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<mods:title id="D4CD1F7AEB6D12D72A0A6B0957C78949">Four new species of the genus Eophileurus Arrow, 1908 (Coleoptera: Scarabaeidae Dynastinae) from Thailand and Vietnam, with notes on some Indochinese species</mods:title>
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<taxonomicName id="4C704D0EE85E176F7EFDFF31695AA2D3" authority="Prell, 1913" authorityName="Prell" authorityYear="1913" box="[151,547,223,250]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="confinis">
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<emphasis id="B904EA9FE85E176F7EFDFF316AEFA2D3" bold="true" box="[151,406,223,250]" italics="true" pageId="3" pageNumber="454">Eophileurus confinis</emphasis>
<bibRefCitation id="EFE14B7CE85E176F7FF4FF31695AA2D3" author="Prell, H." box="[414,547,223,249]" pageId="3" pageNumber="454" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
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<subSubSection id="C36A6506E85E176F7EFDFEED6AF6A337" box="[151,399,259,285]" pageId="3" pageNumber="454" type="description">
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(
<figureCitation id="134B2A08E85E176F7EF5FEED6A63A337" box="[159,282,259,285]" captionStart-0="FIGURES 217" captionStart-1="FIGURES 1824" captionStart-2="FIGURES 2531" captionStart-3="FIGURES 3237" captionStartId-0="4.[151,264,1909,1933]" captionStartId-1="5.[151,264,1516,1540]" captionStartId-2="6.[151,264,1464,1488]" captionStartId-3="7.[151,264,1982,2006]" captionTargetBox-0="[222,1366,186,1882]" captionTargetBox-1="[161,1417,211,1471]" captionTargetBox-2="[154,1434,194,1428]" captionTargetBox-3="[198,1395,649,1953]" captionTargetId-0="figure-18@4.[212,1375,181,1885]" captionTargetId-1="figure-18@5.[151,1435,199,1492]" captionTargetId-2="figure-18@6.[151,1435,181,1441]" captionTargetId-3="figure-253@7.[183,1404,633,1959]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURES 217. Habitus of Eophileurus confinis Prell, 1913. 2, male (Mangshi); 3, male (Nabang); 4, female (Rongshuwang); 5, male (Mohan); 67, 11, male (Bo Kluea); 89, female (Jinghong); 10, male (Phrao); 12, male (Bao Loc); 13, female (Bao Loc); 14, male (Khao Kho); 15, male (Umphang); 16, male (Kraburi); 17, female (Kraburi); 26, 8, 1017, dorsal view; 7, 9, ventral view." captionText-1="FIGURES 1824. Holotype of Eophileurus confinis Prell, 1913 (images ©MTD/Olaf Jäger). 18, habitus, dorsal; 19, habitus, lateral; 20, habitus, ventral; 21, parameres, ventral; 22, parameres, apical; 23, parameres, lateral; 24, labels." captionText-2="FIGURES 2531. Holotype of Eophileurus decipiens Prell, 1913 (images ©MTD/Olaf Jäger). 25, habitus, dorsal; 26, habitus, lateral; 27, habitus, ventral; 28, parameres, ventral; 29, parameres, apical; 30, parameres, lateral; 31, labels." captionText-3="FIGURES 3237. Lectotype of Eophileurus siamensis Arrow, 1914 (images by Keita Matsumoto ©Natural History Museum, London). 32, habitus, dorsal; 33, habitus, lateral; 34, habitus, ventral; 35, parameres, apical; 36, parameres, lateral; 37, labels." figureDoi-0="http://doi.org/10.5281/zenodo.6853946" figureDoi-1="http://doi.org/10.5281/zenodo.6853950" figureDoi-2="http://doi.org/10.5281/zenodo.6853952" figureDoi-3="http://doi.org/10.5281/zenodo.6853954" httpUri-0="https://zenodo.org/record/6853946/files/figure.png" httpUri-1="https://zenodo.org/record/6853950/files/figure.png" httpUri-2="https://zenodo.org/record/6853952/files/figure.png" httpUri-3="https://zenodo.org/record/6853954/files/figure.png" pageId="3" pageNumber="454">Figs. 237</figureCitation>
,
<figureCitation id="134B2A08E85E176F7F4FFEED6AFEA337" box="[293,391,259,285]" captionStart-0="FIGURES 117146" captionStart-1="FIGURES 147152" captionStart-2="FIGURES 153164" captionStart-3="FIGURES 165184" captionStartId-0="27.[151,264,1803,1827]" captionStartId-1="28.[151,264,1316,1340]" captionStartId-2="29.[151,264,1313,1337]" captionStartId-3="30.[151,264,1476,1500]" captionTargetBox-0="[169,1424,203,1763]" captionTargetBox-1="[159,1415,199,1278]" captionTargetBox-2="[152,1435,181,1289]" captionTargetBox-3="[157,1431,189,1441]" captionTargetId-0="figure-18@27.[151,1435,181,1780]" captionTargetId-1="figure-18@28.[151,1435,181,1292]" captionTargetId-2="figure-18@29.[151,1435,181,1289]" captionTargetId-3="figure-18@30.[151,1435,181,1453]" captionTargetPageId-0="27" captionTargetPageId-1="28" captionTargetPageId-2="29" captionTargetPageId-3="30" captionText-0="FIGURES 117146. Parameres of Eophileurus confinis Prell, 1913. 117120 from Nabang; 121122, from Lancangjiang Bridge; 123124 from Jinghong; 125128 from Wiang Papao; 129130 from Mueang Chiang Rai; 131132 from Phrao; 133 134 from Axan Mt.; 135136 from Bao Loc; 137138 from Bo Kluea; 139140 from Khao Kho; 141142 from Umphang; 143144 from Bangkok; 145146 from Kraburi; odd number ones, front view; even number ones, lateral view; 117124 without scale bar, the rest with scale bar = 2 mm." captionText-1="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." captionText-2="FIGURES 153164. Parameres of Eophileurus confinis Prell, 1913. 153154 from Lancangjiang Bridge; 155156 from Jinghong; 157158 from Wiang Papao; 159160 from Khao Kho; 161162 from Bangkok; 163164 from Kraburi; odd number ones, front view; even number ones, lateral view; not in scale." captionText-3="FIGURES 165184. Parameres of Eophileurus Arrow, 1908 species. 165166, E. confinis Prell, 1913 (Bao Loc); 167168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169172, E. minor Yang &amp; Pathomwattananurak, new species, 169170, holotype, 171172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175176, holotype, 177178, paratype (Mu Si), 179180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181182, holotype, 183184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6853980" figureDoi-1="http://doi.org/10.5281/zenodo.6853982" figureDoi-2="http://doi.org/10.5281/zenodo.6853984" figureDoi-3="http://doi.org/10.5281/zenodo.6853988" httpUri-0="https://zenodo.org/record/6853980/files/figure.png" httpUri-1="https://zenodo.org/record/6853982/files/figure.png" httpUri-2="https://zenodo.org/record/6853984/files/figure.png" httpUri-3="https://zenodo.org/record/6853988/files/figure.png" pageId="3" pageNumber="454">117166</figureCitation>
)
</paragraph>
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<treatmentCitation id="0AD1109CE85E176F7EFDFEA66975A34A" author="Prell, H." box="[151,524,328,352]" page="114" pageId="3" pageNumber="454" year="1913">
<taxonomicName id="4C704D0EE85E176F7EFDFEA66975A34A" authority="Prell, 1913: 114" authorityName="Prell" authorityPageNumber="114" authorityYear="1913" box="[151,524,328,352]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE85E176F7EFDFEA66A11A34A" box="[151,360,328,352]" italics="true" pageId="3" pageNumber="454">Eophileurus confinis</emphasis>
<bibRefCitation id="EFE14B7CE85E176F7F07FEA66975A34A" author="Prell, H." box="[365,524,328,352]" pageId="3" pageNumber="454" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913: 114</bibRefCitation>
</taxonomicName>
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(
<typeStatus id="54CB882FE85E176F7C72FEA6693CA34A" box="[536,581,328,352]" pageId="3" pageNumber="454">type</typeStatus>
locality: “
<collectingCountry id="F367761DE85E176F7CC1FEA66994A34A" box="[683,749,328,352]" name="China" pageId="3" pageNumber="454">China</collectingCountry>
:
<collectingRegion id="49B4F86FE85E176F7C9FFEA66820A34A" box="[757,857,328,352]" country="China" name="Shanghai" pageId="3" pageNumber="454">Shanghai</collectingRegion>
”), plates 12, fig. 8 (parameres);
<treatmentCitation id="0AD1109CE85E176F7ACAFEA66E38A34A" author="Arrow, G. J." box="[1184,1345,328,352]" page="83" pageId="3" pageNumber="454" year="1937">
<bibRefCitation id="EFE14B7CE85E176F7ACAFEA66E38A34A" author="Arrow, G. J." box="[1184,1345,328,352]" pageId="3" pageNumber="454" pagination="1 - 124" refId="ref16830" refString="Arrow, G. J. (1937) Scarabaeidae: Dynastinae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Pars 156. W. Junk, Gravenhage, pp. 1 - 124." type="book chapter" year="1937">Arrow 1937: 83</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AD1109CE85E176F7B21FEA66A54A354" author="Endrodi, S." page="103" pageId="3" pageNumber="454" year="1977">
<bibRefCitation id="EFE14B7CE85E176F7B21FEA66A54A354" author="Endrodi, S." pageId="3" pageNumber="454" pagination="93 - 134" refId="ref16934" refString="Endrodi, S. (1977) Monographie de Dynastinae VIII Tribus Phileurini. Entomologische Abhandlungen Staatlichen Museum fur Tierkunde Dresden, 41 (4), 93 - 134. https: // doi. org / 10.1515 / 9783112546161" type="journal article" year="1977">Endrődi 1977: 103</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F367761DE85E176F7F50FE886AF3A354" box="[314,394,358,382]" name="Vietnam" pageId="3" pageNumber="454">Annam</collectingCountry>
: Phuc-son; Combodja;
<collectingCountry id="F367761DE85E176F7C1EFE8869D5A354" box="[628,684,358,382]" name="Thailand" pageId="3" pageNumber="454">Siam</collectingCountry>
);
<treatmentCitation id="0AD1109CE85E176F7CD7FE886805A354" author="Endrodi, S." box="[701,892,358,382]" page="666" pageId="3" pageNumber="454" year="1985">
<bibRefCitation id="EFE14B7CE85E176F7CD7FE886805A354" author="Endrodi, S." box="[701,892,358,382]" pageId="3" pageNumber="454" refId="ref17001" refString="Endrodi, S. (1985) The Dynastinae of the World. W. Junk, Dordrecht, 800 pp." type="book" year="1985">Endrődi 1985: 666</bibRefCitation>
</treatmentCitation>
, fig. 2009 (parameres);
<treatmentCitation id="0AD1109CE85E176F7A07FE886E62A354" author="Zhang, Y. - W." box="[1133,1307,358,382]" page="179" pageId="3" pageNumber="454" year="1991">
<bibRefCitation id="EFE14B7CE85E176F7A07FE886E62A354" author="Zhang, Y. - W." box="[1133,1307,358,382]" pageId="3" pageNumber="454" pagination="173 - 188" refId="ref17718" refString="Zhang, Y. - W. (1991) Revision of the family Dynastidae from China (Coleoptera: Scarabaeoidea). Scientific Treatise on Systematic and Evolutionary Zoology, 1, 173 - 188." type="journal article" year="1991">Zhang 1991: 179</bibRefCitation>
</treatmentCitation>
, fig. 3 (inner claw), figs. 45 (parameres);
<treatmentCitation id="0AD1109CE85E176F7F9FFE6B69DCA3B7" author="Krajcik, M." box="[501,677,389,413]" page="63" pageId="3" pageNumber="454" year="2005">
<bibRefCitation id="EFE14B7CE85E176F7F9FFE6B69DCA3B7" author="Krajcik, M." box="[501,677,389,413]" pageId="3" pageNumber="454" pagination="1 - 122" refId="ref17162" refString="Krajcik, M. (2005) Dynastinae of the World Checklist (Coleoptera, Scarabaeidae, Dynastinae). Animma. x, Supplement 2, 1 - 122." type="journal article" year="2005">Krajčik 2005: 63</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AD1109CE85E176F7CDAFE6B688BA3B7" author="Yamaya, S. &amp; Muramoto, R." box="[688,1010,389,413]" page="25" pageId="3" pageNumber="454" year="2008">
<bibRefCitation id="EFE14B7CE85E176F7CDAFE6B688BA3B7" author="Yamaya, S. &amp; Muramoto, R." box="[688,1010,389,413]" pageId="3" pageNumber="454" pagination="1 - 34" refId="ref17677" refString="Yamaya, S. &amp; Muramoto, R. (2008) On the genus Eophileurus Arrow, 1908 (Scarabaeidae, Dynastinae), with descriptions of five new species. Kogane, Supplement 2, 1 - 34." type="journal article" year="2008">Yamaya &amp; Muramoto 2008: 25</bibRefCitation>
</treatmentCitation>
, figs. 4344 (male habitus), fig. 45 (male pronotum), fig. 46 (male elytra), figs. 4748 (parameres);
<treatmentCitation id="0AD1109CE85E176F7D7FFE4A68A9A396" author="Krajcik, M." box="[789,976,420,444]" page="101" pageId="3" pageNumber="454" year="2012">
<bibRefCitation id="EFE14B7CE85E176F7D7FFE4A68A9A396" author="Krajcik, M." box="[789,976,420,444]" pageId="3" pageNumber="454" pagination="1 - 278" refId="ref17193" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. x, Supplement 5, 1 - 278." type="journal article" year="2012">Krajčik 2012: 101</bibRefCitation>
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;
<treatmentCitation id="0AD1109CE85E176F7DB0FE4A6FF5A391" author="Li, J. &amp; Zhang, X. &amp; Lin, L. &amp; Gao, M." box="[986,1164,419,444]" page="67" pageId="3" pageNumber="454" year="2014">
<bibRefCitation id="EFE14B7CE85E176F7DB0FE4A6FF5A391" author="Li, J. &amp; Zhang, X. &amp; Lin, L. &amp; Gao, M." box="[986,1164,419,444]" pageId="3" pageNumber="454" pagination="65 - 68" refId="ref17274" refString="Li, J., Zhang, X., Lin, L. &amp; Gao, M. (2014) Checklist of the Subfamily Dynastinae of China. Le Coleopteriste, 17 (1), 65 - 68." type="journal article" year="2014">
Li
<emphasis id="B904EA9FE85E176F7D9DFE4A6F52A396" box="[1015,1067,420,444]" italics="true" pageId="3" pageNumber="454">et al.</emphasis>
2014: 67
</bibRefCitation>
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;
<treatmentCitation id="0AD1109CE85E176F7AFDFE4A6E07A396" author="Jakl, S. &amp; Zidek, J." box="[1175,1406,419,444]" page="19" pageId="3" pageNumber="454" year="2015">
<bibRefCitation id="EFE14B7CE85E176F7AFDFE4A6E07A396" author="Jakl, S. &amp; Zidek, J." box="[1175,1406,419,444]" pageId="3" pageNumber="454" pagination="14 - 26" refId="ref17117" refString="Jakl, S. &amp; Zidek, J. (2015) Contribution to the knowledge of Oriental and Austro-Papuan Eophileurus Arrow, with descriptions of two new species. Folia Heyrovskyana, Series A, 23 (2), 14 - 26." type="journal article" year="2015">Jákl &amp; Zídek 2015: 19</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCF368DE85E176F7EFDFE2C6E23A3D3" blockId="3.[151,1437,328,720]" pageId="3" pageNumber="454">
<treatmentCitationGroup id="AB6011A3E85E176F7EFDFE2C6E23A3D3" pageId="3" pageNumber="454">
<treatmentCitation id="0AD1109CE85E176F7EFDFE2C6951A3F0" author="Prell, H." box="[151,552,450,474]" page="115" pageId="3" pageNumber="454" year="1913">
<taxonomicName id="4C704D0EE85E176F7EFDFE2C6951A3F0" authority="Prell, 1913: 115" authorityName="Prell" authorityPageNumber="115" authorityYear="1913" box="[151,552,450,474]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="decipiens">
<emphasis id="B904EA9FE85E176F7EFDFE2C6A02A3F0" box="[151,379,450,474]" italics="true" pageId="3" pageNumber="454">Eophileurus decipiens</emphasis>
<bibRefCitation id="EFE14B7CE85E176F7FE9FE2C6951A3F0" author="Prell, H." box="[387,552,450,474]" pageId="3" pageNumber="454" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913: 115</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54CB882FE85E176F7C5DFE2C691DA3F0" box="[567,612,450,474]" pageId="3" pageNumber="454">type</typeStatus>
locality: “Casamanca, Franz,
<collectingCountry id="F367761DE85E176F7DF7FE2C688AA3F0" box="[925,1011,450,474]" name="Senegal" pageId="3" pageNumber="454">Senegal</collectingCountry>
”), plates 12, fig. 13 (parameres);
<treatmentCitation id="0AD1109CE85E176F7B30FE2C6A5AA3D3" author="Arrow, G. J." page="84" pageId="3" pageNumber="454" year="1937">
<bibRefCitation id="EFE14B7CE85E176F7B30FE2C6A5AA3D3" author="Arrow, G. J." pageId="3" pageNumber="454" pagination="1 - 124" refId="ref16830" refString="Arrow, G. J. (1937) Scarabaeidae: Dynastinae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Pars 156. W. Junk, Gravenhage, pp. 1 - 124." type="book chapter" year="1937">Arrow 1937: 84</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AD1109CE85E176F7F44FE0F6AA6A3D3" author="Endrodi, S." box="[302,479,481,505]" page="95" pageId="3" pageNumber="454" year="1977">
<bibRefCitation id="EFE14B7CE85E176F7F44FE0F6AA6A3D3" author="Endrodi, S." box="[302,479,481,505]" pageId="3" pageNumber="454" pagination="93 - 134" refId="ref16934" refString="Endrodi, S. (1977) Monographie de Dynastinae VIII Tribus Phileurini. Entomologische Abhandlungen Staatlichen Museum fur Tierkunde Dresden, 41 (4), 93 - 134. https: // doi. org / 10.1515 / 9783112546161" type="journal article" year="1977">Endrődi 1977: 95</bibRefCitation>
</treatmentCitation>
(synonymized with
<taxonomicName id="4C704D0EE85E176F7CC5FE0F6883A3D3" authority="Prell, 1913" authorityName="Prell" authorityYear="1913" box="[687,1018,481,505]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE85E176F7CC5FE0F68F8A3D3" box="[687,897,481,505]" italics="true" pageId="3" pageNumber="454">Eophileurus confinis</emphasis>
<bibRefCitation id="EFE14B7CE85E176F7DE2FE0F6883A3D3" author="Prell, H." box="[904,1018,481,505]" pageId="3" pageNumber="454" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
</taxonomicName>
);
<treatmentCitation id="0AD1109CE85E176F7A67FE0F6FC1A3D3" author="Krajcik, M." box="[1037,1208,481,505]" page="63" pageId="3" pageNumber="454" year="2005">
<bibRefCitation id="EFE14B7CE85E176F7A67FE0F6FC1A3D3" author="Krajcik, M." box="[1037,1208,481,505]" pageId="3" pageNumber="454" pagination="1 - 122" refId="ref17162" refString="Krajcik, M. (2005) Dynastinae of the World Checklist (Coleoptera, Scarabaeidae, Dynastinae). Animma. x, Supplement 2, 1 - 122." type="journal article" year="2005">Krajčik 2005: 63</bibRefCitation>
</treatmentCitation>
(in synonymy).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCF368DE85E176F7EFDFDEE6E05A01C" blockId="3.[151,1437,328,720]" pageId="3" pageNumber="454">
<treatmentCitationGroup id="AB6011A3E85E176F7EFDFDEE6E05A01C" pageId="3" pageNumber="454">
<treatmentCitation id="0AD1109CE85E176F7EFDFDEE6939A03D" author="Arrow, G. J." box="[151,576,511,536]" page="261" pageId="3" pageNumber="454" year="1914">
<taxonomicName id="4C704D0EE85E176F7EFDFDEE6939A03D" authority="Arrow, 1914: 261" authorityName="Arrow" authorityPageNumber="261" authorityYear="1914" box="[151,576,511,536]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="B904EA9FE85E176F7EFDFDEE6A06A03D" box="[151,383,512,536]" italics="true" pageId="3" pageNumber="454">Eophileurus siamensis</emphasis>
<bibRefCitation id="EFE14B7CE85E176F7FE2FDEE6939A03D" author="Arrow, G. J." box="[392,576,511,536]" pageId="3" pageNumber="454" pagination="257 - 276" refId="ref16776" refString="Arrow, G. J. (1914) Some further notes on Lamellicorn beetles of the sub-family Dynastinae. The Annals and Magazine of natural History, including Zoology, Botany and Geology, 8 (14), 257 - 276. https: // doi. org / 10.1080 / 00222931408693571" type="journal article" year="1914">Arrow, 1914: 261</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54CB882FE85E176F7C3AFDEE6904A032" box="[592,637,512,536]" pageId="3" pageNumber="454">type</typeStatus>
locality: “
<collectingCountry id="F367761DE85E176F7C86FDEE685CA032" box="[748,805,512,536]" name="Thailand" pageId="3" pageNumber="454">Siam</collectingCountry>
:
<collectingRegion id="49B4F86FE85E176F7D58FDEE68E9A032" box="[818,912,512,536]" country="Thailand" name="Krung Thep Mahanakhon" pageId="3" pageNumber="454">Bangkok</collectingRegion>
, Chantabon”), plate 13, fig. 7 (parameres);
<treatmentCitation id="0AD1109CE85E176F7B30FDEE6A5AA01C" author="Arrow, G. J." page="84" pageId="3" pageNumber="454" year="1937">
