603 lines
68 KiB
XML
603 lines
68 KiB
XML
<document id="E9CAE05A7843DA485ECCF35DF7210BF0" ID-DOI="10.1649/0010-065x-69.3.463" ID-ISSN="1938-4394" ID-Zenodo-Dep="10106050" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630175225603" checkinUser="diego" docAuthor="Filippini, V., Galante, E. & Micó, E." docDate="2015" docId="03CFA3486B64FFC4FD74F9D8586CFC92" docLanguage="en" docName="0010-065X-69.3.463.pdf.imf" docOrigin="The Coleopterists Bulletin 69 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-69.3.463" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Anomala cyclops Filippini, Galante and Mico, new species" docType="treatment" docVersion="3" lastPageNumber="469" masterDocId="FFF6DB306B66FFC3FFBEFFE95A73FFC7" masterDocTitle="Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica" masterLastPageNumber="476" masterPageNumber="463" pageNumber="465" updateTime="1699636354092" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E577003A927803D4733D2C08244052E9">Descriptions of New Species ofAnomalaSamouelle (Coleoptera: Scarabaeidae: Rutelinae) from Costa Rica</mods:title>
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<mods:namePart id="78BCAAE26C18FC0A5517A75AEE66F714">Filippini, V.</mods:namePart>
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<mods:namePart id="A28312183B1AD116991198D5A6F5607A">Galante, E.</mods:namePart>
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<mods:namePart id="B4607F75FDC4F0339D3CB47B61831973">Micó, E.</mods:namePart>
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<mods:title id="8B8646709BCFE5759D3641CE2DF5B478">The Coleopterists Bulletin</mods:title>
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<mods:date id="ED7C216510BDC7F908210FA6C7CFBA0C">2015</mods:date>
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<mods:number id="3F47DDB28DBD45B186D130CCA2F72AA8">2015-09-30</mods:number>
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<mods:number id="6662EEA74D5C23182835CAF548103940">69</mods:number>
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<subSubSection id="C37C41D56B64FFC1FD74F9D85983F9A5" pageId="2" pageNumber="465" type="nomenclature">
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<paragraph id="8BD9125E6B64FFC1FD74F9D85983F9A5" blockId="2.[714,1186,1585,1660]" pageId="2" pageNumber="465">
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<emphasis id="B912CE4C6B64FFC1FD74F9D85983F9A5" bold="true" pageId="2" pageNumber="465">
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<taxonomicName id="4C6669DD6B64FFC1FD74F9D85EEEF980" authority="Filippini, Galante and Mico" authorityName="Filippini, Galante and Mico" box="[714,1181,1585,1607]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="465" phylum="Arthropoda" rank="species" species="cyclops" status="sp. nov.">
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<emphasis id="B912CE4C6B64FFC1FD74F9D85901F980" bold="true" box="[714,882,1585,1607]" italics="true" pageId="2" pageNumber="465">Anomala cyclops</emphasis>
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Filippini, Galante and Micó
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</taxonomicName>
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,
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<taxonomicNameLabel id="A22173376B64FFC1FCC5F9A45983F9A5" box="[891,1008,1613,1634]" pageId="2" pageNumber="465" rank="species">new species</taxonomicNameLabel>
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<subSubSection id="C37C41D56B64FFC1FCFEF98E5E58F9BB" box="[832,1067,1639,1660]" pageId="2" pageNumber="465" type="description">
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<paragraph id="8BD9125E6B64FFC1FCFEF98E5E58F9BB" blockId="2.[714,1186,1585,1660]" box="[832,1067,1639,1660]" pageId="2" pageNumber="465">
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(
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<figureCitation id="135D0EDB6B64FFC1FCF9F98E59FDF9BB" box="[839,910,1639,1660]" captionStart="Figs" captionStartId="2.[197,237,1274,1293]" captionTargetBox="[193,1167,186,1250]" captionTargetId="figure-199@2.[178,1188,183,1261]" captionTargetPageId="2" captionText="Figs. 1–8. Anomala species holotypes, habitus. 1) A. coffea; 2) A. cyclops; 3) A. divisa; 4) A. mersa; 5) A. mesosticta;" figureDoi="http://doi.org/10.5281/zenodo.10106052" httpUri="https://zenodo.org/record/10106052/files/figure.png" pageId="2" pageNumber="465">Figs. 2</figureCitation>
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,
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<figureCitation id="135D0EDB6B64FFC1FC24F98E59C7F9BB" box="[922,948,1639,1660]" captionStart="Figs" captionStartId="3.[175,216,1111,1130]" captionTargetBox="[154,1163,186,1089]" captionTargetId="figure-261@3.[151,1173,183,1093]" captionTargetPageId="3" captionText="Figs. 9–16. Anomala species, protibiae. 9) A. coffea (male: holotype; female: paratype, Estación Pitilla, Costa Rica); 10) A. cyclops (male: holotype; female: paratype, Los Almendros, Costa Rica); 11) A. divisa (male: holotype; female: paratype, Estación Biológica Las Alturas, Costa Rica); 12) A. mersa (male: paratype, Sector Palo Verde, Costa Rica); 13) A. mesosticta (male: holotype; female: paratype, Rio Sardinas, Costa Rica); 14) A. robiginosa (male: holotype; female: paratype, Santa Rosa National Park, Costa Rica); 15) A. strigodermoides (male: holotype; female: paratype, Catarata río Buenavista, Alajuela); 16) A. unilineata (holotype). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106054" httpUri="https://zenodo.org/record/10106054/files/figure.png" pageId="2" pageNumber="465">10</figureCitation>
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,
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<figureCitation id="135D0EDB6B64FFC1FC7EF98E59A9F9BB" box="[960,986,1639,1660]" captionStart="Figs" captionStartId="4.[197,237,1207,1226]" captionTargetBox="[176,1181,187,1184]" captionTargetId="figure-197@4.[173,1195,183,1197]" captionTargetPageId="4" captionText="Figs. 17–24. Anomala species, aedeagi, lateral view (top), dorsal view (bottom left), ventral view (bottom right). 17) A. coffea (paratype, Estación Pitilla, Costa Rica); 18) A. cyclops (holotype); 19) A. divisa (holotype); 20) A. mersa (holotype); 21) A. mesosticta (holotype); 22) A. robiginosa (paratype, Zarcero, Costa Rica); 23) A. strigodermoides (holotype); 24) A. unilineata (paratype, Santa Rosa National Park, Costa Rica). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106056" httpUri="https://zenodo.org/record/10106056/files/figure.png" pageId="2" pageNumber="465">18</figureCitation>
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,
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<figureCitation id="135D0EDB6B64FFC1FC58F98E5E73F9BB" box="[998,1024,1639,1660]" captionStart="Figs" captionStartId="5.[175,215,1638,1657]" captionTargetBox="[178,1140,195,1606]" captionTargetId="figure-1@5.[168,1156,183,1627]" captionTargetPageId="5" captionText="Figs. 25–33. Anomala species, genitalia. Endophalli: 25) A. coffea (paratype, Estación Pitilla, Costa Rica); 26) A. cyclops (paratype, Finca Jenny, Costa Rica); 27) A. divisa (holotype); 28) A. mersa (paratype, Sector Palo Verde, Costa Rica); 29) A. mesosticta (holotype); 30) A. robiginosa (holotype); 31) A. strigodermoides (holotype); 32) A. unilineata (holotype). Aedeagus: 33) A. sticticoptera (lateral view, Cuyotenango, Guatemala, MNHUB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106058" httpUri="https://zenodo.org/record/10106058/files/figure.png" pageId="2" pageNumber="465">26</figureCitation>
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,
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<figureCitation id="135D0EDB6B64FFC1FBB2F98E5E56F9BB" box="[1036,1061,1639,1660]" captionStart="Figs" captionStartId="6.[197,237,1016,1035]" captionTargetBox="[178,1183,206,993]" captionTargetId="figure-344@6.[173,1193,183,1019]" captionTargetPageId="6" captionText="Figs. 34–36. Anomala species, distribution maps. 34) Distribution of A. coffea, A. mersa, and A. robiginosa;" figureDoi="http://doi.org/10.5281/zenodo.10106060" httpUri="https://zenodo.org/record/10106060/files/figure.png" pageId="2" pageNumber="465">35</figureCitation>
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)
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<subSubSection id="C37C41D56B64FFC5FD65F97B5800F9E5" lastPageId="6" lastPageNumber="469" pageId="2" pageNumber="465" type="materials_examined">
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<emphasis id="B912CE4C6B64FFC1FD65F97B5901F960" bold="true" box="[731,882,1682,1703]" pageId="2" pageNumber="465">Type Material.</emphasis>
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<typeStatus id="54DDACFC6B64FFC1FCC1F97B59A8F960" box="[895,987,1682,1703]" pageId="2" pageNumber="465" type="holotype">Holotype</typeStatus>
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:
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"Cerro El Hacha,
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,
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<locationDeviation id="9A6B0B7C6B64FFC1FCAFF945599CF906" box="[785,1007,1708,1729]" location="La Cruz" pageId="2" pageNumber="465">
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SE de La Cruz
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,
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<location id="8EB944856B64FFC1FC45F9455E5DF906" LSID="urn:lsid:plazi:treatment:03CFA3486B64FFC4FD74F9D8586CFC92:8EB944856B64FFC1FC45F9455E5DF906" box="[1019,1070,1708,1729]" name="Prov." pageId="2" pageNumber="465" stateProvince="Guanacaste">Prov.</location>
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<collectingRegion id="49A2DCBC6B64FFC1FB8BF9455ED6F906" box="[1077,1189,1708,1729]" country="Costa Rica" name="Guanacaste" pageId="2" pageNumber="465">Guanacaste</collectingRegion>
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<caption id="DF1942D66B65FFC0FF11FBBE591AFB25" ID-DOI="http://doi.org/10.5281/zenodo.10106054" ID-Zenodo-Dep="10106054" httpUri="https://zenodo.org/record/10106054/files/figure.png" pageId="3" pageNumber="466" startId="3.[175,216,1111,1130]" targetBox="[154,1163,186,1089]" targetPageId="3">
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<paragraph id="8BD9125E6B65FFC0FF11FBBE591AFB25" blockId="3.[151,1173,1111,1250]" pageId="3" pageNumber="466">
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<emphasis id="B912CE4C6B65FFC0FF11FBBE5B64FBAD" bold="true" box="[175,279,1111,1130]" pageId="3" pageNumber="466">Figs. 9–16.</emphasis>
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<taxonomicName id="4C6669DD6B65FFC0FE93FBBE5BB0FBAD" box="[301,451,1111,1130]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B912CE4C6B65FFC0FE93FBBE5B09FBAD" box="[301,378,1111,1130]" italics="true" pageId="3" pageNumber="466">Anomala</emphasis>
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species
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, protibiae.
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<emphasis id="B912CE4C6B65FFC0FD96FBBE5848FBAD" bold="true" box="[552,571,1111,1130]" pageId="3" pageNumber="466">9)</emphasis>
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<taxonomicName id="4C6669DD6B65FFC0FDFDFBBE58E1FBAD" box="[579,658,1111,1130]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="coffea">
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<emphasis id="B912CE4C6B65FFC0FDFDFBBE58E1FBAD" box="[579,658,1111,1130]" italics="true" pageId="3" pageNumber="466">A. coffea</emphasis>
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</taxonomicName>
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||
(male: holotype; female: paratype, Estación Pitilla, Costa Rica);
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||
<emphasis id="B912CE4C6B65FFC0FF6CFB865A82FB45" bold="true" box="[210,241,1135,1154]" pageId="3" pageNumber="466">10)</emphasis>
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<taxonomicName id="4C6669DD6B65FFC0FF47FB865B22FB45" box="[249,337,1135,1154]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="cyclops">
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||
<emphasis id="B912CE4C6B65FFC0FF47FB865B22FB45" box="[249,337,1135,1154]" italics="true" pageId="3" pageNumber="466">A. cyclops</emphasis>
|
||
</taxonomicName>
|
||
(male: holotype; female: paratype, Los Almendros, Costa Rica);
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||
<emphasis id="B912CE4C6B65FFC0FC31FB8659D8FB45" bold="true" box="[911,939,1135,1154]" pageId="3" pageNumber="466">11)</emphasis>
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<taxonomicName id="4C6669DD6B65FFC0FC0DFB86598CFB45" box="[947,1023,1135,1154]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="divisa">
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||
<emphasis id="B912CE4C6B65FFC0FC0DFB86598CFB45" box="[947,1023,1135,1154]" italics="true" pageId="3" pageNumber="466">A. divisa</emphasis>
|
||
</taxonomicName>
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||
(male: holotype; female: paratype, Estación Biológica Las Alturas, Costa Rica);
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||
<emphasis id="B912CE4C6B65FFC0FD09FB6E58A6FB5D" bold="true" box="[695,725,1159,1178]" pageId="3" pageNumber="466">12)</emphasis>
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||
<taxonomicName id="4C6669DD6B65FFC0FD62FB6F595BFB5E" box="[732,808,1158,1177]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="mersa">
|
||
<emphasis id="B912CE4C6B65FFC0FD62FB6F595BFB5E" box="[732,808,1158,1177]" italics="true" pageId="3" pageNumber="466">A. mersa</emphasis>
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||
</taxonomicName>
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||
(male: paratype, Sector Palo Verde, Costa Rica);
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||
<emphasis id="B912CE4C6B65FFC0FF6EFB765A9DFB75" bold="true" box="[208,238,1183,1202]" pageId="3" pageNumber="466">13)</emphasis>
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<taxonomicName id="4C6669DD6B65FFC0FF4DFB765B10FB75" box="[243,355,1183,1202]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="mesosticta">
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||
<emphasis id="B912CE4C6B65FFC0FF4DFB765B10FB75" box="[243,355,1183,1202]" italics="true" pageId="3" pageNumber="466">A. mesosticta</emphasis>
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||
</taxonomicName>
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||
(male: holotype; female: paratype, Rio Sardinas, Costa Rica);
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<emphasis id="B912CE4C6B65FFC0FCCEFB7659FCFB75" bold="true" box="[880,911,1183,1202]" pageId="3" pageNumber="466">14)</emphasis>
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<taxonomicName id="4C6669DD6B65FFC0FC2DFB765E76FB75" box="[915,1029,1183,1202]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="robiginosa">
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||
<emphasis id="B912CE4C6B65FFC0FC2DFB765E76FB75" box="[915,1029,1183,1202]" italics="true" pageId="3" pageNumber="466">A. robiginosa</emphasis>
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</taxonomicName>
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(male: holotype; female: paratype, Santa Rosa National Park, Costa Rica);
|
||
<emphasis id="B912CE4C6B65FFC0FD28FB5E58C6FB0D" bold="true" box="[662,693,1207,1226]" pageId="3" pageNumber="466">15)</emphasis>
|
||
<taxonomicName id="4C6669DD6B65FFC0FD03FB5F5912FB0E" box="[701,865,1206,1225]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="strigodermoides">
|
||
<emphasis id="B912CE4C6B65FFC0FD03FB5F5912FB0E" box="[701,865,1206,1225]" italics="true" pageId="3" pageNumber="466">A. strigodermoides</emphasis>
|
||
</taxonomicName>
|
||
(male: holotype; female: paratype, Catarata río Buenavista, Alajuela);
|
||
<emphasis id="B912CE4C6B65FFC0FE79FB265B95FB25" bold="true" box="[455,486,1231,1250]" pageId="3" pageNumber="466">16)</emphasis>
|
||
<taxonomicName id="4C6669DD6B65FFC0FE53FB275828FB26" box="[493,603,1230,1249]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="466" phylum="Arthropoda" rank="species" species="unilineata">
|
||
<emphasis id="B912CE4C6B65FFC0FE53FB275828FB26" box="[493,603,1230,1249]" italics="true" pageId="3" pageNumber="466">A. unilineata</emphasis>
|
||
</taxonomicName>
|
||
(holotype). Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD9125E6B65FFC0FF29FAF75EE0F990" blockId="3.[151,639,1310,1730]" lastBlockId="3.[685,1173,1310,1729]" pageId="3" pageNumber="466">
|
||
<materialsCitation id="3B0E18036B65FFC0FF29FAF75BBBF9FB" collectingDate="1991-06" collectingDateMax="1992-06-25" collectingDateMin="1991-06" collectionCode="MNCR" collectorName="P. N. Sta. Rosa & D. H. Janzen & W. Hallwachs" country="Costa Rica" elevation="300" location="Guanacaste" pageId="3" pageNumber="466" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Guanacaste" typeStatus="paratype">
|
||
Costa Rica,
|
||
<date id="FFD8349E6B65FFC0FEB0FAF75BF6FAF4" box="[270,389,1310,1331]" pageId="3" pageNumber="466" value="1992-06-25">
|
||
<collectingDate id="EF9CCD766B65FFC0FEB0FAF75BF6FAF4" box="[270,389,1310,1331]" pageId="3" pageNumber="466" value="1992-06-25">25 jun 1992</collectingDate>
|
||
</date>
|
||
, III curso Parataxon. L-N 329200,368000 / INBIOCRI000755645" deposited at
|
||
<collectionCode id="ED778A9B6B65FFC0FF62FABA5B56FAAF" box="[220,293,1363,1384]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="3" pageNumber="466">MNCR</collectionCode>
|
||
.
|
||
<typeStatus id="54DDACFC6B65FFC0FE8FFABA5BFEFAAF" box="[305,397,1363,1384]" pageId="3" pageNumber="466" type="paratype">Paratypes</typeStatus>
|
||
(9):
|
||
<specimenCount id="9D60D9D76B65FFC0FE01FABA5BAEFAAF" box="[447,477,1362,1384]" pageId="3" pageNumber="466" type="male">1♂</specimenCount>
|
||
"Est. Sta. Rosa,
|
||
<quantity id="4C9EBFBB6B65FFC0FF29FA845AA9FA45" box="[151,218,1389,1410]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="466" unit="m" value="300.0">
|
||
<elevation id="004BF56D6B65FFC0FF29FA845AA9FA45" box="[151,218,1389,1410]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="466" unit="m" value="300.0">300 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectorName id="269377886B65FFC0FF58FA845BF2FA45" box="[230,385,1389,1410]" pageId="3" pageNumber="466">P. N. Sta. Rosa</collectorName>
|
||
, Prov.
|
||
<collectingRegion id="49A2DCBC6B65FFC0FE77FA845849FA45" box="[457,570,1389,1410]" country="Costa Rica" name="Guanacaste" pageId="3" pageNumber="466">Guanacaste</collectingRegion>
|
||
,
|
||
<collectingCountry id="F37152CE6B65FFC0FDF9FA845AB6FA5A" name="Costa Rica" pageId="3" pageNumber="466">Costa Rica</collectingCountry>
|
||
,
|
||
<date id="FFD8349E6B65FFC0FF6FFA615B00FA5A" box="[209,371,1416,1437]" pageId="3" pageNumber="466" value="1992-06-03" valueMax="1992-06-12" valueMin="1992-06-03">
|
||
<collectingDate id="EF9CCD766B65FFC0FF6FFA615B00FA5A" box="[209,371,1416,1437]" pageId="3" pageNumber="466" value="1992-06-03" valueMax="1992-06-12" valueMin="1992-06-03">3 a 12 jun 1992</collectingDate>
|
||
</date>
|
||
, III Curso Parataxon. L- N 313000,359800" / INBIOCRI000458888";
|
||
<specimenCount id="9D60D9D76B65FFC0FD93FA4B5838FA7F" box="[557,587,1442,1464]" pageId="3" pageNumber="466" type="female">1♀</specimenCount>
|
||
"Est. Sta. Rosa,
|
||
<quantity id="4C9EBFBB6B65FFC0FEBAFA545B35FA15" box="[260,326,1469,1490]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="466" unit="m" value="300.0">
|
||
<elevation id="004BF56D6B65FFC0FEBAFA545B35FA15" box="[260,326,1469,1490]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="466" unit="m" value="300.0">300 m</elevation>
|
||
</quantity>
|
||
, P. N.
|
||
<collectingRegion id="49A2DCBC6B65FFC0FE33FA545B8EFA15" box="[397,509,1469,1490]" country="Costa Rica" name="Guanacaste" pageId="3" pageNumber="466">Guanacaste</collectingRegion>
|
||
, Prov. Guan,
|
||
<collectingCountry id="F37152CE6B65FFC0FF29FA315B5DFA2A" box="[151,302,1496,1517]" name="Costa Rica" pageId="3" pageNumber="466">COSTA RICA</collectingCountry>
|
||
.
|
||
<collectorName id="269377886B65FFC0FE82FA315BB1FA2A" box="[316,450,1496,1517]" pageId="3" pageNumber="466">D. H. Janzen</collectorName>
|
||
&
|
||
<collectorName id="269377886B65FFC0FE54FA315809FA2A" box="[490,634,1496,1517]" pageId="3" pageNumber="466">W. Hallwachs</collectorName>
|
||
,
|
||
<date id="FFD8349E6B65FFC0FF29FA1B5A8FF9C0" box="[151,252,1522,1543]" pageId="3" pageNumber="466" value="1991-06">
|
||
<collectingDate id="EF9CCD766B65FFC0FF29FA1B5A8FF9C0" box="[151,252,1522,1543]" pageId="3" pageNumber="466" value="1991-06">Jun 1991</collectingDate>
|
||
</date>
|
||
, L- N 313000,359800 / INBIOCRI 000590279;
|
||
<specimenCount id="9D60D9D76B65FFC0FEAFF9E45B5CF9E5" box="[273,303,1548,1570]" pageId="3" pageNumber="466" type="male">1♂</specimenCount>
|
||
<specimenCount id="9D60D9D76B65FFC0FE86F9E45B25F9E5" box="[312,342,1548,1570]" pageId="3" pageNumber="466" type="female">1♀</specimenCount>
|
||
"Est. Sta. Rosa,
|
||
<quantity id="4C9EBFBB6B65FFC0FE41F9E45832F9E5" box="[511,577,1549,1570]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="466" unit="m" value="300.0">
|
||
<elevation id="004BF56D6B65FFC0FE41F9E45832F9E5" box="[511,577,1549,1570]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="466" unit="m" value="300.0">300 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectorName id="269377886B65FFC0FDF3F9E45B73F9FB" pageId="3" pageNumber="466">P. N. Sta. Rosa</collectorName>
|
||
, Prov.
|
||
<collectingRegion id="49A2DCBC6B65FFC0FEF1F9CE5BBBF9FB" box="[335,456,1575,1596]" country="Costa Rica" name="Guanacaste" pageId="3" pageNumber="466">Guanacaste</collectingRegion>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B0E18036B65FFC0FE69F9CE5EFCF990" collectingDate="1992-06-01" collectingDateMax="1994-05-03" collectingDateMin="1992-06-01" collectionCode="MNCR" collectorName="A. M. Mora & E. Lopez & E. Araya. & C. Cano" country="Costa Rica" elevation="100" location="Universidad de Alicante" pageId="3" pageNumber="466" specimenCode="CEUA00106197, CEUA00106196" specimenCount="106" specimenCount-female="2" specimenCount-male="3" stateProvince="Guanacaste" typeStatus="paratype">
|
||
Costa Rica,
|
||
<date id="FFD8349E6B65FFC0FDE2F9CE5B63F990" pageId="3" pageNumber="466" value="1992-06-01" valueMax="1992-06-12" valueMin="1992-06-01">
|
||
<collectingDate id="EF9CCD766B65FFC0FDE2F9CE5B63F990" pageId="3" pageNumber="466" value="1992-06-01" valueMax="1992-06-12" valueMin="1992-06-01">1 a 12 jun 1992</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="269377886B65FFC0FEA3F9AB5BE6F990" box="[285,405,1602,1623]" pageId="3" pageNumber="466">A. M. Mora</collectorName>
|
||
, L-N 313000,359800" / INBIOCRI000912716 and
|
||
<specimenCode id="DBC0BA256B65FFC0FE14F9B55821F9B6" box="[426,594,1628,1649]" collectionCode="CEUA" pageId="3" pageNumber="466">CEUA00106197</specimenCode>
|
||
;
|
||
<specimenCount id="9D60D9D76B65FFC0FDE1F9B5580DF9B5" box="[607,638,1628,1650]" pageId="3" pageNumber="466" type="female">1♀</specimenCount>
|
||
"Los Almendros, P. N.
|
||
<collectingRegion id="49A2DCBC6B65FFC0FEC1F99E5B9DF94B" box="[383,494,1655,1676]" country="Costa Rica" name="Guanacaste" pageId="3" pageNumber="466">Guanacaste</collectingRegion>
|
||
, Prov. Guana.,
|
||
<collectingCountry id="F37152CE6B65FFC0FF29F97B5B54F960" box="[151,295,1682,1703]" name="Costa Rica" pageId="3" pageNumber="466">COSTA RICA</collectingCountry>
|
||
.
|
||
<date id="FFD8349E6B65FFC0FE8CF97B5BA8F960" box="[306,475,1681,1703]" pageId="3" pageNumber="466" value="1993-05-12" valueMax="1993-05-31" valueMin="1993-05-12">
|
||
<collectingDate id="EF9CCD766B65FFC0FE8CF97B5BA8F960" box="[306,475,1681,1703]" pageId="3" pageNumber="466" value="1993-05-12" valueMax="1993-05-31" valueMin="1993-05-12">12–31 May 1993</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="269377886B65FFC0FE56F97B5831F960" box="[488,578,1682,1703]" pageId="3" pageNumber="466">E. Lopez</collectorName>
|
||
, L- N 334800, 369800 / INBIOCRI001168088";
|
||
<specimenCount id="9D60D9D76B65FFC0FDE1F945580DF905" box="[607,638,1708,1730]" pageId="3" pageNumber="466" type="male">1♂</specimenCount>
|
||
"Fca. Jenny,
|
||
<quantity id="4C9EBFBB6B65FFC0FC99FAF75910FAF4" box="[807,867,1310,1331]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="466" unit="km" value="30.0">30 km</quantity>
|
||
N de Liberia, P. N.
|
||
<collectingRegion id="49A2DCBC6B65FFC0FB9DFAF75EE3FAF4" box="[1059,1168,1310,1331]" country="Costa Rica" name="Guanacaste" pageId="3" pageNumber="466">Guanacaste</collectingRegion>
|
||
, Prov. Guan.,
|
||
<collectingCountry id="F37152CE6B65FFC0FC86FAD159BCFA8A" box="[824,975,1336,1357]" name="Costa Rica" pageId="3" pageNumber="466">COSTA RICA</collectingCountry>
|
||
.
|
||
<date id="FFD8349E6B65FFC0FC62FAD15EE3FA8A" box="[988,1168,1336,1358]" pageId="3" pageNumber="466" value="1993-05-25" valueMax="1993-05-29" valueMin="1993-05-25">
|
||
<collectingDate id="EF9CCD766B65FFC0FC62FAD15EE3FA8A" box="[988,1168,1336,1358]" pageId="3" pageNumber="466" value="1993-05-25" valueMax="1993-05-29" valueMin="1993-05-25">25–29 May 1993</collectingDate>
|
||
</date>
|
||
.
|
||
<collectorName id="269377886B65FFC0FD13FABA5972FAAF" box="[685,769,1363,1384]" pageId="3" pageNumber="466">E. Araya.</collectorName>
|
||
L-N 316200,364400 / INBIOCRI001183834";
|
||
<specimenCount id="9D60D9D76B65FFC0FD13FA8458BFFA44" box="[685,716,1389,1411]" pageId="3" pageNumber="466" type="male">2♂</specimenCount>
|
||
"Finca Jenny,
|
||
<quantity id="4C9EBFBB6B65FFC0FCCCFA8459C5FA45" box="[882,950,1389,1410]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="466" unit="km" value="30.0">30 km</quantity>
|
||
N de Liberia, P. N.
|
||
<collectingRegion id="49A2DCBC6B65FFC0FD13FA61596BFA5A" box="[685,792,1416,1437]" country="Costa Rica" name="Guanacaste" pageId="3" pageNumber="466">Guanacaste</collectingRegion>
|
||
, Prov. Guan.,
|
||
<collectingCountry id="F37152CE6B65FFC0FC26FA615E51FA5A" box="[920,1058,1416,1437]" name="Costa Rica" pageId="3" pageNumber="466">COSTA RICA</collectingCountry>
|
||
.
|
||
<date id="FFD8349E6B65FFC0FB94FA615893FA70" pageId="3" pageNumber="466" value="1993-05-20" valueMax="1993-05-24" valueMin="1993-05-20">
|
||
<collectingDate id="EF9CCD766B65FFC0FB94FA615893FA70" pageId="3" pageNumber="466" value="1993-05-20" valueMax="1993-05-24" valueMin="1993-05-20">20–24 May 1993</collectingDate>
|
||
</date>
|
||
.
|
||
<collectorName id="269377886B65FFC0FD57FA4B5936FA70" box="[745,837,1442,1463]" pageId="3" pageNumber="466">E. Araya.</collectorName>
|
||
L-N 316200,364400" / INBIOCRI 001184178 and
|
||
<specimenCode id="DBC0BA256B65FFC0FC84FA5459A0FA15" box="[826,979,1469,1490]" collectionCode="CEUA" pageId="3" pageNumber="466">CEUA00106196</specimenCode>
|
||
;
|
||
<specimenCount id="9D60D9D76B65FFC0FC64FA545984FA14" box="[986,1015,1469,1491]" pageId="3" pageNumber="466" type="female">1♀</specimenCount>
|
||
"Est. Murcielago,
|
||
<quantity id="4C9EBFBB6B65FFC0FD13FA315893FA2A" box="[685,736,1496,1517]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="466" unit="km" value="8.0">8 km</quantity>
|
||
SO. de Cuajiniquil, P. N.
|
||
<collectingRegion id="49A2DCBC6B65FFC0FC58FA315E25FA2A" box="[998,1110,1496,1517]" country="Costa Rica" name="Guanacaste" pageId="3" pageNumber="466">Guanacaste</collectingRegion>
|
||
, Prov. Guana,
|
||
<collectingCountry id="F37152CE6B65FFC0FD4AFA1B590EF9C0" box="[756,893,1522,1543]" name="Costa Rica" pageId="3" pageNumber="466">COSTA RICA</collectingCountry>
|
||
.
|
||
<specimenCount id="9D60D9D76B65FFC0FC3BFA1B59B5F9C0" box="[901,966,1522,1543]" pageId="3" pageNumber="466" type="generic">
|
||
<quantity id="4C9EBFBB6B65FFC0FC3BFA1B59B1F9C0" box="[901,962,1522,1543]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="466" unit="m" value="100.0">
|
||
<elevation id="004BF56D6B65FFC0FC3BFA1B59B1F9C0" box="[901,962,1522,1543]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="466" unit="m" value="100.0">100 m</elevation>
|
||
</quantity>
|
||
.
|
||
</specimenCount>
|
||
<date id="FFD8349E6B65FFC0FC74FA1B5E48F9C0" box="[970,1083,1522,1543]" pageId="3" pageNumber="466" value="1994-05-03">
|
||
<collectingDate id="EF9CCD766B65FFC0FC74FA1B5E48F9C0" box="[970,1083,1522,1543]" pageId="3" pageNumber="466" value="1994-05-03">3 May 1994</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="269377886B65FFC0FBFDFA1B5EE3F9C0" box="[1091,1168,1522,1543]" pageId="3" pageNumber="466">C. Cano</collectorName>
|
||
, L N 320300_347200 # 2907 / INBIOCRI001891273". Seven
|
||
<typeStatus id="54DDACFC6B65FFC0FD51F9CE593AF9FB" box="[751,841,1575,1596]" pageId="3" pageNumber="466" type="paratype">paratypes</typeStatus>
|
||
are deposited at
|
||
<collectionCode id="ED778A9B6B65FFC0FC4CF9CE5E48F9FB" box="[1010,1083,1575,1596]" country="Costa Rica" httpUri="http://grbio.org/cool/6yis-bi95" name="Museo Nacional de Costa Rica" pageId="3" pageNumber="466">MNCR</collectionCode>
|
||
, and
|
||
<specimenCount id="9D60D9D76B65FFC0FBCEF9CE5975F990" pageId="3" pageNumber="466" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
|
||
are deposited at
|
||
<location id="8EB944856B65FFC0FC16F9AB5EFCF990" LSID="urn:lsid:plazi:treatment:03CFA3486B64FFC4FD74F9D8586CFC92:8EB944856B65FFC0FC16F9AB5EFCF990" box="[936,1167,1602,1623]" country="Costa Rica" name="Universidad de Alicante" pageId="3" pageNumber="466" stateProvince="Guanacaste">Universidad de Alicante</location>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD9125E6B65FFC0FD7BF9B55EE7F906" blockId="3.[685,1173,1310,1729]" pageId="3" pageNumber="466">
|
||
<emphasis id="B912CE4C6B65FFC0FD7BF9B5595AF9B6" bold="true" box="[709,809,1628,1649]" pageId="3" pageNumber="466">
|
||
<typeStatus id="54DDACFC6B65FFC0FD7BF9B55957F9B6" box="[709,804,1628,1649]" pageId="3" pageNumber="466" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Male (
|
||
<figureCitation id="135D0EDB6B65FFC0FCC4F9B559CFF9B6" box="[890,956,1628,1649]" captionStart="Figs" captionStartId="2.[197,237,1274,1293]" captionTargetBox="[193,1167,186,1250]" captionTargetId="figure-199@2.[178,1188,183,1261]" captionTargetPageId="2" captionText="Figs. 1–8. Anomala species holotypes, habitus. 1) A. coffea; 2) A. cyclops; 3) A. divisa; 4) A. mersa; 5) A. mesosticta;" figureDoi="http://doi.org/10.5281/zenodo.10106052" httpUri="https://zenodo.org/record/10106052/files/figure.png" pageId="3" pageNumber="466">Fig. 2</figureCitation>
|
||
). Body oval. Length
|
||
<quantity id="4C9EBFBB6B65FFC0FD13F99E5979F94B" box="[685,778,1655,1676]" metricMagnitude="-3" metricUnit="m" metricValue="9.82" pageId="3" pageNumber="466" unit="mm" value="9.82">9.82 mm</quantity>
|
||
, width
|
||
<quantity id="4C9EBFBB6B65FFC0FCE6F99E59C6F94B" box="[856,949,1655,1676]" metricMagnitude="-3" metricUnit="m" metricValue="5.09" pageId="3" pageNumber="466" unit="mm" value="5.09">5.09 mm</quantity>
|
||
. Head reddish brown. Pronotum ochre with a pentagonal, dark brown macula on disc, with apex pointing posteriorly,
|
||
</paragraph>
|
||
<caption id="DF1942D66B62FFC7FF7BFB5E5E7EFAD6" ID-DOI="http://doi.org/10.5281/zenodo.10106056" ID-Zenodo-Dep="10106056" httpUri="https://zenodo.org/record/10106056/files/figure.png" pageId="4" pageNumber="467" startId="4.[197,237,1207,1226]" targetBox="[176,1181,187,1184]" targetPageId="4">
|
||
<paragraph id="8BD9125E6B62FFC7FF7BFB5E5E7EFAD6" blockId="4.[173,1194,1206,1297]" pageId="4" pageNumber="467">
|
||
<emphasis id="B912CE4C6B62FFC7FF7BFB5E5B46FB0D" bold="true" box="[197,309,1206,1226]" pageId="4" pageNumber="467">Figs. 17–24.</emphasis>
|
||
<taxonomicName id="4C6669DD6B62FFC7FEF5FB5F5BADFB0D" box="[331,478,1206,1226]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B912CE4C6B62FFC7FEF5FB5F5BE4FB0E" box="[331,407,1206,1225]" italics="true" pageId="4" pageNumber="467">Anomala</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, aedeagi, lateral view (top), dorsal view (bottom left), ventral view (bottom right).
|
||
<emphasis id="B912CE4C6B62FFC7FF13FB265AB8FB25" bold="true" box="[173,203,1231,1250]" pageId="4" pageNumber="467">17)</emphasis>
|
||
<taxonomicName id="4C6669DD6B62FFC7FF6FFB275B6FFB26" box="[209,284,1230,1249]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="coffea">
|
||
<emphasis id="B912CE4C6B62FFC7FF6FFB275B6FFB26" box="[209,284,1230,1249]" italics="true" pageId="4" pageNumber="467">A. coffea</emphasis>
|
||
</taxonomicName>
|
||
(paratype, Estación Pitilla, Costa Rica);
|
||
<emphasis id="B912CE4C6B62FFC7FDC4FB2658EBFB25" bold="true" box="[634,664,1231,1250]" pageId="4" pageNumber="467">18)</emphasis>
|
||
<taxonomicName id="4C6669DD6B62FFC7FD20FB275886FB26" box="[670,757,1230,1249]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="cyclops">
|
||
<emphasis id="B912CE4C6B62FFC7FD20FB275886FB26" box="[670,757,1230,1249]" italics="true" pageId="4" pageNumber="467">A. cyclops</emphasis>
|
||
</taxonomicName>
|
||
(holotype);
|
||
<emphasis id="B912CE4C6B62FFC7FCE1FB26590DFB25" bold="true" box="[863,894,1231,1250]" pageId="4" pageNumber="467">19)</emphasis>
|
||
<taxonomicName id="4C6669DD6B62FFC7FC3AFB2759BDFB26" box="[900,974,1230,1249]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="divisa">
|
||
<emphasis id="B912CE4C6B62FFC7FC3AFB2759BDFB26" box="[900,974,1230,1249]" italics="true" pageId="4" pageNumber="467">A. divisa</emphasis>
|
||
</taxonomicName>
|
||
(holotype);
|
||
<emphasis id="B912CE4C6B62FFC7FB86FB265E25FB25" bold="true" box="[1080,1110,1231,1250]" pageId="4" pageNumber="467">20)</emphasis>
|
||
<taxonomicName id="4C6669DD6B62FFC7FBE3FB275EDAFB26" box="[1117,1193,1230,1249]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="mersa">
|
||
<emphasis id="B912CE4C6B62FFC7FBE3FB275EDAFB26" box="[1117,1193,1230,1249]" italics="true" pageId="4" pageNumber="467">A. mersa</emphasis>
|
||
</taxonomicName>
|
||
(holotype);
|
||
<emphasis id="B912CE4C6B62FFC7FEADFB0F5B41FB3E" bold="true" box="[275,306,1254,1273]" pageId="4" pageNumber="467">21)</emphasis>
|
||
<taxonomicName id="4C6669DD6B62FFC7FE87FB0F5BDFFB3E" box="[313,428,1254,1273]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="mesosticta">
|
||
<emphasis id="B912CE4C6B62FFC7FE87FB0F5BDFFB3E" box="[313,428,1254,1273]" italics="true" pageId="4" pageNumber="467">A. mesosticta</emphasis>
|
||
</taxonomicName>
|
||
(holotype);
|
||
<emphasis id="B912CE4C6B62FFC7FDA5FB0F584AFB3E" bold="true" box="[539,569,1254,1273]" pageId="4" pageNumber="467">22)</emphasis>
|
||
<taxonomicName id="4C6669DD6B62FFC7FDFFFB0F58C4FB3E" box="[577,695,1254,1273]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="robiginosa">
|
||
<emphasis id="B912CE4C6B62FFC7FDFFFB0F58C4FB3E" box="[577,695,1254,1273]" italics="true" pageId="4" pageNumber="467">A. robiginosa</emphasis>
|
||
</taxonomicName>
|
||
(paratype, Zarcero, Costa Rica);
|
||
<emphasis id="B912CE4C6B62FFC7FC60FB0F598EFB3E" bold="true" box="[990,1021,1254,1273]" pageId="4" pageNumber="467">23)</emphasis>
|
||
<taxonomicName id="4C6669DD6B62FFC7FBBAFB0F5EDAFB3E" box="[1028,1193,1254,1273]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="strigodermoides">
|
||
<emphasis id="B912CE4C6B62FFC7FBBAFB0F5EDAFB3E" box="[1028,1193,1254,1273]" italics="true" pageId="4" pageNumber="467">A. strigodermoides</emphasis>
|
||
</taxonomicName>
|
||
(holotype);
|
||
<emphasis id="B912CE4C6B62FFC7FEAFFB175B5CFAD6" bold="true" box="[273,303,1278,1297]" pageId="4" pageNumber="467">24)</emphasis>
|
||
<taxonomicName id="4C6669DD6B62FFC7FE88FB175BD6FAD6" box="[310,421,1278,1297]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="467" phylum="Arthropoda" rank="species" species="unilineata">
|
||
<emphasis id="B912CE4C6B62FFC7FE88FB175BD6FAD6" box="[310,421,1278,1297]" italics="true" pageId="4" pageNumber="467">A. unilineata</emphasis>
|
||
</taxonomicName>
|
||
(paratype, Santa Rosa National Park, Costa Rica). Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD9125E6B62FFC7FF13FABA5EDBF906" blockId="4.[173,660,1363,1729]" lastBlockId="4.[707,1194,1363,1729]" pageId="4" pageNumber="467">
|
||
and a small macula on lateral foveae. Scutellum dark brown. Elytra ochre with dark brown sides, expanding at base, on median and apical parts of suture. Pygidium and venter ochre. Legs ochre with reddish tarsi and apical part of tibiae.
|
||
<emphasis id="B912CE4C6B62FFC7FDEBFA5458E0FA15" bold="true" box="[597,659,1469,1490]" pageId="4" pageNumber="467">Head:</emphasis>
|
||
Clypeus trapezoidal, surface densely punctatereticulate, flat. Anterior margin straight, clypeus thin in frontal view, width/length ratio 2.42. Frons punctate, less densely than clypeus, with a triangular concavity. Ocular canthus long, thin, with rounded apex. Ratio of interocular width/width of eye 2.9. Ratio antennal funiculus/club 0.85.
|
||
<emphasis id="B912CE4C6B62FFC7FF13F97B5B6CF960" bold="true" box="[173,287,1682,1703]" pageId="4" pageNumber="467">Pronotum:</emphasis>
|
||
Subtrapezoidal. Ratio width/length 1.75. Lateral margins angulated at 1/3 of pronotal length from the anterior margins. Anterior angles quadrate, posterior angles obtuse. Basal margin sinuate, subapical bead complete. Surface with deep, dense punctures.
|
||
<emphasis id="B912CE4C6B62FFC7FC12FA4A5E6BFA7F" bold="true" box="[940,1048,1443,1464]" pageId="4" pageNumber="467">Scutellum:</emphasis>
|
||
Subtriangular, with rounded sides and acute apex. Ratio width/ length 1.4. Surface sparsely punctate.
|
||
<emphasis id="B912CE4C6B62FFC7FBE2FA315EDAFA2A" bold="true" box="[1116,1193,1496,1517]" pageId="4" pageNumber="467">Elytra:</emphasis>
|
||
With costae defined by regular rows of punctures. Subsutural interstice with 2 rows of sparse punctures, 2
|
||
<superScript id="7C13BF166B62FFC7FCB2F9CB596DF9F7" attach="none" box="[780,798,1570,1584]" fontSize="6" pageId="4" pageNumber="467">nd</superScript>
|
||
–4
|
||
<superScript id="7C13BF166B62FFC7FC88F9CB5937F9F7" attach="left" box="[822,836,1570,1584]" fontSize="6" pageId="4" pageNumber="467">th</superScript>
|
||
interstices with 1 irregular, secondary stria.
|
||
<emphasis id="B912CE4C6B62FFC7FC9DF9AB59F8F990" bold="true" box="[803,907,1602,1623]" pageId="4" pageNumber="467">Pygidium:</emphasis>
|
||
Sparsely punctate with long setae on posterior margin. Ratio width/length 1.52.
|
||
<emphasis id="B912CE4C6B62FFC7FD43F99E5935F94B" bold="true" box="[765,838,1655,1676]" pageId="4" pageNumber="467">Venter:</emphasis>
|
||
Space between mesocoxae narrow, slightly convex. Mesometasternal suture welldefined at base of mesocoxae. Abdominal sternites
|
||
</paragraph>
|
||
<caption id="DF1942D66B63FFC6FF11F98F5E27F906" ID-DOI="http://doi.org/10.5281/zenodo.10106058" ID-Zenodo-Dep="10106058" httpUri="https://zenodo.org/record/10106058/files/figure.png" pageId="5" pageNumber="468" startId="5.[175,215,1638,1657]" targetBox="[178,1140,195,1606]" targetPageId="5">
|
||
<paragraph id="8BD9125E6B63FFC6FF11F98F5E27F906" blockId="5.[151,1173,1638,1729]" pageId="5" pageNumber="468">
|
||
<emphasis id="B912CE4C6B63FFC6FF11F98F5B6CF9BE" bold="true" box="[175,287,1638,1657]" pageId="5" pageNumber="468">Figs. 25–33.</emphasis>
|
||
<taxonomicName id="4C6669DD6B63FFC6FE8BF98F5BBDF9BE" box="[309,462,1638,1657]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="468" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B912CE4C6B63FFC6FE8BF98F5BF7F9BE" box="[309,388,1638,1657]" italics="true" pageId="5" pageNumber="468">Anomala</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, genitalia. Endophalli:
|
||
<emphasis id="B912CE4C6B63FFC6FD18F98F58B6F9BE" bold="true" box="[678,709,1638,1657]" pageId="5" pageNumber="468">25)</emphasis>
|
||
<taxonomicName id="4C6669DD6B63FFC6FD71F98F5953F9BE" box="[719,800,1638,1657]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="468" phylum="Arthropoda" rank="species" species="coffea">
|
||
<emphasis id="B912CE4C6B63FFC6FD71F98F5953F9BE" box="[719,800,1638,1657]" italics="true" pageId="5" pageNumber="468">A. coffea</emphasis>
|
||
</taxonomicName>
|
||
(paratype, Estación Pitilla, Costa Rica);
|
||
<emphasis id="B912CE4C6B63FFC6FF29F9975AC6F956" bold="true" box="[151,181,1662,1681]" pageId="5" pageNumber="468">26)</emphasis>
|
||
<taxonomicName id="4C6669DD6B63FFC6FF02F9975B61F956" box="[188,274,1662,1681]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="468" phylum="Arthropoda" rank="species" species="cyclops">
|
||
<emphasis id="B912CE4C6B63FFC6FF02F9975B61F956" box="[188,274,1662,1681]" italics="true" pageId="5" pageNumber="468">A. cyclops</emphasis>
|
||
</taxonomicName>
|
||
(paratype, Finca Jenny, Costa Rica);
|
||
<emphasis id="B912CE4C6B63FFC6FDEFF997581DF956" bold="true" box="[593,622,1662,1681]" pageId="5" pageNumber="468">27)</emphasis>
|
||
<taxonomicName id="4C6669DD6B63FFC6FDCBF99758CCF956" box="[629,703,1662,1681]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="468" phylum="Arthropoda" rank="species" species="divisa">
|
||
<emphasis id="B912CE4C6B63FFC6FDCBF99758CCF956" box="[629,703,1662,1681]" italics="true" pageId="5" pageNumber="468">A. divisa</emphasis>
|
||
</taxonomicName>
|
||
(holotype);
|
||
<emphasis id="B912CE4C6B63FFC6FC96F9975936F956" bold="true" box="[808,837,1662,1681]" pageId="5" pageNumber="468">28)</emphasis>
|
||
<taxonomicName id="4C6669DD6B63FFC6FCF2F99759EBF956" box="[844,920,1662,1681]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="468" phylum="Arthropoda" rank="species" species="mersa">
|
||
<emphasis id="B912CE4C6B63FFC6FCF2F99759EBF956" box="[844,920,1662,1681]" italics="true" pageId="5" pageNumber="468">A. mersa</emphasis>
|
||
</taxonomicName>
|
||
(paratype, Sector Palo Verde, Costa Rica);
|
||
<emphasis id="B912CE4C6B63FFC6FEBCF97F5B6DF96E" bold="true" box="[258,286,1686,1705]" pageId="5" pageNumber="468">29)</emphasis>
|
||
<taxonomicName id="4C6669DD6B63FFC6FE9BF97F5BE3F96E" box="[293,400,1686,1705]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="468" phylum="Arthropoda" rank="species" species="mesosticta">
|
||
<emphasis id="B912CE4C6B63FFC6FE9BF97F5BE3F96E" box="[293,400,1686,1705]" italics="true" pageId="5" pageNumber="468">A. mesosticta</emphasis>
|
||
</taxonomicName>
|
||
(holotype);
|
||
<emphasis id="B912CE4C6B63FFC6FE4AF97F5862F96E" bold="true" box="[500,529,1686,1705]" pageId="5" pageNumber="468">30)</emphasis>
|
||
<taxonomicName id="4C6669DD6B63FFC6FDA9F97F58F6F96E" box="[535,645,1686,1705]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="468" phylum="Arthropoda" rank="species" species="robiginosa">
|
||
<emphasis id="B912CE4C6B63FFC6FDA9F97F58F6F96E" box="[535,645,1686,1705]" italics="true" pageId="5" pageNumber="468">A. robiginosa</emphasis>
|
||
</taxonomicName>
|
||
(holotype);
|
||
<emphasis id="B912CE4C6B63FFC6FD56F97F5976F96E" bold="true" box="[744,773,1686,1705]" pageId="5" pageNumber="468">31)</emphasis>
|
||
<taxonomicName id="4C6669DD6B63FFC6FCB5F97F59D7F96E" box="[779,932,1686,1705]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="468" phylum="Arthropoda" rank="species" species="strigodermoides">
|
||
<emphasis id="B912CE4C6B63FFC6FCB5F97F59D7F96E" box="[779,932,1686,1705]" italics="true" pageId="5" pageNumber="468">A. strigodermoides</emphasis>
|
||
</taxonomicName>
|
||
(holotype);
|
||
<emphasis id="B912CE4C6B63FFC6FBB6F97F5E56F96E" bold="true" box="[1032,1061,1686,1705]" pageId="5" pageNumber="468">32)</emphasis>
|
||
<taxonomicName id="4C6669DD6B63FFC6FB95F97F5EE7F96E" box="[1067,1172,1686,1705]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="468" phylum="Arthropoda" rank="species" species="unilineata">
|
||
<emphasis id="B912CE4C6B63FFC6FB95F97F5EE7F96E" box="[1067,1172,1686,1705]" italics="true" pageId="5" pageNumber="468">A. unilineata</emphasis>
|
||
</taxonomicName>
|
||
(holotype). Aedeagus:
|
||
<emphasis id="B912CE4C6B63FFC6FEE8F9475B00F906" bold="true" box="[342,371,1710,1729]" pageId="5" pageNumber="468">33)</emphasis>
|
||
<taxonomicName id="4C6669DD6B63FFC6FEC4F9475B8BF906" authorityName="Blanchard" authorityYear="1851" box="[378,504,1710,1729]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="468" phylum="Arthropoda" rank="species" species="sticticoptera">
|
||
<emphasis id="B912CE4C6B63FFC6FEC4F9475B8BF906" box="[378,504,1710,1729]" italics="true" pageId="5" pageNumber="468">A. sticticoptera</emphasis>
|
||
</taxonomicName>
|
||
(lateral view, Cuyotenango, Guatemala, MNHUB). Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1942D66B60FFC5FF7BFC115EDAFBCC" ID-DOI="http://doi.org/10.5281/zenodo.10106060" ID-Zenodo-Dep="10106060" box="[197,1193,1015,1035]" httpUri="https://zenodo.org/record/10106060/files/figure.png" pageId="6" pageNumber="469" startId="6.[197,237,1016,1035]" targetBox="[178,1183,206,993]" targetPageId="6">
|
||
<paragraph id="8BD9125E6B60FFC5FF7BFC115EDAFBCC" blockId="6.[173,1193,1015,1059]" box="[197,1193,1015,1035]" pageId="6" pageNumber="469">
|
||
<emphasis id="B912CE4C6B60FFC5FF7BFC115B46FBCC" bold="true" box="[197,309,1015,1035]" pageId="6" pageNumber="469">Figs. 34–36.</emphasis>
|
||
<taxonomicName id="4C6669DD6B60FFC5FEF5FC1E5B97FBCC" box="[331,484,1015,1035]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B912CE4C6B60FFC5FEF5FC1E5BE9FBCD" box="[331,410,1015,1034]" italics="true" pageId="6" pageNumber="469">Anomala</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, distribution maps.
|
||
<emphasis id="B912CE4C6B60FFC5FD29FC1158C5FBCC" bold="true" box="[663,694,1016,1035]" pageId="6" pageNumber="469">34)</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="4C6669DD6B60FFC5FCF7FC1E59EBFBCD" box="[841,920,1015,1034]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="coffea">
|
||
<emphasis id="B912CE4C6B60FFC5FCF7FC1E59EBFBCD" box="[841,920,1015,1034]" italics="true" pageId="6" pageNumber="469">A. coffea</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6669DD6B60FFC5FC18FC1E5985FBCD" box="[934,1014,1015,1034]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="mersa">
|
||
<emphasis id="B912CE4C6B60FFC5FC18FC1E5985FBCD" box="[934,1014,1015,1034]" italics="true" pageId="6" pageNumber="469">A. mersa</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C6669DD6B60FFC5FB95FC1E5ED0FBCD" box="[1067,1187,1015,1034]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="robiginosa">
|
||
<emphasis id="B912CE4C6B60FFC5FB95FC1E5ED0FBCD" box="[1067,1187,1015,1034]" italics="true" pageId="6" pageNumber="469">A. robiginosa</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD9125E6B60FFC5FF13FBF95EDAFBE4" blockId="6.[173,1193,1015,1059]" box="[173,1193,1039,1059]" pageId="6" pageNumber="469">
|
||
<emphasis id="B912CE4C6B60FFC5FF13FBF95ABFFBE4" bold="true" box="[173,204,1040,1059]" pageId="6" pageNumber="469">35)</emphasis>
|
||
Distribution of
|
||
<emphasis id="B912CE4C6B60FFC5FEE9FBE65843FBE5" box="[343,560,1039,1058]" italics="true" pageId="6" pageNumber="469">
|
||
<taxonomicName id="4C6669DD6B60FFC5FEE9FBE65BC0FBE5" box="[343,435,1039,1058]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="cyclops">A. cyclops</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6669DD6B60FFC5FE02FBE65843FBE5" box="[444,560,1039,1058]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="mesosticta">A. mesosticta</taxonomicName>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="4C6669DD6B60FFC5FDDFFBE65976FBE5" box="[609,773,1039,1058]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="strigodermoides">
|
||
<emphasis id="B912CE4C6B60FFC5FDDFFBE65976FBE5" box="[609,773,1039,1058]" italics="true" pageId="6" pageNumber="469">A. strigodermoides</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<emphasis id="B912CE4C6B60FFC5FCAFFBF9595CFBE4" bold="true" box="[785,815,1040,1059]" pageId="6" pageNumber="469">36)</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="4C6669DD6B60FFC5FC02FBE65E7BFBE5" box="[956,1032,1039,1058]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="divisa">
|
||
<emphasis id="B912CE4C6B60FFC5FC02FBE65E7BFBE5" box="[956,1032,1039,1058]" italics="true" pageId="6" pageNumber="469">A. divisa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6669DD6B60FFC5FB8DFBE65ED0FBE5" box="[1075,1187,1039,1058]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="unilineata">
|
||
<emphasis id="B912CE4C6B60FFC5FB8DFBE65ED0FBE5" box="[1075,1187,1039,1058]" italics="true" pageId="6" pageNumber="469">A. unilineata</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD9125E6B60FFC5FF13FB8D5B88FA2A" blockId="6.[173,660,1123,1730]" pageId="6" pageNumber="469">
|
||
with 1 row of setae and 3–5 rows of punctures per sternite. Last sternite strigate, with subapical bead thin and sinuate, apical membrane thin.
|
||
<emphasis id="B912CE4C6B60FFC5FDE2FB7058E1FB69" bold="true" box="[604,658,1177,1198]" pageId="6" pageNumber="469">Legs:</emphasis>
|
||
Protibia (
|
||
<figureCitation id="135D0EDB6B60FFC5FEB1FB5A5B2FFB0F" box="[271,348,1203,1224]" captionStart="Figs" captionStartId="3.[175,216,1111,1130]" captionTargetBox="[154,1163,186,1089]" captionTargetId="figure-261@3.[151,1173,183,1093]" captionTargetPageId="3" captionText="Figs. 9–16. Anomala species, protibiae. 9) A. coffea (male: holotype; female: paratype, Estación Pitilla, Costa Rica); 10) A. cyclops (male: holotype; female: paratype, Los Almendros, Costa Rica); 11) A. divisa (male: holotype; female: paratype, Estación Biológica Las Alturas, Costa Rica); 12) A. mersa (male: paratype, Sector Palo Verde, Costa Rica); 13) A. mesosticta (male: holotype; female: paratype, Rio Sardinas, Costa Rica); 14) A. robiginosa (male: holotype; female: paratype, Santa Rosa National Park, Costa Rica); 15) A. strigodermoides (male: holotype; female: paratype, Catarata río Buenavista, Alajuela); 16) A. unilineata (holotype). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106054" httpUri="https://zenodo.org/record/10106054/files/figure.png" pageId="6" pageNumber="469">Fig. 10</figureCitation>
|
||
) tridentate, apical tooth long and curved, middle tooth below internal apex of protibia, triangular in shape, acute, basal tooth sharp. Metatibia fusiform. Ratio length/width 3.16. First external carina well-developed, surface punctate. Internal protarsal claw bifurcate, with upper branch slightly shorter and half width of the lower one, inferior margin sinuate.
|
||
<emphasis id="B912CE4C6B60FFC5FE45FA8458E0FA45" bold="true" box="[507,659,1389,1410]" pageId="6" pageNumber="469">Male genitalia:</emphasis>
|
||
Parameres with wide apex in lateral view (
|
||
<figureCitation id="135D0EDB6B60FFC5FDFFFA6158F4FA5A" box="[577,647,1416,1437]" captionStart="Figs" captionStartId="4.[197,237,1207,1226]" captionTargetBox="[176,1181,187,1184]" captionTargetId="figure-197@4.[173,1195,183,1197]" captionTargetPageId="4" captionText="Figs. 17–24. Anomala species, aedeagi, lateral view (top), dorsal view (bottom left), ventral view (bottom right). 17) A. coffea (paratype, Estación Pitilla, Costa Rica); 18) A. cyclops (holotype); 19) A. divisa (holotype); 20) A. mersa (holotype); 21) A. mesosticta (holotype); 22) A. robiginosa (paratype, Zarcero, Costa Rica); 23) A. strigodermoides (holotype); 24) A. unilineata (paratype, Santa Rosa National Park, Costa Rica). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106056" httpUri="https://zenodo.org/record/10106056/files/figure.png" pageId="6" pageNumber="469">Fig. 18</figureCitation>
|
||
), ventral margin sinuate. Endophallus (
|
||
<figureCitation id="135D0EDB6B60FFC5FDABFA4A5828FA7F" box="[533,603,1443,1464]" captionStart="Figs" captionStartId="5.[175,215,1638,1657]" captionTargetBox="[178,1140,195,1606]" captionTargetId="figure-1@5.[168,1156,183,1627]" captionTargetPageId="5" captionText="Figs. 25–33. Anomala species, genitalia. Endophalli: 25) A. coffea (paratype, Estación Pitilla, Costa Rica); 26) A. cyclops (paratype, Finca Jenny, Costa Rica); 27) A. divisa (holotype); 28) A. mersa (paratype, Sector Palo Verde, Costa Rica); 29) A. mesosticta (holotype); 30) A. robiginosa (holotype); 31) A. strigodermoides (holotype); 32) A. unilineata (holotype). Aedeagus: 33) A. sticticoptera (lateral view, Cuyotenango, Guatemala, MNHUB). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106058" httpUri="https://zenodo.org/record/10106058/files/figure.png" pageId="6" pageNumber="469">Fig. 26</figureCitation>
|
||
) with 1 long, glabrous sacculus. Ejaculatory duct opening frontally on a lateral inflation.
|
||
</paragraph>
|
||
<paragraph id="8BD9125E6B60FFC5FF7BFA1B5800F9E5" blockId="6.[173,660,1123,1730]" pageId="6" pageNumber="469">
|
||
<emphasis id="B912CE4C6B60FFC5FF7BFA1B5B66F9C0" bold="true" box="[197,277,1522,1543]" pageId="6" pageNumber="469">Female.</emphasis>
|
||
Similar to male except protibia (
|
||
<figureCitation id="135D0EDB6B60FFC5FDF4FA1B58FFF9C0" box="[586,652,1522,1543]" captionStart="Figs" captionStartId="3.[175,216,1111,1130]" captionTargetBox="[154,1163,186,1089]" captionTargetId="figure-261@3.[151,1173,183,1093]" captionTargetPageId="3" captionText="Figs. 9–16. Anomala species, protibiae. 9) A. coffea (male: holotype; female: paratype, Estación Pitilla, Costa Rica); 10) A. cyclops (male: holotype; female: paratype, Los Almendros, Costa Rica); 11) A. divisa (male: holotype; female: paratype, Estación Biológica Las Alturas, Costa Rica); 12) A. mersa (male: paratype, Sector Palo Verde, Costa Rica); 13) A. mesosticta (male: holotype; female: paratype, Rio Sardinas, Costa Rica); 14) A. robiginosa (male: holotype; female: paratype, Santa Rosa National Park, Costa Rica); 15) A. strigodermoides (male: holotype; female: paratype, Catarata río Buenavista, Alajuela); 16) A. unilineata (holotype). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10106054" httpUri="https://zenodo.org/record/10106054/files/figure.png" pageId="6" pageNumber="469">Fig. 10</figureCitation>
|
||
) with wider teeth; last sternite with curved apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C41D56B60FFC5FF7BF9CE5E4EFB39" pageId="6" pageNumber="469" type="description">
|
||
<paragraph id="8BD9125E6B60FFC5FF7BF9CE5E4EFB39" blockId="6.[173,660,1123,1730]" lastBlockId="6.[707,1194,1123,1730]" pageId="6" pageNumber="469">
|
||
<emphasis id="B912CE4C6B60FFC5FF7BF9CE5B5AF9FB" bold="true" box="[197,297,1575,1596]" pageId="6" pageNumber="469">Variation.</emphasis>
|
||
Nine
|
||
<typeStatus id="54DDACFC6B60FFC5FECEF9CE5BA0F9FB" box="[368,467,1575,1596]" pageId="6" pageNumber="469" type="paratype">paratypes</typeStatus>
|
||
. Head varies from orange to dark reddish brown; presence of a complete or incomplete reddish brown transversal band on elytra or median expansion on suture; pygidium and venter vary from ochre to light brown. Body length
|
||
<quantity id="4C9EBFBB6B60FFC5FF50F9455BF7F906" box="[238,388,1708,1730]" metricMagnitude="-3" metricUnit="m" metricValue="9.364999999999998" metricValueMax="10.29" metricValueMin="8.44" pageId="6" pageNumber="469" unit="mm" value="9.364999999999998" valueMax="10.29" valueMin="8.44">8.44–10.29 mm</quantity>
|
||
, body width
|
||
<quantity id="4C9EBFBB6B60FFC5FDBAF94558E3F906" box="[516,656,1708,1730]" metricMagnitude="-3" metricUnit="m" metricValue="4.89" metricValueMax="5.37" metricValueMin="4.41" pageId="6" pageNumber="469" unit="mm" value="4.89" valueMax="5.37" valueMin="4.41">4.41–5.37 mm</quantity>
|
||
. Clypeal width/length ratio 2.11–2.54. Ratio of interocular width/width of eye 2.50–2.90. Ratio antennal funiculus/club 0.67–0.85. Pronotal width/ length ratio 1.67–1.78. Scutellar width/length ratio 1.10–1.43. Pygidial width/length ratio 1.48–1.68. Metatibial length/width ratio 2.88–3.28.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C41D56B60FFC5FD65FAEA591DFAAF" pageId="6" pageNumber="469" type="etymology">
|
||
<paragraph id="8BD9125E6B60FFC5FD65FAEA591DFAAF" blockId="6.[707,1194,1123,1730]" pageId="6" pageNumber="469">
|
||
<emphasis id="B912CE4C6B60FFC5FD65FAEA593AFADF" bold="true" box="[731,841,1283,1304]" pageId="6" pageNumber="469">Etymology.</emphasis>
|
||
From the Latin noun
|
||
<emphasis id="B912CE4C6B60FFC5FBA0FAEA5ED0FADE" box="[1054,1187,1283,1305]" italics="true" pageId="6" pageNumber="469">Cyclops, -opis</emphasis>
|
||
, a one-eyed mythological creature, to be treated as a noun in apposition, for the single macula on the pronotum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C41D56B60FFC5FD65FA8459ADFA7F" pageId="6" pageNumber="469" type="distribution">
|
||
<paragraph id="8BD9125E6B60FFC5FD65FA8459ADFA7F" blockId="6.[707,1194,1123,1730]" pageId="6" pageNumber="469">
|
||
<emphasis id="B912CE4C6B60FFC5FD65FA84592EFA45" bold="true" box="[731,861,1389,1410]" pageId="6" pageNumber="469">Distribution.</emphasis>
|
||
<taxonomicName id="4C6669DD6B60FFC5FCD5FA845E52FA44" box="[875,1057,1389,1411]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="cyclops">
|
||
<emphasis id="B912CE4C6B60FFC5FCD5FA845E52FA44" box="[875,1057,1389,1411]" italics="true" pageId="6" pageNumber="469">Anomala cyclops</emphasis>
|
||
</taxonomicName>
|
||
inhabits the dry forest in northeastern
|
||
<collectingCountry id="F37152CE6B60FFC5FC73FA615E4DFA5A" box="[973,1086,1416,1437]" name="Costa Rica" pageId="6" pageNumber="469">Costa Rica</collectingCountry>
|
||
, from
|
||
<quantity id="4C9EBFBB6B60FFC5FB3AFA615968FA7F" metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="6" pageNumber="469" unit="m" value="200.0" valueMax="300.0" valueMin="100.0">100 to 300 m</quantity>
|
||
elevation (
|
||
<figureCitation id="135D0EDB6B60FFC5FC37FA4A59A1FA7F" box="[905,978,1443,1464]" captionStart="Figs" captionStartId="6.[197,237,1016,1035]" captionTargetBox="[178,1183,206,993]" captionTargetId="figure-344@6.[173,1193,183,1019]" captionTargetPageId="6" captionText="Figs. 34–36. Anomala species, distribution maps. 34) Distribution of A. coffea, A. mersa, and A. robiginosa;" figureDoi="http://doi.org/10.5281/zenodo.10106060" httpUri="https://zenodo.org/record/10106060/files/figure.png" pageId="6" pageNumber="469">Fig. 35</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37C41D56B60FFC4FD65FA54586CFC92" lastPageId="7" lastPageNumber="470" pageId="6" pageNumber="469" type="diagnosis">
|
||
<paragraph id="8BD9125E6B60FFC4FD65FA54586CFC92" blockId="6.[707,1194,1123,1730]" lastBlockId="7.[151,639,194,853]" lastPageId="7" lastPageNumber="470" pageId="6" pageNumber="469">
|
||
<emphasis id="B912CE4C6B60FFC5FD65FA545937FA15" bold="true" box="[731,836,1469,1490]" pageId="6" pageNumber="469">Diagnosis.</emphasis>
|
||
This species is recognized by its medium–small size, ochre color with a median dark macula on the pronotum and a transverse band on the elytra, tridentate protibia, parameres with a wide apex in lateral view and sinuate ventral margin. Species with a similar pronotal pattern and size are
|
||
<taxonomicName id="4C6669DD6B60FFC5FCD3F9B55969F94B" authority="Ohaus, 1897" authorityName="Ohaus" authorityYear="1897" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="boliviana">
|
||
<emphasis id="B912CE4C6B60FFC5FCD3F9B55E56F9B5" box="[877,1061,1628,1650]" italics="true" pageId="6" pageNumber="469">Anomala boliviana</emphasis>
|
||
Ohaus, 1897 (
|
||
<collectingCountry id="F37152CE6B60FFC5FD77F99E5960F94B" box="[713,787,1655,1676]" name="Bolivia" pageId="6" pageNumber="469">Bolivia</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6669DD6B60FFC5FC97F99E5964F960" authority="Bates, 1888" authorityName="Bates" authorityYear="1888" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="subSpecies" species="flavilla" subSpecies="centralis">
|
||
<emphasis id="B912CE4C6B60FFC5FC97F99E5E59F94A" box="[809,1066,1655,1677]" italics="true" pageId="6" pageNumber="469">Anomala flavilla centralis</emphasis>
|
||
Bates, 1888 (
|
||
<collectingCountry id="F37152CE6B60FFC5FD77F97B5962F960" box="[713,785,1682,1703]" name="Mexico" pageId="6" pageNumber="469">Mexico</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C6669DD6B60FFC5FCA1F9785ED6F960" authority="LeConte, 1863" authorityName="LeConte" authorityYear="1863" box="[799,1189,1681,1703]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="centralis">
|
||
<emphasis id="B912CE4C6B60FFC5FCA1F97859B1F960" box="[799,962,1681,1703]" italics="true" pageId="6" pageNumber="469">Anomala centralis</emphasis>
|
||
LeConte, 1863 (
|
||
<collectingCountry id="F37152CE6B60FFC5FBE9F97B5EECF960" box="[1111,1183,1682,1703]" name="Mexico" pageId="6" pageNumber="469">Mexico</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C6669DD6B60FFC5FD4EF9455ED5F906" authority="Bates, 1888" authorityName="Bates" authorityYear="1888" box="[752,1190,1708,1730]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="469" phylum="Arthropoda" rank="species" species="chevrolati">
|
||
<emphasis id="B912CE4C6B60FFC5FD4EF94559CBF905" box="[752,952,1708,1730]" italics="true" pageId="6" pageNumber="469">Anomala chevrolati</emphasis>
|
||
Bates, 1888 (
|
||
<collectingCountry id="F37152CE6B60FFC5FBF3F9455EECF906" box="[1101,1183,1708,1729]" name="Mexico" pageId="6" pageNumber="469">Mexico</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
. However, all these species have a bidentate protibia, whereas
|
||
<taxonomicName id="4C6669DD6B61FFC4FF52FF355B3DFF35" box="[236,334,220,242]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="cyclops">
|
||
<emphasis id="B912CE4C6B61FFC4FF52FF355B3DFF35" box="[236,334,220,242]" italics="true" pageId="7" pageNumber="470">A. cyclops</emphasis>
|
||
</taxonomicName>
|
||
has a tridentate protibia. Moreover,
|
||
<taxonomicName id="4C6669DD6B61FFC4FF6EFF1E5B3BFECA" authorityName="Ohaus" authorityYear="1897" box="[208,328,247,269]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="boliviana">
|
||
<emphasis id="B912CE4C6B61FFC4FF6EFF1E5B3BFECA" box="[208,328,247,269]" italics="true" pageId="7" pageNumber="470">A. boliviana</emphasis>
|
||
</taxonomicName>
|
||
has maculae on the elytra and pygidium, ochre elytral suture, first interstice of elytra wider with 3–4 irregular rows of punctures, and a larger pronotal macula.
|
||
<taxonomicName id="4C6669DD6B61FFC4FE6AFEAF5A9FFEB0" authorityName="Bates" authorityYear="1888" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="470" phylum="Arthropoda" rank="subSpecies" species="flavilla" subSpecies="centralis">
|
||
<emphasis id="B912CE4C6B61FFC4FE6AFEAF5A9FFEB0" italics="true" pageId="7" pageNumber="470">Anomala flavilla centralis</emphasis>
|
||
</taxonomicName>
|
||
has only the elytral suture pigmented, and the parameres are long with an acute and curved apex and the ventral margin acute and pointing posteriorly in lateral view.
|
||
<taxonomicName id="4C6669DD6B61FFC4FE11FE585829FE00" authorityName="LeConte" authorityYear="1863" box="[431,602,433,455]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="centralis">
|
||
<emphasis id="B912CE4C6B61FFC4FE11FE585829FE00" box="[431,602,433,455]" italics="true" pageId="7" pageNumber="470">Anomala centralis</emphasis>
|
||
</taxonomicName>
|
||
has a triangular pronotal macula, ochre elytral suture, and elytra with darker shading on the lateral margins.
|
||
<taxonomicName id="4C6669DD6B61FFC4FF70FDE95BFEFDD1" authorityName="Bates" authorityYear="1888" box="[206,397,512,534]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="chevrolati">
|
||
<emphasis id="B912CE4C6B61FFC4FF70FDE95BFEFDD1" box="[206,397,512,534]" italics="true" pageId="7" pageNumber="470">Anomala chevrolati</emphasis>
|
||
</taxonomicName>
|
||
has an irregular reddish pronotal macula, a wide, dark elytral suture homogeneously wide on the apical half, denser punctures on the elytra, and finer secondary punctures on the elytral surface.
|
||
<taxonomicName id="4C6669DD6B61FFC4FECFFD825AB4FD5C" authority="Ohaus, 1911" authorityName="Ohaus" authorityYear="1911" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="bruchiana">
|
||
<emphasis id="B912CE4C6B61FFC4FECFFD825840FD46" box="[369,563,619,641]" italics="true" pageId="7" pageNumber="470">Anomala bruchiana</emphasis>
|
||
Ohaus, 1911
|
||
</taxonomicName>
|
||
has a tridentate protibia and a small pronotal macula, but it has a triangular shape, ochre elytra with a narrow, dark elytral suture, and slender, long parameres with an acute apex and curved ventral margin.
|
||
<taxonomicName id="4C6669DD6B61FFC4FE85FD19580EFCC2" authority="Bates, 1888" authorityName="Bates" authorityYear="1888" box="[315,637,752,774]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="470" phylum="Arthropoda" rank="species" species="hoepfneri">
|
||
<emphasis id="B912CE4C6B61FFC4FE85FD195B88FCC1" box="[315,507,752,774]" italics="true" pageId="7" pageNumber="470">Anomala hoepfneri</emphasis>
|
||
Bates, 1888
|
||
</taxonomicName>
|
||
has a similar elytral pattern but with a large, irregular pronotal macula, bidentate protibia, and wide and densely punctured elytral interstices.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |