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<document id="E78C37529BE2E2564745B12231E509D1" ID-DOI="10.5852/ejt.2024.921.2413" ID-ISSN="2118-9773" ID-Zenodo-Dep="10590243" ID-ZooBank="80B3C408-DD2C-4373-96AE-15F067D723F8" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1706610496547" checkinUser="plazi" docAuthor="Bockwinkel, Jürgen &amp; Korn, Dieter" docDate="2024" docId="03D987F2292BF642FDCB250F777FF80B" docLanguage="en" docName="EJT.2024.921.1-35.pdf" docOrigin="European Journal of Taxonomy 921 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2413/10641" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Maenioceras Schindewolf 1933" docType="treatment" docVersion="4" lastPageNumber="6" masterDocId="FFE0FF8A292EF647FFFB212F7550FF8C" masterDocTitle="Ammonoids of the Middle Devonian family Maenioceratidae in the Anti-Atlas of Morocco" masterLastPageNumber="35" masterPageNumber="1" pageNumber="6" updateTime="1706631328069" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F8D692A7A3FAB05B09CF8F56FA6AD305">Ammonoids of the Middle Devonian family Maenioceratidae in the Anti-Atlas of Morocco</mods:title>
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<mods:name id="D038DCF3C21258D0AC32E662B39ECACA" type="personal">
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<mods:namePart id="33CEAB2F0B6D01BCAB169A1B87D7AE2C">Bockwinkel, Jürgen</mods:namePart>
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<mods:affiliation id="BA3CEEC4DA5B021E00F2D495C816C4A3">Dechant-Fein-Strasse 22, 51375 Leverkusen, Germany.</mods:affiliation>
<mods:nameIdentifier id="C088ECFB25C1A1CC637EFD4796A8C94E" type="email">j.bockwinkel@t-online.de</mods:nameIdentifier>
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<mods:namePart id="0B18137049EE6011F42A2F06686EB47E">Korn, Dieter</mods:namePart>
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<mods:affiliation id="8D7B8183D4C8E4B75E92C74B9B247A29">Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
<mods:nameIdentifier id="EE91012C2AE5B85F602AFD3686CC44C7" type="email">dieter.korn@mfn.berlin</mods:nameIdentifier>
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<mods:title id="56BEEFD73863C034EB9BFCDC968CC1A7">European Journal of Taxonomy</mods:title>
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<mods:date id="F77F99632210BC1E1A0EEC068A7400F9">2024</mods:date>
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<mods:number id="6B7CB1491AAEEDB3AE523198D0EDA70C">2024-01-30</mods:number>
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<mods:number id="ADDF571F1F8024D0A86D7844C0D2C42C">921</mods:number>
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<subSubSection id="C36A656F292BF642FDCB250F7154FBB6" box="[560,1028,1056,1082]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8BCF36E4292BF642FDCB250F7154FBB6" blockId="5.[560,1028,1056,1082]" box="[560,1028,1056,1082]" pageId="5" pageNumber="6">
<heading id="D0878188292BF642FDCB250F7154FBB6" box="[560,1028,1056,1082]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="6" reason="2">
Genus
<taxonomicName id="4C704D67292BF642FD79250F7154FBB6" ID-CoL="5K26" authority="Schindewolf, 1933" authorityName="Schindewolf" authorityYear="1933" box="[642,1028,1056,1082]" class="Cephalopoda" family="Maenioceratidae" genus="Maenioceras" kingdom="Animalia" order="Agoniatitida" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">
<emphasis id="B904EAF6292BF642FD79250F764EFBB6" bold="true" box="[642,798,1056,1082]" italics="true" pageId="5" pageNumber="6">Maenioceras</emphasis>
Schindewolf, 1933
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36A656F292BF642FF46254D7166FB20" pageId="5" pageNumber="6" type="type_taxon">
<paragraph id="8BCF36E4292BF642FF46254D7408FBF0" blockId="5.[189,344,1122,1148]" box="[189,344,1122,1148]" pageId="5" pageNumber="6">
<heading id="D0878188292BF642FF46254D7408FBF0" bold="true" box="[189,344,1122,1148]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B904EAF6292BF642FF46254D7408FBF0" bold="true" box="[189,344,1122,1148]" pageId="5" pageNumber="6">
<typeStatus id="54CB8846292BF642FF46254D75AAFBF0" box="[189,250,1122,1148]" pageId="5" pageNumber="6">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF36E4292BF642FF4625BD7166FB20" blockId="5.[189,1078,1169,1196]" box="[189,1078,1169,1196]" pageId="5" pageNumber="6">
<taxonomicName id="4C704D67292BF642FF4625BD7668FB20" authority="Sandberger &amp; Sandberger, 1851" authorityName="Sandberger &amp; Sandberger" authorityYear="1851" box="[189,824,1169,1196]" class="Cephalopoda" family="Goniatitidae" genus="Goniatites" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="terebratus">
<emphasis id="B904EAF6292BF642FF4625BD74E6FB20" box="[189,438,1170,1196]" italics="true" pageId="5" pageNumber="6">Goniatites terebratus</emphasis>
Sandberger &amp; Sandberger, 1851
</taxonomicName>
; original designation.
</paragraph>
</subSubSection>
<subSubSection id="C36A656F292BF642FF4625FB703DFAE9" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8BCF36E4292BF642FF4625FB7466FB62" blockId="5.[189,310,1236,1262]" box="[189,310,1236,1262]" pageId="5" pageNumber="6">
<heading id="D0878188292BF642FF4625FB7466FB62" bold="true" box="[189,310,1236,1262]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B904EAF6292BF642FF4625FB7466FB62" bold="true" box="[189,310,1236,1262]" pageId="5" pageNumber="6">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF36E4292BF642FF46242C703DFAE9" blockId="5.[189,1399,1283,1382]" pageId="5" pageNumber="6">
Genus of the family
<taxonomicName id="4C704D67292BF642FE4A242C7724FA91" authorityName="Bogoslovsky" authorityYear="1958" box="[433,628,1283,1309]" class="Cephalopoda" family="Maenioceratidae" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Mollusca" rank="family">Maenioceratidae</taxonomicName>
with ventrolateral grooves; shell usually without constrictions or internal thickenings. External lobe very wide or extremely wide; E
<subScript id="17F434A1292BF642FC2B24157689FAC4" attach="left" box="[976,985,1338,1352]" fontSize="6" pageId="5" pageNumber="6">2</subScript>
lobe usually very shallow, broadly rounded; ventrolateral saddle narrowly rounded or subangular; lateral lobe V-shaped, angular or blunt.
</paragraph>
</subSubSection>
<subSubSection id="C36A656F292BF642FF4624A176FAF926" pageId="5" pageNumber="6" type="taxon_list">
<paragraph id="8BCF36E4292BF642FF4624A174D9FA2B" blockId="5.[189,393,1421,1448]" box="[189,393,1421,1448]" pageId="5" pageNumber="6">
<heading id="D0878188292BF642FF4624A174D9FA2B" bold="true" box="[189,393,1421,1448]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B904EAF6292BF642FF4624A174D9FA2B" bold="true" box="[189,393,1421,1448]" pageId="5" pageNumber="6">Included species</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF36E4292BF642FF4624927037F9CF" blockId="5.[189,1399,1469,1603]" pageId="5" pageNumber="6">
Rhenish Mountains (
<bibRefCitation id="EFE14B15292BF642FE4F2492767DFA54" author="Sandberger G. &amp; Sandberger F." box="[436,813,1469,1496]" pageId="5" pageNumber="6" refId="ref17186" refString="Sandberger G. &amp; Sandberger F. 1850 - 1856. Die Versteinerungen des rheinischen Schichtensystems in Nassau. Mit einer kurzgefassten Geognosie dieses Gebietes und mit steter Berucksichtigung analoger Schichten anderer Lander. Kreidel &amp; Niedner, Wiesbaden. https: // doi. org / 10.5962 / bhl. title. 52349" type="book" year="1850">Sandberger &amp; Sandberger 1850</bibRefCitation>
–1856;
<bibRefCitation id="EFE14B15292BF642FC7D24927111FA54" author="Holzapfel E." box="[902,1089,1469,1496]" pageId="5" pageNumber="6" pagination="1 - 460" refId="ref16671" refString="Holzapfel E. 1895. Das Obere Miteldevon (Schichten mit Stringocephalus Burtini und Maeneceras terebratum) im Rheinischen Gebirge. Abhandlungen der Koniglich Preussischen geologischen Landesanstalt, Neue Folge 16: 1 - 460. Available from https: // www. biodiversitylibrary. org / page / 35736605 [accessed 1 Dec. 2023]." type="journal article" year="1895">Holzapfel 1895</bibRefCitation>
;
<bibRefCitation id="EFE14B15292BF642FBB524927022FA5B" author="Ebbighausen V. &amp; Becker R. T. &amp; Bockwinkel J. &amp; Aboussalam S. Z." box="[1102,1394,1469,1495]" pageId="5" pageNumber="6" pagination="157 - 172" refId="ref16469" refString="Ebbighausen V., Becker R. T., Bockwinkel J. &amp; Aboussalam S. Z. 2007. Givetian (Middle Devonian) brachiopod-goniatite-correlation in the Dra Valley (Anti-Atlas, Morocco) and Bergisch Gladbach- Paffrath Syncline (Rhenish Massif, Germany). Geological Society of London, Special Publications 278: 157 - 172. https: // doi. org / 10.1144 / SP 278.7" type="journal article" year="2007">
Ebbighausen
<emphasis id="B904EAF6292BF642FB142491707BFA5B" box="[1263,1323,1469,1495]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
2007
</bibRefCitation>
;
<bibRefCitation id="EFE14B15292BF642FF4524CE74B9FA77" author="Korn D. &amp; Bockwinkel J." box="[190,489,1505,1531]" pageId="5" pageNumber="6" pagination="1 - 79" refId="ref16990" refString="Korn D. &amp; Bockwinkel J. 2021. The pharciceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea). European Journal of Taxonomy 771: 1 - 79. https: // doi. org / 10.5852 / ejt. 2021.771.1503" type="journal article" year="2021">Korn &amp;Bockwinkel 2021</bibRefCitation>
;this paper):
<taxonomicName id="4C704D67292BF642FD8A24CE718EFA70" authority="Sandberger &amp; Sandberger, 1851" authorityName="Sandberger &amp; Sandberger" authorityYear="1851" box="[625,1246,1505,1532]" class="Cephalopoda" family="Goniatitidae" genus="Goniatites" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="terebratus">
<emphasis id="B904EAF6292BF642FD8A24CE763AFA77" box="[625,874,1505,1531]" italics="true" pageId="5" pageNumber="6">Goniatites terebratus</emphasis>
Sandberger &amp; Sandberger, 1851
</taxonomicName>
;
<taxonomicName id="4C704D67292BF642FB1F24CD749CF9AC" authority="Holzapfel, 1895" authorityName="Holzapfel" authorityYear="1895" class="Cephalopoda" family="Sporadoceratidae" genus="Maeneceras" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="tenue">
<emphasis id="B904EAF6292BF642FB1F24CD75ADF993" italics="true" pageId="5" pageNumber="6">Maeneceras tenue</emphasis>
<bibRefCitation id="EFE14B15292BF642FEF3272A749CF9AC" author="Holzapfel E." box="[264,460,1541,1568]" pageId="5" pageNumber="6" pagination="1 - 460" refId="ref16671" refString="Holzapfel E. 1895. Das Obere Miteldevon (Schichten mit Stringocephalus Burtini und Maeneceras terebratum) im Rheinischen Gebirge. Abhandlungen der Koniglich Preussischen geologischen Landesanstalt, Neue Folge 16: 1 - 460. Available from https: // www. biodiversitylibrary. org / page / 35736605 [accessed 1 Dec. 2023]." type="journal article" year="1895">Holzapfel, 1895</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C704D67292BF642FE27272976F8F9AC" authority="Holzapfel, 1895" authorityName="Holzapfel" authorityYear="1895" box="[476,936,1541,1568]" class="Cephalopoda" family="Sporadoceratidae" genus="Maeneceras" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="decheni">
<emphasis id="B904EAF6292BF642FE2727297788F993" box="[476,728,1541,1567]" italics="true" pageId="5" pageNumber="6">Maeneceras Decheni</emphasis>
<bibRefCitation id="EFE14B15292BF642FD18272A76F8F9AC" author="Holzapfel E." box="[739,936,1541,1568]" pageId="5" pageNumber="6" pagination="1 - 460" refId="ref16671" refString="Holzapfel E. 1895. Das Obere Miteldevon (Schichten mit Stringocephalus Burtini und Maeneceras terebratum) im Rheinischen Gebirge. Abhandlungen der Koniglich Preussischen geologischen Landesanstalt, Neue Folge 16: 1 - 460. Available from https: // www. biodiversitylibrary. org / page / 35736605 [accessed 1 Dec. 2023]." type="journal article" year="1895">Holzapfel, 1895</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C704D67292BF642FC4327297494F9CF" authority="Ebbighausen, Becker &amp; Bockwinkel" authorityName="Ebbighausen, Becker &amp; Bockwinkel" class="Cephalopoda" family="Maenioceratidae" genus="Maenioceras" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="heinorum">
<emphasis id="B904EAF6292BF642FC432729719CF993" box="[952,1228,1541,1567]" italics="true" pageId="5" pageNumber="6">Maenioceras heinorum</emphasis>
Ebbighausen, Becker &amp; Bockwinkel
</taxonomicName>
in
<bibRefCitation id="EFE14B15292BF642FE122706765FF9CF" author="Ebbighausen V. &amp; Becker R. T. &amp; Bockwinkel J. &amp; Aboussalam S. Z." box="[489,783,1577,1603]" pageId="5" pageNumber="6" pagination="157 - 172" refId="ref16469" refString="Ebbighausen V., Becker R. T., Bockwinkel J. &amp; Aboussalam S. Z. 2007. Givetian (Middle Devonian) brachiopod-goniatite-correlation in the Dra Valley (Anti-Atlas, Morocco) and Bergisch Gladbach- Paffrath Syncline (Rhenish Massif, Germany). Geological Society of London, Special Publications 278: 157 - 172. https: // doi. org / 10.1144 / SP 278.7" type="journal article" year="2007">
Ebbighausen
<emphasis id="B904EAF6292BF642FD73270577EBF9CF" box="[648,699,1577,1603]" italics="true" pageId="5" pageNumber="6">et al</emphasis>
., 2007
</bibRefCitation>
;
<taxonomicName id="4C704D67292BF642FCE027057033F9CF" authority="Korn &amp; Bockwinkel, 2021" authorityName="Korn &amp; Bockwinkel" authorityYear="2021" box="[795,1379,1577,1603]" class="Cephalopoda" family="Maenioceratidae" genus="Maenioceras" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="ornatum">
<emphasis id="B904EAF6292BF642FCE02705714FF9CF" box="[795,1055,1578,1603]" italics="true" pageId="5" pageNumber="6">Maenioceras ornatum</emphasis>
<bibRefCitation id="EFE14B15292BF642FBDD27067033F9CF" author="Korn D. &amp; Bockwinkel J." box="[1062,1379,1577,1603]" pageId="5" pageNumber="6" pagination="1 - 79" refId="ref16990" refString="Korn D. &amp; Bockwinkel J. 2021. The pharciceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea). European Journal of Taxonomy 771: 1 - 79. https: // doi. org / 10.5852 / ejt. 2021.771.1503" type="journal article" year="2021">Korn &amp; Bockwinkel, 2021</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCF36E4292BF642FF46274476FAF926" blockId="5.[189,1399,1643,1706]" pageId="5" pageNumber="6">
Anti-Atlas (this paper):
<taxonomicName id="4C704D67292BF642FE1227437611F909" authority="Bockwinkel &amp; Korn, 2024" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[489,833,1643,1669]" class="Cephalopoda" family="Maenioceratidae" genus="Maenioceras" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="afroterebratum" status="sp. nov.">
<emphasis id="B904EAF6292BF642FE1227437611F909" box="[489,833,1643,1669]" italics="true" pageId="5" pageNumber="6">Maenioceras afroterebratum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237578D292BF642FCB4274376E3F90A" box="[847,947,1644,1670]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
;
<taxonomicName id="4C704D67292BF642FC3D27437050F909" authority="Bockwinkel &amp; Korn, 2024" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[966,1280,1643,1669]" class="Cephalopoda" family="Maenioceratidae" genus="Maenioceras" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="mzerrebense" status="sp. nov.">
<emphasis id="B904EAF6292BF642FC3D27437050F909" box="[966,1280,1643,1669]" italics="true" pageId="5" pageNumber="6">Maenioceras mzerrebense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237578D292BF642FAF527437022F90A" box="[1294,1394,1644,1670]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
;
<taxonomicName id="4C704D67292BF642FF4627BF748FF925" authority="Bockwinkel &amp; Korn, 2024" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[189,479,1679,1705]" class="Cephalopoda" family="Maenioceratidae" genus="Maenioceras" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="oufranense" status="sp. nov.">
<emphasis id="B904EAF6292BF642FF4627BF748FF925" box="[189,479,1679,1705]" italics="true" pageId="5" pageNumber="6">Maenioceras oufranense</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237578D292BF642FE1C27BF7715F926" box="[487,581,1680,1706]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
;
<taxonomicName id="4C704D67292BF642FDAA27BF7617F925" authority="Bockwinkel &amp; Korn, 2024" authorityName="Bockwinkel &amp; Korn" authorityYear="2024" box="[593,839,1679,1705]" class="Cephalopoda" family="Maenioceratidae" genus="Maenioceras" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="beckeri" status="sp. nov.">
<emphasis id="B904EAF6292BF642FDAA27BF7617F925" box="[593,839,1679,1705]" italics="true" pageId="5" pageNumber="6">Maenioceras beckeri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237578D292BF642FCB427BF76FAF926" box="[847,938,1680,1706]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C36A656F292BF642FF4627FE777FF80B" pageId="5" pageNumber="6" type="discussion">
<paragraph id="8BCF36E4292BF642FF4627FE747FF967" blockId="5.[189,303,1745,1771]" box="[189,303,1745,1771]" pageId="5" pageNumber="6">
<heading id="D0878188292BF642FF4627FE747FF967" bold="true" box="[189,303,1745,1771]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B904EAF6292BF642FF4627FE747FF967" bold="true" box="[189,303,1745,1771]" pageId="5" pageNumber="6">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF36E4292BF642FF46262E777FF80B" blockId="5.[188,1399,1793,1927]" pageId="5" pageNumber="6">
The North African records of
<taxonomicName id="4C704D67292BF642FDE0262E77E4F896" authorityName="Schindewolf" authorityYear="1933" box="[539,692,1793,1818]" class="Cephalopoda" family="Maenioceratidae" genus="Maenioceras" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Mollusca" rank="genus">
<emphasis id="B904EAF6292BF642FDE0262E77E4F896" box="[539,692,1793,1818]" italics="true" pageId="5" pageNumber="6">Maenioceras</emphasis>
</taxonomicName>
needed revision. Increasing knowledge led to the result that the North African species differ from those described from material of the Rhenish Mountains, although there is close resemblance between some of the species. With respect to the morphological differences, new species are described here.
</paragraph>
</subSubSection>
</treatment>
</document>
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