<bibRefCitation id="EFE14B7CE85E176F7B30FDEE6A5AA01C" author="Arrow, G. J." pageId="3" pageNumber="454" pagination="1 - 124" refId="ref16830" refString="Arrow, G. J. (1937) Scarabaeidae: Dynastinae. In: Schenkling, S. (Ed.), Coleopterorum Catalogus. Pars 156. W. Junk, Gravenhage, pp. 1 - 124." type="book chapter" year="1937">Arrow 1937: 84</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AD1109CE85E176F7F44FDF06A9CA01C" author="Endrodi, S." box="[302,485,542,566]" page="95" pageId="3" pageNumber="454" year="1977">
<bibRefCitation id="EFE14B7CE85E176F7F44FDF06A9CA01C" author="Endrodi, S." box="[302,485,542,566]" pageId="3" pageNumber="454" pagination="93 - 134" refId="ref16934" refString="Endrodi, S. (1977) Monographie de Dynastinae VIII Tribus Phileurini. Entomologische Abhandlungen Staatlichen Museum fur Tierkunde Dresden, 41 (4), 93 - 134. https: // doi. org / 10.1515 / 9783112546161" type="journal article" year="1977">Endrődi, 1977: 95</bibRefCitation>
</treatmentCitation>
(synonymized with of
<taxonomicName id="4C704D0EE85E176F7CBBFDF06F65A01C" authority="Prell, 1913" authorityName="Prell" authorityYear="1913" box="[721,1052,542,566]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE85E176F7CBBFDF068DAA01C" box="[721,931,542,566]" italics="true" pageId="3" pageNumber="454">Eophileurus confinis</emphasis>
<bibRefCitation id="EFE14B7CE85E176F7DC3FDF06F65A01C" author="Prell, H." box="[937,1052,542,566]" pageId="3" pageNumber="454" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
</taxonomicName>
);
<treatmentCitation id="0AD1109CE85E176F7A44FDF06FA3A01C" author="Krajcik, M." box="[1070,1242,542,566]" page="64" pageId="3" pageNumber="454" year="2005">
<bibRefCitation id="EFE14B7CE85E176F7A44FDF06FA3A01C" author="Krajcik, M." box="[1070,1242,542,566]" pageId="3" pageNumber="454" pagination="1 - 122" refId="ref17162" refString="Krajcik, M. (2005) Dynastinae of the World Checklist (Coleoptera, Scarabaeidae, Dynastinae). Animma. x, Supplement 2, 1 - 122." type="journal article" year="2005">Krajčik 2005: 64</bibRefCitation>
</treatmentCitation>
(in synonymy).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCF368DE85E176F7EFDFDD36A57A0B8" blockId="3.[151,1437,328,720]" pageId="3" pageNumber="454">
<treatmentCitationGroup id="AB6011A3E85E176F7EFDFDD36A57A0B8" pageId="3" pageNumber="454">
<treatmentCitation id="0AD1109CE85E176F7EFDFDD369A1A07F" author="Yamaya, S. &amp; Muramoto, R." box="[151,728,573,597]" page="10" pageId="3" pageNumber="454" year="2008">
<taxonomicName id="4C704D0EE85E176F7EFDFDD369A1A07F" authority="Yamaya &amp; Muramoto, 2008: 10" authorityName="Yamaya &amp; Muramoto" authorityPageNumber="10" authorityYear="2008" box="[151,728,573,597]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="takakuwai">
<emphasis id="B904EA9FE85E176F7EFDFDD36AFDA07F" box="[151,388,573,597]" italics="true" pageId="3" pageNumber="454">Eophileurus takakuwai</emphasis>
<bibRefCitation id="EFE14B7CE85E176F7FE6FDD369A1A07F" author="Yamaya, S. &amp; Muramoto, R." box="[396,728,573,597]" pageId="3" pageNumber="454" pagination="1 - 34" refId="ref17677" refString="Yamaya, S. &amp; Muramoto, R. (2008) On the genus Eophileurus Arrow, 1908 (Scarabaeidae, Dynastinae), with descriptions of five new species. Kogane, Supplement 2, 1 - 34." type="journal article" year="2008">Yamaya &amp; Muramoto, 2008: 10</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(
<typeStatus id="54CB882FE85E176F7C82FDD3686CA07F" box="[744,789,573,597]" pageId="3" pageNumber="454">type</typeStatus>
locality: “Dawana,
<collectingCountry id="F367761DE85E176F7D8DFDD36F29A07F" box="[999,1104,573,597]" name="Myanmar" pageId="3" pageNumber="454">Myanmar</collectingCountry>
”), figs. 167168 (male habitus), fig. 169 (male pronotum), 170 (male elytra), figs. 171172 (parameres);
<treatmentCitation id="0AD1109CE85E176F7DDBFDB26F16A05E" author="Krajcik, M." box="[945,1135,604,628]" page="101" pageId="3" pageNumber="454" year="2012">
<bibRefCitation id="EFE14B7CE85E176F7DDBFDB26F16A05E" author="Krajcik, M." box="[945,1135,604,628]" pageId="3" pageNumber="454" pagination="1 - 278" refId="ref17193" refString="Krajcik, M. (2012) Checklist of the World Scarabaeoidea. Animma. x, Supplement 5, 1 - 278." type="journal article" year="2012">Krajčik 2012: 101</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AD1109CE85E176F7A10FDB26E1CA05E" author="Jakl, S. &amp; Zidek, J." box="[1146,1381,603,628]" page="19" pageId="3" pageNumber="454" year="2015">
<bibRefCitation id="EFE14B7CE85E176F7A10FDB26E1CA05E" author="Jakl, S. &amp; Zidek, J." box="[1146,1381,603,628]" pageId="3" pageNumber="454" pagination="14 - 26" refId="ref17117" refString="Jakl, S. &amp; Zidek, J. (2015) Contribution to the knowledge of Oriental and Austro-Papuan Eophileurus Arrow, with descriptions of two new species. Folia Heyrovskyana, Series A, 23 (2), 14 - 26." type="journal article" year="2015">Jákl &amp; Zídek 2015: 19</bibRefCitation>
</treatmentCitation>
;
<emphasis id="B904EA9FE85E176F7B1BFDB26A57A0B8" bold="true" pageId="3" pageNumber="454">new synonym.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCF368DE85E176F7EFDFD776896A0FA" blockId="3.[151,1437,328,720]" pageId="3" pageNumber="454">
<taxonomicName id="4C704D0EE85E176F7EFDFD776A28A09B" authorityName="Endrodi" authorityYear="1978" box="[151,337,665,689]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B904EA9FE85E176F7EFDFD776A28A09B" box="[151,337,665,689]" italics="true" pageId="3" pageNumber="454">Eophileurus malyi</emphasis>
</taxonomicName>
(nec Endrődi):
<bibRefCitation id="EFE14B7CE85E176F7F84FD77685EA09B" author="Yamaya, S. &amp; Muramoto, R." box="[494,807,665,689]" pageId="3" pageNumber="454" pagination="1 - 34" refId="ref17677" refString="Yamaya, S. &amp; Muramoto, R. (2008) On the genus Eophileurus Arrow, 1908 (Scarabaeidae, Dynastinae), with descriptions of five new species. Kogane, Supplement 2, 1 - 34." type="journal article" year="2008">Yamaya &amp; Muramoto 2008: 28</bibRefCitation>
(
<collectingCountry id="F367761DE85E176F7D5EFD7768EBA09B" box="[820,914,665,689]" name="Thailand" pageId="3" pageNumber="454">Thailand</collectingCountry>
,
<collectingCountry id="F367761DE85E176F7DF6FD7768A8A09B" box="[924,977,665,689]" name="Laos" pageId="3" pageNumber="454">Laos</collectingCountry>
), figs. 105106 (male habitus), fig. 107 (male pronotum), fig. 108 (male elytra), figs. 109110 (parameres). Misidentification.
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E85E17697EFDFD176A53A5DD" lastPageId="5" lastPageNumber="456" pageId="3" pageNumber="454" type="materials_examined">
<paragraph id="8BCF368DE85E176F7EFDFD176E11A625" blockId="3.[151,1437,760,2011]" pageId="3" pageNumber="454">
<emphasis id="B904EA9FE85E176F7EFDFD176F7EA138" bold="true" box="[151,1031,760,787]" pageId="3" pageNumber="454">
Type material examined.
<typeStatus id="54CB882FE85E176F7FB0FD17693EA139" box="[474,583,761,787]" pageId="3" pageNumber="454" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C704D0EE85E176F7C10FD166F7EA138" authority="Prell, 1913" authorityName="Prell" authorityYear="1913" box="[634,1031,760,787]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE85E176F7C10FD16680AA138" bold="true" box="[634,883,760,787]" italics="true" pageId="3" pageNumber="454">Eophileurus confinis</emphasis>
<bibRefCitation id="EFE14B7CE85E176F7DEBFD176F7EA138" author="Prell, H." box="[897,1031,760,787]" pageId="3" pageNumber="454" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
</taxonomicName>
</emphasis>
(
<specimenCount id="9D76FD04E85E176F7A74FD176F4CA139" box="[1054,1077,761,787]" pageId="3" pageNumber="454" type="male"></specimenCount>
,
<collectionCode id="ED61AE48E85E176F7A22FD176FF0A139" box="[1096,1161,761,787]" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="3" pageNumber="454">MTD</collectionCode>
,
<figureCitation id="134B2A08E85E176F7AF1FD166E50A139" box="[1179,1321,760,787]" captionStart="FIGURES 1824" captionStartId="5.[151,264,1516,1540]" captionTargetBox="[161,1417,211,1471]" captionTargetId="figure-18@5.[151,1435,199,1492]" captionTargetPageId="5" captionText="FIGURES 1824. Holotype of Eophileurus confinis Prell, 1913 (images ©MTD/Olaf Jäger). 18, habitus, dorsal; 19, habitus, lateral; 20, habitus, ventral; 21, parameres, ventral; 22, parameres, apical; 23, parameres, lateral; 24, labels." figureDoi="http://doi.org/10.5281/zenodo.6853950" httpUri="https://zenodo.org/record/6853950/files/figure.png" pageId="3" pageNumber="454">Figs. 1824</figureCitation>
), labeled: “Shangkai // Typus // Coll. C. Felsche Kauf 20, 1918 // Staatl. Museum für Tierkunde Dresden”, examined through photographs provided by Olaf Jäger.
<emphasis id="B904EA9FE85E176F7C47FCAF68B1A170" bold="true" box="[557,968,832,859]" pageId="3" pageNumber="454">
<typeStatus id="54CB882FE85E176F7C47FCAF69E3A171" box="[557,666,833,859]" pageId="3" pageNumber="454" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C704D0EE85E176F7CD1FCAF68B1A170" authority="Prell, 1913" authorityName="Prell" authorityYear="1913" box="[699,968,832,859]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="decipiens">
<emphasis id="B904EA9FE85E176F7CD1FCAF683CA171" bold="true" box="[699,837,832,859]" italics="true" pageId="3" pageNumber="454">E. decipiens</emphasis>
<bibRefCitation id="EFE14B7CE85E176F7D21FCAF68B1A170" author="Prell, H." box="[843,968,832,859]" pageId="3" pageNumber="454" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
</taxonomicName>
</emphasis>
(
<specimenCount id="9D76FD04E85E176F7DBCFCAF6894A171" box="[982,1005,833,859]" pageId="3" pageNumber="454" type="male"></specimenCount>
,
<collectionCode id="ED61AE48E85E176F7D9DFCAF6F41A171" box="[1015,1080,833,859]" country="Germany" httpUri="http://grbio.org/cool/4j3t-w40g" name="Museum of Zoology Senckenberg Dresden" pageId="3" pageNumber="454">MTD</collectionCode>
,
<figureCitation id="134B2A08E85E176F7A28FCAE6FB1A171" box="[1090,1224,832,859]" captionStart="FIGURES 2531" captionStartId="6.[151,264,1464,1488]" captionTargetBox="[154,1434,194,1428]" captionTargetId="figure-18@6.[151,1435,181,1441]" captionTargetPageId="6" captionText="FIGURES 2531. Holotype of Eophileurus decipiens Prell, 1913 (images ©MTD/Olaf Jäger). 25, habitus, dorsal; 26, habitus, lateral; 27, habitus, ventral; 28, parameres, ventral; 29, parameres, apical; 30, parameres, lateral; 31, labels." figureDoi="http://doi.org/10.5281/zenodo.6853952" httpUri="https://zenodo.org/record/6853952/files/figure.png" pageId="3" pageNumber="454">Figs. 2531</figureCitation>
), labeled: “
<collectingCountry id="F367761DE85E176F7B2BFCAE6EE5A170" box="[1345,1436,832,858]" name="Senegal" pageId="3" pageNumber="454">Senegal</collectingCountry>
// Casamanca // Typus // Coll. C. Felsche Kauf 20, 1918 // Staatl. Museum für Tierkunde Dresden”, examined through photographs provided by Olaf Jäger.
<emphasis id="B904EA9FE85E176F7C5FFC676881A188" bold="true" box="[565,1016,904,931]" pageId="3" pageNumber="454">
<typeStatus id="54CB882FE85E176F7C5FFC6769D2A189" box="[565,683,905,931]" pageId="3" pageNumber="454" type="lectotype">Lectotype</typeStatus>
of
<taxonomicName id="4C704D0EE85E176F7CA5FC676881A188" authority="Arrow, 1914" authorityName="Arrow" authorityYear="1914" box="[719,1016,904,931]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="454" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="B904EA9FE85E176F7CA5FC676819A188" bold="true" box="[719,864,904,931]" italics="true" pageId="3" pageNumber="454">E. siamensis</emphasis>
<bibRefCitation id="EFE14B7CE85E176F7D0FFC666881A188" author="Arrow, G. J." box="[869,1016,904,930]" pageId="3" pageNumber="454" pagination="257 - 276" refId="ref16776" refString="Arrow, G. J. (1914) Some further notes on Lamellicorn beetles of the sub-family Dynastinae. The Annals and Magazine of natural History, including Zoology, Botany and Geology, 8 (14), 257 - 276. https: // doi. org / 10.1080 / 00222931408693571" type="journal article" year="1914">Arrow, 1914</bibRefCitation>
</taxonomicName>
</emphasis>
(hereby designated;
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<figureCitation id="134B2A08E85E176F7B0FFC666B98A1ED" captionStart="FIGURES 3237" captionStartId="7.[151,264,1982,2006]" captionTargetBox="[198,1395,649,1953]" captionTargetId="figure-253@7.[183,1404,633,1959]" captionTargetPageId="7" captionText="FIGURES 3237. Lectotype of Eophileurus siamensis Arrow, 1914 (images by Keita Matsumoto ©Natural History Museum, London). 32, habitus, dorsal; 33, habitus, lateral; 34, habitus, ventral; 35, parameres, apical; 36, parameres, lateral; 37, labels." figureDoi="http://doi.org/10.5281/zenodo.6853954" httpUri="https://zenodo.org/record/6853954/files/figure.png" pageId="3" pageNumber="454">Figs. 3237</figureCitation>
), labeled: “SYN-TYPE //
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. // Bangkok. S. S. Flower. 98-48. //
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<emphasis id="B904EA9FE85E176F7A67FC426E70A1EC" box="[1037,1289,940,966]" italics="true" pageId="3" pageNumber="454">Eophileurus siamensis</emphasis>
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type arrow //
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014380700”, an additional red label will be provided labeling: “
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des. Yang &amp; Pathomwattananurak 2022”, examined through photographs provided by Keita Matsumoto.
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25.
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13.
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19.
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4.
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1315.
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<collectingMethod id="52314E9AE85E176F7D98FB1E6F06A721" box="[1010,1151,1264,1291]" pageId="3" pageNumber="454">by light trap</collectingMethod>
,
<collectorName id="2685535BE85E176F7AE1FB1E6E66A720" box="[1163,1311,1264,1290]" pageId="3" pageNumber="454">Jian-Yue Qiu</collectorName>
&amp;
<collectorName id="2685535BE85E176F7B2EFB1F6EE5A721" box="[1348,1436,1265,1291]" pageId="3" pageNumber="454">Hao Xu</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B183CD0E85E176F7EBAFAFB6A5DA778" ID-GBIF-Occurrence="3863916363" collectingDate="2009-06-01" collectingDateMax="2009-06-02" collectingDateMin="2009-06-01" collectionCode="CWXB, VI" collectorName="WenXuan Bi" country="China" county="Mengla County" elevation="600" location="Menglun Town" municipality="Xipian" pageId="3" pageNumber="454" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D76FD04E85E176F7EBAFAFB6B8AA705" box="[208,243,1301,1327]" pageId="3" pageNumber="454" type="male">1♂</specimenCount>
(
<collectionCode id="ED61AE48E85E176F7F6EFAFB6A24A705" box="[260,349,1301,1327]" pageId="3" pageNumber="454">CWXB</collectionCode>
),
<collectingMunicipality id="6BABACF7E85E176F7F18FAFA6AB8A704" box="[370,449,1300,1326]" pageId="3" pageNumber="454">Xipian</collectingMunicipality>
,
<location id="8EAF6056E85E176F7FA7FAFA6905A705" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7FA7FAFA6905A705" box="[461,636,1300,1327]" country="China" county="Mengla County" municipality="Xipian" name="Menglun Town" pageId="3" pageNumber="454">Menglun Town</location>
,
<collectingCounty id="62AE4E01E85E176F7CE2FAFA6843A705" box="[648,826,1300,1327]" pageId="3" pageNumber="454">Mengla County</collectingCounty>
,
<location id="8EAF6056E85E176F7D2CFAFA6F4FA704" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7D2CFAFA6F4FA704" box="[838,1078,1300,1326]" country="China" county="Mengla County" municipality="Xipian" name="Xishuangbanna Pref." pageId="3" pageNumber="454">Xishuangbanna Pref.</location>
, alt.
<quantity id="4C889B68E85E176F7A07FAFA6FCEA705" box="[1133,1207,1300,1327]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="454" unit="m" value="600.0">
<elevation id="005DD1BEE85E176F7A07FAFA6FCEA705" box="[1133,1207,1300,1327]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="454" unit="m" value="600.0">600 m</elevation>
</quantity>
,
<date id="FFCE104DE85E176F7AAEFAFB6E2DA705" box="[1220,1364,1301,1327]" pageId="3" pageNumber="454" value="2009-06-01" valueMax="2009-06-02" valueMin="2009-06-01">
<collectingDate id="EF8AE9A5E85E176F7AAEFAFB6E2DA705" box="[1220,1364,1301,1327]" pageId="3" pageNumber="454" value="2009-06-01" valueMax="2009-06-02" valueMin="2009-06-01">
12.
<collectionCode id="ED61AE48E85E176F7A9AFAFB6E68A705" box="[1264,1297,1301,1327]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="454">VI</collectionCode>
.2009
</collectingDate>
</date>
,
<collectorName id="2685535BE85E176F7B0AFAFB6B8BA778" pageId="3" pageNumber="454">WenXuan Bi</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B183CD0E85E176F7F47FAD768A7A75C" ID-GBIF-Occurrence="3863916387" collectingDate="2021-04-20" collectionCode="CQZY" collectorName="Ce Hui" country="China" county="Menghai County" elevation="761" latitude="21.938622" location="Manbonanga" longLatPrecision="1" longitude="100.661804" municipality="Near National Highway" pageId="3" pageNumber="454" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D76FD04E85E176F7F47FAD76A28A779" box="[301,337,1337,1363]" pageId="3" pageNumber="454" type="male">1♂</specimenCount>
,
<specimenCount id="9D76FD04E85E176F7F36FAD76A06A779" box="[348,383,1337,1363]" pageId="3" pageNumber="454" type="female">1♀</specimenCount>
(
<collectionCode id="ED61AE48E85E176F7FE1FAD76AA4A779" box="[395,477,1337,1363]" pageId="3" pageNumber="454">CQZY</collectionCode>
),
<geoCoordinate id="EE44504AE85E176F7F84FAD669FEA778" box="[494,647,1336,1362]" degrees="21.938621" direction="north" orientation="latitude" pageId="3" pageNumber="454" precision="1" value="21.938622">21.938621°N</geoCoordinate>
,
<geoCoordinate id="EE44504AE85E176F7CFAFAD6684AA778" box="[656,819,1336,1362]" degrees="100.661806" direction="east" orientation="longitude" pageId="3" pageNumber="454" precision="1" value="100.661804">100.661806°E</geoCoordinate>
,
<collectingMunicipality id="6BABACF7E85E176F7D57FAD76F3DA778" box="[829,1092,1336,1363]" pageId="3" pageNumber="454">Near National Highway</collectingMunicipality>
214,
<quantity id="4C889B68E85E176F7A17FAD66FCDA778" box="[1149,1204,1336,1363]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="454" unit="km" value="5.0">5 km</quantity>
east to
<location id="8EAF6056E85E176F7B69FAD66EE1A778" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7B69FAD66EE1A778" box="[1283,1432,1336,1362]" country="China" county="Menghai County" latitude="21.938622" longLatPrecision="1" longitude="100.661804" municipality="Near National Highway" name="Manbonanga" pageId="3" pageNumber="454">Manbonanga</location>
,
<collectingCounty id="62AE4E01E85E176F7EFDFAB26A2CA75D" box="[151,341,1372,1399]" pageId="3" pageNumber="454">Menghai County</collectingCounty>
,
<location id="8EAF6056E85E176F7F35FAB26935A75C" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7F35FAB26935A75C" box="[351,588,1372,1398]" country="China" county="Menghai County" latitude="21.938622" longLatPrecision="1" longitude="100.661804" municipality="Near National Highway" name="Xishuangbanna Pref." pageId="3" pageNumber="454">Xishuangbanna Pref.</location>
, alt.
