plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/CF/A3/03CFA3486B6FFFC9FF13FE6F5B5EFE9B.xml
maintenance bc2564723d added 03
2024-06-21 12:22:17 +02:00

380 lines
45 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="0F465576712FE684FE4CDA2ED770281B" ID-DOI="10.1649/0010-065x-69.3.463" ID-ISSN="1938-4394" ID-Zenodo-Dep="10106050" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630175225603" checkinUser="diego" docAuthor="Filippini, V., Galante, E. &amp; Micó, E." docDate="2015" docId="03CFA3486B6FFFC9FF13FE6F5B5EFE9B" docLanguage="en" docName="0010-065X-69.3.463.pdf.imf" docOrigin="The Coleopterists Bulletin 69 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-69.3.463" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Anomala mesosticta Filippini, Galante, and Mico, new species" docType="treatment" docVersion="3" lastPageNumber="472" masterDocId="FFF6DB306B66FFC3FFBEFFE95A73FFC7" masterDocTitle="Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica" masterLastPageNumber="476" masterPageNumber="463" pageNumber="472" updateTime="1699636354092" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="815487115C10DC4E5B1798FFD0193C1F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7982BBD0365AA3EF43E8EC154184BE3D">
<mods:title id="EABD6B78EC6BF1B372DE4A62600CC731">Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica</mods:title>
</mods:titleInfo>
<mods:name id="CA6E7A53FCB1C85A62F38FEC698E53D9" type="personal">
<mods:role id="09E62B5308DCAD80A0909697D9A734B9">
<mods:roleTerm id="E41E4D4E7508EEEB517F64067DF82C61">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DC55D2A61A26E89456C3779533030150">Filippini, V.</mods:namePart>
</mods:name>
<mods:name id="1845D3FFA9458EAF5D49689BEE2A8F13" type="personal">
<mods:role id="96FDB7D0BD7EFA052C9B127ADDEFD1A9">
<mods:roleTerm id="2395D9FDC298585984DE012ED5797D81">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C94B97BFC6C46DBCE7D3AE5AECFEF58F">Galante, E.</mods:namePart>
</mods:name>
<mods:name id="FCBF157E05EDC6FABAB5640D3C98584A" type="personal">
<mods:role id="A4596D7E7D7ED888491747B0DB9DEB9D">
<mods:roleTerm id="A81737661FAC88A6532224E7DA8F670C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B74682134D940E98FBD60D51C2287AA5">Micó, E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="80E46A3517BDA4171E34C7693E31787F">text</mods:typeOfResource>
<mods:relatedItem id="B1B571B164D7417D4A6B53FFFF0735CB" type="host">
<mods:titleInfo id="A9489CC186C58039EE728701891D0FD3">
<mods:title id="FD6B2341F642CA8E77AECD919DCD8058">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="91D735C5E5B632B28D7FC0A62540D435">
<mods:date id="4011EF391C0AA6CADEEA10A4055B663A">2015</mods:date>
<mods:detail id="C9C70D4A1095BA5B5FA1ABBA1DD73792" type="pubDate">
<mods:number id="162172CE9168E92C3B36BDE9B609DE41">2015-09-30</mods:number>
</mods:detail>
<mods:detail id="88D3C8DF55920AAE372C4F0304FC8983" type="volume">
<mods:number id="97EDCC76BD2FA86FB733D00920D24B09">69</mods:number>
</mods:detail>
<mods:detail id="A3AFF7A3A9540D2518D0DEB38EB0EB79" type="issue">
<mods:number id="79D43EBEC6212D4FA62AE312E0814F9B">3</mods:number>
</mods:detail>
<mods:extent id="DC318B28D999FD91F1CEF0864A70DB16" unit="page">
<mods:start id="D8907676AA08663107958086DDE64CBE">463</mods:start>
<mods:end id="BE168A27D70EC664810D32F2244C47B8">476</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="02B4F1309099E54F3605BCEE280D1C07">
<mods:url id="79681A0A6F7141EEF419947196786841">http://dx.doi.org/10.1649/0010-065x-69.3.463</mods:url>
</mods:location>
<mods:classification id="D045410FD312EE466271B414897CDD8C">journal article</mods:classification>
<mods:identifier id="6B933C4710F7858CF8FC07D5B475C608" type="DOI">10.1649/0010-065x-69.3.463</mods:identifier>
<mods:identifier id="597B2AF571A35829BCE5B0BF2370ED5C" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="4810DC926FF29BCF460652BD78A93206" type="Zenodo-Dep">10106050</mods:identifier>
<mods:identifier id="1DA4DBF7EF9748FB6B990D0E9820559E" type="ZooBank">9A0330AD-EEF7-4A80-8D80-1B6116BD856F</mods:identifier>
</mods:mods>
<treatment id="03CFA3486B6FFFC9FF13FE6F5B5EFE9B" LSID="urn:lsid:plazi:treatment:03CFA3486B6FFFC9FF13FE6F5B5EFE9B" httpUri="http://treatment.plazi.org/id/03CFA3486B6FFFC9FF13FE6F5B5EFE9B" lastPageId="10" lastPageNumber="472" pageId="9" pageNumber="472">
<subSubSection id="C37C41D56B6FFFCAFF13FE6F5B94FE71" pageId="9" pageNumber="472" type="nomenclature">
<paragraph id="8BD9125E6B6FFFCAFF13FE6F5B94FE71" blockId="9.[173,616,390,465]" pageId="9" pageNumber="472">
<heading id="D091A5326B6FFFCAFF13FE6F5B94FE71" centered="true" fontSize="10" level="2" pageId="9" pageNumber="472" reason="2">
<emphasis id="B912CE4C6B6FFFCAFF13FE6F5B94FE71" bold="true" pageId="9" pageNumber="472">
<taxonomicName id="4C6669DD6B6FFFCAFF13FE6F5B17FE71" authority="Filippini, Galante, and Mico" authorityName="Filippini, Galante, and Mico" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="472" phylum="Arthropoda" rank="species" species="mesosticta" status="sp. nov.">
<emphasis id="B912CE4C6B6FFFCAFF13FE6F5B07FE5B" bold="true" box="[173,372,390,412]" italics="true" pageId="9" pageNumber="472">Anomala mesosticta</emphasis>
Filippini, Galante, and Micó
</taxonomicName>
,
<taxonomicNameLabel id="A22173376B6FFFCAFECFFE485B94FE71" box="[369,487,417,438]" pageId="9" pageNumber="472" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6FFFCAFEABFE555873FE16" box="[277,512,444,465]" pageId="9" pageNumber="472" type="description">
<paragraph id="8BD9125E6B6FFFCAFEABFE555873FE16" blockId="9.[173,616,390,465]" box="[277,512,444,465]" pageId="9" pageNumber="472">
(
