C39F419FD60FC4CFD1C" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[849,978,655,681]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kammeri">
<emphasis id="B9010172CC2C4C39F419FD60FC4CFD1C" box="[849,978,655,681]" italics="true" pageId="29" pageNumber="30">L. kammeri</emphasis>
</taxonomicName>
the aedeagus is entirely straight and the sacculus ends near the middle of the ventral side of the valva. The latter character state occurs also in
<taxonomicName id="4C75A6E3CC2C4C39F25FFD5CFA03FD79" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[1303,1437,691,716]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="stueningi">
<emphasis id="B9010172CC2C4C39F25FFD5CFA03FD79" box="[1303,1437,691,716]" italics="true" pageId="29" pageNumber="30">L. stueningi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C75A6E3CC2C4C39F781FD38FE34FD44" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[201,426,727,753]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="utaheidenreichae">
<emphasis id="B9010172CC2C4C39F781FD38FE34FD44" box="[201,426,727,753]" italics="true" pageId="29" pageNumber="30">L. utaheidenreichae</emphasis>
</taxonomicName>
. In
<taxonomicName id="4C75A6E3CC2C4C39F693FD38FDFCFD45" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[475,610,727,752]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="stueningi">
<emphasis id="B9010172CC2C4C39F693FD38FDFCFD45" box="[475,610,727,752]" italics="true" pageId="29" pageNumber="30">L. stueningi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C75A6E3CC2C4C39F5D5FD38FCE0FD44" authorityName="Lehmann &amp; Zahiri &amp; Husemann" authorityYear="2023" box="[669,894,727,753]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="utaheidenreichae">
<emphasis id="B9010172CC2C4C39F5D5FD38FCE0FD44" box="[669,894,727,753]" italics="true" pageId="29" pageNumber="30">L. utaheidenreichae</emphasis>
</taxonomicName>
the vinculum is broader in front of the saccus, forming a sclerotized plate. This plate is also present, but much less pronounced, in
<taxonomicName id="4C75A6E3CC2C4C39F37BFD14FB2FFCA0" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1075,1201,763,789]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kammeri">
<emphasis id="B9010172CC2C4C39F37BFD14FB2FFCA0" box="[1075,1201,763,789]" italics="true" pageId="29" pageNumber="30">L. kammeri</emphasis>
</taxonomicName>
.
<taxonomicName id="4C75A6E3CC2C4C39F3F6FD14FA02FCA0" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[1214,1436,763,789]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kammeri">
<emphasis id="B9010172CC2C4C39F3F6FD14FA02FCA0" box="[1214,1436,763,789]" italics="true" pageId="29" pageNumber="30">Lukeniana kammeri</emphasis>
</taxonomicName>
has at leaset three characters unique among
<taxonomicName id="4C75A6E3CC2C4C39F533FCF0FCD2FC8C" box="[635,844,799,825]" pageId="29" pageNumber="30">
<emphasis id="B9010172CC2C4C39F533FCF0FD6FFC8C" box="[635,753,799,825]" italics="true" pageId="29" pageNumber="30">Lukeniana</emphasis>
species
</taxonomicName>
: (i) the valva is large, almost 1.5 the size of one uncus lobe; (ii) the saccus is unusually small, less than half the size of the juxta; and (iii) the saccus is lunule-shaped.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC2C4C39F78FFC89FE77FBEC" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8BCADD60CC2C4C39F78FFC89FB9BFB80" blockId="29.[151,1437,151,1257]" pageId="29" pageNumber="30">
<emphasis id="B9010172CC2C4C39F78FFC89FEFFFC35" bold="true" box="[199,353,870,896]" pageId="29" pageNumber="30">Distribution.</emphasis>
<taxonomicName id="4C75A6E3CC2C4C39F623FC88FDD3FC34" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[363,589,871,897]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kammeri">
<emphasis id="B9010172CC2C4C39F623FC88FDD3FC34" box="[363,589,871,897]" italics="true" pageId="29" pageNumber="30">Lukeniana kammeri</emphasis>
</taxonomicName>
is known only from Thika (102′S, 3707′E; elevation
<emphasis id="B9010172CC2C4C39F3F6FC88FB41FC35" box="[1214,1247,871,896]" italics="true" pageId="29" pageNumber="30">ca.</emphasis>
<quantity id="4C8D7085CC2C4C39F3A1FC89FA09FC34" box="[1257,1431,870,897]" metricMagnitude="3" metricUnit="m" metricValue="1.4845" metricValueMax="1.518" metricValueMin="1.451" pageId="29" pageNumber="30" unit="m" value="1484.5" valueMax="1518.0" valueMin="1451.0">
<geoCoordinate id="EE41BBA7CC2C4C39F3A1FC89FAB9FC34" box="[1257,1319,870,897]" degrees="1,451" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="55" value="1.451">1,451</geoCoordinate>
<geoCoordinate id="EE41BBA7CC2C4C39F26FFC89FAEBFC34" box="[1319,1397,870,897]" degrees="1,518" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="55" value="-1.518">–1,518</geoCoordinate>
m
</quantity>
, average annual rainfall
<quantity id="4C8D7085CC2C4C39F6E8FC65FE61FC10" box="[416,511,906,933]" metricMagnitude="-1" metricUnit="m" metricValue="9.5" pageId="29" pageNumber="30" unit="mm" value="950.0">950 mm</quantity>
)
<emphasis id="B9010172CC2C4C39F547FC64FDAEFC11" box="[527,560,907,932]" italics="true" pageId="29" pageNumber="30">ca.</emphasis>
<quantity id="4C8D7085CC2C4C39F570FC64FDE1FC10" box="[568,639,907,933]" metricMagnitude="4" metricUnit="m" metricValue="3.8" pageId="29" pageNumber="30" unit="km" value="38.0">38 km</quantity>
northeast of Nairobi, near the confluence of the Thika and Chania rivers,
<emphasis id="B9010172CC2C4C39F78DFC40FF79FC7D" box="[197,231,943,968]" italics="true" pageId="29" pageNumber="30">ca.</emphasis>
<quantity id="4C8D7085CC2C4C39F7A5FC40FEAAFC7C" box="[237,308,943,969]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="29" pageNumber="30" unit="km" value="70.0">70 km</quantity>
south of Mount
<collectingCountry id="F3629DF0CC2C4C39F6A5FC40FDA8FC7C" box="[493,566,943,969]" name="Kenya" pageId="29" pageNumber="30">Kenya</collectingCountry>
and
<emphasis id="B9010172CC2C4C39F523FC40FD13FC7D" box="[619,653,943,968]" italics="true" pageId="29" pageNumber="30">ca.</emphasis>
<quantity id="4C8D7085CC2C4C39F5DBFC41FD44FC7C" box="[659,730,942,969]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="29" pageNumber="30" unit="km" value="50.0">50 km</quantity>
east of the southern
<collectingCountry id="F3629DF0CC2C4C39F488FC40FB97FC7C" box="[960,1033,943,969]" name="Kenya" pageId="29" pageNumber="30">Kenya</collectingCountry>
Rift of the EARS, in central
<collectingCountry id="F3629DF0CC2C4C39F204FC40FA07FC7C" box="[1356,1433,943,969]" name="Kenya" pageId="29" pageNumber="30">Kenya</collectingCountry>
. Apart from small forest/woodland remnants (
<bibRefCitation id="EFE4A091CC2C4C39F5EBFC3CFC6DFC58" author="Malombe, I. &amp; Mutangah, J. G." box="[675,1011,979,1005]" pageId="29" pageNumber="30" refId="ref80689" refString="Malombe, I. &amp; Mutangah, J. G. (2004) Assessment of conservation value of natural habitat relics in Thika District. Final Report. National Museums of Kenya, Nairobi, v + 21 pp." type="book" year="2004">Malombe &amp; Mutangah 2004</bibRefCitation>
), most of the natural vegetation was largely destroyed through agriculture, horticulture, and the establishment of sisal, and coffee and pineapple plantations.
<taxonomicName id="4C75A6E3CC2C4C39F794FBF4FE25FB80" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[220,443,1051,1077]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kammeri">
<emphasis id="B9010172CC2C4C39F794FBF4FE25FB80" box="[220,443,1051,1077]" italics="true" pageId="29" pageNumber="30">Lukeniana kammeri</emphasis>
</taxonomicName>
is to be considered an Afromontane linking species.
</paragraph>
<paragraph id="8BCADD60CC2C4C39F78FFBD1FE77FBEC" blockId="29.[151,1437,151,1257]" box="[199,489,1086,1113]" pageId="29" pageNumber="30">
<emphasis id="B9010172CC2C4C39F78FFBD1FEB4FBED" bold="true" box="[199,298,1086,1112]" pageId="29" pageNumber="30">Habitat.</emphasis>
See Appendix 1.
</paragraph>
</subSubSection>
<subSubSection id="C36F8EEBCC2C4C39F78FFB8CFC4CFB5C" pageId="29" pageNumber="30" type="etymology">
<paragraph id="8BCADD60CC2C4C39F78FFB8CFC4CFB5C" blockId="29.[151,1437,151,1257]" pageId="29" pageNumber="30">
<emphasis id="B9010172CC2C4C39F78FFB8CFED0FBC8" bold="true" box="[199,334,1123,1149]" pageId="29" pageNumber="30">Etymology.</emphasis>
<taxonomicName id="4C75A6E3CC2C4C39F61EFB8CFDA8FBC8" authorityName="Lehmann, Zahiri &amp; Husemann" authorityYear="2023" box="[342,566,1123,1149]" class="Insecta" family="Metarbelidae" genus="Lukeniana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kammeri">
<emphasis id="B9010172CC2C4C39F61EFB8CFDA8FBC8" box="[342,566,1123,1149]" italics="true" pageId="29" pageNumber="30">Lukeniana kammeri</emphasis>
</taxonomicName>
is named in honour of the machinist Karl Wilhelm Günther Kammer (1927‒ 2018). The first author is grateful to Günther for walks in the forest close to his house at the Schacksdorf Airport where we discovered a large population (often&gt;
<specimenCount id="9D7316E9CC2C4C39F5F5FB45FCCBFB70" box="[701,853,1194,1221]" pageId="29" pageNumber="30" type="generic">50 specimens</specimenCount>
per day) of the rare butterfly
<taxonomicName id="4C75A6E3CC2C4C39F3E9FB44FEB8FB5C" authority="(Linnaeus, 1771)" baseAuthorityName="Linnaeus" baseAuthorityYear="1771" class="Insecta" family="Nymphalidae" genus="Pyronia" kingdom="Animalia" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="tithonus">
<emphasis id="B9010172CC2C4C39F3E9FB44FAC5FB70" box="[1185,1371,1195,1221]" italics="true" pageId="29" pageNumber="30">Pyronia tithonus</emphasis>
(
<bibRefCitation id="EFE4A091CC2C4C39F222FB44FE80FB5C" author="Linnaeus, C." pageId="29" pageNumber="30" pagination="537" refId="ref80420" refString="Linnaeus, C. (1771) Tithonus PAPILIO. In: Linne, C. A. (Ed.) Mantissa Plantarum - Altera Generum editionis VI &amp; Specierum editionis II. Regni Animalis Appendix. Animalia. Insecta. Impensis Direct. Laurentii Salvii, Holmiae, pp. 537." type="book chapter" year="1771">Linnaeus, 1771</bibRefCitation>
)
</taxonomicName>
during 1974–1982. (See Appendix 1 for the full etymology).
</paragraph>
</subSubSection>
</treatment>
</document>
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