<quantity id="4C889B68E85E176F7CEAFAB269B1A75D" box="[640,712,1372,1399]" metricMagnitude="2" metricUnit="m" metricValue="7.609999999999999" pageId="3" pageNumber="454" unit="m" value="761.0">
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</quantity>
,
<date id="FFCE104DE85E176F7CB9FAB36828A75D" box="[723,849,1373,1399]" pageId="3" pageNumber="454" value="2021-04-20">
<collectingDate id="EF8AE9A5E85E176F7CB9FAB36828A75D" box="[723,849,1373,1399]" pageId="3" pageNumber="454" value="2021-04-20">20.IV.2021</collectingDate>
</date>
,
<collectorName id="2685535BE85E176F7D36FAB368D5A75C" box="[860,940,1372,1399]" pageId="3" pageNumber="454">Ce Hui</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B183CD0E85E176F7D83FAB36E16A7B0" ID-GBIF-Occurrence="3863916379" collectingDate="2009-05-04" collectionCode="SHNU, V" collectorName="Jia-Yao Hu &amp; Zi-Wei Yin" country="China" county="Nabanhe Nature Reserve" elevation="700" location="Jinghong City" municipality="Manfei" pageId="3" pageNumber="454" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D76FD04E85E176F7D83FAB36F75A75D" box="[1001,1036,1373,1399]" pageId="3" pageNumber="454" type="female">1♀</specimenCount>
(
<collectionCode id="ED61AE48E85E176F7A70FAB36F15A75D" box="[1050,1132,1373,1399]" pageId="3" pageNumber="454">SHNU</collectionCode>
),
<collectingCounty id="62AE4E01E85E176F7A15FAB26EE1A75D" box="[1151,1432,1372,1399]" pageId="3" pageNumber="454">Nabanhe Nature Reserve</collectingCounty>
,
<collectingMunicipality id="6BABACF7E85E176F7EFDFA6E6B92A7B0" box="[151,235,1408,1434]" pageId="3" pageNumber="454">Manfei</collectingMunicipality>
,
<location id="8EAF6056E85E176F7E9FFA6E6AEAA7B0" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7E9FFA6E6AEAA7B0" box="[245,403,1408,1434]" country="China" county="Nabanhe Nature Reserve" municipality="Manfei" name="Jinghong City" pageId="3" pageNumber="454">Jinghong City</location>
,
<location id="8EAF6056E85E176F7FF7FA6E69F3A7B0" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7FF7FA6E69F3A7B0" box="[413,650,1408,1434]" country="China" county="Nabanhe Nature Reserve" municipality="Manfei" name="Xishuangbanna Pref." pageId="3" pageNumber="454">Xishuangbanna Pref.</location>
, alt.
<quantity id="4C889B68E85E176F7CD7FA6F687DA7B1" box="[701,772,1409,1435]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="454" unit="m" value="700.0">
<elevation id="005DD1BEE85E176F7CD7FA6F687DA7B1" box="[701,772,1409,1435]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="454" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<date id="FFCE104DE85E176F7D65FA6F68FCA7B1" box="[783,901,1409,1435]" pageId="3" pageNumber="454" value="2009-05-04">
<collectingDate id="EF8AE9A5E85E176F7D65FA6F68FCA7B1" box="[783,901,1409,1435]" pageId="3" pageNumber="454" value="2009-05-04">
04.
<collectionCode id="ED61AE48E85E176F7D5AFA6F683CA7B1" box="[816,837,1409,1435]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="454" type="Museum">V</collectionCode>
.2009
</collectingDate>
</date>
,
<collectingMethod id="52314E9AE85E176F7DE5FA6E6F61A7B1" box="[911,1048,1408,1435]" pageId="3" pageNumber="454">by light trap</collectingMethod>
,
<collectorName id="2685535BE85E176F7A48FA6E6FE6A7B1" box="[1058,1183,1408,1435]" pageId="3" pageNumber="454">Jia-Yao Hu</collectorName>
&amp;
<collectorName id="2685535BE85E176F7AAAFA6E6E45A7B0" box="[1216,1340,1408,1434]" pageId="3" pageNumber="454">Zi-Wei Yin</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B183CD0E85E176F7B13FA6F6E28A794" ID-GBIF-Occurrence="3863916374" collectingDate="2017-06-01" collectionCode="CQZY, VI" collectorName="Zi-Chun Xiong" country="China" county="Near Lancangjiang Brigde" elevation="1400" location="Lincang City" municipality="Yun county" pageId="3" pageNumber="454" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D76FD04E85E176F7B13FA6F6EE5A7B1" box="[1401,1436,1409,1435]" pageId="3" pageNumber="454" type="female">1♀</specimenCount>
(
<collectionCode id="ED61AE48E85E176F7EF5FA4B6B88A795" box="[159,241,1445,1471]" pageId="3" pageNumber="454">CQZY</collectionCode>
),
<collectingCounty id="62AE4E01E85E176F7F6EFA4B6949A794" box="[260,560,1444,1471]" pageId="3" pageNumber="454">Near Lancangjiang Brigde</collectingCounty>
,
<collectingMunicipality id="6BABACF7E85E176F7C50FA4B69C2A795" box="[570,699,1445,1471]" pageId="3" pageNumber="454">Yun county</collectingMunicipality>
,
<location id="8EAF6056E85E176F7CADFA4A6823A794" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7CADFA4A6823A794" box="[711,858,1444,1470]" country="China" county="Near Lancangjiang Brigde" municipality="Yun county" name="Lincang City" pageId="3" pageNumber="454">Lincang City</location>
, alt.
<quantity id="4C889B68E85E176F7DFAFA4B689EA795" box="[912,999,1445,1471]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="454" unit="m" value="1400.0">
<elevation id="005DD1BEE85E176F7DFAFA4B689EA795" box="[912,999,1445,1471]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="454" unit="m" value="1400.0">1400 m</elevation>
</quantity>
,
<date id="FFCE104DE85E176F7D99FA4B6F1EA795" box="[1011,1127,1445,1471]" pageId="3" pageNumber="454" value="2017-06-01">
<collectingDate id="EF8AE9A5E85E176F7D99FA4B6F1EA795" box="[1011,1127,1445,1471]" pageId="3" pageNumber="454" value="2017-06-01">
1.
<collectionCode id="ED61AE48E85E176F7A6AFA4B6F5BA795" box="[1024,1058,1445,1471]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="454">VI</collectionCode>
.2017
</collectingDate>
</date>
,
<collectorName id="2685535BE85E176F7A18FA4A6E67A794" box="[1138,1310,1444,1470]" pageId="3" pageNumber="454">Zi-Chun Xiong</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B183CD0E85E176F7B37FA4B6924A42C" ID-GBIF-Occurrence="3863916368" collectingDate="2017-06-01" collectionCode="CQZY, VI" collectorName="Zi-Chun Xiong" country="China" county="Lancangjiang Brigde" elevation="1400" location="Lincang City" municipality="Yun county" pageId="3" pageNumber="454" specimenCode="F1" specimenCount="4" specimenCount-female="1" specimenCount-male="3">
<specimenCount id="9D76FD04E85E176F7B37FA4B6EE1A795" box="[1373,1432,1445,1471]" pageId="3" pageNumber="454" type="male">3♂♂</specimenCount>
,
<specimenCount id="9D76FD04E85E176F7EFDFA276BC3A7C9" box="[151,186,1481,1507]" pageId="3" pageNumber="454" type="female">1♀</specimenCount>
(
<collectionCode id="ED61AE48E85E176F7EA1FA276A64A7C9" box="[203,285,1481,1507]" pageId="3" pageNumber="454">CQZY</collectionCode>
),
<specimenCode id="DBD69EF6E85E176F7F58FA276A36A7C9" box="[306,335,1481,1507]" collectionCode="CQZY" pageId="3" pageNumber="454">F1</specimenCode>
generation of female from locality close to
<collectingCounty id="62AE4E01E85E176F7D20FA266F43A7C8" box="[842,1082,1480,1506]" pageId="3" pageNumber="454">Lancangjiang Brigde</collectingCounty>
,
<collectingMunicipality id="6BABACF7E85E176F7A2CFA276FB0A7C9" box="[1094,1225,1481,1507]" pageId="3" pageNumber="454">Yun county</collectingMunicipality>
,
<location id="8EAF6056E85E176F7ABCFA266E15A7C8" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7ABCFA266E15A7C8" box="[1238,1388,1480,1506]" country="China" county="Lancangjiang Brigde" municipality="Yun county" name="Lincang City" pageId="3" pageNumber="454">Lincang City</location>
, alt.
<quantity id="4C889B68E85E176F7EFDFA036B96A42D" box="[151,239,1517,1543]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="454" unit="m" value="1400.0">
<elevation id="005DD1BEE85E176F7EFDFA036B96A42D" box="[151,239,1517,1543]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="454" unit="m" value="1400.0">1400 m</elevation>
</quantity>
,
<date id="FFCE104DE85E176F7E96FA036A09A42D" box="[252,368,1517,1543]" pageId="3" pageNumber="454" value="2017-06-01">
<collectingDate id="EF8AE9A5E85E176F7E96FA036A09A42D" box="[252,368,1517,1543]" pageId="3" pageNumber="454" value="2017-06-01">
1.
<collectionCode id="ED61AE48E85E176F7F63FA036A52A42D" box="[265,299,1517,1543]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="454">VI</collectionCode>
.2017
</collectingDate>
</date>
,
<collectorName id="2685535BE85E176F7F17FA026953A42C" box="[381,554,1516,1542]" pageId="3" pageNumber="454">Zi-Chun Xiong</collectorName>
leg.
</materialsCitation>
, adult emerged in
<date id="FFCE104DE85E176F7D5AFA0368DAA42D" box="[816,931,1517,1543]" pageId="3" pageNumber="454" value="2017-08">VIII.2017</date>
;
<materialsCitation id="3B183CD0E85E176F7DC5FA03680FA400" ID-GBIF-Occurrence="3863916324" collectingDate="2017-05-11" collectionCode="CQZY, V" collectorName="Yu-Tang Wang" country="China" county="Yingjiang County" location="Nabang Town" municipality="Rongshuwang" pageId="3" pageNumber="454" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D76FD04E85E176F7DC5FA0368ABA42D" box="[943,978,1517,1543]" pageId="3" pageNumber="454" type="male">1♂</specimenCount>
(
<collectionCode id="ED61AE48E85E176F7D88FA036F4DA42D" box="[994,1076,1517,1543]" pageId="3" pageNumber="454">CQZY</collectionCode>
),
<collectingMunicipality id="6BABACF7E85E176F7A22FA026F92A42C" box="[1096,1259,1516,1542]" pageId="3" pageNumber="454">Rongshuwang</collectingMunicipality>
,
<location id="8EAF6056E85E176F7A9DFA026EE1A42D" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7A9DFA026EE1A42D" box="[1271,1432,1516,1543]" country="China" county="Yingjiang County" municipality="Rongshuwang" name="Nabang Town" pageId="3" pageNumber="454">Nabang Town</location>
,
<collectingCounty id="62AE4E01E85E176F7EFDF9FE6A1AA401" box="[151,355,1552,1579]" pageId="3" pageNumber="454">Yingjiang County</collectingCounty>
,
<location id="8EAF6056E85E176F7F05F9FE6973A400" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7F05F9FE6973A400" box="[367,522,1552,1578]" country="China" county="Yingjiang County" municipality="Rongshuwang" name="Dehong Pref." pageId="3" pageNumber="454">Dehong Pref.</location>
,
<date id="FFCE104DE85E176F7C7DF9FF69F2A401" box="[535,651,1553,1579]" pageId="3" pageNumber="454" value="2017-05-11">
<collectingDate id="EF8AE9A5E85E176F7C7DF9FF69F2A401" box="[535,651,1553,1579]" pageId="3" pageNumber="454" value="2017-05-11">
11.
<collectionCode id="ED61AE48E85E176F7C5EF9FF6933A401" box="[564,586,1553,1579]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="454" type="Museum">V</collectionCode>
.2017
</collectingDate>
</date>
,
<collectorName id="2685535BE85E176F7CF2F9FF6838A401" box="[664,833,1553,1579]" pageId="3" pageNumber="454">Yu-Tang Wang</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B183CD0E85E176F7DEEF9FF69E6A464" ID-GBIF-Occurrence="3863916325" collectingDate="2009-05-19" collectionCode="CQZY, V" collectorName="Hai-Cheng Shan" country="China" county="Yingjiang County" location="Dehong Pref." municipality="Nabang Town" pageId="3" pageNumber="454" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D76FD04E85E176F7DEEF9FF68DEA401" box="[900,935,1553,1579]" pageId="3" pageNumber="454" type="male">1♂</specimenCount>
(
<collectionCode id="ED61AE48E85E176F7DD2F9FF6F73A401" box="[952,1034,1553,1579]" pageId="3" pageNumber="454">CQZY</collectionCode>
),
<collectingMunicipality id="6BABACF7E85E176F7A75F9FE6FB8A401" box="[1055,1217,1552,1579]" pageId="3" pageNumber="454">Nabang Town</collectingMunicipality>
,
<collectingCounty id="62AE4E01E85E176F7AA7F9FE6EE0A401" box="[1229,1433,1552,1579]" pageId="3" pageNumber="454">Yingjiang County</collectingCounty>
,
<location id="8EAF6056E85E176F7EFDF9DA6A57A464" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7EFDF9DA6A57A464" box="[151,302,1588,1614]" country="China" county="Yingjiang County" municipality="Nabang Town" name="Dehong Pref." pageId="3" pageNumber="454">Dehong Pref.</location>
,
<date id="FFCE104DE85E176F7F5DF9DB6AD5A465" box="[311,428,1589,1615]" pageId="3" pageNumber="454" value="2009-05-19">
<collectingDate id="EF8AE9A5E85E176F7F5DF9DB6AD5A465" box="[311,428,1589,1615]" pageId="3" pageNumber="454" value="2009-05-19">
19.
<collectionCode id="ED61AE48E85E176F7F3FF9DB6A12A465" box="[341,363,1589,1615]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="454" type="Museum">V</collectionCode>
.2009
</collectingDate>
</date>
,
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leg.
</materialsCitation>
;
<materialsCitation id="3B183CD0E85E176F7CC3F9DB6FB4A458" ID-GBIF-Occurrence="3863916320" collectingDate="2016-09-22" collectionCode="CQZY" collectorName="Xiao-Dong Yang" country="China" county="Yingjiang County" elevation="530" location="Nabang Town" municipality="Nabang Hydropower Station" pageId="3" pageNumber="454" specimenCount="1" specimenCount-male="1">
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),
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<location id="8EAF6056E85E176F7E9DF9B66AE9A458" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7E9DF9B66AE9A458" box="[247,400,1624,1650]" country="China" county="Yingjiang County" municipality="Nabang Hydropower Station" name="Dehong Pref." pageId="3" pageNumber="454">Dehong Pref.</location>
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<quantity id="4C889B68E85E176F7FACF9B66969A459" box="[454,528,1624,1651]" metricMagnitude="2" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="454" unit="m" value="530.0">
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,
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<collectingDate id="EF8AE9A5E85E176F7C76F9B669E7A458" box="[540,670,1624,1650]" pageId="3" pageNumber="454" value="2016-09-22">22.IX.2016</collectingDate>
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, collected during the night,
<collectorName id="2685535BE85E176F7DB1F9B66FE0A459" box="[987,1177,1624,1651]" pageId="3" pageNumber="454">Xiao-Dong Yang</collectorName>
leg.
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;
<materialsCitation id="3B183CD0E85E176F7AB0F9B76BBAA490" ID-GBIF-Occurrence="3863916334" collectingDate="2019-10-26" collectionCode="CQZY" collectorName="Chen Zhang" country="China" county="Mangshi County" elevation="827" latitude="24.36375" location="Dehong Pref." longLatPrecision="1" longitude="98.476" municipality="Rescue Centre" pageId="3" pageNumber="454" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
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<specimenCount id="9D76FD04E85E176F7B66F9B76E57A459" box="[1292,1326,1625,1651]" pageId="3" pageNumber="454" type="female">1♀</specimenCount>
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),
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<collectingMunicipality id="6BABACF7E85E176F7F8FF99369F5A4BD" box="[485,652,1661,1687]" pageId="3" pageNumber="454">Rescue Centre</collectingMunicipality>
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<location id="8EAF6056E85E176F7D08F9926885A4BC" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7D08F9926885A4BC" box="[866,1020,1660,1686]" country="China" county="Mangshi County" latitude="24.36375" longLatPrecision="1" longitude="98.476" municipality="Rescue Centre" name="Dehong Pref." pageId="3" pageNumber="454">Dehong Pref.</location>
, alt.
<quantity id="4C889B68E85E176F7A5EF9936F07A4BD" box="[1076,1150,1661,1687]" metricMagnitude="2" metricUnit="m" metricValue="8.27" pageId="3" pageNumber="454" unit="m" value="827.0">
<elevation id="005DD1BEE85E176F7A5EF9936F07A4BD" box="[1076,1150,1661,1687]" metricMagnitude="2" metricUnit="m" metricValue="8.27" pageId="3" pageNumber="454" unit="m" value="827.0">827 m</elevation>
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,
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<collectingDate id="EF8AE9A5E85E176F7AE1F9926E7CA4BC" box="[1163,1285,1660,1686]" pageId="3" pageNumber="454" value="2019-10-26">26.X.2019</collectingDate>
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,
<collectorName id="2685535BE85E176F7B7BF9926EE4A4BC" box="[1297,1437,1660,1686]" pageId="3" pageNumber="454">Chen Zhang</collectorName>
leg.
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;
<materialsCitation id="3B183CD0E85E176F7EA5F94F695EA4F4" ID-GBIF-Occurrence="3863916351" collectingDate="2017-10-05" collectionCode="CQZY" collectorName="Yi-Zhou Liu" country="China" county="Yingjiang County" elevation="931" latitude="24.667591" location="Nabang Town" longLatPrecision="1" longitude="97.59848" municipality="Rongshuwang" pageId="3" pageNumber="454" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D76FD04E85E176F7EA5F94F6B8BA491" box="[207,242,1697,1723]" pageId="3" pageNumber="454" type="female">1♀</specimenCount>
(
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),
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,
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,
<collectingMunicipality id="6BABACF7E85E176F7CC1F94E6837A490" box="[683,846,1696,1722]" pageId="3" pageNumber="454">Rongshuwang</collectingMunicipality>
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<location id="8EAF6056E85E176F7D33F94E6881A491" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7D33F94E6881A491" box="[857,1016,1696,1723]" country="China" county="Yingjiang County" latitude="24.667591" longLatPrecision="1" longitude="97.59848" municipality="Rongshuwang" name="Nabang Town" pageId="3" pageNumber="454">Nabang Town</location>
,
<collectingCounty id="62AE4E01E85E176F7A69F94E6FB5A491" box="[1027,1228,1696,1723]" pageId="3" pageNumber="454">Yingjiang County</collectingCounty>
,
<location id="8EAF6056E85E176F7ABCF94E6E16A490" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7ABCF94E6E16A490" box="[1238,1391,1696,1722]" country="China" county="Yingjiang County" latitude="24.667591" longLatPrecision="1" longitude="97.59848" municipality="Rongshuwang" name="Dehong Pref." pageId="3" pageNumber="454">Dehong Pref.</location>
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<quantity id="4C889B68E85E176F7EFDF92B6B98A4F5" box="[151,225,1733,1759]" metricMagnitude="2" metricUnit="m" metricValue="9.309999999999999" pageId="3" pageNumber="454" unit="m" value="931.0">
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,
<date id="FFCE104DE85E176F7E84F92A6A20A4F5" box="[238,345,1732,1759]" pageId="3" pageNumber="454" value="2017-10-05">
<collectingDate id="EF8AE9A5E85E176F7E84F92A6A20A4F5" box="[238,345,1732,1759]" pageId="3" pageNumber="454" value="2017-10-05">5.X.2017</collectingDate>
</date>
,
<collectorName id="2685535BE85E176F7F0FF92A6A8AA4F4" box="[357,499,1732,1758]" pageId="3" pageNumber="454">Yi-Zhou Liu</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B183CD0E85E176F7C5EF92B6F58A528" ID-GBIF-Occurrence="3863916347" collectingDate="2018-06" collectingDateMax="2018-07-31" collectingDateMin="2018-06" collectionCode="CQZY, VI" collectorName="Wei-Zong Yang" country="China" county="Yingjiang County" elevation="1200" location="Hulukou" municipality="Nabang Secondary Hydropower Station" pageId="3" pageNumber="454" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D76FD04E85E176F7C5EF92B692EA4F5" box="[564,599,1733,1759]" pageId="3" pageNumber="454" type="male">1♂</specimenCount>
(
<collectionCode id="ED61AE48E85E176F7C0DF92B69C0A4F5" box="[615,697,1733,1759]" pageId="3" pageNumber="454">CQZY</collectionCode>
),
<collectingMunicipality id="6BABACF7E85E176F7CA7F92A6FEBA4F4" box="[717,1170,1732,1758]" pageId="3" pageNumber="454">Nabang Secondary Hydropower Station</collectingMunicipality>
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<location id="8EAF6056E85E176F7AF5F92A6E7DA4F4" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7AF5F92A6E7DA4F4" box="[1183,1284,1732,1758]" country="China" county="Yingjiang County" municipality="Nabang Secondary Hydropower Station" name="Hulukou" pageId="3" pageNumber="454">Hulukou</location>
,
<location id="8EAF6056E85E176F7B7AF92A6EE1A4F5" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7B7AF92A6EE1A4F5" box="[1296,1432,1732,1759]" country="China" county="Yingjiang County" municipality="Nabang Secondary Hydropower Station" name="Xima Town" pageId="3" pageNumber="454">Xima Town</location>
,
<collectingCounty id="62AE4E01E85E176F7EFDF9066A18A529" box="[151,353,1768,1795]" pageId="3" pageNumber="454">Yingjiang County</collectingCounty>
,
<location id="8EAF6056E85E176F7F06F906697CA528" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7F06F906697CA528" box="[364,517,1768,1794]" country="China" county="Yingjiang County" municipality="Nabang Secondary Hydropower Station" name="Dehong Pref." pageId="3" pageNumber="454">Dehong Pref.</location>
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<quantity id="4C889B68E85E176F7C50F90769EBA529" box="[570,658,1769,1795]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="454" unit="m" value="1200.0">
<elevation id="005DD1BEE85E176F7C50F90769EBA529" box="[570,658,1769,1795]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="454" unit="m" value="1200.0">1200 m</elevation>
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,
<date id="FFCE104DE85E176F7CF7F9076848A529" box="[669,817,1769,1795]" pageId="3" pageNumber="454" value="2018-06" valueMax="2018-07-31" valueMin="2018-06-00">
<collectingDate id="EF8AE9A5E85E176F7CF7F9076848A529" box="[669,817,1769,1795]" pageId="3" pageNumber="454" value="2018-06" valueMax="2018-07-31" valueMin="2018-06-00">
<collectionCode id="ED61AE48E85E176F7CF7F90769C5A529" box="[669,700,1769,1795]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="3" pageNumber="454">VI</collectionCode>
VII.2018
</collectingDate>
</date>
,
<collectorName id="2685535BE85E176F7D57F9066897A529" box="[829,1006,1768,1795]" pageId="3" pageNumber="454">Wei-Zong Yang</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B183CD0E85E176F7A47F906681DA50C" ID-GBIF-Occurrence="3863916329" collectingDate="2021-09" collectionCode="CWT" collectorName="Shasawat Sirita" country="Thailand" location="Huay Pla Kang Village" pageId="3" pageNumber="454" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Rai">
<emphasis id="B904EA9FE85E176F7A47F9066FDDA528" bold="true" box="[1069,1188,1768,1794]" pageId="3" pageNumber="454">
<collectingCountry id="F367761DE85E176F7A47F9066FE7A528" box="[1069,1182,1768,1794]" name="Thailand" pageId="3" pageNumber="454">Thailand</collectingCountry>
:
</emphasis>
<specimenCount id="9D76FD04E85E176F7AC1F9076FB7A529" box="[1195,1230,1769,1795]" pageId="3" pageNumber="454" type="male">1♂</specimenCount>
(
<collectionCode id="ED61AE48E85E176F7AB7F9076E58A529" box="[1245,1313,1769,1795]" pageId="3" pageNumber="454">CWT</collectionCode>
),
<location id="8EAF6056E85E176F7B5EF9076A48A50C" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7B5EF9076A48A50C" country="Thailand" name="Huay Pla Kang Village" pageId="3" pageNumber="454" stateProvince="Chiang Rai">Huay Pla Kang Village</location>
,
<collectingRegion id="49B4F86FE85E176F7F57F8E2697CA50D" box="[317,517,1804,1831]" country="Thailand" name="Chiang Rai" pageId="3" pageNumber="454">Chiang Rai Prov.</collectingRegion>
,
<date id="FFCE104DE85E176F7C78F8E36908A50D" box="[530,625,1805,1831]" pageId="3" pageNumber="454" value="2021-09">
<collectingDate id="EF8AE9A5E85E176F7C78F8E36908A50D" box="[530,625,1805,1831]" pageId="3" pageNumber="454" value="2021-09">IX.2021</collectingDate>
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,
<collectorName id="2685535BE85E176F7C15F8E26856A50C" box="[639,815,1804,1830]" pageId="3" pageNumber="454">Shasawat Sirita</collectorName>
leg.
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;
<materialsCitation id="3B183CD0E85E176F7D18F8E3691DA560" ID-GBIF-Occurrence="3863916377" collectingDate="2021-10-08" collectionCode="CWT" collectorName="Shasawat Sirita" country="Thailand" location="Huay Pla Kang Village" pageId="3" pageNumber="454" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiang Rai">
<specimenCount id="9D76FD04E85E176F7D18F8E368EFA50D" box="[882,918,1805,1831]" pageId="3" pageNumber="454" type="male">1♂</specimenCount>
,
<specimenCount id="9D76FD04E85E176F7DCFF8E368B1A50D" box="[933,968,1805,1831]" pageId="3" pageNumber="454" type="female">1♀</specimenCount>
(
<collectionCode id="ED61AE48E85E176F7DB3F8E36F64A50D" box="[985,1053,1805,1831]" pageId="3" pageNumber="454">CWT</collectionCode>
),
<location id="8EAF6056E85E176F7A58F8E36E47A50C" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7A58F8E36E47A50C" box="[1074,1342,1804,1831]" country="Thailand" name="Huay Pla Kang Village" pageId="3" pageNumber="454" stateProvince="Chiang Rai">Huay Pla Kang Village</location>
,
<collectingRegion id="49B4F86FE85E176F7B21F8E26A7BA561" country="Thailand" name="Chiang Rai" pageId="3" pageNumber="454">Chiang Rai Prov.</collectingRegion>
,
<date id="FFCE104DE85E176F7F67F8DF6A01A561" box="[269,376,1841,1867]" pageId="3" pageNumber="454" value="2021-10-08">
<collectingDate id="EF8AE9A5E85E176F7F67F8DF6A01A561" box="[269,376,1841,1867]" pageId="3" pageNumber="454" value="2021-10-08">8.X.2021</collectingDate>
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,
<collectorName id="2685535BE85E176F7FE9F8DE6948A560" box="[387,561,1840,1866]" pageId="3" pageNumber="454">Shasawat Sirita</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B183CD0E85E176F7C1AF8DF6AC1A544" ID-GBIF-Occurrence="3863916313" collectingDate="2014-09" collectionCode="CWT" collectorName="W. Pathomwattananurak" country="Thailand" location="Huai Chomphu Subdist." pageId="3" pageNumber="454" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Chiang Rai">
<specimenCount id="9D76FD04E85E176F7C1AF8DF69EDA561" box="[624,660,1841,1867]" pageId="3" pageNumber="454" type="male">1♂</specimenCount>
,
<specimenCount id="9D76FD04E85E176F7CCBF8DF69BDA561" box="[673,708,1841,1867]" pageId="3" pageNumber="454" type="female">1♀</specimenCount>
(
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),
<location id="8EAF6056E85E176F7D40F8DE6F42A560" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7D40F8DE6F42A560" box="[810,1083,1840,1866]" country="Thailand" name="Huai Chomphu Subdist." pageId="3" pageNumber="454" stateProvince="Chiang Rai">Huai Chomphu Subdist.</location>
,
<collectingRegion id="49B4F86FE85E176F7A2FF8DE6E70A561" box="[1093,1289,1840,1867]" country="Thailand" name="Chiang Rai" pageId="3" pageNumber="454">Chiang Rai Prov.</collectingRegion>
,
<date id="FFCE104DE85E176F7B79F8DF6E0BA561" box="[1299,1394,1841,1867]" pageId="3" pageNumber="454" value="2014-09">
<collectingDate id="EF8AE9A5E85E176F7B79F8DF6E0BA561" box="[1299,1394,1841,1867]" pageId="3" pageNumber="454" value="2014-09">IX.2014</collectingDate>
</date>
,
<collectorName id="2685535BE85E176F7B17F8DF6AFAA544" pageId="3" pageNumber="454">W. Pathomwattananurak</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B183CD0E85E176F7FACF8BB6E11A544" ID-GBIF-Occurrence="3863916307" box="[454,1384,1876,1903]" collectionCode="CWT" country="Thailand" location="Wiang Papao Dist." pageId="3" pageNumber="454" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Chiang Rai">
<specimenCount id="9D76FD04E85E176F7FACF8BB6979A545" box="[454,512,1877,1903]" pageId="3" pageNumber="454" type="male">2♂♂</specimenCount>
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(
<collectionCode id="ED61AE48E85E176F7C2BF8BB69FCA545" box="[577,645,1877,1903]" pageId="3" pageNumber="454">CWT</collectionCode>
),
<location id="8EAF6056E85E176F7CF2F8BA6809A544" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85E176F7CF2F8BA6809A544" box="[664,880,1876,1903]" country="Thailand" name="Wiang Papao Dist." pageId="3" pageNumber="454" stateProvince="Chiang Rai">Wiang Papao Dist.</location>
,
<collectingRegion id="49B4F86FE85E176F7D17F8BA6F3AA545" box="[893,1091,1876,1903]" country="Thailand" name="Chiang Rai" pageId="3" pageNumber="454">Chiang Rai Prov.</collectingRegion>
, 2019, local collector leg
</materialsCitation>
.;
<materialsCitation id="3B183CD0E85E176F7B13F8BB68F6A5B8" ID-GBIF-Occurrence="3863916382" collectingDate="2019-08" collectionCode="CWT" collectorName="Local" country="Thailand" location="Phrao Dist." pageId="3" pageNumber="454" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai">
<specimenCount id="9D76FD04E85E176F7B13F8BB6EE5A545" box="[1401,1436,1877,1903]" pageId="3" pageNumber="454" type="male">1♂</specimenCount>
(
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.;
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<collectingDate id="EF8AE9A5E85E176F7D5AF82E68D7A5F1" box="[816,942,1984,2011]" pageId="3" pageNumber="454" value="2015-07-07">7.VII.2015</collectingDate>
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leg.
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<caption id="DF0F6605E85917687EFDF89B6A62A5D3" ID-DOI="http://doi.org/10.5281/zenodo.6853946" ID-Zenodo-Dep="6853946" httpUri="https://zenodo.org/record/6853946/files/figure.png" pageId="4" pageNumber="455" startId="4.[151,264,1909,1933]" targetBox="[222,1366,186,1882]" targetPageId="4">
<paragraph id="8BCF368DE85917687EFDF89B6A62A5D3" blockId="4.[151,1436,1909,2041]" pageId="4" pageNumber="455">
<emphasis id="B904EA9FE85917687EFDF89B6A3EA5A7" bold="true" box="[151,327,1909,1933]" pageId="4" pageNumber="455">FIGURES 217.</emphasis>
Habitus of
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<emphasis id="B904EA9FE85917687FD6F89B69F4A5A7" box="[444,653,1909,1933]" italics="true" pageId="4" pageNumber="455">Eophileurus confinis</emphasis>
<bibRefCitation id="EFE14B7CE85917687CF9F89B687CA5A7" author="Prell, H." box="[659,773,1909,1933]" pageId="4" pageNumber="455" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
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. 2, male (Mangshi); 3, male (Nabang); 4, female (Rongshuwang); 5, male (Mohan); 67, 11, male (Bo Kluea); 89, female (Jinghong); 10, male (Phrao); 12, male (Bao Loc); 13, female (Bao Loc); 14, male (Khao Kho); 15, male (Umphang); 16, male (Kraburi); 17, female (Kraburi); 26, 8, 1017, dorsal view; 7, 9, ventral view.
</paragraph>
</caption>
<caption id="DF0F6605E85817697EFDFA026FAFA402" ID-DOI="http://doi.org/10.5281/zenodo.6853950" ID-Zenodo-Dep="6853950" httpUri="https://zenodo.org/record/6853950/files/figure.png" pageId="5" pageNumber="456" startId="5.[151,264,1516,1540]" targetBox="[161,1417,211,1471]" targetPageId="5">
<paragraph id="8BCF368DE85817697EFDFA026FAFA402" blockId="5.[151,1436,1516,1576]" pageId="5" pageNumber="456">
<emphasis id="B904EA9FE85817697EFDFA026A2CA42E" bold="true" box="[151,341,1516,1540]" pageId="5" pageNumber="456">FIGURES 1824.</emphasis>
Holotype of
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<emphasis id="B904EA9FE85817697FB4FA0269CBA42E" box="[478,690,1516,1540]" italics="true" pageId="5" pageNumber="456">Eophileurus confinis</emphasis>
<bibRefCitation id="EFE14B7CE85817697CD3FA026853A42E" author="Prell, H." box="[697,810,1516,1540]" pageId="5" pageNumber="456" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
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(images ©MTD/Olaf Jäger). 18, habitus, dorsal; 19, habitus, lateral; 20, habitus, ventral; 21, parameres, ventral; 22, parameres, apical; 23, parameres, lateral; 24, labels.
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<paragraph id="8BCF368DE85817697EADF9BF6A53A5DD" blockId="5.[151,1437,1617,2039]" pageId="5" pageNumber="456">
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,
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</date>
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<collectingMunicipality id="6BABACF7E85817697DC8F99B6F28A4A5" box="[930,1105,1653,1679]" pageId="5" pageNumber="456">Umphang Dist.</collectingMunicipality>
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<collectingDate id="EF8AE9A5E85817697ABDF99B6E20A4A5" box="[1239,1369,1653,1679]" pageId="5" pageNumber="456" value="2020-11-17">17.XI.2020</collectingDate>
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, local collector leg.;
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<location id="8EAF6056E85817697F81F977692AA499" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85817697F81F977692AA499" box="[491,595,1689,1715]" name="Bangkok" pageId="5" pageNumber="456">Bangkok</location>
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,
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<date id="FFCE104DE85817697FE1F9536974A4F2" box="[395,525,1725,1752]" pageId="5" pageNumber="456" value="2021-09-25">
<collectingDate id="EF8AE9A5E85817697FE1F9536974A4F2" box="[395,525,1725,1752]" pageId="5" pageNumber="456" value="2021-09-25">25.IX.2021</collectingDate>
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(
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, local collector leg.;
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,
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<collectingMunicipality id="6BABACF7E85817697DE5F90F6F32A4D1" box="[911,1099,1761,1787]" pageId="5" pageNumber="456">Lang Suan Dist.</collectingMunicipality>
,
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,
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<collectingDate id="EF8AE9A5E85817697B49F90F6EE1A4D6" box="[1315,1432,1761,1788]" pageId="5" pageNumber="456" value="2021-05-15">
15.
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.2021
</collectingDate>
</date>
</materialsCitation>
,
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<collectorName id="2685535BE85817697EFDF8EB6A02A535" box="[151,379,1797,1823]" pageId="5" pageNumber="456">Uraiwan Abpamano</collectorName>
leg.;
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,
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(
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),
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generation of female from
<collectingMunicipality id="6BABACF7E85817697A67F8EB6FB2A535" box="[1037,1227,1797,1823]" pageId="5" pageNumber="456">Lang Suan Dist.</collectingMunicipality>
,
<location id="8EAF6056E85817697AB3F8EB6EE0A535" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85817697AB3F8EB6EE0A535" box="[1241,1433,1797,1823]" municipality="Lang Suan Dist." name="Chumporn Prov." pageId="5" pageNumber="456">Chumporn Prov.</location>
,
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<collectingDate id="EF8AE9A5E85817697EFDF8C76A74A56E" box="[151,269,1833,1860]" pageId="5" pageNumber="456" value="2021-05-15">
15.
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.2021
</collectingDate>
</date>
</materialsCitation>
,
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<collectorName id="2685535BE85817697F73F8C76A82A569" box="[281,507,1833,1859]" pageId="5" pageNumber="456">Uraiwan Abpamano</collectorName>
leg., adult emerged in
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<collectingDate id="EF8AE9A5E85817697D6AF8C76818A569" box="[768,865,1833,1859]" pageId="5" pageNumber="456" value="2021-09">IX.2021</collectingDate>
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;
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<collectingCountry id="F367761DE85817697D07F8C768AEA569" box="[877,983,1833,1859]" name="Vietnam" pageId="5" pageNumber="456">Vietnam</collectingCountry>
:
</emphasis>
<specimenCount id="9D76FD04E85817697D8EF8C76F7EA569" box="[996,1031,1833,1859]" pageId="5" pageNumber="456" type="male">1♂</specimenCount>
(
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)
</materialsCitation>
,
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<location id="8EAF6056E85817697A17F8C76E6FA569" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85817697A17F8C76E6FA569" box="[1149,1302,1833,1859]" country="Vietnam" name="Bao Loc City" pageId="5" pageNumber="456" stateProvince="Lam Dong">Bao Loc City</location>
,
<collectingRegion id="49B4F86FE85817697B49F8C76BACA54D" country="Vietnam" name="Lam Dong" pageId="5" pageNumber="456">Lam Dong Prov.</collectingRegion>
,
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<collectingDate id="EF8AE9A5E85817697E88F8A36A41A54D" box="[226,312,1869,1895]" pageId="5" pageNumber="456" value="2019-10">X.2019</collectingDate>
</date>
, local collector leg.;
<specimenCount id="9D76FD04E85817697C41F8A36937A54D" box="[555,590,1869,1895]" pageId="5" pageNumber="456" type="female">1♀</specimenCount>
(
<collectionCode id="ED61AE48E85817697C0AF8A369CBA54D" box="[608,690,1869,1895]" pageId="5" pageNumber="456">CQZY</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B183CD0E85817697CADF8A36A5CA5A1" ID-GBIF-Occurrence="3863916364" collectingDate="2017-08" collectionCode="CQZY" country="Vietnam" location="Bao Loc City" pageId="5" pageNumber="456" specimenCount="1" specimenCount-female="1" stateProvince="Lam Dong">
<location id="8EAF6056E85817697CADF8A3681DA54D" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85817697CADF8A3681DA54D" box="[711,868,1869,1895]" country="Vietnam" name="Bao Loc City" pageId="5" pageNumber="456" stateProvince="Lam Dong">Bao Loc City</location>
,
<collectingRegion id="49B4F86FE85817697D18F8A36F4DA54D" box="[882,1076,1869,1895]" country="Vietnam" name="Lam Dong" pageId="5" pageNumber="456">Lam Dong Prov.</collectingRegion>
,
<date id="FFCE104DE85817697A2AF8A36FCBA54D" box="[1088,1202,1869,1895]" pageId="5" pageNumber="456" value="2017-08">
<collectingDate id="EF8AE9A5E85817697A2AF8A36FCBA54D" box="[1088,1202,1869,1895]" pageId="5" pageNumber="456" value="2017-08">VIII.2017</collectingDate>
</date>
, local collector leg.;
<specimenCount id="9D76FD04E85817697EFDF89F6BC3A5A1" box="[151,186,1905,1931]" pageId="5" pageNumber="456" type="female">1♀</specimenCount>
(
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)
</materialsCitation>
,
<materialsCitation id="3B183CD0E85817697F58F89F6F96A5A1" ID-GBIF-Occurrence="3863916341" box="[306,1263,1905,1931]" collectingDate="2018-05" collectionCode="V, CQZY" country="Vietnam" location="Bao Lam Dist." municipality="Dambri" pageId="5" pageNumber="456" specimenCount="1" specimenCount-male="1" stateProvince="Lam Dong">
<collectingMunicipality id="6BABACF7E85817697F58F89F6AF3A5A1" box="[306,394,1905,1931]" pageId="5" pageNumber="456">Dambri</collectingMunicipality>
,
<location id="8EAF6056E85817697FF2F89F693AA5A1" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85817697FF2F89F693AA5A1" box="[408,579,1905,1931]" country="Vietnam" municipality="Dambri" name="Bao Lam Dist." pageId="5" pageNumber="456" stateProvince="Lam Dong">Bao Lam Dist.</location>
,
<collectingRegion id="49B4F86FE85817697C3AF89F6868A5A1" box="[592,785,1905,1931]" country="Vietnam" name="Lam Dong" pageId="5" pageNumber="456">Lam Dong Prov.</collectingRegion>
,
<date id="FFCE104DE85817697D77F89F6816A5A1" box="[797,879,1905,1931]" pageId="5" pageNumber="456" value="2018-05">
<collectingDate id="EF8AE9A5E85817697D77F89F6816A5A1" box="[797,879,1905,1931]" pageId="5" pageNumber="456" value="2018-05">
<collectionCode id="ED61AE48E85817697D77F89F684BA5A1" box="[797,818,1905,1931]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="456" type="Museum">V</collectionCode>
.2018
</collectingDate>
</date>
, local collector leg.;
<specimenCount id="9D76FD04E85817697A0BF89F6FFDA5A1" box="[1121,1156,1905,1931]" pageId="5" pageNumber="456" type="male">1♂</specimenCount>
(
<collectionCode id="ED61AE48E85817697AFFF89F6F9EA5A1" box="[1173,1255,1905,1931]" pageId="5" pageNumber="456">CQZY</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B183CD0E85817697A96F89F6B80A5F9" ID-GBIF-Occurrence="3863916327" collectingDate="2019-03" collectionCode="CQZY" country="Vietnam" elevation="1300" latitude="15.964556" location="Tay Giang Dist." longLatPrecision="1" longitude="107.43906" municipality="Axan Mt." pageId="5" pageNumber="456" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Quang Nam">
<geoCoordinate id="EE44504AE85817697A96F89F6EE1A5A6" box="[1276,1432,1905,1932]" degrees="15" direction="north" minutes="57" orientation="latitude" pageId="5" pageNumber="456" precision="1" seconds="52.4" value="15.964556">15°57'52.4&quot;N</geoCoordinate>
,
<geoCoordinate id="EE44504AE85817697EFDF87B6A47A585" box="[151,318,1941,1967]" degrees="107" direction="east" minutes="26" orientation="longitude" pageId="5" pageNumber="456" precision="1" seconds="20.6" value="107.43906">107°26'20.6&quot;E</geoCoordinate>
,
<collectingMunicipality id="6BABACF7E85817697F23F87B6AC0A585" box="[329,441,1941,1967]" pageId="5" pageNumber="456">Axan Mt.</collectingMunicipality>
,
<location id="8EAF6056E85817697FAFF87B6904A585" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85817697FAFF87B6904A585" box="[453,637,1941,1967]" country="Vietnam" latitude="15.964556" longLatPrecision="1" longitude="107.43906" municipality="Axan Mt." name="Tay Giang Dist." pageId="5" pageNumber="456" stateProvince="Quang Nam">Tay Giang Dist.</location>
,
<collectingRegion id="49B4F86FE85817697CE0F87B6823A585" box="[650,858,1941,1967]" country="Vietnam" name="Quang Nam" pageId="5" pageNumber="456">Quang Nam Prov.</collectingRegion>
, alt.
<quantity id="4C889B68E85817697DF8F87B6892A585" box="[914,1003,1941,1967]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="456" unit="m" value="1300.0">
<elevation id="005DD1BEE85817697DF8F87B6892A585" box="[914,1003,1941,1967]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="456" unit="m" value="1300.0">1300 m</elevation>
</quantity>
,
<date id="FFCE104DE85817697D92F87B6F2FA585" box="[1016,1110,1941,1967]" pageId="5" pageNumber="456" value="2019-03">
<collectingDate id="EF8AE9A5E85817697D92F87B6F2FA585" box="[1016,1110,1941,1967]" pageId="5" pageNumber="456" value="2019-03">III.2019</collectingDate>
</date>
, local collector leg.;
<specimenCount id="9D76FD04E85817697B2CF87B6E12A585" box="[1350,1387,1941,1967]" pageId="5" pageNumber="456" type="male">1♂</specimenCount>
,
<specimenCount id="9D76FD04E85817697B13F87B6EE5A585" box="[1401,1436,1941,1967]" pageId="5" pageNumber="456" type="female">1♀</specimenCount>
(
<collectionCode id="ED61AE48E85817697EF5F8576B88A5F9" box="[159,241,1977,2003]" pageId="5" pageNumber="456">CQZY</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B183CD0E85817697F6EF8576A5FA5DD" ID-GBIF-Occurrence="3863916355" collectingDate="2019-07" country="Vietnam" elevation="1300" latitude="15.964556" location="Tay Giang Dist." longLatPrecision="1" longitude="107.43906" municipality="Axan Mt." pageId="5" pageNumber="456" specimenCount="1" stateProvince="Quang Nam">
<geoCoordinate id="EE44504AE85817697F6EF8576AD9A5FE" box="[260,416,1977,2004]" degrees="15" direction="north" minutes="57" orientation="latitude" pageId="5" pageNumber="456" precision="1" seconds="52.4" value="15.964556">15°57'52.4&quot;N</geoCoordinate>
,
<geoCoordinate id="EE44504AE85817697FC1F857692BA5F9" box="[427,594,1977,2003]" degrees="107" direction="east" minutes="26" orientation="longitude" pageId="5" pageNumber="456" precision="1" seconds="20.6" value="107.43906">107°26'20.6&quot;E</geoCoordinate>
,
<collectingMunicipality id="6BABACF7E85817697C36F85769B3A5F9" box="[604,714,1977,2003]" pageId="5" pageNumber="456">Axan Mt.</collectingMunicipality>
,
<location id="8EAF6056E85817697CBFF85768F3A5F9" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85817697CBFF85768F3A5F9" box="[725,906,1977,2003]" country="Vietnam" latitude="15.964556" longLatPrecision="1" longitude="107.43906" municipality="Axan Mt." name="Tay Giang Dist." pageId="5" pageNumber="456" stateProvince="Quang Nam">Tay Giang Dist.</location>
,
<collectingRegion id="49B4F86FE85817697DFFF8576F18A5F9" box="[917,1121,1977,2003]" country="Vietnam" name="Quang Nam" pageId="5" pageNumber="456">Quang Nam Prov.</collectingRegion>
, alt.
<quantity id="4C889B68E85817697AFCF8576F9EA5F9" box="[1174,1255,1977,2003]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="456" unit="m" value="1300.0">
<elevation id="005DD1BEE85817697AFCF8576F9EA5F9" box="[1174,1255,1977,2003]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="456" unit="m" value="1300.0">1300m</elevation>
</quantity>
,
<date id="FFCE104DE85817697A98F8576E22A5F9" box="[1266,1371,1977,2003]" pageId="5" pageNumber="456" value="2019-07">
<collectingDate id="EF8AE9A5E85817697A98F8576E22A5F9" box="[1266,1371,1977,2003]" pageId="5" pageNumber="456" value="2019-07">VII.2019</collectingDate>
</date>
, local collector leg
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF0F6605E85B176A7EFDFA566FAFA7DE" ID-DOI="http://doi.org/10.5281/zenodo.6853952" ID-Zenodo-Dep="6853952" httpUri="https://zenodo.org/record/6853952/files/figure.png" pageId="6" pageNumber="457" startId="6.[151,264,1464,1488]" targetBox="[154,1434,194,1428]" targetPageId="6">
<paragraph id="8BCF368DE85B176A7EFDFA566FAFA7DE" blockId="6.[151,1436,1464,1525]" pageId="6" pageNumber="457">
<emphasis id="B904EA9FE85B176A7EFDFA566A2DA7FA" bold="true" box="[151,340,1464,1488]" pageId="6" pageNumber="457">FIGURES 2531.</emphasis>
Holotype of
<taxonomicName id="4C704D0EE85B176A7FB0FA56684BA7FB" authority="Prell, 1913" authorityName="Prell" authorityYear="1913" box="[474,818,1464,1489]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="decipiens">
<emphasis id="B904EA9FE85B176A7FB0FA5669C4A7FA" box="[474,701,1464,1488]" italics="true" pageId="6" pageNumber="457">Eophileurus decipiens</emphasis>
<bibRefCitation id="EFE14B7CE85B176A7CA9FA56684BA7FB" author="Prell, H." box="[707,818,1464,1489]" pageId="6" pageNumber="457" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
</taxonomicName>
(images ©MTD/Olaf Jäger). 25, habitus, dorsal; 26, habitus, lateral; 27, habitus, ventral; 28, parameres, ventral; 29, parameres, apical; 30, parameres, lateral; 31, labels.
</paragraph>
</caption>
<subSubSection id="C36A6506E85B17647EADF9CC6F91A5D0" lastPageId="8" lastPageNumber="459" pageId="6" pageNumber="457" type="distribution">
<paragraph id="8BCF368DE85B176A7EADF9CC68ADA417" blockId="6.[151,1437,1570,2029]" box="[199,980,1570,1597]" pageId="6" pageNumber="457">
<emphasis id="B904EA9FE85B176A7EADF9CC6A18A416" bold="true" box="[199,353,1570,1596]" pageId="6" pageNumber="457">Distribution.</emphasis>
<collectingCountry id="F367761DE85B176A7F0DF9CD6AA7A417" box="[359,478,1571,1597]" name="Cambodia" pageId="6" pageNumber="457">Cambodia</collectingCountry>
;
<collectingCountry id="F367761DE85B176A7F83F9CD6956A417" box="[489,559,1571,1597]" name="China" pageId="6" pageNumber="457">China</collectingCountry>
;
<collectingCountry id="F367761DE85B176A7C51F9CD690CA417" box="[571,629,1571,1597]" name="Laos" pageId="6" pageNumber="457">Laos</collectingCountry>
;
<collectingCountry id="F367761DE85B176A7CEAF9CD6988A417" box="[640,753,1571,1597]" name="Myanmar" pageId="6" pageNumber="457">Myanmar</collectingCountry>
;
<collectingCountry id="F367761DE85B176A7C91F9CD681BA417" box="[763,866,1571,1597]" name="Thailand" pageId="6" pageNumber="457">Thailand</collectingCountry>
;
<collectingCountry id="F367761DE85B176A7D07F9CD68A9A417" box="[877,976,1571,1597]" name="Vietnam" pageId="6" pageNumber="457">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="8BCF368DE85B176A7EADF9A8687AA513" blockId="6.[151,1437,1570,2029]" pageId="6" pageNumber="457">
<emphasis id="B904EA9FE85B176A7EADF9A86A33A44A" bold="true" box="[199,330,1606,1632]" pageId="6" pageNumber="457">Variations.</emphasis>
<taxonomicName id="4C704D0EE85B176A7F3AF9A969C0A44B" authority="Prell, 1913" authorityName="Prell" authorityYear="1913" box="[336,697,1607,1633]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="457" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE85B176A7F3AF9A96941A44B" box="[336,568,1607,1633]" italics="true" pageId="6" pageNumber="457">Eophileurus confinis</emphasis>
<bibRefCitation id="EFE14B7CE85B176A7C54F9A969C0A44B" author="Prell, H." box="[574,697,1607,1633]" pageId="6" pageNumber="457" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
</taxonomicName>
is a widely distributed species, currently recorded from
<collectingCountry id="F367761DE85B176A7B40F9A96E14A44B" box="[1322,1389,1607,1633]" name="China" pageId="6" pageNumber="457">China</collectingCountry>
and Indochina (
<figureCitation id="134B2A08E85B176A7F7DF9856A03A4AF" box="[279,378,1643,1669]" captionStart="FIGURE 197" captionStartId="32.[151,250,1456,1480]" captionTargetBox="[151,1435,181,1432]" captionTargetId="figure-18@32.[151,1435,181,1432]" captionTargetPageId="32" captionText="FIGURE 197. Map of the currently known distributions of Eophileurus Arrow, 1908 species involved in this paper excluding E. felschei Prell, 1913. Imprecise locality record excluded from the map." figureDoi="http://doi.org/10.5281/zenodo.6853994" httpUri="https://zenodo.org/record/6853994/files/figure.png" pageId="6" pageNumber="457">Fig. 197</figureCitation>
). The external characters (
<figureCitation id="134B2A08E85B176A7CC9F9856865A4AF" box="[675,796,1643,1669]" captionStart="FIGURES 217" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[222,1366,186,1882]" captionTargetId="figure-18@4.[212,1375,181,1885]" captionTargetPageId="4" captionText="FIGURES 217. Habitus of Eophileurus confinis Prell, 1913. 2, male (Mangshi); 3, male (Nabang); 4, female (Rongshuwang); 5, male (Mohan); 67, 11, male (Bo Kluea); 89, female (Jinghong); 10, male (Phrao); 12, male (Bao Loc); 13, female (Bao Loc); 14, male (Khao Kho); 15, male (Umphang); 16, male (Kraburi); 17, female (Kraburi); 26, 8, 1017, dorsal view; 7, 9, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853946" httpUri="https://zenodo.org/record/6853946/files/figure.png" pageId="6" pageNumber="457">Figs. 217</figureCitation>
) of the specimens are similar regardless of the localities. The shape of the parameres (
<figureCitation id="134B2A08E85B176A7F8DF96169F7A483" box="[487,654,1679,1705]" captionStart-0="FIGURES 117146" captionStart-1="FIGURES 147152" captionStart-2="FIGURES 153164" captionStart-3="FIGURES 165184" captionStartId-0="27.[151,264,1803,1827]" captionStartId-1="28.[151,264,1316,1340]" captionStartId-2="29.[151,264,1313,1337]" captionStartId-3="30.[151,264,1476,1500]" captionTargetBox-0="[169,1424,203,1763]" captionTargetBox-1="[159,1415,199,1278]" captionTargetBox-2="[152,1435,181,1289]" captionTargetBox-3="[157,1431,189,1441]" captionTargetId-0="figure-18@27.[151,1435,181,1780]" captionTargetId-1="figure-18@28.[151,1435,181,1292]" captionTargetId-2="figure-18@29.[151,1435,181,1289]" captionTargetId-3="figure-18@30.[151,1435,181,1453]" captionTargetPageId-0="27" captionTargetPageId-1="28" captionTargetPageId-2="29" captionTargetPageId-3="30" captionText-0="FIGURES 117146. Parameres of Eophileurus confinis Prell, 1913. 117120 from Nabang; 121122, from Lancangjiang Bridge; 123124 from Jinghong; 125128 from Wiang Papao; 129130 from Mueang Chiang Rai; 131132 from Phrao; 133 134 from Axan Mt.; 135136 from Bao Loc; 137138 from Bo Kluea; 139140 from Khao Kho; 141142 from Umphang; 143144 from Bangkok; 145146 from Kraburi; odd number ones, front view; even number ones, lateral view; 117124 without scale bar, the rest with scale bar = 2 mm." captionText-1="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." captionText-2="FIGURES 153164. Parameres of Eophileurus confinis Prell, 1913. 153154 from Lancangjiang Bridge; 155156 from Jinghong; 157158 from Wiang Papao; 159160 from Khao Kho; 161162 from Bangkok; 163164 from Kraburi; odd number ones, front view; even number ones, lateral view; not in scale." captionText-3="FIGURES 165184. Parameres of Eophileurus Arrow, 1908 species. 165166, E. confinis Prell, 1913 (Bao Loc); 167168, E. heyrovskyi Kral &amp; Strnad, 1992 (Longli); 169172, E. minor Yang &amp; Pathomwattananurak, new species, 169170, holotype, 171172, paratype (Dong Tien, images ©Ming-Zhi Zhao); 173174, E. prelli Yang &amp; Pathomwattananurak, new species, holotype; 174180, E. quadratifovealis Yang &amp; Pathomwattananurak, new species, 175176, holotype, 177178, paratype (Mu Si), 179180, paratype (Chatabon, images by Keita Matsumoto ©Natural History Museum, London); 181184, E. sidereus Yang &amp; Pathomwattananurak, new species, 181182, holotype, 183184, paratype; odd number ones, front view; even number ones, lateral view; scale bar = 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6853980" figureDoi-1="http://doi.org/10.5281/zenodo.6853982" figureDoi-2="http://doi.org/10.5281/zenodo.6853984" figureDoi-3="http://doi.org/10.5281/zenodo.6853988" httpUri-0="https://zenodo.org/record/6853980/files/figure.png" httpUri-1="https://zenodo.org/record/6853982/files/figure.png" httpUri-2="https://zenodo.org/record/6853984/files/figure.png" httpUri-3="https://zenodo.org/record/6853988/files/figure.png" pageId="6" pageNumber="457">Figs. 117166</figureCitation>
), on the other hand, exhibits significant variations both individually and geographically, while the overall structure of the parameres being the same. Following the definitions of parts of the parameres in the material and method section, the following paragraphs demonstrate the individual and geographical variations of this species, how one geographical form transitions to another one, and why the various morphologies of parameres belong to the same species.
</paragraph>
<paragraph id="8BCF368DE85B176B7EADF8AD6A8BA04B" blockId="6.[151,1437,1570,2029]" lastBlockId="7.[151,1437,150,609]" lastPageId="7" lastPageNumber="458" pageId="6" pageNumber="457">
Regarding the individual variations: while examining
<specimenCount id="9D76FD04E85B176A7D4DF8AD68E9A577" box="[807,912,1859,1885]" pageId="6" pageNumber="457" type="male">six males</specimenCount>
which were raised from eggs laid by the same female (the female was collected from the wild, without mating with any male after its collection), it is discovered that their parameres (
<figureCitation id="134B2A08E85B176A7FECF8656950A58C" box="[390,553,1931,1958]" captionStart="FIGURES 147152" captionStartId="28.[151,264,1316,1340]" captionTargetBox="[159,1415,199,1278]" captionTargetId="figure-18@28.[151,1435,181,1292]" captionTargetPageId="28" captionText="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853982" httpUri="https://zenodo.org/record/6853982/files/figure.png" pageId="6" pageNumber="457">Figs. 147152</figureCitation>
) show rather large variations. The most distinct variations can be seen on BN, LP, apex of LP and apices of the parameres, although other parts, such as the slope of LIM and the overall slimness of the parameres also show slight variations. BN have both the left part overlapping the right part (
<figureCitation id="134B2A08E85B176B7A99F83D6B85A29B" captionStart="FIGURES 147152" captionStartId="28.[151,264,1316,1340]" captionTargetBox="[159,1415,199,1278]" captionTargetId="figure-18@28.[151,1435,181,1292]" captionTargetPageId="28" captionText="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853982" httpUri="https://zenodo.org/record/6853982/files/figure.png" lastPageId="7" lastPageNumber="458" pageId="6" pageNumber="457">Figs. 147, 148, 151, 152</figureCitation>
) and vice versa (
<figureCitation id="134B2A08E85A176B7FA9FF796912A29B" box="[451,619,150,177]" captionStart="FIGURES 147152" captionStartId="28.[151,264,1316,1340]" captionTargetBox="[159,1415,199,1278]" captionTargetId="figure-18@28.[151,1435,181,1292]" captionTargetPageId="28" captionText="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853982" httpUri="https://zenodo.org/record/6853982/files/figure.png" pageId="7" pageNumber="458">Figs. 149, 151</figureCitation>
); the lower margin of BN can also be seen distinctly extend downward (
<figureCitation id="134B2A08E85A176B7EF5FF556AF9A2FF" box="[159,384,186,213]" captionStart="FIGURES 147152" captionStartId="28.[151,264,1316,1340]" captionTargetBox="[159,1415,199,1278]" captionTargetId="figure-18@28.[151,1435,181,1292]" captionTargetPageId="28" captionText="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853982" httpUri="https://zenodo.org/record/6853982/files/figure.png" pageId="7" pageNumber="458">Figs. 148, 149, 151</figureCitation>
) or rather horizontal (
<figureCitation id="134B2A08E85A176B7C14FF556818A2FF" box="[638,865,186,213]" captionStart="FIGURES 147152" captionStartId="28.[151,264,1316,1340]" captionTargetBox="[159,1415,199,1278]" captionTargetId="figure-18@28.[151,1435,181,1292]" captionTargetPageId="28" captionText="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853982" httpUri="https://zenodo.org/record/6853982/files/figure.png" pageId="7" pageNumber="458">Figs. 147, 150, 152</figureCitation>
). LP present larger variations: some with rounded outer-most point (
<figureCitation id="134B2A08E85A176B7F0FFF316933A2D3" box="[357,586,222,249]" captionStart="FIGURES 147152" captionStartId="28.[151,264,1316,1340]" captionTargetBox="[159,1415,199,1278]" captionTargetId="figure-18@28.[151,1435,181,1292]" captionTargetPageId="28" captionText="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853982" httpUri="https://zenodo.org/record/6853982/files/figure.png" pageId="7" pageNumber="458">Figs. 147, 149, 152</figureCitation>
), while one almost forming a small individual process (
<figureCitation id="134B2A08E85A176B7AB8FF316E39A2D3" box="[1234,1344,222,249]" captionStart="FIGURES 147152" captionStartId="28.[151,264,1316,1340]" captionTargetBox="[159,1415,199,1278]" captionTargetId="figure-18@28.[151,1435,181,1292]" captionTargetPageId="28" captionText="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853982" httpUri="https://zenodo.org/record/6853982/files/figure.png" pageId="7" pageNumber="458">Figs. 151</figureCitation>
, similar situation can be seen from parameres of
<taxonomicName id="4C704D0EE85A176B7C03FEED699BA337" authorityName="Prell" authorityYear="1913" box="[617,738,259,285]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE85A176B7C03FEED699BA337" box="[617,738,259,285]" italics="true" pageId="7" pageNumber="458">E. confinis</emphasis>
</taxonomicName>
from other areas, such as in
<figureCitation id="134B2A08E85A176B7A5AFEED6FB5A337" box="[1072,1228,258,285]" captionStart="FIGURES 117146" captionStartId="27.[151,264,1803,1827]" captionTargetBox="[169,1424,203,1763]" captionTargetId="figure-18@27.[151,1435,181,1780]" captionTargetPageId="27" captionText="FIGURES 117146. Parameres of Eophileurus confinis Prell, 1913. 117120 from Nabang; 121122, from Lancangjiang Bridge; 123124 from Jinghong; 125128 from Wiang Papao; 129130 from Mueang Chiang Rai; 131132 from Phrao; 133 134 from Axan Mt.; 135136 from Bao Loc; 137138 from Bo Kluea; 139140 from Khao Kho; 141142 from Umphang; 143144 from Bangkok; 145146 from Kraburi; odd number ones, front view; even number ones, lateral view; 117124 without scale bar, the rest with scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853980" httpUri="https://zenodo.org/record/6853980/files/figure.png" pageId="7" pageNumber="458">Fig. 125, 131</figureCitation>
), and some lies in between (
<figureCitation id="134B2A08E85A176B7F6FFEC96AD7A36B" box="[261,430,294,321]" captionStart="FIGURES 147152" captionStartId="28.[151,264,1316,1340]" captionTargetBox="[159,1415,199,1278]" captionTargetId="figure-18@28.[151,1435,181,1292]" captionTargetPageId="28" captionText="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853982" httpUri="https://zenodo.org/record/6853982/files/figure.png" pageId="7" pageNumber="458">Figs. 148, 150</figureCitation>
); the apices of LP all point inward, but can be upward (
<figureCitation id="134B2A08E85A176B7A45FEC96FEBA36B" box="[1071,1170,295,321]" captionStart="FIGURES 147152" captionStartId="28.[151,264,1316,1340]" captionTargetBox="[159,1415,199,1278]" captionTargetId="figure-18@28.[151,1435,181,1292]" captionTargetPageId="28" captionText="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853982" httpUri="https://zenodo.org/record/6853982/files/figure.png" pageId="7" pageNumber="458">Fig. 148</figureCitation>
), horizontal (
<figureCitation id="134B2A08E85A176B7B40FEC96B82A34F" captionStart="FIGURES 147152" captionStartId="28.[151,264,1316,1340]" captionTargetBox="[159,1415,199,1278]" captionTargetId="figure-18@28.[151,1435,181,1292]" captionTargetPageId="28" captionText="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853982" httpUri="https://zenodo.org/record/6853982/files/figure.png" pageId="7" pageNumber="458">Figs. 149, 151, 152</figureCitation>
) or downward (
<figureCitation id="134B2A08E85A176B7FD8FEA56920A34F" box="[434,601,330,357]" captionStart="FIGURES 147152" captionStartId="28.[151,264,1316,1340]" captionTargetBox="[159,1415,199,1278]" captionTargetId="figure-18@28.[151,1435,181,1292]" captionTargetPageId="28" captionText="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853982" httpUri="https://zenodo.org/record/6853982/files/figure.png" pageId="7" pageNumber="458">Figs. 147, 150</figureCitation>
). The apices of the parameres are either slightly enlarged (
<figureCitation id="134B2A08E85A176B7A98FEA56BBDA3A3" captionStart="FIGURES 147152" captionStartId="28.[151,264,1316,1340]" captionTargetBox="[159,1415,199,1278]" captionTargetId="figure-18@28.[151,1435,181,1292]" captionTargetPageId="28" captionText="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853982" httpUri="https://zenodo.org/record/6853982/files/figure.png" pageId="7" pageNumber="458">Figs. 147, 148, 152</figureCitation>
), parallel-sided (
<figureCitation id="134B2A08E85A176B7FEAFE81695BA3A3" box="[384,546,366,393]" captionStart="FIGURES 147152" captionStartId="28.[151,264,1316,1340]" captionTargetBox="[159,1415,199,1278]" captionTargetId="figure-18@28.[151,1435,181,1292]" captionTargetPageId="28" captionText="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853982" httpUri="https://zenodo.org/record/6853982/files/figure.png" pageId="7" pageNumber="458">Figs. 149, 151</figureCitation>
) or slightly contracted (
<figureCitation id="134B2A08E85A176B7D44FE8168E9A3A3" box="[814,912,366,393]" captionStart="FIGURES 147152" captionStartId="28.[151,264,1316,1340]" captionTargetBox="[159,1415,199,1278]" captionTargetId="figure-18@28.[151,1435,181,1292]" captionTargetPageId="28" captionText="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853982" httpUri="https://zenodo.org/record/6853982/files/figure.png" pageId="7" pageNumber="458">Fig. 150</figureCitation>
). Some LIM have larger slope (
<figureCitation id="134B2A08E85A176B7A98FE816EEDA3A3" box="[1266,1428,366,393]" captionStart="FIGURES 147152" captionStartId="28.[151,264,1316,1340]" captionTargetBox="[159,1415,199,1278]" captionTargetId="figure-18@28.[151,1435,181,1292]" captionTargetPageId="28" captionText="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853982" httpUri="https://zenodo.org/record/6853982/files/figure.png" pageId="7" pageNumber="458">Figs. 148, 151</figureCitation>
) and the remaining are with smaller slope (
<figureCitation id="134B2A08E85A176B7C04FE7D68FAA387" box="[622,899,402,429]" captionStart="FIGURES 147152" captionStartId="28.[151,264,1316,1340]" captionTargetBox="[159,1415,199,1278]" captionTargetId="figure-18@28.[151,1435,181,1292]" captionTargetPageId="28" captionText="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853982" httpUri="https://zenodo.org/record/6853982/files/figure.png" pageId="7" pageNumber="458">Figs. 147, 149, 150, 152</figureCitation>
). Some parameres are overall slenderer than the others (the set of
<figureCitation id="134B2A08E85A176B7F35FE596AC7A3FB" box="[351,446,439,465]" captionStart="FIGURES 147152" captionStartId="28.[151,264,1316,1340]" captionTargetBox="[159,1415,199,1278]" captionTargetId="figure-18@28.[151,1435,181,1292]" captionTargetPageId="28" captionText="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853982" httpUri="https://zenodo.org/record/6853982/files/figure.png" pageId="7" pageNumber="458">Fig. 147</figureCitation>
versus
<figureCitation id="134B2A08E85A176B7C7CFE59699AA3FB" box="[534,739,438,465]" captionStart="FIGURES 147152" captionStartId="28.[151,264,1316,1340]" captionTargetBox="[159,1415,199,1278]" captionTargetId="figure-18@28.[151,1435,181,1292]" captionTargetPageId="28" captionText="FIGURES 147152. Parameres of Eophileurus confinis Prell, 1913 from male offspring of the same wild female; not to scale." figureDoi="http://doi.org/10.5281/zenodo.6853982" httpUri="https://zenodo.org/record/6853982/files/figure.png" pageId="7" pageNumber="458">Figs. 151 and 152</figureCitation>
being the most distinct). It is therefore demonstrated that the parameres of
<taxonomicName id="4C704D0EE85A176B7F58FE356AD5A3DF" authorityName="Prell" authorityYear="1913" box="[306,428,475,501]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE85A176B7F58FE356AD5A3DF" box="[306,428,475,501]" italics="true" pageId="7" pageNumber="458">E. confinis</emphasis>
</taxonomicName>
have significant individual variations. This can be seen much extremer while examining males from the same area (same habitat), such as sets in
<figureCitation id="134B2A08E85A176B7D71FE1168C3A033" box="[795,954,511,537]" captionStart="FIGURES 117146" captionStartId="27.[151,264,1803,1827]" captionTargetBox="[169,1424,203,1763]" captionTargetId="figure-18@27.[151,1435,181,1780]" captionTargetPageId="27" captionText="FIGURES 117146. Parameres of Eophileurus confinis Prell, 1913. 117120 from Nabang; 121122, from Lancangjiang Bridge; 123124 from Jinghong; 125128 from Wiang Papao; 129130 from Mueang Chiang Rai; 131132 from Phrao; 133 134 from Axan Mt.; 135136 from Bao Loc; 137138 from Bo Kluea; 139140 from Khao Kho; 141142 from Umphang; 143144 from Bangkok; 145146 from Kraburi; odd number ones, front view; even number ones, lateral view; 117124 without scale bar, the rest with scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853980" httpUri="https://zenodo.org/record/6853980/files/figure.png" pageId="7" pageNumber="458">Figs. 117118</figureCitation>
versus 119120 and
<figureCitation id="134B2A08E85A176B7AC1FE116E34A033" box="[1195,1357,510,537]" captionStart="FIGURES 117146" captionStartId="27.[151,264,1803,1827]" captionTargetBox="[169,1424,203,1763]" captionTargetId="figure-18@27.[151,1435,181,1780]" captionTargetPageId="27" captionText="FIGURES 117146. Parameres of Eophileurus confinis Prell, 1913. 117120 from Nabang; 121122, from Lancangjiang Bridge; 123124 from Jinghong; 125128 from Wiang Papao; 129130 from Mueang Chiang Rai; 131132 from Phrao; 133 134 from Axan Mt.; 135136 from Bao Loc; 137138 from Bo Kluea; 139140 from Khao Kho; 141142 from Umphang; 143144 from Bangkok; 145146 from Kraburi; odd number ones, front view; even number ones, lateral view; 117124 without scale bar, the rest with scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853980" httpUri="https://zenodo.org/record/6853980/files/figure.png" pageId="7" pageNumber="458">Figs. 125126</figureCitation>
versus 127128. Overall, this led to a conclusion that LP bear the most distinct individual variations, as seen in both the offspring set and the local sets.
</paragraph>
<caption id="DF0F6605E85A176B7EFDF8506EF6A5D0" ID-DOI="http://doi.org/10.5281/zenodo.6853954" ID-Zenodo-Dep="6853954" httpUri="https://zenodo.org/record/6853954/files/figure.png" pageId="7" pageNumber="458" startId="7.[151,264,1982,2006]" targetBox="[198,1395,649,1953]" targetPageId="7">
<paragraph id="8BCF368DE85A176B7EFDF8506EF6A5D0" blockId="7.[151,1436,1982,2042]" pageId="7" pageNumber="458">
<emphasis id="B904EA9FE85A176B7EFDF8506A2CA5FC" bold="true" box="[151,341,1982,2006]" pageId="7" pageNumber="458">FIGURES 3237.</emphasis>
Lectotype of
<taxonomicName id="4C704D0EE85A176B7F89F8506836A5FC" authority="Arrow, 1914" authorityName="Arrow" authorityYear="1914" box="[483,847,1982,2007]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="458" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="B904EA9FE85A176B7F89F85069B1A5FC" box="[483,712,1982,2006]" italics="true" pageId="7" pageNumber="458">Eophileurus siamensis</emphasis>
<bibRefCitation id="EFE14B7CE85A176B7CA5F8516836A5FC" author="Arrow, G. J." box="[719,847,1982,2007]" pageId="7" pageNumber="458" pagination="257 - 276" refId="ref16776" refString="Arrow, G. J. (1914) Some further notes on Lamellicorn beetles of the sub-family Dynastinae. The Annals and Magazine of natural History, including Zoology, Botany and Geology, 8 (14), 257 - 276. https: // doi. org / 10.1080 / 00222931408693571" type="journal article" year="1914">Arrow, 1914</bibRefCitation>
</taxonomicName>
(images by Keita Matsumoto ©Natural History Museum, London). 32, habitus, dorsal; 33, habitus, lateral; 34, habitus, ventral; 35, parameres, apical; 36, parameres, lateral; 37, labels.
</paragraph>
</caption>
<paragraph id="8BCF368DE85517647EADFF796E0BA386" blockId="8.[151,1437,151,2042]" pageId="8" pageNumber="459">
Viewing in a larger geographical scale, it is seen that males with extremely similar or identical external characters (
<figureCitation id="134B2A08E85517647EF5FF546A60A2FF" box="[159,281,186,213]" captionStart="FIGURES 217" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[222,1366,186,1882]" captionTargetId="figure-18@4.[212,1375,181,1885]" captionTargetPageId="4" captionText="FIGURES 217. Habitus of Eophileurus confinis Prell, 1913. 2, male (Mangshi); 3, male (Nabang); 4, female (Rongshuwang); 5, male (Mohan); 67, 11, male (Bo Kluea); 89, female (Jinghong); 10, male (Phrao); 12, male (Bao Loc); 13, female (Bao Loc); 14, male (Khao Kho); 15, male (Umphang); 16, male (Kraburi); 17, female (Kraburi); 26, 8, 1017, dorsal view; 7, 9, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853946" httpUri="https://zenodo.org/record/6853946/files/figure.png" pageId="8" pageNumber="459">Figs. 217</figureCitation>
) but with differently shaped parameres (
<figureCitation id="134B2A08E85517647C8CFF5468F0A2FE" box="[742,905,186,212]" captionStart="FIGURES 117146" captionStartId="27.[151,264,1803,1827]" captionTargetBox="[169,1424,203,1763]" captionTargetId="figure-18@27.[151,1435,181,1780]" captionTargetPageId="27" captionText="FIGURES 117146. Parameres of Eophileurus confinis Prell, 1913. 117120 from Nabang; 121122, from Lancangjiang Bridge; 123124 from Jinghong; 125128 from Wiang Papao; 129130 from Mueang Chiang Rai; 131132 from Phrao; 133 134 from Axan Mt.; 135136 from Bao Loc; 137138 from Bo Kluea; 139140 from Khao Kho; 141142 from Umphang; 143144 from Bangkok; 145146 from Kraburi; odd number ones, front view; even number ones, lateral view; 117124 without scale bar, the rest with scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853980" httpUri="https://zenodo.org/record/6853980/files/figure.png" pageId="8" pageNumber="459">Figs. 117146</figureCitation>
) can be found throughout the distribution. The general morphology of the parameres is usually similar when comparing males from the adjacent regions, while those from males of two opposite ends of the distribution are usually highly different from each other. In the next few paragraphs, the authors will demonstrate how the morphology of the parameres of the northern-most recorded individual (excluding the
<typeStatus id="54CB882FE85517647FD3FEA56A90A34F" box="[441,489,331,357]" pageId="8" pageNumber="459">type</typeStatus>
locality “
<collectingRegion id="49B4F86FE85517647C3CFEA469BAA34E" box="[598,707,330,356]" country="China" name="Shanghai" pageId="8" pageNumber="459">Shanghai</collectingRegion>
” of
<taxonomicName id="4C704D0EE85517647C99FEA56815A34E" authorityName="Prell" authorityYear="1913" box="[755,876,330,356]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE85517647C99FEA56815A34E" box="[755,876,330,356]" italics="true" pageId="8" pageNumber="459">E. confinis</emphasis>
</taxonomicName>
which is doubtful) gradually transitions to that of the southern-most recorded individual by showing parameres of four intermediate individuals linking the two in a geographically continuous line where the localities of the specimens have similarly distant gaps between them.
</paragraph>
<paragraph id="8BCF368DE85517647EADFE5B6E74A1EC" blockId="8.[151,1437,151,2042]" pageId="8" pageNumber="459">
<materialsCitation id="3B183CD0E85517647EADFE5B6E70A1EC" ID-GBIF-Occurrence="3863916306" collectionCode="LP, BN" country="China" county="The" location="Jinghong" municipality="Lancangjiang Bridge" pageId="8" pageNumber="459" specimenCount="1" stateProvince="Krung Thep Mahanakhon">
<collectingCounty id="62AE4E01E85517647EADFE5B6B8AA3E5" box="[199,243,437,463]" pageId="8" pageNumber="459">The</collectingCounty>
northern-most three individuals (L1:
<collectingMunicipality id="6BABACF7E85517647CC6FE5B68E7A3E5" box="[684,926,437,463]" pageId="8" pageNumber="459">Lancangjiang Bridge</collectingMunicipality>
, central Yunnan,
<collectingCountry id="F367761DE85517647A00FE5B6FC9A3E5" box="[1130,1200,437,463]" name="China" pageId="8" pageNumber="459">China</collectingCountry>
,
<figureCitation id="134B2A08E85517647AD5FE5B6E1FA3FA" box="[1215,1382,437,464]" captionStart="FIGURES 153164" captionStartId="29.[151,264,1313,1337]" captionTargetBox="[152,1435,181,1289]" captionTargetId="figure-18@29.[151,1435,181,1289]" captionTargetPageId="29" captionText="FIGURES 153164. Parameres of Eophileurus confinis Prell, 1913. 153154 from Lancangjiang Bridge; 155156 from Jinghong; 157158 from Wiang Papao; 159160 from Khao Kho; 161162 from Bangkok; 163164 from Kraburi; odd number ones, front view; even number ones, lateral view; not in scale." figureDoi="http://doi.org/10.5281/zenodo.6853984" httpUri="https://zenodo.org/record/6853984/files/figure.png" pageId="8" pageNumber="459">Figs. 153154</figureCitation>
; L
<figureCitation id="134B2A08E85517647BE3FE586EE1A3FA" box="[1417,1432,438,464]" captionStart="FIGURES 217" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[222,1366,186,1882]" captionTargetId="figure-18@4.[212,1375,181,1885]" captionTargetPageId="4" captionText="FIGURES 217. Habitus of Eophileurus confinis Prell, 1913. 2, male (Mangshi); 3, male (Nabang); 4, female (Rongshuwang); 5, male (Mohan); 67, 11, male (Bo Kluea); 89, female (Jinghong); 10, male (Phrao); 12, male (Bao Loc); 13, female (Bao Loc); 14, male (Khao Kho); 15, male (Umphang); 16, male (Kraburi); 17, female (Kraburi); 26, 8, 1017, dorsal view; 7, 9, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853946" httpUri="https://zenodo.org/record/6853946/files/figure.png" pageId="8" pageNumber="459">2</figureCitation>
:
<location id="8EAF6056E85517647EFDFE376A78A3D9" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85517647EFDFE376A78A3D9" box="[151,257,473,499]" country="China" county="The" municipality="Lancangjiang Bridge" name="Jinghong" pageId="8" pageNumber="459" stateProvince="Krung Thep Mahanakhon">Jinghong</location>
, southern Yunnan,
<collectingCountry id="F367761DE85517647F89FE376950A3D9" box="[483,553,473,499]" name="China" pageId="8" pageNumber="459">China</collectingCountry>
,
<figureCitation id="134B2A08E85517647C52FE376999A3DE" box="[568,736,473,500]" captionStart="FIGURES 153164" captionStartId="29.[151,264,1313,1337]" captionTargetBox="[152,1435,181,1289]" captionTargetId="figure-18@29.[151,1435,181,1289]" captionTargetPageId="29" captionText="FIGURES 153164. Parameres of Eophileurus confinis Prell, 1913. 153154 from Lancangjiang Bridge; 155156 from Jinghong; 157158 from Wiang Papao; 159160 from Khao Kho; 161162 from Bangkok; 163164 from Kraburi; odd number ones, front view; even number ones, lateral view; not in scale." figureDoi="http://doi.org/10.5281/zenodo.6853984" httpUri="https://zenodo.org/record/6853984/files/figure.png" pageId="8" pageNumber="459">Figs. 155156</figureCitation>
; L
<figureCitation id="134B2A08E85517647D6FFE34686AA3DE" box="[773,787,474,500]" captionStart="FIGURES 217" captionStartId="4.[151,264,1909,1933]" captionTargetBox="[222,1366,186,1882]" captionTargetId="figure-18@4.[212,1375,181,1885]" captionTargetPageId="4" captionText="FIGURES 217. Habitus of Eophileurus confinis Prell, 1913. 2, male (Mangshi); 3, male (Nabang); 4, female (Rongshuwang); 5, male (Mohan); 67, 11, male (Bo Kluea); 89, female (Jinghong); 10, male (Phrao); 12, male (Bao Loc); 13, female (Bao Loc); 14, male (Khao Kho); 15, male (Umphang); 16, male (Kraburi); 17, female (Kraburi); 26, 8, 1017, dorsal view; 7, 9, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6853946" httpUri="https://zenodo.org/record/6853946/files/figure.png" pageId="8" pageNumber="459">3</figureCitation>
:
<location id="8EAF6056E85517647D48FE3768C7A3DE" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85517647D48FE3768C7A3DE" box="[802,958,473,500]" country="China" county="The" municipality="Lancangjiang Bridge" name="Wiang Papao" pageId="8" pageNumber="459" stateProvince="Krung Thep Mahanakhon">Wiang Papao</location>
, northern
<collectingCountry id="F367761DE85517647A52FE376FE7A3D9" box="[1080,1182,473,499]" name="Thailand" pageId="8" pageNumber="459">Thailand</collectingCountry>
,
<figureCitation id="134B2A08E85517647AC4FE376E2DA3DE" box="[1198,1364,473,500]" captionStart="FIGURES 153164" captionStartId="29.[151,264,1313,1337]" captionTargetBox="[152,1435,181,1289]" captionTargetId="figure-18@29.[151,1435,181,1289]" captionTargetPageId="29" captionText="FIGURES 153164. Parameres of Eophileurus confinis Prell, 1913. 153154 from Lancangjiang Bridge; 155156 from Jinghong; 157158 from Wiang Papao; 159160 from Khao Kho; 161162 from Bangkok; 163164 from Kraburi; odd number ones, front view; even number ones, lateral view; not in scale." figureDoi="http://doi.org/10.5281/zenodo.6853984" httpUri="https://zenodo.org/record/6853984/files/figure.png" pageId="8" pageNumber="459">Figs. 157158</figureCitation>
) have similarly shaped parameres.
<location id="8EAF6056E85517647FB0FE13697FA03D" LSID="urn:lsid:plazi:treatment:03D9879BE85E17657EFDFF316A09A61F:8EAF6056E85517647FB0FE13697FA03D" box="[474,518,509,535]" country="China" county="The" municipality="Lancangjiang Bridge" name="The" pageId="8" pageNumber="459" stateProvince="Krung Thep Mahanakhon">The</location>
most distinct difference is on the shape of
<collectionCode id="ED61AE48E85517647D84FE106F76A032" box="[1006,1039,510,536]" country="Russia" name="Laboratory of Palaeontology" pageId="8" pageNumber="459">LP</collectionCode>
, which is expected due to the large individual variations. Another noticeable difference between them is that
<collectionCode id="ED61AE48E85517647D8CFDCC6F74A016" box="[998,1037,546,572]" pageId="8" pageNumber="459">BN</collectionCode>
are vertically long in L1, whereas slightly shorter in L2 and even shorter in L3. The general structure of these parameres can be summarized as:
<collectionCode id="ED61AE48E85517647B1FFDA86EE5A04A" box="[1397,1436,582,608]" pageId="8" pageNumber="459">BN</collectionCode>
rather long and wide, with lower margin slightly upward; ULM almost straight; WP almost at middle; LIM mostly vertical, parallel to each other. The next form can be seen in L4 (Khao Kho, northern-central
<collectingCountry id="F367761DE85517647AD9FD636E60A08D" box="[1203,1305,653,679]" name="Thailand" pageId="8" pageNumber="459">Thailand</collectingCountry>
,
<figureCitation id="134B2A08E85517647B4FFD636BBCA0E1" captionStart="FIGURES 153164" captionStartId="29.[151,264,1313,1337]" captionTargetBox="[152,1435,181,1289]" captionTargetId="figure-18@29.[151,1435,181,1289]" captionTargetPageId="29" captionText="FIGURES 153164. Parameres of Eophileurus confinis Prell, 1913. 153154 from Lancangjiang Bridge; 155156 from Jinghong; 157158 from Wiang Papao; 159160 from Khao Kho; 161162 from Bangkok; 163164 from Kraburi; odd number ones, front view; even number ones, lateral view; not in scale." figureDoi="http://doi.org/10.5281/zenodo.6853984" httpUri="https://zenodo.org/record/6853984/files/figure.png" pageId="8" pageNumber="459">Figs. 159 160</figureCitation>
). Its
<collectionCode id="ED61AE48E85517647E95FD5F6A5FA0E1" box="[255,294,689,715]" pageId="8" pageNumber="459">BN</collectionCode>
is relatively shorter and narrower than the northern ones, the lower margin is parallel, even slightly extended downward; ULM is distinctly more rounded, especially near base; WP is slightly closer to the base; BP is larger and wider than the northern ones and LIM has a gap between the upper most part that is distinctly larger than the gap between the apices (the slope is larger). Going more south, the individual from L5 (
<collectingRegion id="49B4F86FE85517647AB0FCF26E3DA11C" box="[1242,1348,796,822]" country="Thailand" name="Krung Thep Mahanakhon" pageId="8" pageNumber="459">Bangkok</collectingRegion>
, central
<collectingCountry id="F367761DE85517647EFDFCAE6B84A170" box="[151,253,832,858]" name="Thailand" pageId="8" pageNumber="459">Thailand</collectingCountry>
,
<figureCitation id="134B2A08E85517647F6DFCAE6ADEA170" box="[263,423,832,858]" captionStart="FIGURES 153164" captionStartId="29.[151,264,1313,1337]" captionTargetBox="[152,1435,181,1289]" captionTargetId="figure-18@29.[151,1435,181,1289]" captionTargetPageId="29" captionText="FIGURES 153164. Parameres of Eophileurus confinis Prell, 1913. 153154 from Lancangjiang Bridge; 155156 from Jinghong; 157158 from Wiang Papao; 159160 from Khao Kho; 161162 from Bangkok; 163164 from Kraburi; odd number ones, front view; even number ones, lateral view; not in scale." figureDoi="http://doi.org/10.5281/zenodo.6853984" httpUri="https://zenodo.org/record/6853984/files/figure.png" pageId="8" pageNumber="459">Figs. 161162</figureCitation>
) has much smaller and narrower
<collectionCode id="ED61AE48E85517647D78FCAF6845A171" box="[786,828,833,859]" pageId="8" pageNumber="459">BN</collectionCode>
, with the lower margin extending distinctly downward; ULM distinctly rounded; WP even closer to the base; BP significantly larger and wider; LIM with much larger slope. Finally, the southern-most individual from L6 (Kraburi, southern
<collectingCountry id="F367761DE85517647DE3FC666896A188" box="[905,1007,904,930]" name="Thailand" pageId="8" pageNumber="459">Thailand</collectingCountry>
,
<figureCitation id="134B2A08E85517647D97FC666FD9A188" box="[1021,1184,904,930]" captionStart="FIGURES 153164" captionStartId="29.[151,264,1313,1337]" captionTargetBox="[152,1435,181,1289]" captionTargetId="figure-18@29.[151,1435,181,1289]" captionTargetPageId="29" captionText="FIGURES 153164. Parameres of Eophileurus confinis Prell, 1913. 153154 from Lancangjiang Bridge; 155156 from Jinghong; 157158 from Wiang Papao; 159160 from Khao Kho; 161162 from Bangkok; 163164 from Kraburi; odd number ones, front view; even number ones, lateral view; not in scale." figureDoi="http://doi.org/10.5281/zenodo.6853984" httpUri="https://zenodo.org/record/6853984/files/figure.png" pageId="8" pageNumber="459">Figs. 163164</figureCitation>
) has similarly shaped
<collectionCode id="ED61AE48E85517647EFDFC426BB8A1EC" box="[151,193,940,966]" pageId="8" pageNumber="459">BN</collectionCode>
; even more rounded ULM; WP closer to base; much larger BP and LIM with slightly larger slope
</materialsCitation>
.
</paragraph>
<paragraph id="8BCF368DE85517647EADFC2169C5A7B2" blockId="8.[151,1437,151,2042]" pageId="8" pageNumber="459">
Thus, following the demonstration, it can be seen that except LP having different shape and no trace of any transition (which are expected due to the significant individual variations, even none of them has the exact same shape in the
<specimenCount id="9D76FD04E85517647F4CFBF96ABEA61B" box="[294,455,1047,1073]" pageId="8" pageNumber="459" type="generic">six specimens</specimenCount>
), the remaining parts is shown to have clear and gradual morphological transition from one to the other: from L1 to L6, BN goes smaller and narrower, with lower margin gradually extended more downward; ULM from almost straight to gradually more rounded; WP gradually closer to base in proportion to the length of the parameres; BP gradually larger and wider; LIM from vertical and parallel, to gradually having larger slope (larger gap near the opening than between the apices). The morphology of each parameres (except the outermost two) is intermediate to its adjacent others. Thus, based on this transition, most parts of the parameres is shown to have significant geographical variations, but can be linked via intermediate forms. The basic structure, however, is the same regardless of the variations: base of the parameres with a pair of overlapping notches; widest point excluding the lateral processes are close to the middle and the lateral margins are with a pair of outward processes which turn inward close to the middle, these processes can be rounded, angulate, or even with a small process-like protuberance at its outer-most point.
</paragraph>
<paragraph id="8BCF368DE85517647EADFA4C6F91A5D0" blockId="8.[151,1437,151,2042]" pageId="8" pageNumber="459">
Under these criteria, other geographical forms can be included, as those illustrated in
<figureCitation id="134B2A08E85517647AF8FA4C6E4FA796" box="[1170,1334,1442,1468]" captionStart="FIGURES 117146" captionStartId="27.[151,264,1803,1827]" captionTargetBox="[169,1424,203,1763]" captionTargetId="figure-18@27.[151,1435,181,1780]" captionTargetPageId="27" captionText="FIGURES 117146. Parameres of Eophileurus confinis Prell, 1913. 117120 from Nabang; 121122, from Lancangjiang Bridge; 123124 from Jinghong; 125128 from Wiang Papao; 129130 from Mueang Chiang Rai; 131132 from Phrao; 133 134 from Axan Mt.; 135136 from Bao Loc; 137138 from Bo Kluea; 139140 from Khao Kho; 141142 from Umphang; 143144 from Bangkok; 145146 from Kraburi; odd number ones, front view; even number ones, lateral view; 117124 without scale bar, the rest with scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853980" httpUri="https://zenodo.org/record/6853980/files/figure.png" pageId="8" pageNumber="459">Figs. 117146</figureCitation>
. In those geographical forms, some (usually at the outer region of the distribution) cannot be shown delicate transition to other forms as the above demonstration yet. This is due to the lack of specimens from certain areas, forming gaps between known populations. For the Vietnamese specimens (
<figureCitation id="134B2A08E85517647C8FF9E368F3A40D" box="[741,906,1549,1575]" captionStart="FIGURES 117146" captionStartId="27.[151,264,1803,1827]" captionTargetBox="[169,1424,203,1763]" captionTargetId="figure-18@27.[151,1435,181,1780]" captionTargetPageId="27" captionText="FIGURES 117146. Parameres of Eophileurus confinis Prell, 1913. 117120 from Nabang; 121122, from Lancangjiang Bridge; 123124 from Jinghong; 125128 from Wiang Papao; 129130 from Mueang Chiang Rai; 131132 from Phrao; 133 134 from Axan Mt.; 135136 from Bao Loc; 137138 from Bo Kluea; 139140 from Khao Kho; 141142 from Umphang; 143144 from Bangkok; 145146 from Kraburi; odd number ones, front view; even number ones, lateral view; 117124 without scale bar, the rest with scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853980" httpUri="https://zenodo.org/record/6853980/files/figure.png" pageId="8" pageNumber="459">Figs. 133136</figureCitation>
), which are separated from the remaining ones by a large gap of eastern
<collectingCountry id="F367761DE85517647FC2F9DF6977A461" box="[424,526,1585,1611]" name="Thailand" pageId="8" pageNumber="459">Thailand</collectingCountry>
, the whole of
<collectingCountry id="F367761DE85517647CCDF9DC69A4A466" box="[679,733,1586,1612]" name="Laos" pageId="8" pageNumber="459">Laos</collectingCountry>
and
<collectingCountry id="F367761DE85517647D7BF9DF68FEA461" box="[785,903,1585,1611]" name="Cambodia" pageId="8" pageNumber="459">Cambodia</collectingCountry>
, and northern
<collectingCountry id="F367761DE85517647A49F9DF6FFFA461" box="[1059,1158,1585,1611]" name="Vietnam" pageId="8" pageNumber="459">Vietnam</collectingCountry>
.And the western Yunnan population (
<figureCitation id="134B2A08E85517647F75F9BB6ABDA445" box="[287,452,1621,1647]" captionStart="FIGURES 117146" captionStartId="27.[151,264,1803,1827]" captionTargetBox="[169,1424,203,1763]" captionTargetId="figure-18@27.[151,1435,181,1780]" captionTargetPageId="27" captionText="FIGURES 117146. Parameres of Eophileurus confinis Prell, 1913. 117120 from Nabang; 121122, from Lancangjiang Bridge; 123124 from Jinghong; 125128 from Wiang Papao; 129130 from Mueang Chiang Rai; 131132 from Phrao; 133 134 from Axan Mt.; 135136 from Bao Loc; 137138 from Bo Kluea; 139140 from Khao Kho; 141142 from Umphang; 143144 from Bangkok; 145146 from Kraburi; odd number ones, front view; even number ones, lateral view; 117124 without scale bar, the rest with scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853980" httpUri="https://zenodo.org/record/6853980/files/figure.png" pageId="8" pageNumber="459">Figs. 117120</figureCitation>
), although having the central Yunnan individuals not too far away as of direct distance, the riverside from which the parent individual was collected does not link to the area where the western Yunnan population distributes, but directly connects to southern Yunnan. The transition form of that population is speculated to be around eastern
<collectingCountry id="F367761DE85517647FE0F92F6A80A4F1" box="[394,505,1729,1755]" name="Myanmar" pageId="8" pageNumber="459">Myanmar</collectingCountry>
. The Umphang individual (
<figureCitation id="134B2A08E85517647D5EF92F68A0A4F1" box="[820,985,1729,1755]" captionStart="FIGURES 117146" captionStartId="27.[151,264,1803,1827]" captionTargetBox="[169,1424,203,1763]" captionTargetId="figure-18@27.[151,1435,181,1780]" captionTargetPageId="27" captionText="FIGURES 117146. Parameres of Eophileurus confinis Prell, 1913. 117120 from Nabang; 121122, from Lancangjiang Bridge; 123124 from Jinghong; 125128 from Wiang Papao; 129130 from Mueang Chiang Rai; 131132 from Phrao; 133 134 from Axan Mt.; 135136 from Bao Loc; 137138 from Bo Kluea; 139140 from Khao Kho; 141142 from Umphang; 143144 from Bangkok; 145146 from Kraburi; odd number ones, front view; even number ones, lateral view; 117124 without scale bar, the rest with scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853980" httpUri="https://zenodo.org/record/6853980/files/figure.png" pageId="8" pageNumber="459">Figs. 141142</figureCitation>
) is highly similar to the more southern individuals and the northern-central individual, but due to the lack of specimen between them, the transition cannot yet be shown. Again, based on the basic structure being the same and the external characters being highly similar and rather stable, they are treated as
<taxonomicName id="4C704D0EE85517647C56F8C369CCA56C" authorityName="Prell" authorityYear="1913" box="[572,693,1836,1862]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE85517647C56F8C369CCA56C" box="[572,693,1836,1862]" italics="true" pageId="8" pageNumber="459">E. confinis</emphasis>
</taxonomicName>
. For the intraspecific relation, the authors follow
<bibRefCitation id="EFE14B7CE85517647A85F8C26EE5A56C" author="Endrodi, S." box="[1263,1436,1836,1862]" pageId="8" pageNumber="459" pagination="93 - 134" refId="ref16934" refString="Endrodi, S. (1977) Monographie de Dynastinae VIII Tribus Phileurini. Entomologische Abhandlungen Staatlichen Museum fur Tierkunde Dresden, 41 (4), 93 - 134. https: // doi. org / 10.1515 / 9783112546161" type="journal article" year="1977">Endrődi (1977)</bibRefCitation>
treating all geographical forms as the same species without separating out any subspecies. This is due to the large individual variations of the parameres, there seem not to be a very stable local form, only overall geographical forms which include many individuals with differently shaped but overall structurally similar parameres. These forms also do not present clear boundary between their adjacent forms. Most forms are linked to each other in a web-shaped relation, thus making it impossible to cut any part of the distribution out to form infraspecific taxa.
</paragraph>
</subSubSection>
<subSubSection id="C36A6506E85417657EADFF796A09A61F" pageId="9" pageNumber="460" type="discussion">
<paragraph id="8BCF368DE85417657EADFF7968E3A387" blockId="9.[151,1437,151,1077]" pageId="9" pageNumber="460">
<emphasis id="B904EA9FE85417657EADFF796A42A29B" bold="true" box="[199,315,151,177]" pageId="9" pageNumber="460">Remarks.</emphasis>
<bibRefCitation id="EFE14B7CE85417657F29FF796AB0A29B" author="Prell, H." box="[323,457,151,177]" pageId="9" pageNumber="460" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell (1913)</bibRefCitation>
named
<taxonomicName id="4C704D0EE85417657C49FF7968F6A29B" authority="Prell, 1913" authorityName="Prell" authorityYear="1913" box="[547,911,151,177]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE85417657C49FF796874A29B" box="[547,781,151,177]" italics="true" pageId="9" pageNumber="460">Eophileurus confinis</emphasis>
<bibRefCitation id="EFE14B7CE85417657D7EFF7968F6A29B" author="Prell, H." box="[788,911,151,177]" pageId="9" pageNumber="460" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54CB882FE85417657DF4FF796F7BA29B" box="[926,1026,151,177]" pageId="9" pageNumber="460" type="holotype">holotype</typeStatus>
<figureCitation id="134B2A08E85417657A60FF796FE9A29B" box="[1034,1168,151,177]" captionStart="FIGURES 1824" captionStartId="5.[151,264,1516,1540]" captionTargetBox="[161,1417,211,1471]" captionTargetId="figure-18@5.[151,1435,199,1492]" captionTargetPageId="5" captionText="FIGURES 1824. Holotype of Eophileurus confinis Prell, 1913 (images ©MTD/Olaf Jäger). 18, habitus, dorsal; 19, habitus, lateral; 20, habitus, ventral; 21, parameres, ventral; 22, parameres, apical; 23, parameres, lateral; 24, labels." figureDoi="http://doi.org/10.5281/zenodo.6853950" httpUri="https://zenodo.org/record/6853950/files/figure.png" pageId="9" pageNumber="460">Figs. 1824</figureCitation>
) and
<taxonomicName id="4C704D0EE85417657AA5FF796BB6A2FF" authority="Prell, 1913" authorityName="Prell" authorityYear="1913" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="decipiens">
<emphasis id="B904EA9FE85417657AA5FF796E20A29B" box="[1231,1369,151,177]" italics="true" pageId="9" pageNumber="460">E. decipiens</emphasis>
<bibRefCitation id="EFE14B7CE85417657B0AFF796BB6A2FF" author="Prell, H." pageId="9" pageNumber="460" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell, 1913</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54CB882FE85417657EB5FF556A3AA2FF" box="[223,323,187,213]" pageId="9" pageNumber="460" type="holotype">holotype</typeStatus>
<figureCitation id="134B2A08E85417657F20FF556AA8A2FF" box="[330,465,186,213]" captionStart="FIGURES 2531" captionStartId="6.[151,264,1464,1488]" captionTargetBox="[154,1434,194,1428]" captionTargetId="figure-18@6.[151,1435,181,1441]" captionTargetPageId="6" captionText="FIGURES 2531. Holotype of Eophileurus decipiens Prell, 1913 (images ©MTD/Olaf Jäger). 25, habitus, dorsal; 26, habitus, lateral; 27, habitus, ventral; 28, parameres, ventral; 29, parameres, apical; 30, parameres, lateral; 31, labels." figureDoi="http://doi.org/10.5281/zenodo.6853952" httpUri="https://zenodo.org/record/6853952/files/figure.png" pageId="9" pageNumber="460">Figs. 2531</figureCitation>
) each based on
<specimenCount id="9D76FD04E85417657CEDFF55685CA2FF" box="[647,805,187,213]" pageId="9" pageNumber="460" type="generic">one specimen</specimenCount>
. He noticed the high similarity between the two species and mentioned the possibility of a infraspecific relation between them.
<bibRefCitation id="EFE14B7CE85417657D8DFF316FFEA2D3" author="Arrow, G. J." box="[999,1159,223,249]" pageId="9" pageNumber="460" pagination="257 - 276" refId="ref16776" refString="Arrow, G. J. (1914) Some further notes on Lamellicorn beetles of the sub-family Dynastinae. The Annals and Magazine of natural History, including Zoology, Botany and Geology, 8 (14), 257 - 276. https: // doi. org / 10.1080 / 00222931408693571" type="journal article" year="1914">Arrow (1914)</bibRefCitation>
described
<taxonomicName id="4C704D0EE85417657B61FF316A5FA337" authority="Arrow, 1914" authorityName="Arrow" authorityYear="1914" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="B904EA9FE85417657B61FF316EE5A2D2" box="[1291,1436,223,248]" italics="true" pageId="9" pageNumber="460">E. siamensis</emphasis>
<bibRefCitation id="EFE14B7CE85417657EFDFEED6A5FA337" author="Arrow, G. J." box="[151,294,259,285]" pageId="9" pageNumber="460" pagination="257 - 276" refId="ref16776" refString="Arrow, G. J. (1914) Some further notes on Lamellicorn beetles of the sub-family Dynastinae. The Annals and Magazine of natural History, including Zoology, Botany and Geology, 8 (14), 257 - 276. https: // doi. org / 10.1080 / 00222931408693571" type="journal article" year="1914">Arrow, 1914</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54CB882FE85417657F5DFEED6AD9A337" box="[311,416,259,285]" pageId="9" pageNumber="460" type="lectotype">lectotype</typeStatus>
<figureCitation id="134B2A08E85417657FC2FEED6949A337" box="[424,560,259,285]" captionStart="FIGURES 3237" captionStartId="7.[151,264,1982,2006]" captionTargetBox="[198,1395,649,1953]" captionTargetId="figure-253@7.[183,1404,633,1959]" captionTargetPageId="7" captionText="FIGURES 3237. Lectotype of Eophileurus siamensis Arrow, 1914 (images by Keita Matsumoto ©Natural History Museum, London). 32, habitus, dorsal; 33, habitus, lateral; 34, habitus, ventral; 35, parameres, apical; 36, parameres, lateral; 37, labels." figureDoi="http://doi.org/10.5281/zenodo.6853954" httpUri="https://zenodo.org/record/6853954/files/figure.png" pageId="9" pageNumber="460">Figs. 3237</figureCitation>
) without mentioning
<bibRefCitation id="EFE14B7CE85417657D4DFEED68BBA337" author="Prell, H." box="[807,962,259,285]" pageId="9" pageNumber="460" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prells (1913)</bibRefCitation>
species. In the revisional work involving
<taxonomicName id="4C704D0EE85417657EFDFEC96AA2A36B" authority=", Endrodi (1977)" authorityName=", Endrodi" authorityYear="1977" box="[151,475,295,321]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="genus">
<emphasis id="B904EA9FE85417657EFDFEC96A59A36B" box="[151,288,295,321]" italics="true" pageId="9" pageNumber="460">Eophileurus</emphasis>
,
<bibRefCitation id="EFE14B7CE85417657F45FEC96AA2A36B" author="Endrodi, S." box="[303,475,295,321]" pageId="9" pageNumber="460" pagination="93 - 134" refId="ref16934" refString="Endrodi, S. (1977) Monographie de Dynastinae VIII Tribus Phileurini. Entomologische Abhandlungen Staatlichen Museum fur Tierkunde Dresden, 41 (4), 93 - 134. https: // doi. org / 10.1515 / 9783112546161" type="journal article" year="1977">Endrődi (1977)</bibRefCitation>
</taxonomicName>
examined many additional specimens and treated both
<taxonomicName id="4C704D0EE85417657A24FEC96FA1A36B" authorityName="Prell" authorityYear="1913" box="[1102,1240,295,321]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="decipiens">
<emphasis id="B904EA9FE85417657A24FEC96FA1A36B" box="[1102,1240,295,321]" italics="true" pageId="9" pageNumber="460">E. decipiens</emphasis>
</taxonomicName>
and
<taxonomicName id="4C704D0EE85417657B7AFEC96EE5A36A" authorityName="Arrow" authorityYear="1914" box="[1296,1436,295,320]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="B904EA9FE85417657B7AFEC96EE5A36A" box="[1296,1436,295,320]" italics="true" pageId="9" pageNumber="460">E. siamensis</emphasis>
</taxonomicName>
as junior synonyms of
<taxonomicName id="4C704D0EE85417657FFDFEA56969A34F" authorityName="Prell" authorityYear="1913" box="[407,528,331,357]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE85417657FFDFEA56969A34F" box="[407,528,331,357]" italics="true" pageId="9" pageNumber="460">E. confinis</emphasis>
</taxonomicName>
. The authors agree with the treatment by Endrődi based on material examined in this paper.
<taxonomicName id="4C704D0EE85417657F7AFE81682DA3A3" authority="Yamaya &amp; Muramoto, 2008" authorityName="Yamaya &amp; Muramoto" authorityYear="2008" box="[272,852,367,393]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="takakuwai">
<emphasis id="B904EA9FE85417657F7AFE81696AA3A3" box="[272,531,367,393]" italics="true" pageId="9" pageNumber="460">Eophileurus takakuwai</emphasis>
<bibRefCitation id="EFE14B7CE85417657C72FE81682DA3A3" author="Yamaya, S. &amp; Muramoto, R." box="[536,852,367,393]" pageId="9" pageNumber="460" pagination="1 - 34" refId="ref17677" refString="Yamaya, S. &amp; Muramoto, R. (2008) On the genus Eophileurus Arrow, 1908 (Scarabaeidae, Dynastinae), with descriptions of five new species. Kogane, Supplement 2, 1 - 34." type="journal article" year="2008">Yamaya &amp; Muramoto, 2008</bibRefCitation>
</taxonomicName>
, whose characters also lie within the variations of
<taxonomicName id="4C704D0EE85417657BEEFE816B89A387" authorityName="Prell" authorityYear="1913" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE85417657BEEFE816B89A387" italics="true" pageId="9" pageNumber="460">E. confinis</emphasis>
</taxonomicName>
, is thereby treated as a junior synonym of
<taxonomicName id="4C704D0EE85417657CA7FE7D683CA387" authorityName="Prell" authorityYear="1913" box="[717,837,403,429]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE85417657CA7FE7D683CA387" box="[717,837,403,429]" italics="true" pageId="9" pageNumber="460">E. confinis</emphasis>
</taxonomicName>
herein.
</paragraph>
<paragraph id="8BCF368DE85417657EADFE596FFFA1E3" blockId="9.[151,1437,151,1077]" pageId="9" pageNumber="460">
Several historical issues remain for
<taxonomicName id="4C704D0EE85417657C0BFE5969A3A3FB" authorityName="Prell" authorityYear="1913" box="[609,730,439,465]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE85417657C0BFE5969A3A3FB" box="[609,730,439,465]" italics="true" pageId="9" pageNumber="460">E. confinis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C704D0EE85417657C80FE59680CA3FB" authorityName="Prell" authorityYear="1913" box="[746,885,439,465]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="decipiens">
<emphasis id="B904EA9FE85417657C80FE59680CA3FB" box="[746,885,439,465]" italics="true" pageId="9" pageNumber="460">E. decipiens</emphasis>
</taxonomicName>
and
<taxonomicName id="4C704D0EE85417657DC5FE596F45A3FA" authorityName="Arrow" authorityYear="1914" box="[943,1084,439,464]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="B904EA9FE85417657DC5FE596F45A3FA" box="[943,1084,439,464]" italics="true" pageId="9" pageNumber="460">E. siamensis</emphasis>
</taxonomicName>
. Firstly, the type locality of
<taxonomicName id="4C704D0EE85417657BEEFE596B89A3DF" authorityName="Prell" authorityYear="1913" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE85417657BEEFE596B89A3DF" italics="true" pageId="9" pageNumber="460">E. confinis</emphasis>
</taxonomicName>
is unclear since the locality label of the
<typeStatus id="54CB882FE85417657CC5FE356868A3DF" box="[687,785,475,501]" pageId="9" pageNumber="460" type="holotype">holotype</typeStatus>
(
<figureCitation id="134B2A08E85417657D4AFE356809A3DF" box="[800,880,475,501]" captionStart="FIGURES 1824" captionStartId="5.[151,264,1516,1540]" captionTargetBox="[161,1417,211,1471]" captionTargetId="figure-18@5.[151,1435,199,1492]" captionTargetPageId="5" captionText="FIGURES 1824. Holotype of Eophileurus confinis Prell, 1913 (images ©MTD/Olaf Jäger). 18, habitus, dorsal; 19, habitus, lateral; 20, habitus, ventral; 21, parameres, ventral; 22, parameres, apical; 23, parameres, lateral; 24, labels." figureDoi="http://doi.org/10.5281/zenodo.6853950" httpUri="https://zenodo.org/record/6853950/files/figure.png" pageId="9" pageNumber="460">Fig. 24</figureCitation>
) states “Shangkai” (possibly referring to a village in Manipur,
<collectingCountry id="F367761DE85417657F4BFE116A25A033" box="[289,348,511,537]" name="India" pageId="9" pageNumber="460">India</collectingCountry>
) while original publication stated “
<collectingCountry id="F367761DE85417657C8EFE116850A033" box="[740,809,511,537]" name="China" pageId="9" pageNumber="460">China</collectingCountry>
(
<collectingRegion id="49B4F86FE85417657D52FE1168DCA033" box="[824,933,511,537]" country="China" name="Shanghai" pageId="9" pageNumber="460">Shanghai</collectingRegion>
)” instead. It is possible that Prell misread the label or Felsche personally informed its origin of
<collectingCountry id="F367761DE85417657CD6FDCD6986A017" box="[700,767,547,573]" name="China" pageId="9" pageNumber="460">China</collectingCountry>
to Prell. Overall, the exact situation is now untraceable. The conclusion cannot be drawn currently because no specimen has been discovered from both localities subsequently. Until further discovery is made, the authors suggest that the type locality should be considered as “
<collectingRegion id="49B4F86FE85417657ABCFD856E3AA0AF" box="[1238,1347,619,645]" country="China" name="Shanghai" pageId="9" pageNumber="460">Shanghai</collectingRegion>
,
<collectingCountry id="F367761DE85417657B27FD856EEBA0AF" box="[1357,1426,619,645]" name="China" pageId="9" pageNumber="460">China</collectingCountry>
” as originally stated in
<bibRefCitation id="EFE14B7CE85417657FE7FD61696FA083" author="Prell, H." box="[397,534,655,681]" pageId="9" pageNumber="460" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell (1913)</bibRefCitation>
, although it is unlikely that a
<taxonomicName id="4C704D0EE85417657D0AFD616F38A083" box="[864,1089,655,681]" pageId="9" pageNumber="460">
<emphasis id="B904EA9FE85417657D0AFD616890A083" box="[864,1001,655,681]" italics="true" pageId="9" pageNumber="460">Eophileurus</emphasis>
species
</taxonomicName>
mainly known from Indochina could distribute to
<collectingRegion id="49B4F86FE85417657F05FD5D6AA2A0E7" box="[367,475,691,717]" country="China" name="Shanghai" pageId="9" pageNumber="460">Shanghai</collectingRegion>
. Secondly, the type locality of
<taxonomicName id="4C704D0EE85417657D54FD5D68B1A0E7" authorityName="Prell" authorityYear="1913" box="[830,968,691,717]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="decipiens">
<emphasis id="B904EA9FE85417657D54FD5D68B1A0E7" box="[830,968,691,717]" italics="true" pageId="9" pageNumber="460">E. decipiens</emphasis>
</taxonomicName>
is “Casamanca”, which is clearly falsely labeled (
<bibRefCitation id="EFE14B7CE85417657E9FFD396A15A0DB" author="Prell, H." box="[245,364,727,753]" pageId="9" pageNumber="460" pagination="103 - 124" refId="ref17521" refString="Prell, H. (1913) Beitrage zur Kenntnis der Dynastinen. viii. Ueber das Genus Eophilurus Arrow. Memoires de la Societe Entomologique de Belgique, 22, 103 - 124." type="journal article" year="1913">Prell 1913</bibRefCitation>
;
<bibRefCitation id="EFE14B7CE85417657F1DFD39696BA0DB" author="Endrodi, S." box="[375,530,727,753]" pageId="9" pageNumber="460" pagination="93 - 134" refId="ref16934" refString="Endrodi, S. (1977) Monographie de Dynastinae VIII Tribus Phileurini. Entomologische Abhandlungen Staatlichen Museum fur Tierkunde Dresden, 41 (4), 93 - 134. https: // doi. org / 10.1515 / 9783112546161" type="journal article" year="1977">Endrődi 1977</bibRefCitation>
). Considering the high similarity of the parameres of the
<typeStatus id="54CB882FE85417657AEEFD396F9FA0DB" box="[1156,1254,727,753]" pageId="9" pageNumber="460" type="holotype">holotype</typeStatus>
(
<figureCitation id="134B2A08E85417657A9EFD396E00A0DB" box="[1268,1401,727,753]" captionStart="FIGURES 2531" captionStartId="6.[151,264,1464,1488]" captionTargetBox="[154,1434,194,1428]" captionTargetId="figure-18@6.[151,1435,181,1441]" captionTargetPageId="6" captionText="FIGURES 2531. Holotype of Eophileurus decipiens Prell, 1913 (images ©MTD/Olaf Jäger). 25, habitus, dorsal; 26, habitus, lateral; 27, habitus, ventral; 28, parameres, ventral; 29, parameres, apical; 30, parameres, lateral; 31, labels." figureDoi="http://doi.org/10.5281/zenodo.6853952" httpUri="https://zenodo.org/record/6853952/files/figure.png" pageId="9" pageNumber="460">Figs. 2830</figureCitation>
) to those of the population from northern
<collectingCountry id="F367761DE85417657C54FD1569DBA13F" box="[574,674,763,789]" name="Thailand" pageId="9" pageNumber="460">Thailand</collectingCountry>
(
<figureCitation id="134B2A08E85417657CDAFD15682DA13F" box="[688,852,762,789]" captionStart="FIGURES 117146" captionStartId="27.[151,264,1803,1827]" captionTargetBox="[169,1424,203,1763]" captionTargetId="figure-18@27.[151,1435,181,1780]" captionTargetPageId="27" captionText="FIGURES 117146. Parameres of Eophileurus confinis Prell, 1913. 117120 from Nabang; 121122, from Lancangjiang Bridge; 123124 from Jinghong; 125128 from Wiang Papao; 129130 from Mueang Chiang Rai; 131132 from Phrao; 133 134 from Axan Mt.; 135136 from Bao Loc; 137138 from Bo Kluea; 139140 from Khao Kho; 141142 from Umphang; 143144 from Bangkok; 145146 from Kraburi; odd number ones, front view; even number ones, lateral view; 117124 without scale bar, the rest with scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.6853980" httpUri="https://zenodo.org/record/6853980/files/figure.png" pageId="9" pageNumber="460">Figs. 125132</figureCitation>
), the possible origin of the
<typeStatus id="54CB882FE85417657AEBFD156F9AA13F" box="[1153,1251,763,789]" pageId="9" pageNumber="460" type="holotype">holotype</typeStatus>
could be around that area. Lastly,
<taxonomicName id="4C704D0EE85417657F39FCF16916A113" authority="Arrow, 1914" authorityName="Arrow" authorityYear="1914" box="[339,623,799,825]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="B904EA9FE85417657F39FCF16AA7A112" box="[339,478,799,824]" italics="true" pageId="9" pageNumber="460">E. siamensis</emphasis>
<bibRefCitation id="EFE14B7CE85417657F89FCF16916A113" author="Arrow, G. J." box="[483,623,799,825]" pageId="9" pageNumber="460" pagination="257 - 276" refId="ref16776" refString="Arrow, G. J. (1914) Some further notes on Lamellicorn beetles of the sub-family Dynastinae. The Annals and Magazine of natural History, including Zoology, Botany and Geology, 8 (14), 257 - 276. https: // doi. org / 10.1080 / 00222931408693571" type="journal article" year="1914">Arrow, 1914</bibRefCitation>
</taxonomicName>
was described from “
<collectingRegion id="49B4F86FE85417657D37FCF168BEA113" box="[861,967,799,825]" country="Thailand" name="Krung Thep Mahanakhon" pageId="9" pageNumber="460">Bangkok</collectingRegion>
” and “Chantabon” based on
<specimenCount id="9D76FD04E85417657B6EFCF16EE0A113" box="[1284,1433,799,825]" pageId="9" pageNumber="460" type="generic" typeStatus="syntypes">two syntypes</specimenCount>
. The originally dissected
<typeStatus id="54CB882FE85417657FDEFCAD6972A177" box="[436,523,835,861]" pageId="9" pageNumber="460" type="syntype">syntype</typeStatus>
(from
<collectingRegion id="49B4F86FE85417657C35FCAD69BEA177" box="[607,711,835,861]" country="Thailand" name="Krung Thep Mahanakhon" pageId="9" pageNumber="460">Bangkok</collectingRegion>
,
<figureCitation id="134B2A08E85417657CBFFCAD6827A177" box="[725,862,835,861]" captionStart="FIGURES 3237" captionStartId="7.[151,264,1982,2006]" captionTargetBox="[198,1395,649,1953]" captionTargetId="figure-253@7.[183,1404,633,1959]" captionTargetPageId="7" captionText="FIGURES 3237. Lectotype of Eophileurus siamensis Arrow, 1914 (images by Keita Matsumoto ©Natural History Museum, London). 32, habitus, dorsal; 33, habitus, lateral; 34, habitus, ventral; 35, parameres, apical; 36, parameres, lateral; 37, labels." figureDoi="http://doi.org/10.5281/zenodo.6853954" httpUri="https://zenodo.org/record/6853954/files/figure.png" pageId="9" pageNumber="460">Figs. 3237</figureCitation>
) matches the original description and is therefore designated as the
<typeStatus id="54CB882FE85417657F03FC896AAAA1AB" box="[361,467,871,897]" pageId="9" pageNumber="460" type="lectotype">lectotype</typeStatus>
. The other
<typeStatus id="54CB882FE85417657C31FC8969CBA1AB" box="[603,690,871,897]" pageId="9" pageNumber="460" type="syntype">syntype</typeStatus>
(from Chantabon) was dissected by Keita Matsumoto (BMNH) and proved to be
<taxonomicName id="4C704D0EE85417657F36FC65694DA18F" authority="Yang &amp; Pathomwattananurak, 2022" authorityName="Yang &amp; Pathomwattananurak" authorityYear="2022" box="[348,564,907,933]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="quadratifovealis" status="sp. nov.">
<emphasis id="B904EA9FE85417657F36FC65694DA18F" box="[348,564,907,933]" italics="true" pageId="9" pageNumber="460">E. quadratifovealis</emphasis>
</taxonomicName>
,
<emphasis id="B904EA9FE85417657C28FC6569AAA18E" bold="true" box="[578,723,906,933]" pageId="9" pageNumber="460">
<taxonomicNameLabel id="A23757E4E85417657C28FC6569B6A18E" box="[578,719,906,933]" pageId="9" pageNumber="460" rank="species">new species</taxonomicNameLabel>
.
</emphasis>
The spelling of “Chantabon” is likely a typo of “Chantaboon”, currently known as
<collectingRegion id="49B4F86FE85417657F1FFC41697BA1E3" box="[373,514,943,969]" country="Thailand" name="Chanthaburi" pageId="9" pageNumber="460">Chanthaburi</collectingRegion>
, where the
<typeStatus id="54CB882FE85417657CE8FC41699DA1E3" box="[642,740,943,969]" pageId="9" pageNumber="460" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C704D0EE85417657D63FC416899A1E3" authorityName="Yang &amp; Pathomwattananurak" authorityYear="2022" box="[777,992,943,969]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="quadratifovealis">
<emphasis id="B904EA9FE85417657D63FC416899A1E3" box="[777,992,943,969]" italics="true" pageId="9" pageNumber="460">E. quadratifovealis</emphasis>
</taxonomicName>
was collected.
</paragraph>
<paragraph id="8BCF368DE85417657EADFC3D6A09A61F" blockId="9.[151,1437,151,1077]" pageId="9" pageNumber="460">
Additionally, the authors noticed that
<taxonomicName id="4C704D0EE85417657C0DFC3D69A7A1C7" authorityName="Prell" authorityYear="1913" box="[615,734,979,1005]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="confinis">
<emphasis id="B904EA9FE85417657C0DFC3D69A7A1C7" box="[615,734,979,1005]" italics="true" pageId="9" pageNumber="460">E. confinis</emphasis>
</taxonomicName>
from
<collectingCountry id="F367761DE85417657D75FC3D68FAA1C7" box="[799,899,979,1005]" name="Thailand" pageId="9" pageNumber="460">Thailand</collectingCountry>
and
<collectingCountry id="F367761DE85417657DDCFC3D6895A1C7" box="[950,1004,979,1005]" name="Laos" pageId="9" pageNumber="460">Laos</collectingCountry>
was misidentified as
<taxonomicName id="4C704D0EE85417657AB3FC3D6BB6A63B" authority="Endrodi, 1978" authorityName="Endrodi" authorityYear="1978" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B904EA9FE85417657AB3FC3D6E4CA1C7" box="[1241,1333,979,1005]" italics="true" pageId="9" pageNumber="460">E. malyi</emphasis>
<bibRefCitation id="EFE14B7CE85417657B51FC3D6BB6A63B" author="Endrodi, S." pageId="9" pageNumber="460" pagination="183 - 185" refId="ref16975" refString="Endrodi, S. (1978) Zwei neue Eophileurus Arten aus Thailand. Annales historico-naturales Musei nationalis hungarici, 70, 183 - 185." type="journal article" year="1978">Endrődi, 1978</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54CB882FE85417657EB4FC196A69A63B" box="[222,272,1015,1041]" pageId="9" pageNumber="460">type</typeStatus>
material examined) by
<bibRefCitation id="EFE14B7CE85417657C7DFC196819A63B" author="Yamaya, S. &amp; Muramoto, R." box="[535,864,1015,1041]" pageId="9" pageNumber="460" pagination="1 - 34" refId="ref17677" refString="Yamaya, S. &amp; Muramoto, R. (2008) On the genus Eophileurus Arrow, 1908 (Scarabaeidae, Dynastinae), with descriptions of five new species. Kogane, Supplement 2, 1 - 34." type="journal article" year="2008">Yamaya &amp; Muramoto (2008)</bibRefCitation>
. Additional comments are presented in the remarks section of
<taxonomicName id="4C704D0EE85417657F66FBF56A10A61F" authorityName="Endrodi" authorityYear="1978" box="[268,361,1051,1077]" class="Insecta" family="Dynastidae" genus="Eophileurus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="malyi">
<emphasis id="B904EA9FE85417657F66FBF56A10A61F" box="[268,361,1051,1077]" italics="true" pageId="9" pageNumber="460">E. malyi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>