<figureCitation id="135D0EDB6B6FFFCAFEA2FE555B11FE16" box="[284,354,444,465]" captionStart="Figs" captionStartId="2.[197,237,1274,1293]" captionTargetBox="[193,1167,186,1250]" captionTargetId="figure-199@2.[178,1188,183,1261]" captionTargetPageId="2" captionText="Figs. 1–8. Anomala species holotypes, habitus. 1) A. coffea; 2) A. cyclops; 3) A. divisa; 4) A. mersa; 5) A. mesosticta;" figureDoi="http://doi.org/10.5281/zenodo.10106052" httpUri="https://zenodo.org/record/10106052/files/figure.png" pageId="9" pageNumber="472">Figs. 5</figureCitation>
,
<figureCitation id="135D0EDB6B6FFFCAFED1FE555BFAFE16" box="[367,393,444,465]" captionStart="Figs" captionStartId="3.[175,216,1111,1130]" captionTargetBox="[154,1163,186,1089]" captionTargetId="figure-261@3.[151,1173,183,1093]" captionTargetPageId="3" captionText="Figs. 9–16. Anomala species, protibiae. 9) A. coffea (male: holotype; female: paratype, Estación Pitilla, Costa Rica); 10) A. cyclops (male: holotype; female: paratype, Los Almendros, Costa Rica); 11) A. divisa (male: holotype; female: paratype, Estación Biológica Las Alturas, Costa Rica); 12) A. mersa (male: paratype, Sector Palo Verde, Costa Rica); 13) A. mesosticta (male: holotype; female: paratype, Rio Sardinas, Costa Rica); 14) A. robiginosa (male: holotype; female: paratype, Santa Rosa National Park, Costa Rica); 15) A. strigodermoides (male: holotype; female: paratype, Catarata río Buenavista, Alajuela); 16) A. unilineata (holotype). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106054" httpUri="https://zenodo.org/record/10106054/files/figure.png" pageId="9" pageNumber="472">13</figureCitation>
,
<figureCitation id="135D0EDB6B6FFFCAFE2BFE555BDCFE16" box="[405,431,444,465]" captionStart="Figs" captionStartId="4.[197,237,1207,1226]" captionTargetBox="[176,1181,187,1184]" captionTargetId="figure-197@4.[173,1195,183,1197]" captionTargetPageId="4" captionText="Figs. 17–24. Anomala species, aedeagi, lateral view (top), dorsal view (bottom left), ventral view (bottom right). 17) A. coffea (paratype, Estación Pitilla, Costa Rica); 18) A. cyclops (holotype); 19) A. divisa (holotype); 20) A. mersa (holotype); 21) A. mesosticta (holotype); 22) A. robiginosa (paratype, Zarcero, Costa Rica); 23) A. strigodermoides (holotype); 24) A. unilineata (paratype, Santa Rosa National Park, Costa Rica). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106056" httpUri="https://zenodo.org/record/10106056/files/figure.png" pageId="9" pageNumber="472">21</figureCitation>
,
<figureCitation id="135D0EDB6B6FFFCAFE05FE555BA6FE16" box="[443,469,444,465]" captionStart="Figs" captionStartId="5.[175,215,1638,1657]" captionTargetBox="[178,1140,195,1606]" captionTargetId="figure-1@5.[168,1156,183,1627]" captionTargetPageId="5" captionText="Figs. 25–33. Anomala species, genitalia. Endophalli: 25) A. coffea (paratype, Estación Pitilla, Costa Rica); 26) A. cyclops (paratype, Finca Jenny, Costa Rica); 27) A. divisa (holotype); 28) A. mersa (paratype, Sector Palo Verde, Costa Rica); 29) A. mesosticta (holotype); 30) A. robiginosa (holotype); 31) A. strigodermoides (holotype); 32) A. unilineata (holotype). Aedeagus: 33) A. sticticoptera (lateral view, Cuyotenango, Guatemala, MNHUB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106058" httpUri="https://zenodo.org/record/10106058/files/figure.png" pageId="9" pageNumber="472">29</figureCitation>
,
<figureCitation id="135D0EDB6B6FFFCAFE5EFE555B8AFE16" box="[480,505,444,465]" captionStart="Figs" captionStartId="6.[197,237,1016,1035]" captionTargetBox="[178,1183,206,993]" captionTargetId="figure-344@6.[173,1193,183,1019]" captionTargetPageId="6" captionText="Figs. 34–36. Anomala species, distribution maps. 34) Distribution of A. coffea, A. mersa, and A. robiginosa;" figureDoi="http://doi.org/10.5281/zenodo.10106060" httpUri="https://zenodo.org/record/10106060/files/figure.png" pageId="9" pageNumber="472">35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6FFFCAFF11FE0F595FFBC8" pageId="9" pageNumber="472" type="materials_examined">
<paragraph id="8BD9125E6B6FFFCAFF11FE0F5861FB69" blockId="9.[151,639,486,1729]" pageId="9" pageNumber="472">
<emphasis id="B912CE4C6B6FFFCAFF11FE0F5B4DFE3C" bold="true" box="[175,318,486,507]" pageId="9" pageNumber="472">Type Material.</emphasis>
<materialsCitation id="3B0E18036B6FFFCAFEF5FE0F5BBDFD8C" collectingDate="1993-03-20" collectingDateMax="1993-03-27" collectingDateMin="1993-03-20" collectionCode="MNCR" collectorName="F. Araya." country="Costa Rica" elevation="30" location="Los Arbolitos" pageId="9" pageNumber="472" specimenCount="31" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54DDACFC6B6FFFCAFEF5FE0F5BD7FE3C" box="[331,420,486,507]" pageId="9" pageNumber="472" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D60D9D76B6FFFCAFE14FE0F5BCFFE3B" box="[426,444,486,508]" pageId="9" pageNumber="472" type="male">♂</specimenCount>
&quot;
<location id="8EB944856B6FFFCAFE79FE0F5834FE3C" LSID="urn:lsid:plazi:treatment:03CFA3486B6FFFC9FF13FE6F5B5EFE9B:8EB944856B6FFFCAFE79FE0F5834FE3C" box="[455,583,486,507]" country="Costa Rica" name="Los Arbolitos" pageId="9" pageNumber="472">Los Arbolitos</location>
, Prov. Hered.
<collectingCountry id="F37152CE6B6FFFCAFF60FDE85B1FFDD1" box="[222,364,513,534]" name="Costa Rica" pageId="9" pageNumber="472">COSTA RICA</collectingCountry>
.
<specimenCount id="9D60D9D76B6FFFCAFEC9FDE85BDBFDD1" box="[375,424,513,534]" pageId="9" pageNumber="472" type="generic">
<quantity id="4C9EBFBB6B6FFFCAFEC9FDE85BD7FDD1" box="[375,420,513,534]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="472" unit="m" value="30.0">
<elevation id="004BF56D6B6FFFCAFEC9FDE85BD7FDD1" box="[375,420,513,534]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="472" unit="m" value="30.0">30m</elevation>
</quantity>
.
</specimenCount>
<date id="FFD8349E6B6FFFCAFE11FDE85811FDD1" box="[431,610,512,534]" pageId="9" pageNumber="472" value="1993-03-20" valueMax="1993-03-27" valueMin="1993-03-20">
<collectingDate id="EF9CCD766B6FFFCAFE11FDE85811FDD1" box="[431,610,512,534]" pageId="9" pageNumber="472" value="1993-03-20" valueMax="1993-03-27" valueMin="1993-03-20">20–27 MAR 1993</collectingDate>
</date>
.
<collectorName id="269377886B6FFFCAFDD2FDE85AAAFDF7" pageId="9" pageNumber="472">F. Araya.</collectorName>
L_N_291400_536100 #1952 / INBIOCRI 001677111&quot; deposited at
<collectionCode id="ED778A9B6B6FFFCAFE38FDDF5BBDFD8C" box="[390,462,566,587]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="9" pageNumber="472">MNCR</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B0E18036B6FFFCAFE66FDDF5809FD72" collectingDate="1993-03-09" collectingDateMax="1993-03-30" collectingDateMin="1993-03-09" collectionCode="CEUA" collectorName="M. Ortiz." country="Costa Rica" elevation="70" location="Amubri" municipality="Prov." pageId="9" pageNumber="472" specimenCode="CEUA00106193" specimenCount="4" specimenCount-male="4" stateProvince="Limon" typeStatus="paratype">
<typeStatus id="54DDACFC6B6FFFCAFE66FDDF5843FD8C" box="[472,560,566,587]" pageId="9" pageNumber="472" type="paratype">Paratypes</typeStatus>
(9):
<specimenCount id="9D60D9D76B6FFFCAFDDEFDDF580DFD8C" box="[608,638,565,587]" pageId="9" pageNumber="472" type="male">2♂</specimenCount>
&quot;La Virgen de Sarapiquí, prov. Here.,
<collectingCountry id="F37152CE6B6FFFCAFE4FFDB85808FDA1" box="[497,635,593,614]" name="Costa Rica" pageId="9" pageNumber="472">COSTA RICA</collectingCountry>
.
<date id="FFD8349E6B6FFFCAFF29FD825B46FD47" box="[151,309,619,640]" pageId="9" pageNumber="472" value="1993-03-09" valueMax="1993-03-30" valueMin="1993-03-09">
<collectingDate id="EF9CCD766B6FFFCAFF29FD825B46FD47" box="[151,309,619,640]" pageId="9" pageNumber="472" value="1993-03-09" valueMax="1993-03-30" valueMin="1993-03-09">9 a 30 mar 1993</collectingDate>
</date>
.
<collectorName id="269377886B6FFFCAFE81FD825BE5FD47" box="[319,406,619,640]" pageId="9" pageNumber="472">M. Ortiz.</collectorName>
L_N_263950_521050&quot; / INBIOCRI001299837 and
<specimenCode id="DBC0BA256B6FFFCAFE17FD6F5821FD5C" box="[425,594,646,667]" collectionCode="CEUA" pageId="9" pageNumber="472">CEUA00106193</specimenCode>
;
<specimenCount id="9D60D9D76B6FFFCAFDE1FD6F580DFD5C" box="[607,638,645,667]" pageId="9" pageNumber="472" type="male">2♂</specimenCount>
&quot;
<location id="8EB944856B6FFFCAFF1EFD495A86FD72" LSID="urn:lsid:plazi:treatment:03CFA3486B6FFFC9FF13FE6F5B5EFE9B:8EB944856B6FFFCAFF1EFD495A86FD72" box="[160,245,672,693]" country="Costa Rica" municipality="Prov." name="Amubri" pageId="9" pageNumber="472" stateProvince="Limon">Amubri</location>
,
<collectingMunicipality id="6BBD88246B6FFFCAFEBDFD495B4AFD72" box="[259,313,672,693]" pageId="9" pageNumber="472">Prov.</collectingMunicipality>
<collectingRegion id="49A2DCBC6B6FFFCAFEFDFD495BF9FD72" box="[323,394,672,693]" country="Costa Rica" name="Limon" pageId="9" pageNumber="472">Limon</collectingRegion>
,
<collectingCountry id="F37152CE6B6FFFCAFE26FD495840FD72" box="[408,563,672,693]" name="Costa Rica" pageId="9" pageNumber="472">COSTA RICA</collectingCountry>
.
<quantity id="4C9EBFBB6B6FFFCAFDFFFD495809FD72" box="[577,634,672,693]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="472" unit="m" value="70.0">
<specimenCount id="9D60D9D76B6FFFCAFDFFFD495809FD72" box="[577,634,672,693]" pageId="9" pageNumber="472" type="generic">
<elevation id="004BF56D6B6FFFCAFDFFFD495809FD72" box="[577,634,672,693]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="472" unit="m" value="70.0">70 m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
<materialsCitation id="3B0E18036B6FFFCAFDC4FD495B72FCC2" collectingDate="1993-11-08" collectingDateMax="1993-11-27" collectingDateMin="1993-11-08" collectionCode="CEUA" collectorName="Gallardo" country="Costa Rica" pageId="9" pageNumber="472" specimenCode="CEUA00106192" specimenCount="1" stateProvince="Limon" typeStatus="paratype">
.
<date id="FFD8349E6B6FFFCAFF29FD525B32FD17" box="[151,321,698,720]" pageId="9" pageNumber="472" value="1993-11-08" valueMax="1993-11-27" valueMin="1993-11-08">
<collectingDate id="EF9CCD766B6FFFCAFF29FD525B32FD17" box="[151,321,698,720]" pageId="9" pageNumber="472" value="1993-11-08" valueMax="1993-11-27" valueMin="1993-11-08">08–27 nov 1993</collectingDate>
</date>
, G. M.
<collectorName id="269377886B6FFFCAFE2FFD525B9EFD17" box="[401,493,699,720]" pageId="9" pageNumber="472">Gallardo</collectorName>
, L S 385500_ 578100 # 2461&quot; /
<specimenCode id="DBC0BA256B6FFFCAFEFBFD3C5B96FD2D" box="[325,485,725,746]" collectionCode="CEUA" pageId="9" pageNumber="472">CEUA00106192</specimenCode>
and INBIOCRI 001957606
</materialsCitation>
;
<materialsCitation id="3B0E18036B6FFFCAFEB2FD195BCEFCFD" collectingDate="1987-03-24" country="Costa Rica" location="Reserva" municipality="Cerere Rio Cerere" pageId="9" pageNumber="472" specimenCount="1" specimenCount-female="1" stateProvince="Limon" typeStatus="paratype">
<specimenCount id="9D60D9D76B6FFFCAFEB2FD195B59FCC1" box="[268,298,752,774]" pageId="9" pageNumber="472" type="female">1♀</specimenCount>
&quot;
<collectingRegion id="49A2DCBC6B6FFFCAFE87FD195B09FCC2" box="[313,378,752,773]" country="Costa Rica" name="Limon" pageId="9" pageNumber="472">Limon</collectingRegion>
,
<collectingCountry id="F37152CE6B6FFFCAFE3BFD195B99FCC2" box="[389,490,752,773]" name="Costa Rica" pageId="9" pageNumber="472">Costa Rica</collectingCountry>
<location id="8EB944856B6FFFCAFE4CFD195848FCC2" LSID="urn:lsid:plazi:treatment:03CFA3486B6FFFC9FF13FE6F5B5EFE9B:8EB944856B6FFFCAFE4CFD195848FCC2" box="[498,571,752,773]" country="Costa Rica" municipality="Cerere Rio Cerere" name="Reserva" pageId="9" pageNumber="472" stateProvince="Limon">Reserva</location>
Hitoy-
<collectingMunicipality id="6BBD88246B6FFFCAFF29FCE35B33FCD8" box="[151,320,778,799]" pageId="9" pageNumber="472">Cerere Rio Cerere</collectingMunicipality>
100 msnm
<date id="FFD8349E6B6FFFCAFE0FFCE3584DFCD8" box="[433,574,778,799]" pageId="9" pageNumber="472" value="1987-03-24">
<collectingDate id="EF9CCD766B6FFFCAFE0FFCE3584DFCD8" box="[433,574,778,799]" pageId="9" pageNumber="472" value="1987-03-24">24 marzo 1987</collectingDate>
</date>
Angel Solis / INBIOCRI002517679&quot;
</materialsCitation>
;
<materialsCitation id="3B0E18036B6FFFCAFE75FCCC5868FC4D" collectingDate="1993-02-18" collectingDateMax="1993-02-30" collectingDateMin="1993-02-18" collectorName="R. N. F. S. Barra" country="Costa Rica" elevation="10" location="Rio Sardinas" municipality="Prov." pageId="9" pageNumber="472" specimenCount="1" specimenCount-female="1" stateProvince="Limon" typeStatus="paratype">
<specimenCount id="9D60D9D76B6FFFCAFE75FCCC5B9AFCFC" box="[459,489,805,827]" pageId="9" pageNumber="472" type="female">1♀</specimenCount>
&quot;
<location id="8EB944856B6FFFCAFE45FCCC5809FCFD" LSID="urn:lsid:plazi:treatment:03CFA3486B6FFFC9FF13FE6F5B5EFE9B:8EB944856B6FFFCAFE45FCCC5809FCFD" box="[507,634,805,826]" country="Costa Rica" municipality="Prov." name="Rio Sardinas" pageId="9" pageNumber="472" stateProvince="Limon">Rio Sardinas</location>
,
<quantity id="4C9EBFBB6B6FFFCAFF29FCA95ABDFC92" box="[151,206,832,853]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="472" unit="m" value="10.0">
<elevation id="004BF56D6B6FFFCAFF29FCA95ABDFC92" box="[151,206,832,853]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="472" unit="m" value="10.0">10 m</elevation>
</quantity>
,
<collectorName id="269377886B6FFFCAFF65FCA95B1BFC92" box="[219,360,832,853]" pageId="9" pageNumber="472">R.N.F.S. Barra</collectorName>
del Colorado,
<collectingMunicipality id="6BBD88246B6FFFCAFE41FCA95842FC92" box="[511,561,832,853]" pageId="9" pageNumber="472">Prov.</collectingMunicipality>
<collectingRegion id="49A2DCBC6B6FFFCAFD84FCA95808FC92" box="[570,635,832,853]" country="Costa Rica" name="Limon" pageId="9" pageNumber="472">Limon</collectingRegion>
,
<collectingCountry id="F37152CE6B6FFFCAFF29FCB35B54FCA8" box="[151,295,858,879]" name="Costa Rica" pageId="9" pageNumber="472">COSTA RICA</collectingCountry>
.
<date id="FFD8349E6B6FFFCAFE8DFCB35B92FCA8" box="[307,481,858,879]" pageId="9" pageNumber="472" value="1993-02-18" valueMax="1993-02-30" valueMin="1993-02-18">
<collectingDate id="EF9CCD766B6FFFCAFE8DFCB35B92FCA8" box="[307,481,858,879]" pageId="9" pageNumber="472" value="1993-02-18" valueMax="1993-02-30" valueMin="1993-02-18">18 a 30 feb 1993</collectingDate>
</date>
. F. Araya, L-N- 291500, 564700 / INBIOCRI001297396&quot;
</materialsCitation>
;
<materialsCitation id="3B0E18036B6FFFCAFD98FC9C5809FC1E" collectingDate="1994-05-09" collectingDateMax="1994-05-14" collectingDateMin="1994-05-09" collectorName="G. Carballo" country="Costa Rica" county="Prov." elevation="150" location="Hitoy Cerere" municipality="Valle La Estrella" pageId="9" pageNumber="472" specimenCount="201" specimenCount-male="1" stateProvince="Limon" typeStatus="paratype">
<specimenCount id="9D60D9D76B6FFFCAFD98FC9C5830FC4D" box="[550,579,884,906]" pageId="9" pageNumber="472" type="male">1♂</specimenCount>
&quot;R.B.
<location id="8EB944856B6FFFCAFF29FC665B7CFC63" LSID="urn:lsid:plazi:treatment:03CFA3486B6FFFC9FF13FE6F5B5EFE9B:8EB944856B6FFFCAFF29FC665B7CFC63" box="[151,271,911,932]" country="Costa Rica" county="Prov." municipality="Valle La Estrella" name="Hitoy Cerere" pageId="9" pageNumber="472" stateProvince="Limon">Hitoy Cerere</location>
,
<collectingMunicipality id="6BBD88246B6FFFCAFEA9FC665BDCFC63" box="[279,431,911,932]" pageId="9" pageNumber="472">Valle La Estrella</collectingMunicipality>
,
<collectingCounty id="62B86AD26B6FFFCAFE09FC665B9BFC63" box="[439,488,911,932]" pageId="9" pageNumber="472">Prov.</collectingCounty>
<collectingRegion id="49A2DCBC6B6FFFCAFE55FC665858FC63" box="[491,555,911,932]" country="Costa Rica" name="Limon" pageId="9" pageNumber="472">Limon</collectingRegion>
,
<collectingCountry id="F37152CE6B6FFFCAFD8DFC665AA2FC78" name="Costa Rica" pageId="9" pageNumber="472">COSTA RICA</collectingCountry>
.
<quantity id="4C9EBFBB6B6FFFCAFF64FC435B4CFC78" box="[218,319,937,959]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="9" pageNumber="472" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">
<elevation id="004BF56D6B6FFFCAFF64FC435B4CFC78" box="[218,319,937,959]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="9" pageNumber="472" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">
100–
<specimenCount id="9D60D9D76B6FFFCAFEB6FC435B4CFC78" box="[264,319,938,959]" pageId="9" pageNumber="472" type="generic">200m</specimenCount>
</elevation>
</quantity>
.
<date id="FFD8349E6B6FFFCAFEF9FC435BA8FC78" box="[327,475,937,959]" pageId="9" pageNumber="472" value="1994-05-09" valueMax="1994-05-14" valueMin="1994-05-09">
<collectingDate id="EF9CCD766B6FFFCAFEF9FC435BA8FC78" box="[327,475,937,959]" pageId="9" pageNumber="472" value="1994-05-09" valueMax="1994-05-14" valueMin="1994-05-09">9–14 May 1994</collectingDate>
</date>
,
<collectorName id="269377886B6FFFCAFE5AFC435823FC78" box="[484,592,938,959]" pageId="9" pageNumber="472">G. Carballo</collectorName>
, L N 643400_184600 # 2856 / INBIOCRI001798864&quot;
</materialsCitation>
;
<materialsCitation id="3B0E18036B6FFFCAFF29FC365BDCFBEE" collectingDate="1993-10-12" collectingDateMax="1993-10-31" collectingDateMin="1993-10-12" collectorName="G. Gallardo" country="Costa Rica" elevation="70" location="Amubri" municipality="Prov." pageId="9" pageNumber="472" specimenCount="71" specimenCount-male="1" stateProvince="Limon" typeStatus="paratype">
<specimenCount id="9D60D9D76B6FFFCAFF29FC365AC6FC32" box="[151,181,991,1013]" pageId="9" pageNumber="472" type="male">1♂</specimenCount>
&quot;
<location id="8EB944856B6FFFCAFF78FC365B67FC33" LSID="urn:lsid:plazi:treatment:03CFA3486B6FFFC9FF13FE6F5B5EFE9B:8EB944856B6FFFCAFF78FC365B67FC33" box="[198,276,991,1012]" country="Costa Rica" municipality="Prov." name="Amubri" pageId="9" pageNumber="472" stateProvince="Limon">Amubri</location>
,
<collectingMunicipality id="6BBD88246B6FFFCAFE9EFC365B20FC33" box="[288,339,991,1012]" pageId="9" pageNumber="472">Prov.</collectingMunicipality>
<collectingRegion id="49A2DCBC6B6FFFCAFEE4FC365BEFFC33" box="[346,412,991,1012]" country="Costa Rica" name="Limon" pageId="9" pageNumber="472">Limon</collectingRegion>
,
<collectingCountry id="F37152CE6B6FFFCAFE16FC36584AFC33" box="[424,569,991,1012]" name="Costa Rica" pageId="9" pageNumber="472">COSTA RICA</collectingCountry>
.
<specimenCount id="9D60D9D76B6FFFCAFDFBFC36580DFC33" box="[581,638,991,1012]" pageId="9" pageNumber="472" type="generic">
<quantity id="4C9EBFBB6B6FFFCAFDFBFC365809FC33" box="[581,634,991,1012]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="472" unit="m" value="70.0">
<elevation id="004BF56D6B6FFFCAFDFBFC365809FC33" box="[581,634,991,1012]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="472" unit="m" value="70.0">70 m</elevation>
</quantity>
.
</specimenCount>
<date id="FFD8349E6B6FFFCAFF29FC135B5CFBC8" box="[151,303,1017,1039]" pageId="9" pageNumber="472" value="1993-10-12" valueMax="1993-10-31" valueMin="1993-10-12">
<collectingDate id="EF9CCD766B6FFFCAFF29FC135B5CFBC8" box="[151,303,1017,1039]" pageId="9" pageNumber="472" value="1993-10-12" valueMax="1993-10-31" valueMin="1993-10-12">12–31 Oct 1993</collectingDate>
</date>
.
<collectorName id="269377886B6FFFCAFE86FC135BD5FBC8" box="[312,422,1018,1039]" pageId="9" pageNumber="472">G. Gallardo</collectorName>
, L S 385500_578000 # 2407 / INBIOCRI001644531&quot;
</materialsCitation>
;
<materialsCitation id="3B0E18036B6FFFCAFE09FBFD5BCCFB53" collectingDate="1989-04" collectionCode="MNCR" collectorName="R. Aguilar &amp; J. Solano" country="Costa Rica" county="Prov." location="Cuatro Esquinas" municipality="Tortuguero" pageId="9" pageNumber="472" specimenCount="1" specimenCount-male="1" stateProvince="Limon" typeStatus="paratype">
<specimenCount id="9D60D9D76B6FFFCAFE09FBFD5BA6FBED" box="[439,469,1044,1066]" pageId="9" pageNumber="472" type="male">1♂</specimenCount>
&quot;
<location id="8EB944856B6FFFCAFE5FFBFD5808FBEE" LSID="urn:lsid:plazi:treatment:03CFA3486B6FFFC9FF13FE6F5B5EFE9B:8EB944856B6FFFCAFE5FFBFD5808FBEE" box="[481,635,1044,1065]" country="Costa Rica" county="Prov." municipality="Tortuguero" name="Cuatro Esquinas" pageId="9" pageNumber="472" stateProvince="Limon">Cuatro Esquinas</location>
, P. N.
<collectingMunicipality id="6BBD88246B6FFFCAFF74FBC65B47FB83" box="[202,308,1071,1092]" pageId="9" pageNumber="472">Tortuguero</collectingMunicipality>
,
<collectingCounty id="62B86AD26B6FFFCAFE80FBC65B03FB83" box="[318,368,1071,1092]" pageId="9" pageNumber="472">Prov.</collectingCounty>
<collectingRegion id="49A2DCBC6B6FFFCAFECAFBC65BC6FB83" box="[372,437,1071,1092]" country="Costa Rica" name="Limon" pageId="9" pageNumber="472">Limon</collectingRegion>
,
<collectingCountry id="F37152CE6B6FFFCAFE00FBC6583FFB83" box="[446,588,1071,1092]" name="Costa Rica" pageId="9" pageNumber="472">COSTA RICA</collectingCountry>
. 0 m,
<date id="FFD8349E6B6FFFCAFF29FBA05B71FB99" box="[151,258,1097,1118]" pageId="9" pageNumber="472" value="1989-04">
<collectingDate id="EF9CCD766B6FFFCAFF29FBA05B71FB99" box="[151,258,1097,1118]" pageId="9" pageNumber="472" value="1989-04">April 1989</collectingDate>
</date>
,
<collectorName id="269377886B6FFFCAFEB2FBA05B01FB99" box="[268,370,1097,1118]" pageId="9" pageNumber="472">R. Aguilar</collectorName>
&amp;
<collectorName id="269377886B6FFFCAFE2FFBA05B98FB99" box="[401,491,1097,1118]" pageId="9" pageNumber="472">J. Solano</collectorName>
, L- N 280000_ 590500 / INBIOCRI000084822&quot;. Seven
<typeStatus id="54DDACFC6B6FFFCAFD9DFB8D580DFBBE" box="[547,638,1124,1145]" pageId="9" pageNumber="472" type="paratype">paratypes</typeStatus>
are deposited at
<collectionCode id="ED778A9B6B6FFFCAFEFAFB965BFDFB53" box="[324,398,1151,1172]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="9" pageNumber="472">MNCR</collectionCode>
, and
</materialsCitation>
<materialsCitation id="3B0E18036B6FFFCAFE77FB96587DFB69" country="Costa Rica" location="Universidad de Alicante" pageId="9" pageNumber="472" specimenCount="1" stateProvince="Limon" typeStatus="paratype">
<specimenCount id="9D60D9D76B6FFFCAFE77FB965826FB53" box="[457,597,1151,1172]" pageId="9" pageNumber="472" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
are deposited at
<location id="8EB944856B6FFFCAFEA7FB70587DFB69" LSID="urn:lsid:plazi:treatment:03CFA3486B6FFFC9FF13FE6F5B5EFE9B:8EB944856B6FFFCAFEA7FB70587DFB69" box="[281,526,1177,1198]" country="Costa Rica" name="Universidad de Alicante" pageId="9" pageNumber="472" stateProvince="Limon">Universidad de Alicante</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BD9125E6B6FFFCAFF11FB5D5EFDFC78" blockId="9.[151,639,486,1729]" lastBlockId="9.[685,1173,194,1729]" pageId="9" pageNumber="472">
<emphasis id="B912CE4C6B6FFFCAFF11FB5D5B60FB0E" bold="true" box="[175,275,1204,1225]" pageId="9" pageNumber="472">
<typeStatus id="54DDACFC6B6FFFCAFF11FB5D5B7DFB0E" box="[175,270,1204,1225]" pageId="9" pageNumber="472" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male (
<figureCitation id="135D0EDB6B6FFFCAFEDAFB5D5BD6FB0E" box="[356,421,1204,1225]" captionStart="Figs" captionStartId="2.[197,237,1274,1293]" captionTargetBox="[193,1167,186,1250]" captionTargetId="figure-199@2.[178,1188,183,1261]" captionTargetPageId="2" captionText="Figs. 1–8. Anomala species holotypes, habitus. 1) A. coffea; 2) A. cyclops; 3) A. divisa; 4) A. mersa; 5) A. mesosticta;" figureDoi="http://doi.org/10.5281/zenodo.10106052" httpUri="https://zenodo.org/record/10106052/files/figure.png" pageId="9" pageNumber="472">Fig. 5</figureCitation>
). Body oval. Length
<quantity id="4C9EBFBB6B6FFFCAFF29FB275A8EFB24" box="[151,253,1230,1251]" metricMagnitude="-2" metricUnit="m" metricValue="1.088" pageId="9" pageNumber="472" unit="mm" value="10.88">10.88 mm</quantity>
, width
<quantity id="4C9EBFBB6B6FFFCAFEF7FB275BD7FB24" box="[329,420,1230,1251]" metricMagnitude="-3" metricUnit="m" metricValue="5.52" pageId="9" pageNumber="472" unit="mm" value="5.52">5.52 mm</quantity>
. Head reddish brown. Pronotum dark brown on disc, with sides and basal median part ochre. Scutellum dark brown with reddish sides. Elytra ochre with dark brown margins, expanding on median and apical parts of suture; punctures of striae pigmented. Pygidium, legs, and venter ochre.
<emphasis id="B912CE4C6B6FFFCAFE9EFA875B2DFA44" bold="true" box="[288,350,1390,1411]" pageId="9" pageNumber="472">Head:</emphasis>
Clypeus trapezoidal, surface densely punctate-reticulate, convex. Anterior margin straight, clypeus thin in frontal view, width/length ratio 2.24. Frons punctate, less densely than clypeus, with a triangular concavity. Ocular canthus long, thin, with rounded apex. Ratio of interocular width/width of eye 2.99. Ratio antennal funiculus/ club 0.73.
<emphasis id="B912CE4C6B6FFFCAFF41F9CE5B1AF9FB" bold="true" box="[255,361,1575,1596]" pageId="9" pageNumber="472">Pronotum:</emphasis>
Subtrapezoidal. Ratio width/ length 1.7. Lateral margins angulated at 1/3 of pronotal length from the anterior margins. Anterior angles quadrate, posterior angles obtuse. Basal margin sinuate, subapical bead complete. Surface with deep dense punctures.
<emphasis id="B912CE4C6B6FFFCAFDB0F945580EF906" bold="true" box="[526,637,1708,1729]" pageId="9" pageNumber="472">Scutellum:</emphasis>
Subtriangular, with rounded sides and acute apex. Ratio width/length 1.38. Surface densely punctate.
<emphasis id="B912CE4C6B6FFFCAFD13FF1E5888FECB" bold="true" box="[685,763,247,268]" pageId="9" pageNumber="472">Elytra:</emphasis>
With costae defined by regular rows of punctures. Subsutural interstice with 2–4 rows of sparse punctures, 2
<superScript id="7C13BF166B6FFFCAFC26FECE59D8FEF2" attach="right" box="[920,939,295,309]" fontSize="6" pageId="9" pageNumber="472">nd</superScript>
–4
<superScript id="7C13BF166B6FFFCAFC7AFECE59A1FEF2" attach="left" box="[964,978,295,309]" fontSize="6" pageId="9" pageNumber="472">th</superScript>
interstices with 1 irregular, secondary stria. Surface covered with secondary punctures.
<emphasis id="B912CE4C6B6FFFCAFC3AFE8B599EFEB0" bold="true" box="[900,1005,354,375]" pageId="9" pageNumber="472">Pygidium:</emphasis>
With coalescing punctures and long setae on posterior margin. Ratio width/length 1.66.
<emphasis id="B912CE4C6B6FFFCAFC15FE7E5986FE6B" bold="true" box="[939,1013,407,428]" pageId="9" pageNumber="472">Venter:</emphasis>
Space between mesocoxae narrow, slightly convex. Mesometasternal suture well-defined at base of mesocoxae. Abdominal sternites with 1 row of setae and 3–5 rows of punctures per sternite. Sixth abdominal sternite slightly raised, as a bump. Last sternite punctate, with subapical bead thick and sinuate, apical membrane wide.
<emphasis id="B912CE4C6B6FFFCAFC90FDB8591BFDA1" bold="true" box="[814,872,593,614]" pageId="9" pageNumber="472">Legs:</emphasis>
Protibia (
<figureCitation id="135D0EDB6B6FFFCAFC6FFDB85E6FFDA1" box="[977,1052,593,614]" captionStart="Figs" captionStartId="3.[175,216,1111,1130]" captionTargetBox="[154,1163,186,1089]" captionTargetId="figure-261@3.[151,1173,183,1093]" captionTargetPageId="3" captionText="Figs. 9–16. Anomala species, protibiae. 9) A. coffea (male: holotype; female: paratype, Estación Pitilla, Costa Rica); 10) A. cyclops (male: holotype; female: paratype, Los Almendros, Costa Rica); 11) A. divisa (male: holotype; female: paratype, Estación Biológica Las Alturas, Costa Rica); 12) A. mersa (male: paratype, Sector Palo Verde, Costa Rica); 13) A. mesosticta (male: holotype; female: paratype, Rio Sardinas, Costa Rica); 14) A. robiginosa (male: holotype; female: paratype, Santa Rosa National Park, Costa Rica); 15) A. strigodermoides (male: holotype; female: paratype, Catarata río Buenavista, Alajuela); 16) A. unilineata (holotype). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106054" httpUri="https://zenodo.org/record/10106054/files/figure.png" pageId="9" pageNumber="472">Fig. 13</figureCitation>
) tridentate, apical tooth long and curved, middle tooth below internal apex of protibia triangular, acute, basal tooth scarcely developed. Metatibia fusiform. Ratio length/ width 3.23. First external carina well-developed. Surface punctate. Internal protarsal claw bifurcate, with upper branch slightly shorter, as wide as the width of the lower one, inferior margin curved.
<emphasis id="B912CE4C6B6FFFCAFBE0FCE35979FCFD" bold="true" pageId="9" pageNumber="472">Male genitalia:</emphasis>
Parameres short with wide, blunt apex in lateral view (
<figureCitation id="135D0EDB6B6FFFCAFCEEFCA959EFFC92" box="[848,924,832,853]" captionStart="Figs" captionStartId="4.[197,237,1207,1226]" captionTargetBox="[176,1181,187,1184]" captionTargetId="figure-197@4.[173,1195,183,1197]" captionTargetPageId="4" captionText="Figs. 17–24. Anomala species, aedeagi, lateral view (top), dorsal view (bottom left), ventral view (bottom right). 17) A. coffea (paratype, Estación Pitilla, Costa Rica); 18) A. cyclops (holotype); 19) A. divisa (holotype); 20) A. mersa (holotype); 21) A. mesosticta (holotype); 22) A. robiginosa (paratype, Zarcero, Costa Rica); 23) A. strigodermoides (holotype); 24) A. unilineata (paratype, Santa Rosa National Park, Costa Rica). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106056" httpUri="https://zenodo.org/record/10106056/files/figure.png" pageId="9" pageNumber="472">Fig. 21</figureCitation>
), ventral margin sinuate. Endophallus (
<figureCitation id="135D0EDB6B6FFFCAFC91FCB35900FCA8" box="[815,883,858,879]" captionStart="Figs" captionStartId="5.[175,215,1638,1657]" captionTargetBox="[178,1140,195,1606]" captionTargetId="figure-1@5.[168,1156,183,1627]" captionTargetPageId="5" captionText="Figs. 25–33. Anomala species, genitalia. Endophalli: 25) A. coffea (paratype, Estación Pitilla, Costa Rica); 26) A. cyclops (paratype, Finca Jenny, Costa Rica); 27) A. divisa (holotype); 28) A. mersa (paratype, Sector Palo Verde, Costa Rica); 29) A. mesosticta (holotype); 30) A. robiginosa (holotype); 31) A. strigodermoides (holotype); 32) A. unilineata (holotype). Aedeagus: 33) A. sticticoptera (lateral view, Cuyotenango, Guatemala, MNHUB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106058" httpUri="https://zenodo.org/record/10106058/files/figure.png" pageId="9" pageNumber="472">Fig. 29</figureCitation>
) composed of a long sacculus, with a ridge of sclerotized spines at about half the length and a stripe of sclerotized setae at base. Ejaculatory opening lateral, at apex of an inflation.
</paragraph>
<paragraph id="8BD9125E6B6FFFCAFD7BFC2D595FFBC8" blockId="9.[685,1173,194,1729]" pageId="9" pageNumber="472">
<emphasis id="B912CE4C6B6FFFCAFD7BFC2D5965FC1E" bold="true" box="[709,790,964,985]" pageId="9" pageNumber="472">Female.</emphasis>
Similar to male except protibia (
<figureCitation id="135D0EDB6B6FFFCAFBF4FC2D5EFEFC1E" box="[1098,1165,964,985]" captionStart="Figs" captionStartId="3.[175,216,1111,1130]" captionTargetBox="[154,1163,186,1089]" captionTargetId="figure-261@3.[151,1173,183,1093]" captionTargetPageId="3" captionText="Figs. 9–16. Anomala species, protibiae. 9) A. coffea (male: holotype; female: paratype, Estación Pitilla, Costa Rica); 10) A. cyclops (male: holotype; female: paratype, Los Almendros, Costa Rica); 11) A. divisa (male: holotype; female: paratype, Estación Biológica Las Alturas, Costa Rica); 12) A. mersa (male: paratype, Sector Palo Verde, Costa Rica); 13) A. mesosticta (male: holotype; female: paratype, Rio Sardinas, Costa Rica); 14) A. robiginosa (male: holotype; female: paratype, Santa Rosa National Park, Costa Rica); 15) A. strigodermoides (male: holotype; female: paratype, Catarata río Buenavista, Alajuela); 16) A. unilineata (holotype). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106054" httpUri="https://zenodo.org/record/10106054/files/figure.png" pageId="9" pageNumber="472">Fig. 13</figureCitation>
) with longer and wider teeth, last sternite with curved apex.
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6FFFCAFD7BFBFD5E57FAAF" pageId="9" pageNumber="472" type="description">
<paragraph id="8BD9125E6B6FFFCAFD7BFBFD5E57FAAF" blockId="9.[685,1173,194,1729]" pageId="9" pageNumber="472">
<emphasis id="B912CE4C6B6FFFCAFD7BFBFD5959FBEE" bold="true" box="[709,810,1044,1065]" pageId="9" pageNumber="472">Variation.</emphasis>
Nine
<typeStatus id="54DDACFC6B6FFFCAFCD1FBFD59BCFBEE" box="[879,975,1044,1065]" pageId="9" pageNumber="472" type="paratype">paratypes</typeStatus>
. Head and pronotal disc vary from reddish to dark brown; extension of dark maculae on pronotum variable to almost dark with thin, ochre sides; dark lateral bands and median expansion on elytra variable in width; venter varies from ochre to light brown. Body length
<quantity id="4C9EBFBB6B6FFFCAFD53FB5D59FFFB0E" box="[749,908,1203,1225]" metricMagnitude="-2" metricUnit="m" metricValue="1.101" metricValueMax="1.176" metricValueMin="1.026" pageId="9" pageNumber="472" unit="mm" value="11.01" valueMax="11.76" valueMin="10.26">10.26–11.76 mm</quantity>
, body width
<quantity id="4C9EBFBB6B6FFFCAFBB9FB5D5EE2FB0E" box="[1031,1169,1203,1225]" metricMagnitude="-3" metricUnit="m" metricValue="5.949999999999999" metricValueMax="6.38" metricValueMin="5.52" pageId="9" pageNumber="472" unit="mm" value="5.949999999999999" valueMax="6.38" valueMin="5.52">5.52–6.38 mm</quantity>
. Clypeal width/length ratio 2.00–2.24. Ratio of interocular width/width of eye 2.91–3.05. Ratio of antennal funiculus/club 0.73–0.87. Pronotal width/ length ratio 1.62–1.72. Scutellar width/length ratio 1.33–1.49. Pygidial width/length ratio 1.59–1.73. Metatibial length/width ratio 2.79–3.23.
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6FFFCAFD7BFA8759B8FA7F" pageId="9" pageNumber="472" type="etymology">
<paragraph id="8BD9125E6B6FFFCAFD7BFA8759B8FA7F" blockId="9.[685,1173,194,1729]" pageId="9" pageNumber="472">
<emphasis id="B912CE4C6B6FFFCAFD7BFA87594BFA44" bold="true" box="[709,824,1390,1411]" pageId="9" pageNumber="472">Etymology.</emphasis>
From the Greek prefix
<emphasis id="B912CE4C6B6FFFCAFBF0FA845EF1FA44" box="[1102,1154,1389,1411]" italics="true" pageId="9" pageNumber="472">méso</emphasis>
s, middle, and adjective
<emphasis id="B912CE4C6B6FFFCAFC30FA6159BEFA59" box="[910,973,1416,1438]" italics="true" pageId="9" pageNumber="472">stiktós</emphasis>
, maculated, for the central macula on the elytra.
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6FFFCAFD7BFA5459DDF9C0" pageId="9" pageNumber="472" type="distribution">
<paragraph id="8BD9125E6B6FFFCAFD7BFA5459DDF9C0" blockId="9.[685,1173,194,1729]" pageId="9" pageNumber="472">
<emphasis id="B912CE4C6B6FFFCAFD7BFA54593BFA15" bold="true" box="[709,840,1469,1490]" pageId="9" pageNumber="472">Distribution.</emphasis>
<taxonomicName id="4C6669DD6B6FFFCAFCEBFA545E5AFA14" box="[853,1065,1469,1491]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="472" phylum="Arthropoda" rank="species" species="mesosticta">
<emphasis id="B912CE4C6B6FFFCAFCEBFA545E5AFA14" box="[853,1065,1469,1491]" italics="true" pageId="9" pageNumber="472">Anomala mesosticta</emphasis>
</taxonomicName>
occurs in the Caribbean lowlands of
<collectingCountry id="F37152CE6B6FFFCAFC05FA315E5BFA2A" box="[955,1064,1496,1517]" name="Costa Rica" pageId="9" pageNumber="472">Costa Rica</collectingCountry>
, from
<quantity id="4C9EBFBB6B6FFFCAFBD2FA315898F9C0" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="9" pageNumber="472" unit="m" value="100.0" valueMax="200.0" valueMin="0.0">0 to 200 m</quantity>
elevation (
<figureCitation id="135D0EDB6B6FFFCAFCE7FA1B59D0F9C0" box="[857,931,1522,1543]" captionStart="Figs" captionStartId="6.[197,237,1016,1035]" captionTargetBox="[178,1183,206,993]" captionTargetId="figure-344@6.[173,1193,183,1019]" captionTargetPageId="6" captionText="Figs. 34–36. Anomala species, distribution maps. 34) Distribution of A. coffea, A. mersa, and A. robiginosa;" figureDoi="http://doi.org/10.5281/zenodo.10106060" httpUri="https://zenodo.org/record/10106060/files/figure.png" pageId="9" pageNumber="472">Fig. 35</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37C41D56B6FFFC9FD7BF9E45B5EFE9B" lastPageId="10" lastPageNumber="473" pageId="9" pageNumber="472" type="diagnosis">
<paragraph id="8BD9125E6B6FFFC9FD7BF9E45B5EFE9B" blockId="9.[685,1173,194,1729]" lastBlockId="10.[173,659,194,348]" lastPageId="10" lastPageNumber="473" pageId="9" pageNumber="472">
<emphasis id="B912CE4C6B6FFFCAFD7BF9E4595DF9E5" bold="true" box="[709,814,1549,1570]" pageId="9" pageNumber="472">Diagnosis.</emphasis>
This species is recognized by its ochre elytra with dark brown margins expanding medially, tridentate protibia, and parameres short with a wide blunt apex. This species has an elytral pattern similar to that of
<taxonomicName id="4C6669DD6B6FFFCAFC7CF99E59DBF960" authority="Bates, 1888" authorityName="Bates" authorityYear="1888" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="472" phylum="Arthropoda" rank="species" species="denticollis">
<emphasis id="B912CE4C6B6FFFCAFC7CF99E5EE7F94A" box="[962,1172,1655,1677]" italics="true" pageId="9" pageNumber="472">Anomala denticollis</emphasis>
Bates, 1888 (
<collectingCountry id="F37152CE6B6FFFCAFC8AF97B59D2F960" box="[820,929,1682,1703]" name="Guatemala" pageId="9" pageNumber="472">Guatemala</collectingCountry>
)
</taxonomicName>
, and
<taxonomicName id="4C6669DD6B6FFFCAFC5DF9785E2DF960" authorityName="Bates" authorityYear="1888" box="[995,1118,1681,1703]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="472" phylum="Arthropoda" rank="species" species="hoepfneri">
<emphasis id="B912CE4C6B6FFFCAFC5DF9785E2DF960" box="[995,1118,1681,1703]" italics="true" pageId="9" pageNumber="472">A. hoepfneri</emphasis>
</taxonomicName>
. The former species has sinuate lateral margins on the pronotum and black maculae on the calli, which are not present in
<taxonomicName id="4C6669DD6B6CFFC9FEE3FF355BAEFF35" box="[349,477,220,242]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="473" phylum="Arthropoda" rank="species" species="mesosticta">
<emphasis id="B912CE4C6B6CFFC9FEE3FF355BAEFF35" box="[349,477,220,242]" italics="true" pageId="10" pageNumber="473">A. mesosticta</emphasis>
</taxonomicName>
. The latter species is larger, has dark maculae on the basal calli and light-colored lateral elytral margins, and wide interstices on the elytra with several irregular rows of punctures.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink