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<document id="42D575F27D725ADC2C195B26C28BDD4B" ID-CLB-Dataset="22877" ID-DOI="10.5852/ejt.2020.629" ID-GBIF-Dataset="3722aa65-b935-4294-ac20-0d1179da9015" ID-Zenodo-Dep="3762280" ID-ZooBank="ECEBD539-6E3E-45BE-A0CB-264DF3270CC0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587578640316" checkinUser="valdenar" docAuthor="Herbert, David G." docDate="2020" docId="03C78785460FFFC14211422A33F8FA1B" docLanguage="en" docName="ejt-629_herbert.pdf.imf" docOrigin="European Journal of Taxonomy 629" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Afrodonta Melvill &amp; Ponsonby 1908" docType="treatment" docVersion="7" lastPageNumber="4" masterDocId="FFFEFFFD460CFFC2400243213138FFD9" masterDocTitle="Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies" masterLastPageNumber="55" masterPageNumber="1" pageNumber="4" updateTime="1698835565728" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="630D2373FDB95B3E9B4EA5427155A276">Revision of the aperturally dentate Charopidae (Gastropoda: Stylommatophora) of southern Africa - genus Afrodonta s. lat., with description of five new genera, twelve new species and one new subspecies</mods:title>
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<mods:namePart id="80B98DDE84A7653C2F6A7D49F162E618">Herbert, David G.</mods:namePart>
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<mods:affiliation id="199AF6A6433FCDBC598F224645ECC30F">Department of Natural Sciences, National Museum Wales, Cathays Park, Cardiff, CF 10 3 NP, UK. Formerly at KwaZulu-Natal Museum, P. Bag 9070, Pietermaritzburg 3201, South Africa</mods:affiliation>
<mods:nameIdentifier id="B0E138CC143DB1FEF392D36D4607F22C" type="email">phasianella@gmail.com</mods:nameIdentifier>
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<mods:title id="FAA29A33F918001825A9E1EA85217D90">European Journal of Taxonomy</mods:title>
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<mods:date id="AB915108795D32F9B0968EC1B10244F1">2020</mods:date>
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<heading id="D09981FF460FFFC14211422A3518FEFC" box="[531,1056,267,294]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="4" reason="2">
Genus
<taxonomicName id="4C6E4D10460FFFC14264422A3518FEFC" authority="Melvill &amp; Ponsonby, 1908" authorityName="Melvill &amp; Ponsonby" authorityYear="1908" box="[614,1056,267,294]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
<emphasis id="B91AEA81460FFFC14264422A33DAFEFF" bold="true" box="[614,738,267,294]" italics="true" pageId="3" pageNumber="4">Afrodonta</emphasis>
<bibRefCitation id="EFFF4B62460FFFC142EB422A3518FEFC" author="Melvill J. C. &amp; Ponsonby J. H." box="[745,1056,267,293]" pageId="3" pageNumber="4" pagination="129 - 136" refId="ref26977" refString="Melvill J. C. &amp; Ponsonby J. H. 1908. Descriptions of nine terrestrial Mollusca from South Africa. Annals and Magazine of Natural History, Series 8, 1 (2): 129 - 136. https: // doi. org / 10.1080 / 00222930808692373" type="journal article" year="1908">Melvill &amp; Ponsonby, 1908</bibRefCitation>
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<treatmentCitation id="0ACF1082460FFFC140BF42573388FE49" author="Melvill J. C. &amp; Ponsonby J. H." box="[189,688,374,400]" page="133" pageId="3" pageNumber="4" year="1908">
<taxonomicName id="4C6E4D10460FFFC140BF42573388FE49" authority="Melvill &amp; Ponsonby, 1908: 133" authorityName="Melvill &amp; Ponsonby" authorityPageNumber="133" authorityYear="1908" box="[189,688,374,400]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
<emphasis id="B91AEA81460FFFC140BF4257300BFE49" box="[189,307,374,400]" italics="true" pageId="3" pageNumber="4">Afrodonta</emphasis>
<bibRefCitation id="EFFF4B62460FFFC1413842573388FE49" author="Melvill J. C. &amp; Ponsonby J. H." box="[314,688,374,400]" pageId="3" pageNumber="4" pagination="129 - 136" refId="ref26977" refString="Melvill J. C. &amp; Ponsonby J. H. 1908. Descriptions of nine terrestrial Mollusca from South Africa. Annals and Magazine of Natural History, Series 8, 1 (2): 129 - 136. https: // doi. org / 10.1080 / 00222930808692373" type="journal article" year="1908">Melvill &amp; Ponsonby, 1908: 133</bibRefCitation>
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.
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</paragraph>
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<subSubSection id="C3746518460FFFC140BF429A3060FE0D" box="[189,344,442,469]" pageId="3" pageNumber="4" type="type_taxon">
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<emphasis id="B91AEA81460FFFC140BF429A3060FE0D" bold="true" box="[189,344,442,469]" pageId="3" pageNumber="4">
<typeStatus id="54D58831460FFFC140BF429A31C2FE0C" box="[189,250,443,469]" pageId="3" pageNumber="4">Type</typeStatus>
species
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<subSubSection id="C3746518460FFFC140BF42CD347AFDDE" box="[189,1346,492,519]" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="8BD13693460FFFC140BF42CD347AFDDE" blockId="3.[189,1346,492,519]" box="[189,1346,492,519]" pageId="3" pageNumber="4">
<treatmentCitationGroup id="AB7E11BD460FFFC140BF42CD347AFDDE" box="[189,1346,492,519]" pageId="3" pageNumber="4">
<taxonomicName id="4C6E4D10460FFFC140BF42CD3231FDDF" authority="Melvill &amp; Ponsonby, 1908" authorityName="Melvill &amp; Ponsonby" authorityYear="1908" box="[189,777,492,518]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="bilamellaris">
<emphasis id="B91AEA81460FFFC140BF42CD30F0FDDF" box="[189,456,492,518]" italics="true" pageId="3" pageNumber="4">Afrodonta bilamellaris</emphasis>
<bibRefCitation id="EFFF4B62460FFFC141CD42CD3231FDDF" author="Melvill J. C. &amp; Ponsonby J. H." box="[463,777,492,518]" pageId="3" pageNumber="4" pagination="129 - 136" refId="ref26977" refString="Melvill J. C. &amp; Ponsonby J. H. 1908. Descriptions of nine terrestrial Mollusca from South Africa. Annals and Magazine of Natural History, Series 8, 1 (2): 129 - 136. https: // doi. org / 10.1080 / 00222930808692373" type="journal article" year="1908">Melvill &amp; Ponsonby, 1908</bibRefCitation>
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, by subsequent designation (
<treatmentCitation id="0ACF1082460FFFC1445842CD340DFDDE" author="Burnup H. C." box="[1114,1333,492,519]" page="335" pageId="3" pageNumber="4" year="1912">
<bibRefCitation id="EFFF4B62460FFFC1445842CD340DFDDE" author="Burnup H. C." box="[1114,1333,492,519]" pageId="3" pageNumber="4" pagination="333 - 343" refId="ref26463" refString="Burnup H. C. 1912. On Afrodonta Melv. and Pons., with descriptions of new species. Annals of the Natal Museum 2 (3): 333 - 343." type="journal article" year="1912">
Burnup
<date id="FFD01053460FFFC144B942CD35C2FDDF" box="[1211,1274,492,518]" pageId="3" pageNumber="4" value="1912">1912</date>
: 335
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).
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</paragraph>
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<subSubSection id="C3746518460FFFC140BF4111323DFC51" pageId="3" pageNumber="4" type="diagnosis">
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<heading id="D09981FF460FFFC140BF4111300EFD93" bold="true" box="[189,310,560,586]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B91AEA81460FFFC140BF4111300EFD93" bold="true" box="[189,310,560,586]" pageId="3" pageNumber="4">Diagnosis</emphasis>
</heading>
</paragraph>
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Shell very small (max. diameter ±
<quantity id="4C969B76460FFFC142514143338AFDA4" box="[595,690,610,637]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="4" unit="mm" value="1.8">1.8 mm</quantity>
), discoidal to lenticular or almost subglobose; spire slightly to moderately raised; umbilical width narrow to moderate; whorl depth variable and whorls not conspicuously tightly coiled; periphery rounded to somewhat flat-sided, close to mid-whorl; last adult whorl sometimes descendant prior to aperture. Protoconch smooth or microscopically shagreened, with neither axial nor spiral sculpture, but sometimes with a slight subterminal swelling, comprising 0.67– 0.75 whorls beyond apical cap; teleoconch surface texture silky with a sculpture of simple, close-set, microscopic axial riblets of alternating strength, often with even finer spiral threads in their intervals. Aperture lunate; apertural dentition very variable.
</paragraph>
</subSubSection>
<subSubSection id="C3746518460FFFC140BF409233F8FA1B" pageId="3" pageNumber="4" type="discussion">
<paragraph id="8BD13693460FFFC140BF40923017FC14" blockId="3.[189,303,947,973]" box="[189,303,947,973]" pageId="3" pageNumber="4">
<heading id="D09981FF460FFFC140BF40923017FC14" bold="true" box="[189,303,947,973]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B91AEA81460FFFC140BF40923017FC14" bold="true" box="[189,303,947,973]" pageId="3" pageNumber="4">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BD13693460FFFC140BF40C5320AFB92" blockId="3.[189,1399,996,1099]" pageId="3" pageNumber="4">
<taxonomicName id="4C6E4D10460FFFC140BF40C5300BFC27" authorityName="Melvill &amp; Ponsonby" authorityYear="1908" box="[189,307,996,1022]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
<emphasis id="B91AEA81460FFFC140BF40C5300BFC27" box="[189,307,996,1022]" italics="true" pageId="3" pageNumber="4">Afrodonta</emphasis>
</taxonomicName>
comprises a relatively distinct group characterised by the almost smooth protoconch sculpture and silky teleoconch surface consisting of simple riblets of alternating strength. However, shell profile and apertural dentition exhibit considerable variation.
</paragraph>
<paragraph id="8BD13693460FFFC140BF475433F8FA1B" blockId="3.[189,1399,1141,1474]" pageId="3" pageNumber="4">
Some basic observations on the external anatomy and radula of the
<typeStatus id="54D58831460FFFC143B1475732DEFB49" box="[947,998,1142,1168]" pageId="3" pageNumber="4">type</typeStatus>
species and of
<taxonomicName id="4C6E4D10460FFFC1448F4754344FFB56" baseAuthorityName="Burnup" baseAuthorityYear="1912" box="[1165,1399,1141,1167]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="novemlamellaris">
<emphasis id="B91AEA81460FFFC1448F4754344FFB56" box="[1165,1399,1141,1167]" italics="true" pageId="3" pageNumber="4">Af.novemlamellaris</emphasis>
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were provided by Godwin-Austen (in
<bibRefCitation id="EFFF4B62460FFFC1426F47BD32AEFB6F" author="Melvill J. C. &amp; Ponsonby J. H." box="[621,918,1180,1206]" pageId="3" pageNumber="4" pagination="129 - 136" refId="ref26977" refString="Melvill J. C. &amp; Ponsonby J. H. 1908. Descriptions of nine terrestrial Mollusca from South Africa. Annals and Magazine of Natural History, Series 8, 1 (2): 129 - 136. https: // doi. org / 10.1080 / 00222930808692373" type="journal article" year="1908">
Melvill &amp; Ponsonby
<date id="FFD01053460FFFC1435847BD32AEFB6F" box="[858,918,1180,1206]" pageId="3" pageNumber="4" value="1908">1908</date>
</bibRefCitation>
) and
<bibRefCitation id="EFFF4B62460FFFC143D047BD35B2FB6F" author="Connolly M." box="[978,1162,1179,1206]" pageId="3" pageNumber="4" pagination="105 - 220" refId="ref26598" refString="Connolly M. 1925. The non-marine Mollusca of Portuguese East Africa. Transactions of the Royal Society of South Africa 12 (3): 105 - 220. https: // doi. org / 10.1080 / 00359192409519300" type="journal article" year="1925">
Connolly (
<date id="FFD01053460FFFC1444E47BA35B2FB6F" box="[1100,1162,1179,1206]" pageId="3" pageNumber="4" value="1925">1925</date>
</bibRefCitation>
,
<bibRefCitation id="EFFF4B62460FFFC1449147BD35E9FB6F" author="Connolly M." box="[1171,1233,1180,1206]" pageId="3" pageNumber="4" pagination="1 - 660" refId="ref26665" refString="Connolly M. 1939. A monographic survey of South African non-marine Mollusca. Annals of the South African Museum 33: 1 - 660." type="journal article" year="1939">
<date id="FFD01053460FFFC1449147BD35E9FB6F" box="[1171,1233,1180,1206]" pageId="3" pageNumber="4" value="1939">1939</date>
</bibRefCitation>
), respectively. A peripodial groove is present, but a caudal gland is not evident. The radula formula is (6–7)+ (8–9)+1+(8–9)+(6–7), with tricuspid central and lateral teeth, and somewhat pectinate marginals. The jaw is composed of narrow plates set side-by-side (stegognathous). In
<date id="FFD01053460FFFC143E4462F351BFAF0" box="[998,1059,1294,1321]" pageId="3" pageNumber="4" value="1925">1925</date>
, Connolly also described and illustrated the pallial anatomy of
<taxonomicName id="4C6E4D10460FFFC1423F46143210FA96" baseAuthorityName="Burnup" baseAuthorityYear="1912" box="[573,808,1333,1359]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="novemlamellaris">
<emphasis id="B91AEA81460FFFC1423F46143210FA96" box="[573,808,1333,1359]" italics="true" pageId="3" pageNumber="4">Af. novemlamellaris</emphasis>
</taxonomicName>
, noting that the ureter was reflected and included a secondary component running parallel to the rectum.
<bibRefCitation id="EFFF4B62460FFFC14325467A32FEFAAC" author="Solem A." box="[807,966,1371,1397]" pageId="3" pageNumber="4" pagination="342 - 364" refId="ref27176" refString="Solem A. 1970. The land snail genus Afrodonta (Mollusca: Gastropoda: Endodontidae). Annals of the Natal Museum 20: 342 - 364." type="journal article" year="1970">
Solem (
<date id="FFD01053460FFFC14383467A3286FAAC" box="[897,958,1371,1397]" pageId="3" pageNumber="4" value="1970">1970</date>
)
</bibRefCitation>
later commented that the presence of a secondary ureter and the configuration of the heart and kidney set
<taxonomicName id="4C6E4D10460FFFC143DA46A03576FA42" authorityName="Melvill &amp; Ponsonby" authorityYear="1908" box="[984,1102,1409,1435]" class="Gastropoda" family="Endodontidae" genus="Afrodonta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="genus">
<emphasis id="B91AEA81460FFFC143DA46A03576FA42" box="[984,1102,1409,1435]" italics="true" pageId="3" pageNumber="4">Afrodonta</emphasis>
</taxonomicName>
apart from Pacific Island endodontids, which lack a secondary ureter.
</paragraph>
</subSubSection>
</treatment>
</document>
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