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<document id="6EC8896E414BD74C0C706EE31D91B282" ID-CLB-Dataset="24671" ID-DOI="10.11646/zootaxa.4706.3.4" ID-GBIF-Dataset="47f9ebfa-4250-4218-9613-c1d1398ca33b" ID-ISSN="1175-5326" ID-Zenodo-Dep="3568896" ID-ZooBank="29429537-8240-4E24-B926-6F23A7BD151E" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1575968595120" checkinUser="plazi" docAuthor="Farahi, Sara, Shishehbor, Parviz, Nemati, Alireza &amp; Witaliński, Wojciech" docDate="2019" docId="03C71F47FFD8FF9EDABCC0F8FD4DFEF3" docLanguage="en" docName="zootaxa.4706.3.4.pdf" docOrigin="Zootaxa 4706 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trachygamasus karuni Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński 2019, sp. nov." docType="treatment" docVersion="6" lastPageNumber="448" masterDocId="FFFE673FFFD9FF97DA2BC332FFD6FFB0" masterDocTitle="Trachygamasus karuni sp. nov., a new mite species from Iran (Parasitiformes: Parasitidae)" masterLastPageNumber="450" masterPageNumber="439" pageNumber="440" updateTime="1698773684450" updateUser="ExternalLinkService">
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<mods:title id="F3BC123CD1E8144053EE41EB4CBBEC8F">Trachygamasus karuni sp. nov., a new mite species from Iran (Parasitiformes: Parasitidae)</mods:title>
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<mods:namePart id="340A78137E899A7EFE45FDF86616DA45">Farahi, Sara</mods:namePart>
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<mods:namePart id="863FA4226A9CCD816901ECA436C83DEA">Shishehbor, Parviz</mods:namePart>
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<mods:namePart id="9B50158D26B86AA489C7C7D6BB9A17ED">Nemati, Alireza</mods:namePart>
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<mods:namePart id="001EC5B410D9E65FDF24F2FF9713D3E5">Witaliński, Wojciech</mods:namePart>
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<mods:title id="CB8BA53AC1036CA2A641774A5EC8AD16">Zootaxa</mods:title>
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<mods:date id="2627CD676DD507DC58A83046A4B89D4E">2019</mods:date>
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<mods:number id="61990A1D12F1BA4275A800D355A07BF2">2019-12-10</mods:number>
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<mods:number id="B002D72C6EDBEBF6B61BA73D4A865830">4706</mods:number>
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<treatment id="03C71F47FFD8FF9EDABCC0F8FD4DFEF3" ID-GBIF-Taxon="160687974" LSID="urn:lsid:plazi:treatment:03C71F47FFD8FF9EDABCC0F8FD4DFEF3" httpUri="http://treatment.plazi.org/id/03C71F47FFD8FF9EDABCC0F8FD4DFEF3" lastPageId="9" lastPageNumber="448" pageId="1" pageNumber="440">
<subSubSection id="C374FDDAFFD8FF96DABCC0F8FCCFFC55" box="[151,793,969,997]" pageId="1" pageNumber="440" type="nomenclature">
<paragraph id="8BD1AE51FFD8FF96DABCC0F8FCCFFC55" blockId="1.[151,962,969,1068]" box="[151,793,969,997]" pageId="1" pageNumber="440">
<heading id="D099193DFFD8FF96DABCC0F8FCCFFC55" bold="true" box="[151,793,969,997]" fontSize="11" level="1" pageId="1" pageNumber="440" reason="1">
<emphasis id="B91A7243FFD8FF96DABCC0F8FCCFFC55" bold="true" box="[151,793,969,997]" pageId="1" pageNumber="440">
<taxonomicName id="4C6ED5D2FFD8FF96DABCC0F8FD62FC54" authority="Farahi &amp; Witalinski" authorityName="Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński" authorityYear="2019" box="[151,692,969,997]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="440" phylum="Arthropoda" rank="species" species="karuni" status="sp. nov.">
<emphasis id="B91A7243FFD8FF96DABCC0F8FE66FC55" bold="true" box="[151,432,970,997]" italics="true" pageId="1" pageNumber="440">Trachygamasus karuni</emphasis>
Farahi &amp; Witaliński
</taxonomicName>
<taxonomicNameLabel id="A229CF38FFD8FF96D890C0F9FCCFFC55" box="[699,793,971,997]" pageId="1" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C374FDDAFFD8FF96DABCC0DCFC14FB9C" pageId="1" pageNumber="440" type="description">
<paragraph id="8BD1AE51FFD8FF96DABCC0DCFEEAFBB8" blockId="1.[151,962,969,1068]" box="[151,316,1006,1032]" pageId="1" pageNumber="440">
(
<figureCitation id="1355B2D4FFD8FF96DAB4C0DCFEE2FBB8" box="[159,308,1006,1032]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURES 35" captionStart-3="FIGURES 613" captionStart-4="FIGURE 14" captionStart-5="FIGURES 1519" captionStart-6="FIGURES 2024" captionStartId-0="3.[151,250,1781,1806]" captionStartId-1="4.[151,250,1822,1847]" captionStartId-2="5.[151,264,577,602]" captionStartId-3="5.[151,264,1898,1923]" captionStartId-4="6.[151,250,1783,1808]" captionStartId-5="7.[151,264,1924,1949]" captionStartId-6="8.[151,264,1924,1949]" captionTargetBox-0="[251,1340,262,1686]" captionTargetBox-1="[248,1346,216,1731]" captionTargetBox-2="[155,1430,169,554]" captionTargetBox-3="[189,1398,721,1869]" captionTargetBox-4="[293,1195,224,1660]" captionTargetBox-5="[164,1387,178,1897]" captionTargetBox-6="[166,1364,181,1889]" captionTargetId-0="figure@3.[219,1353,237,1705]" captionTargetId-1="figure@4.[235,1358,206,1744]" captionTargetId-2="figure@5.[151,1436,166,558]" captionTargetId-3="figure@5.[183,1405,717,1879]" captionTargetId-4="figure@6.[250,1205,223,1683]" captionTargetId-5="figure@7.[151,1399,166,1904]" captionTargetId-6="figure@8.[151,1376,166,1904]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="8" captionText-0="FIGURE 1. Trachygamasus karuni sp. nov., female. A. Dorsal side of idiosoma, paratype. Dorsal setae and setae series (left-hand side), as well as the dorsal poroids and gland pores (right-hand side) are marked and specifically acknowledged. Pseudoperforated cuticle is marked in the right lower part. B. Seta r3." captionText-1="FIGURE 2. Trachygamasus karuni sp. nov., female. Ventral side of idiosoma, paratype." captionText-2="FIGURES 35. Trachygamasus karuni sp. nov. 3. Anterior part of podonotum in female with anterior ends of peritremes, podonotal setae, poroids idj1 and gland pores gdj2, the last-named with double openings. 4 &amp; 5. Ventral view of gnathosoma in female (Figure 4) and male (Figure 5). Holotype: Figure 4; paratypes: Figures 3 and 5." captionText-3="FIGURES 613. Trachygamasus karuni sp. nov., female. 6. Presternal plates. 7. Genital region showing epigynium and an outline of the endogynium in situ. Setae and pores as in Figure 2. 8. Endogynium, two aspects. 9. Gnathotectum. 10. Chelicera antiaxially. 11. Coxa I ventrally. 12. Ambulacra on leg II (Figure 12 left) and leg III (Figure 12 right). 13. Tibia II, posterolateral view. Holotype: Figures 7 and 8 right; paratypes: Figures 6, 8 left, 913." captionText-4="FIGURE 14. Trachygamasus karuni sp. nov., male. Dorsal side of idiosoma, abbreviations as in Figure 1." captionText-5="FIGURES 1519. Trachygamasus karuni sp. nov., male. 15. Ventral side of idiosoma, abbreviations as in Figure 2. 16. Anterior margin of sternum with genital lamina flanked by presternal plates. 17. Hypostome in ventral perspective showing corniculi, hypostomatic setae, palpcoxa seta and palptrochanter. 18. Gnathotectum, two aspects. 19. Chelicera adaxially." captionText-6="FIGURES 2024. Trachygamasus karuni sp. nov., male, leg II. 20. Femur, genu and tibia in ventral perspective. 21. Femur, genu and tibia posterolaterally. 22. Spur on genu anterolaterally. 23 &amp; 24. Spurs on femur II and genu II, enlarged. Setae anterolateral (al), posteroventral (pv), posterolateral (pl), and posterodorsal (pd) are marked." figureDoi-0="http://doi.org/10.5281/zenodo.3568898" figureDoi-1="http://doi.org/10.5281/zenodo.3568900" figureDoi-2="http://doi.org/10.5281/zenodo.3568902" figureDoi-3="http://doi.org/10.5281/zenodo.3568904" figureDoi-4="http://doi.org/10.5281/zenodo.3568906" figureDoi-5="http://doi.org/10.5281/zenodo.3568908" figureDoi-6="http://doi.org/10.5281/zenodo.3568910" httpUri-0="https://zenodo.org/record/3568898/files/figure.png" httpUri-1="https://zenodo.org/record/3568900/files/figure.png" httpUri-2="https://zenodo.org/record/3568902/files/figure.png" httpUri-3="https://zenodo.org/record/3568904/files/figure.png" httpUri-4="https://zenodo.org/record/3568906/files/figure.png" httpUri-5="https://zenodo.org/record/3568908/files/figure.png" httpUri-6="https://zenodo.org/record/3568910/files/figure.png" pageId="1" pageNumber="440">Figures 124</figureCitation>
)
</paragraph>
<paragraph id="8BD1AE51FFD8FF96DABCC720FC14FB9C" blockId="1.[151,962,969,1068]" box="[151,962,1042,1068]" pageId="1" pageNumber="440">
<uri id="FFFFA253FFD8FF96DABCC720FC14FB9C" box="[151,962,1042,1068]" pageId="1" pageNumber="440">
urn:lsid:zoobank.org:act:
<uuid id="FFC89484FFD8FF96DB84C720FC14FB9C" box="[431,962,1042,1068]" pageId="1" pageNumber="440">396D61D8-4620-4E78-9701-CAA94B4614FD</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C374FDDAFFD8FF96DABCC76BFD7AF9B0" pageId="1" pageNumber="440" type="diagnosis">
<paragraph id="8BD1AE51FFD8FF96DABCC76BFCB5FAFC" blockId="1.[151,1437,1113,2041]" pageId="1" pageNumber="440">
<emphasis id="B91A7243FFD8FF96DABCC76BFE23FBC4" bold="true" box="[151,501,1113,1140]" pageId="1" pageNumber="440">Diagnosis (female and male).</emphasis>
Cuticle well sclerotised with many regularly distributed pits; podonotum with 23 pairs of setae (all four sets of normal setae present except setae
<emphasis id="B91A7243FFD8FF96D95EC74CFC58FB28" box="[885,910,1150,1176]" italics="true" pageId="1" pageNumber="440">r1</emphasis>
), setae
<emphasis id="B91A7243FFD8FF96D9C3C74CFC28FB28" box="[1000,1022,1150,1176]" italics="true" pageId="1" pageNumber="440">j4</emphasis>
,
<emphasis id="B91A7243FFD8FF96DE25C74DFBF1FB29" box="[1038,1063,1151,1177]" italics="true" pageId="1" pageNumber="440">z5</emphasis>
and
<emphasis id="B91A7243FFD8FF96DE4AC74CFBACFB28" box="[1121,1146,1150,1176]" italics="true" pageId="1" pageNumber="440">r3</emphasis>
thickened and terminally pilose, seta
<emphasis id="B91A7243FFD8FF96DB30C790FEE7FB0C" box="[283,305,1186,1212]" italics="true" pageId="1" pageNumber="440">j1</emphasis>
only moderately; opisthonotum with 24 setae and 23 lateral setae unidentifiable on each side, setae
<emphasis id="B91A7243FFD8FF96DABCC7F4FF63FB50" box="[151,181,1222,1248]" italics="true" pageId="1" pageNumber="440">Z1</emphasis>
,
<emphasis id="B91A7243FFD8FF96DAE9C7F4FF36FB50" box="[194,224,1222,1248]" italics="true" pageId="1" pageNumber="440">Z4</emphasis>
thicker and apically pilose; opisthogastric setae
<emphasis id="B91A7243FFD8FF96D8D0C7F4FCF1FB50" box="[763,807,1222,1248]" italics="true" pageId="1" pageNumber="440">JV4</emphasis>
,
<emphasis id="B91A7243FFD8FF96D91FC7F5FCB6FB51" box="[820,864,1223,1249]" italics="true" pageId="1" pageNumber="440">JV5</emphasis>
and
<emphasis id="B91A7243FFD8FF96D9BEC7F4FC12FB50" box="[917,964,1222,1248]" italics="true" pageId="1" pageNumber="440">ZV4</emphasis>
enlarged and terminally pilose, in females setae
<emphasis id="B91A7243FFD8FF96DAFCC7D9FED5FAB5" box="[215,259,1259,1285]" italics="true" pageId="1" pageNumber="440">JV5</emphasis>
and
<emphasis id="B91A7243FFD8FF96DB11C7D8FEBFFAB4" box="[314,361,1258,1284]" italics="true" pageId="1" pageNumber="440">ZV4</emphasis>
located on soft cuticle; gnathotectum trispinate with central prong broader and more solid than the lateral ones which are thin and acute, sometimes curved;
<emphasis id="B91A7243FFD8FF96D971C63CFC54FA98" box="[858,898,1294,1320]" italics="true" pageId="1" pageNumber="440">gv1</emphasis>
gland pores not discernible; sternal poroid
<emphasis id="B91A7243FFD8FF96DF51C63CFA4AFA98" box="[1402,1436,1294,1320]" italics="true" pageId="1" pageNumber="440">
<date id="FFD08891FFD8FF96DF51C63CFA4AFA98" box="[1402,1436,1294,1320]" pageId="1" pageNumber="440">iv3</date>
</emphasis>
circular, pore
<emphasis id="B91A7243FFD8FF96DB19C601FE82FAFD" box="[306,340,1331,1357]" italics="true" pageId="1" pageNumber="440">
<date id="FFD08891FFD8FF96DB19C601FE82FAFD" box="[306,340,1331,1357]" pageId="1" pageNumber="440">iv5</date>
</emphasis>
followed laterally by three-opening
<emphasis id="B91A7243FFD8FF96D8C5C600FCC1FAFC" box="[750,791,1330,1356]" italics="true" pageId="1" pageNumber="440">gv2</emphasis>
gland.
</paragraph>
<paragraph id="8BD1AE51FFD8FF96DAECC664FD0FFA24" blockId="1.[151,1437,1113,2041]" pageId="1" pageNumber="440">Female: epigynium bell-shaped with convex lateral margins and concave posterior margin; endogynium hyaline and poorly visible as axially elongated saccule.</paragraph>
<paragraph id="8BD1AE51FFD8FF96DAECC6ACFD7AF9B0" blockId="1.[151,1437,1113,2041]" pageId="1" pageNumber="440">
Male: presternal platelets large; genital lamina with bar-shaped anterior thickening and a hyaline protrusion directed anteriad; sternal setae
<emphasis id="B91A7243FFD8FF96DBDEC6F0FDC0FA6C" box="[501,534,1474,1500]" italics="true" pageId="1" pageNumber="440">st1</emphasis>
far from the anterior sternal margin; leg II with minute spurs: one on the femur and one on the genu, whereas none on the tibia.
</paragraph>
</subSubSection>
<subSubSection id="C374FDDAFFD8FF95DAECC53BFBA0F8B7" lastPageId="2" lastPageNumber="441" pageId="1" pageNumber="440" type="description">
<paragraph id="8BD1AE51FFD8FF96DAECC53BFD4EF995" blockId="1.[151,1437,1113,2041]" box="[199,664,1545,1573]" pageId="1" pageNumber="440">
<emphasis id="B91A7243FFD8FF96DAECC53BFE8FF994" bold="true" box="[199,345,1545,1572]" pageId="1" pageNumber="440">Description.</emphasis>
<emphasis id="B91A7243FFD8FF96DB4BC538FE62F994" box="[352,436,1546,1572]" italics="true" pageId="1" pageNumber="440">Female</emphasis>
(
<figureCitation id="1355B2D4FFD8FF96DBE9C538FD9DF994" box="[450,587,1546,1572]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURES 35" captionStartId-0="3.[151,250,1781,1806]" captionStartId-1="4.[151,250,1822,1847]" captionStartId-2="5.[151,264,577,602]" captionTargetBox-0="[251,1340,262,1686]" captionTargetBox-1="[248,1346,216,1731]" captionTargetBox-2="[155,1430,169,554]" captionTargetId-0="figure@3.[219,1353,237,1705]" captionTargetId-1="figure@4.[235,1358,206,1744]" captionTargetId-2="figure@5.[151,1436,166,558]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 1. Trachygamasus karuni sp. nov., female. A. Dorsal side of idiosoma, paratype. Dorsal setae and setae series (left-hand side), as well as the dorsal poroids and gland pores (right-hand side) are marked and specifically acknowledged. Pseudoperforated cuticle is marked in the right lower part. B. Seta r3." captionText-1="FIGURE 2. Trachygamasus karuni sp. nov., female. Ventral side of idiosoma, paratype." captionText-2="FIGURES 35. Trachygamasus karuni sp. nov. 3. Anterior part of podonotum in female with anterior ends of peritremes, podonotal setae, poroids idj1 and gland pores gdj2, the last-named with double openings. 4 &amp; 5. Ventral view of gnathosoma in female (Figure 4) and male (Figure 5). Holotype: Figure 4; paratypes: Figures 3 and 5." figureDoi-0="http://doi.org/10.5281/zenodo.3568898" figureDoi-1="http://doi.org/10.5281/zenodo.3568900" figureDoi-2="http://doi.org/10.5281/zenodo.3568902" httpUri-0="https://zenodo.org/record/3568898/files/figure.png" httpUri-1="https://zenodo.org/record/3568900/files/figure.png" httpUri-2="https://zenodo.org/record/3568902/files/figure.png" pageId="1" pageNumber="440">Figures 14</figureCitation>
,
<figureCitation id="1355B2D4FFD8FF96D87CC538FD46F995" box="[599,656,1546,1573]" captionStart="FIGURES 613" captionStartId="5.[151,264,1898,1923]" captionTargetBox="[189,1398,721,1869]" captionTargetId="figure@5.[183,1405,717,1879]" captionTargetPageId="5" captionText="FIGURES 613. Trachygamasus karuni sp. nov., female. 6. Presternal plates. 7. Genital region showing epigynium and an outline of the endogynium in situ. Setae and pores as in Figure 2. 8. Endogynium, two aspects. 9. Gnathotectum. 10. Chelicera antiaxially. 11. Coxa I ventrally. 12. Ambulacra on leg II (Figure 12 left) and leg III (Figure 12 right). 13. Tibia II, posterolateral view. Holotype: Figures 7 and 8 right; paratypes: Figures 6, 8 left, 913." figureDoi="http://doi.org/10.5281/zenodo.3568904" httpUri="https://zenodo.org/record/3568904/files/figure.png" pageId="1" pageNumber="440">613</figureCitation>
)
</paragraph>
<paragraph id="8BD1AE51FFD8FF96DAECC51CFA4CF94C" blockId="1.[151,1437,1113,2041]" pageId="1" pageNumber="440">
<emphasis id="B91A7243FFD8FF96DAECC51CFE51F9F8" box="[199,391,1582,1608]" italics="true" pageId="1" pageNumber="440">Dorsal idiosoma</emphasis>
(
<figureCitation id="1355B2D4FFD8FF96DBB2C51CFDC5F9F8" box="[409,531,1582,1608]" captionStart="FIGURE 1" captionStartId="3.[151,250,1781,1806]" captionTargetBox="[251,1340,262,1686]" captionTargetId="figure@3.[219,1353,237,1705]" captionTargetPageId="3" captionText="FIGURE 1. Trachygamasus karuni sp. nov., female. A. Dorsal side of idiosoma, paratype. Dorsal setae and setae series (left-hand side), as well as the dorsal poroids and gland pores (right-hand side) are marked and specifically acknowledged. Pseudoperforated cuticle is marked in the right lower part. B. Seta r3." figureDoi="http://doi.org/10.5281/zenodo.3568898" httpUri="https://zenodo.org/record/3568898/files/figure.png" pageId="1" pageNumber="440">Figure 1A</figureCitation>
). Idiosoma well sclerotised, brownish and pyriform in outline, 485560 long, 325450 wide (n=6),
<typeStatus id="54D510F3FFD8FF96DBA7C560FE38F9DC" box="[396,494,1618,1644]" pageId="1" pageNumber="440" type="holotype">holotype</typeStatus>
560 long, 405 wide. Podonotum and opisthonotum separated by lines in cuticle behind podonotal setae. Cuticle with many regularly distributed pits. Dorsal setae simple, except
<emphasis id="B91A7243FFD8FF96DE90C544FB06F920" box="[1211,1232,1654,1680]" italics="true" pageId="1" pageNumber="440">j1</emphasis>
,
<emphasis id="B91A7243FFD8FF96DEF5C544FB22F920" box="[1246,1268,1654,1680]" italics="true" pageId="1" pageNumber="440">j4</emphasis>
,
<emphasis id="B91A7243FFD8FF96DF29C544FACDF920" box="[1282,1307,1654,1680]" italics="true" pageId="1" pageNumber="440">r3</emphasis>
,
<emphasis id="B91A7243FFD8FF96DF02C545FA94F921" box="[1321,1346,1655,1681]" italics="true" pageId="1" pageNumber="440">z5</emphasis>
,
<emphasis id="B91A7243FFD8FF96DF64C544FABBF920" box="[1359,1389,1654,1680]" italics="true" pageId="1" pageNumber="440">Z1</emphasis>
and
<emphasis id="B91A7243FFD8FF96DABCC5A8FF63F904" box="[151,181,1690,1716]" italics="true" pageId="1" pageNumber="440">Z4</emphasis>
larger and terminally pilose (
<figureCitation id="1355B2D4FFD8FF96D82FC5A8FDACF904" box="[516,634,1690,1716]" captionStart="FIGURE 1" captionStartId="3.[151,250,1781,1806]" captionTargetBox="[251,1340,262,1686]" captionTargetId="figure@3.[219,1353,237,1705]" captionTargetPageId="3" captionText="FIGURE 1. Trachygamasus karuni sp. nov., female. A. Dorsal side of idiosoma, paratype. Dorsal setae and setae series (left-hand side), as well as the dorsal poroids and gland pores (right-hand side) are marked and specifically acknowledged. Pseudoperforated cuticle is marked in the right lower part. B. Seta r3." figureDoi="http://doi.org/10.5281/zenodo.3568898" httpUri="https://zenodo.org/record/3568898/files/figure.png" pageId="1" pageNumber="440">Figure 1B</figureCitation>
), seta
<emphasis id="B91A7243FFD8FF96D8EFC5A8FD0CF904" box="[708,730,1690,1716]" italics="true" pageId="1" pageNumber="440">j1</emphasis>
only moderately. Length of simple dorsal setae: 1420, larger setae length: 2227 (
<emphasis id="B91A7243FFD8FF96DBABC58CFE43F968" box="[384,405,1726,1752]" italics="true" pageId="1" pageNumber="440">j1</emphasis>
), 2933 (
<emphasis id="B91A7243FFD8FF96D829C58CFDCEF968" box="[514,536,1726,1752]" italics="true" pageId="1" pageNumber="440">j4</emphasis>
), 4247 (
<emphasis id="B91A7243FFD8FF96D8AFC58CFD4BF968" box="[644,669,1726,1752]" italics="true" pageId="1" pageNumber="440">r3</emphasis>
), 3033 (
<emphasis id="B91A7243FFD8FF96D922C58DFCF5F969" box="[777,803,1727,1753]" italics="true" pageId="1" pageNumber="440">z5</emphasis>
), 3134 (
<emphasis id="B91A7243FFD8FF96D9A4C58CFC7AF968" box="[911,940,1726,1752]" italics="true" pageId="1" pageNumber="440">Z1</emphasis>
), 2937 (
<emphasis id="B91A7243FFD8FF96DE32C58CFBE1F968" box="[1049,1079,1726,1752]" italics="true" pageId="1" pageNumber="440">Z4</emphasis>
). Length of peritrematal groove 258275 (
<typeStatus id="54D510F3FFD8FF96DB23C5D0FEBAF94C" box="[264,364,1762,1788]" pageId="1" pageNumber="440" type="holotype">holotype</typeStatus>
275); expanded behind coxa I, ending near podonotal seta
<emphasis id="B91A7243FFD8FF96DE2AC5D0FBCCF94C" box="[1025,1050,1762,1788]" italics="true" pageId="1" pageNumber="440">z1</emphasis>
and poroid
<emphasis id="B91A7243FFD8FF96DE8AC5D0FB1AF94C" box="[1185,1228,1762,1788]" italics="true" pageId="1" pageNumber="440">idj1</emphasis>
(
<figureCitation id="1355B2D4FFD8FF96DEF0C5D0FA93F94C" box="[1243,1349,1762,1788]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1847]" captionTargetBox="[248,1346,216,1731]" captionTargetId="figure@4.[235,1358,206,1744]" captionTargetPageId="4" captionText="FIGURE 2. Trachygamasus karuni sp. nov., female. Ventral side of idiosoma, paratype." figureDoi="http://doi.org/10.5281/zenodo.3568900" httpUri="https://zenodo.org/record/3568900/files/figure.png" pageId="1" pageNumber="440">Figures 2</figureCitation>
and
<figureCitation id="1355B2D4FFD8FF96DF57C5D0FA5AF94C" box="[1404,1420,1762,1788]" captionStart="FIGURES 35" captionStartId="5.[151,264,577,602]" captionTargetBox="[155,1430,169,554]" captionTargetId="figure@5.[151,1436,166,558]" captionTargetPageId="5" captionText="FIGURES 35. Trachygamasus karuni sp. nov. 3. Anterior part of podonotum in female with anterior ends of peritremes, podonotal setae, poroids idj1 and gland pores gdj2, the last-named with double openings. 4 &amp; 5. Ventral view of gnathosoma in female (Figure 4) and male (Figure 5). Holotype: Figure 4; paratypes: Figures 3 and 5." figureDoi="http://doi.org/10.5281/zenodo.3568902" httpUri="https://zenodo.org/record/3568902/files/figure.png" pageId="1" pageNumber="440">3</figureCitation>
).
</paragraph>
<paragraph id="8BD1AE51FFD8FF96DAECC434FE05F83D" blockId="1.[151,1437,1113,2041]" pageId="1" pageNumber="440">
<emphasis id="B91A7243FFD8FF96DAECC434FE53F890" box="[199,389,1798,1824]" italics="true" pageId="1" pageNumber="440">Ventral idiosoma</emphasis>
(
<figureCitation id="1355B2D4FFD8FF96DBB8C434FE23F890" box="[403,501,1798,1824]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1847]" captionTargetBox="[248,1346,216,1731]" captionTargetId="figure@4.[235,1358,206,1744]" captionTargetPageId="4" captionText="FIGURE 2. Trachygamasus karuni sp. nov., female. Ventral side of idiosoma, paratype." figureDoi="http://doi.org/10.5281/zenodo.3568900" httpUri="https://zenodo.org/record/3568900/files/figure.png" pageId="1" pageNumber="440">Figure 2</figureCitation>
). Presternal plates (
<figureCitation id="1355B2D4FFD8FF96D8E5C434FCEEF890" box="[718,824,1798,1824]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1847]" captionTargetBox="[248,1346,216,1731]" captionTargetId="figure@4.[235,1358,206,1744]" captionTargetPageId="4" captionText="FIGURE 2. Trachygamasus karuni sp. nov., female. Ventral side of idiosoma, paratype." figureDoi="http://doi.org/10.5281/zenodo.3568900" httpUri="https://zenodo.org/record/3568900/files/figure.png" pageId="1" pageNumber="440">Figures 2</figureCitation>
and
<figureCitation id="1355B2D4FFD8FF96D946C434FCAAF891" box="[877,892,1798,1825]" captionStart="FIGURES 613" captionStartId="5.[151,264,1898,1923]" captionTargetBox="[189,1398,721,1869]" captionTargetId="figure@5.[183,1405,717,1879]" captionTargetPageId="5" captionText="FIGURES 613. Trachygamasus karuni sp. nov., female. 6. Presternal plates. 7. Genital region showing epigynium and an outline of the endogynium in situ. Setae and pores as in Figure 2. 8. Endogynium, two aspects. 9. Gnathotectum. 10. Chelicera antiaxially. 11. Coxa I ventrally. 12. Ambulacra on leg II (Figure 12 left) and leg III (Figure 12 right). 13. Tibia II, posterolateral view. Holotype: Figures 7 and 8 right; paratypes: Figures 6, 8 left, 913." figureDoi="http://doi.org/10.5281/zenodo.3568904" httpUri="https://zenodo.org/record/3568904/files/figure.png" pageId="1" pageNumber="440">6</figureCitation>
) distinct, with anterior and lateral margins thickened. Tritosternum present, biramous. Sternal shield with moderately visible reticulation in the anterior part and two pairs of circular depressions posteriorly to the sternal poroids
<emphasis id="B91A7243FFD8FF96D967C47CFCB9F8D8" box="[844,879,1870,1896]" italics="true" pageId="1" pageNumber="440">
<date id="FFD08891FFD8FF96D967C47CFCB9F8D8" box="[844,879,1870,1896]" pageId="1" pageNumber="440">iv2</date>
</emphasis>
, and behind
<emphasis id="B91A7243FFD8FF96D9D4C47CFBF6F8D8" box="[1023,1056,1870,1896]" italics="true" pageId="1" pageNumber="440">st4</emphasis>
setae (
<figureCitation id="1355B2D4FFD8FF96DE46C47CFB0EF8D8" box="[1133,1240,1870,1896]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1847]" captionTargetBox="[248,1346,216,1731]" captionTargetId="figure@4.[235,1358,206,1744]" captionTargetPageId="4" captionText="FIGURE 2. Trachygamasus karuni sp. nov., female. Ventral side of idiosoma, paratype." figureDoi="http://doi.org/10.5281/zenodo.3568900" httpUri="https://zenodo.org/record/3568900/files/figure.png" pageId="1" pageNumber="440">Figures 2</figureCitation>
and
<figureCitation id="1355B2D4FFD8FF96DF24C47DFAC9F8D9" box="[1295,1311,1871,1897]" captionStart="FIGURES 613" captionStartId="5.[151,264,1898,1923]" captionTargetBox="[189,1398,721,1869]" captionTargetId="figure@5.[183,1405,717,1879]" captionTargetPageId="5" captionText="FIGURES 613. Trachygamasus karuni sp. nov., female. 6. Presternal plates. 7. Genital region showing epigynium and an outline of the endogynium in situ. Setae and pores as in Figure 2. 8. Endogynium, two aspects. 9. Gnathotectum. 10. Chelicera antiaxially. 11. Coxa I ventrally. 12. Ambulacra on leg II (Figure 12 left) and leg III (Figure 12 right). 13. Tibia II, posterolateral view. Holotype: Figures 7 and 8 right; paratypes: Figures 6, 8 left, 913." figureDoi="http://doi.org/10.5281/zenodo.3568904" httpUri="https://zenodo.org/record/3568904/files/figure.png" pageId="1" pageNumber="440">7</figureCitation>
). Idiosoma margin behind anus convex.
</paragraph>
<paragraph id="8BD1AE51FFD8FF95DAECC4A4FC28FF67" blockId="1.[151,1437,1113,2041]" lastBlockId="2.[151,1437,153,2015]" lastPageId="2" lastPageNumber="441" pageId="1" pageNumber="440">
<emphasis id="B91A7243FFD8FF96DAECC4A4FEBBF800" box="[199,365,1942,1968]" italics="true" pageId="1" pageNumber="440">Genital region</emphasis>
(
<figureCitation id="1355B2D4FFD8FF96DB55C4A4FE3DF800" box="[382,491,1942,1968]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1847]" captionTargetBox="[248,1346,216,1731]" captionTargetId="figure@4.[235,1358,206,1744]" captionTargetPageId="4" captionText="FIGURE 2. Trachygamasus karuni sp. nov., female. Ventral side of idiosoma, paratype." figureDoi="http://doi.org/10.5281/zenodo.3568900" httpUri="https://zenodo.org/record/3568900/files/figure.png" pageId="1" pageNumber="440">Figures 2</figureCitation>
and
<figureCitation id="1355B2D4FFD8FF96D80CC4A5FDE1F801" box="[551,567,1943,1969]" captionStart="FIGURES 613" captionStartId="5.[151,264,1898,1923]" captionTargetBox="[189,1398,721,1869]" captionTargetId="figure@5.[183,1405,717,1879]" captionTargetPageId="5" captionText="FIGURES 613. Trachygamasus karuni sp. nov., female. 6. Presternal plates. 7. Genital region showing epigynium and an outline of the endogynium in situ. Setae and pores as in Figure 2. 8. Endogynium, two aspects. 9. Gnathotectum. 10. Chelicera antiaxially. 11. Coxa I ventrally. 12. Ambulacra on leg II (Figure 12 left) and leg III (Figure 12 right). 13. Tibia II, posterolateral view. Holotype: Figures 7 and 8 right; paratypes: Figures 6, 8 left, 913." figureDoi="http://doi.org/10.5281/zenodo.3568904" httpUri="https://zenodo.org/record/3568904/files/figure.png" pageId="1" pageNumber="440">7</figureCitation>
). Paragynia fused with the sternal shield. Poroids
<emphasis id="B91A7243FFD8FF96DE56C4A4FB76F800" box="[1149,1184,1942,1968]" italics="true" pageId="1" pageNumber="440">
<date id="FFD08891FFD8FF96DE56C4A4FB76F800" box="[1149,1184,1942,1968]" pageId="1" pageNumber="440">iv3</date>
</emphasis>
roundish,
<emphasis id="B91A7243FFD8FF96DF37C4A4FA92F800" box="[1308,1348,1942,1968]" italics="true" pageId="1" pageNumber="440">gv1</emphasis>
absent. Epigynal shield bell-shaped, with convex lateral margins and concave posterior margin; three pairs of thickenings, located in the central part of epigynium, are discernible (
<figureCitation id="1355B2D4FFD8FF96D92EC4ECFCB0F849" box="[773,870,2014,2041]" captionStart="FIGURES 613" captionStartId="5.[151,264,1898,1923]" captionTargetBox="[189,1398,721,1869]" captionTargetId="figure@5.[183,1405,717,1879]" captionTargetPageId="5" captionText="FIGURES 613. Trachygamasus karuni sp. nov., female. 6. Presternal plates. 7. Genital region showing epigynium and an outline of the endogynium in situ. Setae and pores as in Figure 2. 8. Endogynium, two aspects. 9. Gnathotectum. 10. Chelicera antiaxially. 11. Coxa I ventrally. 12. Ambulacra on leg II (Figure 12 left) and leg III (Figure 12 right). 13. Tibia II, posterolateral view. Holotype: Figures 7 and 8 right; paratypes: Figures 6, 8 left, 913." figureDoi="http://doi.org/10.5281/zenodo.3568904" httpUri="https://zenodo.org/record/3568904/files/figure.png" pageId="1" pageNumber="440">Figure 7</figureCitation>
). Endogynium (
<figureCitation id="1355B2D4FFD8FF96DE33C4ECFB55F848" box="[1048,1155,2014,2040]" captionStart="FIGURE 2" captionStartId="4.[151,250,1822,1847]" captionTargetBox="[248,1346,216,1731]" captionTargetId="figure@4.[235,1358,206,1744]" captionTargetPageId="4" captionText="FIGURE 2. Trachygamasus karuni sp. nov., female. Ventral side of idiosoma, paratype." figureDoi="http://doi.org/10.5281/zenodo.3568900" httpUri="https://zenodo.org/record/3568900/files/figure.png" pageId="1" pageNumber="440">Figures 2</figureCitation>
,
<figureCitation id="1355B2D4FFD8FF96DEA5C4EDFB0BF848" box="[1166,1245,2014,2041]" captionStart="FIGURES 613" captionStartId="5.[151,264,1898,1923]" captionTargetBox="[189,1398,721,1869]" captionTargetId="figure@5.[183,1405,717,1879]" captionTargetPageId="5" captionText="FIGURES 613. Trachygamasus karuni sp. nov., female. 6. Presternal plates. 7. Genital region showing epigynium and an outline of the endogynium in situ. Setae and pores as in Figure 2. 8. Endogynium, two aspects. 9. Gnathotectum. 10. Chelicera antiaxially. 11. Coxa I ventrally. 12. Ambulacra on leg II (Figure 12 left) and leg III (Figure 12 right). 13. Tibia II, posterolateral view. Holotype: Figures 7 and 8 right; paratypes: Figures 6, 8 left, 913." figureDoi="http://doi.org/10.5281/zenodo.3568904" httpUri="https://zenodo.org/record/3568904/files/figure.png" pageId="1" pageNumber="440">7 and 8</figureCitation>
) hyaline, weakly visible as an axially oriented saccular structure, with thin arcuate lines located laterally. Poroids
<emphasis id="B91A7243FFDBFF95DEEAC3ABFB35FF03" box="[1217,1251,153,179]" italics="true" pageId="2" pageNumber="441">
<date id="FFD08891FFDBFF95DEEAC3ABFB35FF03" box="[1217,1251,153,179]" pageId="2" pageNumber="441">iv5</date>
</emphasis>
situated at some distance behind the posterior epigynal margin,
<emphasis id="B91A7243FFDBFF95D88EC38FFD1BFF67" box="[677,717,189,215]" italics="true" pageId="2" pageNumber="441">gv2</emphasis>
gland with three openings.
</paragraph>
<paragraph id="8BD1AE51FFDBFF95DAECC3D3FAAFFE3B" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="441">
<emphasis id="B91A7243FFDBFF95DAECC3D3FDD2FF4B" box="[199,516,225,251]" italics="true" pageId="2" pageNumber="441">Setation of ventral idiosoma</emphasis>
. Setae of sternogenital region —
<typeStatus id="54D510F3FFDBFF95D955C3D3FC32FF4B" box="[894,996,225,251]" pageId="2" pageNumber="441" type="holotype">holotype</typeStatus>
: 20 (
<emphasis id="B91A7243FFDBFF95DE30C3D3FBEAFF4B" box="[1051,1084,225,251]" italics="true" pageId="2" pageNumber="441">st1</emphasis>
), 17 (
<emphasis id="B91A7243FFDBFF95DE54C3D3FB49FF4B" box="[1151,1183,225,251]" italics="true" pageId="2" pageNumber="441">st2</emphasis>
), 18 (
<emphasis id="B91A7243FFDBFF95DECFC3D3FAD3FF4B" box="[1252,1285,225,251]" italics="true" pageId="2" pageNumber="441">st3</emphasis>
), 20 (
<emphasis id="B91A7243FFDBFF95DF63C3D3FABFFF4B" box="[1352,1385,225,251]" italics="true" pageId="2" pageNumber="441">st4</emphasis>
), 21 (
<emphasis id="B91A7243FFDBFF95DA8AC237FF17FEAF" box="[161,193,261,287]" italics="true" pageId="2" pageNumber="441">st5</emphasis>
);
<typeStatus id="54D510F3FFDBFF95DAF0C237FE9DFEAF" box="[219,331,261,287]" pageId="2" pageNumber="441" type="paratype">paratypes</typeStatus>
: 1825 (
<emphasis id="B91A7243FFDBFF95DB86C237FE18FEAF" box="[429,462,261,287]" italics="true" pageId="2" pageNumber="441">st1</emphasis>
), 1725 (
<emphasis id="B91A7243FFDBFF95D815C237FD89FEAF" box="[574,607,261,287]" italics="true" pageId="2" pageNumber="441">st2</emphasis>
), 1620 (
<emphasis id="B91A7243FFDBFF95D8E4C237FD26FEAF" box="[719,752,261,287]" italics="true" pageId="2" pageNumber="441">st3</emphasis>
), 1316 (
<emphasis id="B91A7243FFDBFF95D94BC237FC57FEAF" box="[864,897,261,287]" italics="true" pageId="2" pageNumber="441">st4</emphasis>
), 1618 (
<emphasis id="B91A7243FFDBFF95D9DAC237FBC4FEAF" box="[1009,1042,261,287]" italics="true" pageId="2" pageNumber="441">st5</emphasis>
). Opisthogaster setae —
<typeStatus id="54D510F3FFDBFF95DF19C237FA4EFEAF" box="[1330,1432,261,287]" pageId="2" pageNumber="441" type="holotype">holotype</typeStatus>
: 26 (
<emphasis id="B91A7243FFDBFF95DAE8C21BFF38FEF3" box="[195,238,297,323]" italics="true" pageId="2" pageNumber="441">JV1</emphasis>
), 36 (
<emphasis id="B91A7243FFDBFF95DB1AC21BFE8BFEF3" box="[305,349,297,323]" italics="true" pageId="2" pageNumber="441">JV2</emphasis>
), 34 (
<emphasis id="B91A7243FFDBFF95DBB4C21BFE1DFEF3" box="[415,459,297,323]" italics="true" pageId="2" pageNumber="441">JV3</emphasis>
), 41 (
<emphasis id="B91A7243FFDBFF95D826C21BFDEFFEF3" box="[525,569,297,323]" italics="true" pageId="2" pageNumber="441">JV4</emphasis>
), 13 (
<emphasis id="B91A7243FFDBFF95D857C21BFD7CFEF3" box="[636,682,297,323]" italics="true" pageId="2" pageNumber="441">ZV1</emphasis>
), 37 (
<emphasis id="B91A7243FFDBFF95D8C6C21BFCCAFEF3" box="[749,796,297,323]" italics="true" pageId="2" pageNumber="441">ZV2</emphasis>
and
<emphasis id="B91A7243FFDBFF95D97AC21BFC56FEF3" box="[849,896,297,323]" italics="true" pageId="2" pageNumber="441">ZV3</emphasis>
), 16 (
<emphasis id="B91A7243FFDBFF95D9E9C21BFC37FEF3" box="[962,993,297,323]" italics="true" pageId="2" pageNumber="441">Ad</emphasis>
);
<typeStatus id="54D510F3FFDBFF95D9D2C21BFBBFFEF3" box="[1017,1129,297,323]" pageId="2" pageNumber="441" type="paratype">paratypes</typeStatus>
: 2125 (
<emphasis id="B91A7243FFDBFF95DEE3C21BFB25FEF3" box="[1224,1267,297,323]" italics="true" pageId="2" pageNumber="441">JV1</emphasis>
), 2735 (
<emphasis id="B91A7243FFDBFF95DF4BC21BFA5AFEF3" box="[1376,1420,297,323]" italics="true" pageId="2" pageNumber="441">JV2</emphasis>
), 2935 (
<emphasis id="B91A7243FFDBFF95DAC4C27FFECCFED7" box="[239,282,333,359]" italics="true" pageId="2" pageNumber="441">JV3</emphasis>
), 3238 (
<emphasis id="B91A7243FFDBFF95DBA1C27FFE60FED7" box="[394,438,333,359]" italics="true" pageId="2" pageNumber="441">JV4</emphasis>
), 1014 (
<emphasis id="B91A7243FFDBFF95D80EC27FFD85FED7" box="[549,595,333,359]" italics="true" pageId="2" pageNumber="441">ZV1</emphasis>
), 2733 (
<emphasis id="B91A7243FFDBFF95D8E8C27FFD24FED7" box="[707,754,333,359]" italics="true" pageId="2" pageNumber="441">ZV2</emphasis>
), 2634 (
<emphasis id="B91A7243FFDBFF95D949C27FFC46FED7" box="[866,912,333,359]" italics="true" pageId="2" pageNumber="441">ZV3</emphasis>
), 1620 (
<emphasis id="B91A7243FFDBFF95DE2BC27FFBC9FED7" box="[1024,1055,333,359]" italics="true" pageId="2" pageNumber="441">Ad</emphasis>
). Sternal and opisthogastric setae simple except
<emphasis id="B91A7243FFDBFF95DB12C243FEB3FE3B" box="[313,357,369,395]" italics="true" pageId="2" pageNumber="441">JV4</emphasis>
enlarged and terminally pilose. Setae
<emphasis id="B91A7243FFDBFF95D93BC243FCEAFE3B" box="[784,828,369,395]" italics="true" pageId="2" pageNumber="441">JV5</emphasis>
and
<emphasis id="B91A7243FFDBFF95D959C243FC77FE3B" box="[882,929,369,395]" italics="true" pageId="2" pageNumber="441">ZV4</emphasis>
located on soft cuticle and similar to
<emphasis id="B91A7243FFDBFF95DF6DC243FAA4FE3B" box="[1350,1394,369,395]" italics="true" pageId="2" pageNumber="441">JV4</emphasis>
.
</paragraph>
<paragraph id="8BD1AE51FFDBFF95DAECC2A7FED4FD37" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="441">
<taxonomicName id="4C6ED5D2FFDBFF95DAECC2A7FE80FE1F" authorityName="Krefft" authorityYear="1871" box="[199,342,405,431]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="2" pageNumber="441" phylum="Nematoda" rank="genus">
<emphasis id="B91A7243FFDBFF95DAECC2A7FE80FE1F" box="[199,342,405,431]" italics="true" pageId="2" pageNumber="441">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="1355B2D4FFDBFF95DB4CC2A7FE00FE1F" box="[359,470,405,431]" captionStart="FIGURES 35" captionStartId="5.[151,264,577,602]" captionTargetBox="[155,1430,169,554]" captionTargetId="figure@5.[151,1436,166,558]" captionTargetPageId="5" captionText="FIGURES 35. Trachygamasus karuni sp. nov. 3. Anterior part of podonotum in female with anterior ends of peritremes, podonotal setae, poroids idj1 and gland pores gdj2, the last-named with double openings. 4 &amp; 5. Ventral view of gnathosoma in female (Figure 4) and male (Figure 5). Holotype: Figure 4; paratypes: Figures 3 and 5." figureDoi="http://doi.org/10.5281/zenodo.3568902" httpUri="https://zenodo.org/record/3568902/files/figure.png" pageId="2" pageNumber="441">Figures 4</figureCitation>
,
<figureCitation id="1355B2D4FFDBFF95DBCFC2A7FDF7FE1F" box="[484,545,405,431]" captionStart="FIGURES 613" captionStartId="5.[151,264,1898,1923]" captionTargetBox="[189,1398,721,1869]" captionTargetId="figure@5.[183,1405,717,1879]" captionTargetPageId="5" captionText="FIGURES 613. Trachygamasus karuni sp. nov., female. 6. Presternal plates. 7. Genital region showing epigynium and an outline of the endogynium in situ. Setae and pores as in Figure 2. 8. Endogynium, two aspects. 9. Gnathotectum. 10. Chelicera antiaxially. 11. Coxa I ventrally. 12. Ambulacra on leg II (Figure 12 left) and leg III (Figure 12 right). 13. Tibia II, posterolateral view. Holotype: Figures 7 and 8 right; paratypes: Figures 6, 8 left, 913." figureDoi="http://doi.org/10.5281/zenodo.3568904" httpUri="https://zenodo.org/record/3568904/files/figure.png" pageId="2" pageNumber="441">9, 10</figureCitation>
). Gnathotectum trispinate (
<figureCitation id="1355B2D4FFDBFF95D97CC2A7FC6DFE1F" box="[855,955,405,431]" captionStart="FIGURES 613" captionStartId="5.[151,264,1898,1923]" captionTargetBox="[189,1398,721,1869]" captionTargetId="figure@5.[183,1405,717,1879]" captionTargetPageId="5" captionText="FIGURES 613. Trachygamasus karuni sp. nov., female. 6. Presternal plates. 7. Genital region showing epigynium and an outline of the endogynium in situ. Setae and pores as in Figure 2. 8. Endogynium, two aspects. 9. Gnathotectum. 10. Chelicera antiaxially. 11. Coxa I ventrally. 12. Ambulacra on leg II (Figure 12 left) and leg III (Figure 12 right). 13. Tibia II, posterolateral view. Holotype: Figures 7 and 8 right; paratypes: Figures 6, 8 left, 913." figureDoi="http://doi.org/10.5281/zenodo.3568904" httpUri="https://zenodo.org/record/3568904/files/figure.png" pageId="2" pageNumber="441">Figure 9</figureCitation>
), central prong with rounded apex, lateral prongs narrow and sharply pointed. Corniculi conical and short. Hypostome (
<figureCitation id="1355B2D4FFDBFF95DE2CC28BFBBDFE63" box="[1031,1131,441,467]" captionStart="FIGURES 35" captionStartId="5.[151,264,577,602]" captionTargetBox="[155,1430,169,554]" captionTargetId="figure@5.[151,1436,166,558]" captionTargetPageId="5" captionText="FIGURES 35. Trachygamasus karuni sp. nov. 3. Anterior part of podonotum in female with anterior ends of peritremes, podonotal setae, poroids idj1 and gland pores gdj2, the last-named with double openings. 4 &amp; 5. Ventral view of gnathosoma in female (Figure 4) and male (Figure 5). Holotype: Figure 4; paratypes: Figures 3 and 5." figureDoi="http://doi.org/10.5281/zenodo.3568902" httpUri="https://zenodo.org/record/3568902/files/figure.png" pageId="2" pageNumber="441">Figure 4</figureCitation>
): hypognathal groove with 47 rows of denticles, hypostomatic and palpcoxal setae simple, internal posterior hypostomatic setae longer, i.e.
<emphasis id="B91A7243FFDBFF95DABCC130FF67FDAB" box="[151,177,514,539]" italics="true" pageId="2" pageNumber="441">ca</emphasis>
. twice the length of the external posterior setae. Chelicera (
<figureCitation id="1355B2D4FFDBFF95D961C133FC6DFDAB" box="[842,955,513,539]" captionStart="FIGURES 613" captionStartId="5.[151,264,1898,1923]" captionTargetBox="[189,1398,721,1869]" captionTargetId="figure@5.[183,1405,717,1879]" captionTargetPageId="5" captionText="FIGURES 613. Trachygamasus karuni sp. nov., female. 6. Presternal plates. 7. Genital region showing epigynium and an outline of the endogynium in situ. Setae and pores as in Figure 2. 8. Endogynium, two aspects. 9. Gnathotectum. 10. Chelicera antiaxially. 11. Coxa I ventrally. 12. Ambulacra on leg II (Figure 12 left) and leg III (Figure 12 right). 13. Tibia II, posterolateral view. Holotype: Figures 7 and 8 right; paratypes: Figures 6, 8 left, 913." figureDoi="http://doi.org/10.5281/zenodo.3568904" httpUri="https://zenodo.org/record/3568904/files/figure.png" pageId="2" pageNumber="441">Figure 10</figureCitation>
): fixed digit with one tooth in the front and three teeth behind
<emphasis id="B91A7243FFDBFF95DB48C117FE24FD8F" box="[355,498,549,575]" italics="true" pageId="2" pageNumber="441">pilus dentilis</emphasis>
, the posterior tooth divided into two parts, anterior and posterior. Movable digit with three teeth, the proximal one largest. Palp trochanter with seta
<date id="FFD08891FFDBFF95D976C17BFC1DFDD3" box="[861,971,585,611]" pageId="2" pageNumber="441">
<emphasis id="B91A7243FFDBFF95D976C17BFCA1FDD3" box="[861,887,585,611]" italics="true" pageId="2" pageNumber="441">v1</emphasis>
and
<emphasis id="B91A7243FFDBFF95D99BC17BFC1DFDD3" box="[944,971,585,611]" italics="true" pageId="2" pageNumber="441">v2</emphasis>
</date>
simple. Setae
<emphasis id="B91A7243FFDBFF95DE5AC17BFB51FDD3" box="[1137,1159,585,611]" italics="true" pageId="2" pageNumber="441">al</emphasis>
on palp femur and genu spatulate.
</paragraph>
<paragraph id="8BD1AE51FFDBFF95DAECC1A0FADAFC8B" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="441">
<emphasis id="B91A7243FFDBFF95DAECC1A0FF2AFD1B" box="[199,252,658,683]" italics="true" pageId="2" pageNumber="441">Legs</emphasis>
. Coxa I with a thickening indented at the level between setae
<date id="FFD08891FFDBFF95D9EDC1A3FBE2FD1B" box="[966,1076,657,683]" pageId="2" pageNumber="441">
<emphasis id="B91A7243FFDBFF95D9EDC1A3FC36FD1B" box="[966,992,657,683]" italics="true" pageId="2" pageNumber="441">v1</emphasis>
and
<emphasis id="B91A7243FFDBFF95DE31C1A3FBE2FD1B" box="[1050,1076,657,683]" italics="true" pageId="2" pageNumber="441">v2</emphasis>
</date>
(
<figureCitation id="1355B2D4FFDBFF95DE6FC1A3FB63FD1B" box="[1092,1205,657,683]" captionStart="FIGURES 613" captionStartId="5.[151,264,1898,1923]" captionTargetBox="[189,1398,721,1869]" captionTargetId="figure@5.[183,1405,717,1879]" captionTargetPageId="5" captionText="FIGURES 613. Trachygamasus karuni sp. nov., female. 6. Presternal plates. 7. Genital region showing epigynium and an outline of the endogynium in situ. Setae and pores as in Figure 2. 8. Endogynium, two aspects. 9. Gnathotectum. 10. Chelicera antiaxially. 11. Coxa I ventrally. 12. Ambulacra on leg II (Figure 12 left) and leg III (Figure 12 right). 13. Tibia II, posterolateral view. Holotype: Figures 7 and 8 right; paratypes: Figures 6, 8 left, 913." figureDoi="http://doi.org/10.5281/zenodo.3568904" httpUri="https://zenodo.org/record/3568904/files/figure.png" pageId="2" pageNumber="441">Figure 11</figureCitation>
). Ambulacra of legs IIIV characteristic, i.e. anterior pulvilli irregular in shape, but terminally rounded, lateral pulvilli narrow, curved and acuminate (
<figureCitation id="1355B2D4FFDBFF95DB61C1EBFE6AFD43" box="[330,444,729,755]" captionStart="FIGURES 613" captionStartId="5.[151,264,1898,1923]" captionTargetBox="[189,1398,721,1869]" captionTargetId="figure@5.[183,1405,717,1879]" captionTargetPageId="5" captionText="FIGURES 613. Trachygamasus karuni sp. nov., female. 6. Presternal plates. 7. Genital region showing epigynium and an outline of the endogynium in situ. Setae and pores as in Figure 2. 8. Endogynium, two aspects. 9. Gnathotectum. 10. Chelicera antiaxially. 11. Coxa I ventrally. 12. Ambulacra on leg II (Figure 12 left) and leg III (Figure 12 right). 13. Tibia II, posterolateral view. Holotype: Figures 7 and 8 right; paratypes: Figures 6, 8 left, 913." figureDoi="http://doi.org/10.5281/zenodo.3568904" httpUri="https://zenodo.org/record/3568904/files/figure.png" pageId="2" pageNumber="441">Figure 12</figureCitation>
). Leg chaetotaxy normal, all setae smooth and needle-like except for posterolateral and some posterodorsal setae on genu and tibia I, femur II, genu II and tibia II, as well as genu IV and tibia IV which are narrow, but terminally blunt (
<figureCitation id="1355B2D4FFDBFF95DBCBC013FD87FC8B" box="[480,593,801,827]" captionStart="FIGURES 613" captionStartId="5.[151,264,1898,1923]" captionTargetBox="[189,1398,721,1869]" captionTargetId="figure@5.[183,1405,717,1879]" captionTargetPageId="5" captionText="FIGURES 613. Trachygamasus karuni sp. nov., female. 6. Presternal plates. 7. Genital region showing epigynium and an outline of the endogynium in situ. Setae and pores as in Figure 2. 8. Endogynium, two aspects. 9. Gnathotectum. 10. Chelicera antiaxially. 11. Coxa I ventrally. 12. Ambulacra on leg II (Figure 12 left) and leg III (Figure 12 right). 13. Tibia II, posterolateral view. Holotype: Figures 7 and 8 right; paratypes: Figures 6, 8 left, 913." figureDoi="http://doi.org/10.5281/zenodo.3568904" httpUri="https://zenodo.org/record/3568904/files/figure.png" pageId="2" pageNumber="441">Figure 13</figureCitation>
). Other aspects of the leg structure and setation unremarkable.
</paragraph>
<paragraph id="8BD1AE51FFDBFF95DAECC077FE01FCEF" blockId="2.[151,1437,153,2015]" box="[199,471,836,863]" pageId="2" pageNumber="441">
<emphasis id="B91A7243FFDBFF95DAECC077FED7FCEF" box="[199,257,837,863]" italics="true" pageId="2" pageNumber="441">Male</emphasis>
(
<figureCitation id="1355B2D4FFDBFF95DB24C077FEAAFCEF" box="[271,380,836,863]" captionStart="FIGURES 35" captionStartId="5.[151,264,577,602]" captionTargetBox="[155,1430,169,554]" captionTargetId="figure@5.[151,1436,166,558]" captionTargetPageId="5" captionText="FIGURES 35. Trachygamasus karuni sp. nov. 3. Anterior part of podonotum in female with anterior ends of peritremes, podonotal setae, poroids idj1 and gland pores gdj2, the last-named with double openings. 4 &amp; 5. Ventral view of gnathosoma in female (Figure 4) and male (Figure 5). Holotype: Figure 4; paratypes: Figures 3 and 5." figureDoi="http://doi.org/10.5281/zenodo.3568902" httpUri="https://zenodo.org/record/3568902/files/figure.png" pageId="2" pageNumber="441">Figures 5</figureCitation>
,
<figureCitation id="1355B2D4FFDBFF95DBA3C077FE19FCEF" box="[392,463,837,863]" captionStart-0="FIGURE 14" captionStart-1="FIGURES 1519" captionStart-2="FIGURES 2024" captionStartId-0="6.[151,250,1783,1808]" captionStartId-1="7.[151,264,1924,1949]" captionStartId-2="8.[151,264,1924,1949]" captionTargetBox-0="[293,1195,224,1660]" captionTargetBox-1="[164,1387,178,1897]" captionTargetBox-2="[166,1364,181,1889]" captionTargetId-0="figure@6.[250,1205,223,1683]" captionTargetId-1="figure@7.[151,1399,166,1904]" captionTargetId-2="figure@8.[151,1376,166,1904]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionText-0="FIGURE 14. Trachygamasus karuni sp. nov., male. Dorsal side of idiosoma, abbreviations as in Figure 1." captionText-1="FIGURES 1519. Trachygamasus karuni sp. nov., male. 15. Ventral side of idiosoma, abbreviations as in Figure 2. 16. Anterior margin of sternum with genital lamina flanked by presternal plates. 17. Hypostome in ventral perspective showing corniculi, hypostomatic setae, palpcoxa seta and palptrochanter. 18. Gnathotectum, two aspects. 19. Chelicera adaxially." captionText-2="FIGURES 2024. Trachygamasus karuni sp. nov., male, leg II. 20. Femur, genu and tibia in ventral perspective. 21. Femur, genu and tibia posterolaterally. 22. Spur on genu anterolaterally. 23 &amp; 24. Spurs on femur II and genu II, enlarged. Setae anterolateral (al), posteroventral (pv), posterolateral (pl), and posterodorsal (pd) are marked." figureDoi-0="http://doi.org/10.5281/zenodo.3568906" figureDoi-1="http://doi.org/10.5281/zenodo.3568908" figureDoi-2="http://doi.org/10.5281/zenodo.3568910" httpUri-0="https://zenodo.org/record/3568906/files/figure.png" httpUri-1="https://zenodo.org/record/3568908/files/figure.png" httpUri-2="https://zenodo.org/record/3568910/files/figure.png" pageId="2" pageNumber="441">1424</figureCitation>
)
</paragraph>
<paragraph id="8BD1AE51FFDBFF95DAECC05BFE21FB87" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="441">
<emphasis id="B91A7243FFDBFF95DAECC05BFE51FC33" box="[199,391,873,899]" italics="true" pageId="2" pageNumber="441">Dorsal idiosoma</emphasis>
(
<figureCitation id="1355B2D4FFDBFF95DBB2C05BFDDBFC33" box="[409,525,873,899]" captionStart="FIGURE 14" captionStartId="6.[151,250,1783,1808]" captionTargetBox="[293,1195,224,1660]" captionTargetId="figure@6.[250,1205,223,1683]" captionTargetPageId="6" captionText="FIGURE 14. Trachygamasus karuni sp. nov., male. Dorsal side of idiosoma, abbreviations as in Figure 1." figureDoi="http://doi.org/10.5281/zenodo.3568906" httpUri="https://zenodo.org/record/3568906/files/figure.png" pageId="2" pageNumber="441">Figure 14</figureCitation>
).
<typeStatus id="54D510F3FFDBFF95D80EC05BFD44FC33" box="[549,658,873,899]" pageId="2" pageNumber="441" type="paratype">Paratypes</typeStatus>
with holodorsal shield showing poorly discernible border between podonotum and opisthonotum. Idiosoma oval, well sclerotised and brown, 460485 long, 260315 wide, n=5. Dorsal setation as in female: podonotal setae simple except for
<emphasis id="B91A7243FFDBFF95D913C083FC98FC7B" box="[824,846,945,971]" italics="true" pageId="2" pageNumber="441">j1</emphasis>
,
<emphasis id="B91A7243FFDBFF95D976C083FCA5FC7B" box="[861,883,945,971]" italics="true" pageId="2" pageNumber="441">j4</emphasis>
,
<emphasis id="B91A7243FFDBFF95D9AAC083FC4DFC7B" box="[897,923,945,971]" italics="true" pageId="2" pageNumber="441">r3</emphasis>
and
<emphasis id="B91A7243FFDBFF95D9F9C083FC3DFC7B" box="[978,1003,945,971]" italics="true" pageId="2" pageNumber="441">z5</emphasis>
. The length of simple podonotal setae 1218,
<emphasis id="B91A7243FFDBFF95DAC0C0E7FED7FC5F" box="[235,257,981,1007]" italics="true" pageId="2" pageNumber="441">
j
<date id="FFD08891FFDBFF95DADCC0E7FED7FC5F" box="[247,257,981,1007]" pageId="2" pageNumber="441" value="1924-01-18">1</date>
</emphasis>
1824,
<emphasis id="B91A7243FFDBFF95DB77C0E7FEA4FC5F" box="[348,370,981,1007]" italics="true" pageId="2" pageNumber="441">
j
<date id="FFD08891FFDBFF95DB4DC0E7FEA4FC5F" box="[358,370,981,1007]" pageId="2" pageNumber="441" value="1933-04-29">4</date>
</emphasis>
2933,
<emphasis id="B91A7243FFDBFF95DBE5C0E7FE31FC5F" box="[462,487,981,1007]" italics="true" pageId="2" pageNumber="441">r3</emphasis>
3942,
<emphasis id="B91A7243FFDBFF95D869C0E7FD8DFC5F" box="[578,603,981,1007]" italics="true" pageId="2" pageNumber="441">z5</emphasis>
33. Opisthonotal setae as in the female. The length of simple opisthonotal setae is 1316,
<emphasis id="B91A7243FFDBFF95DB69C0CBFEB6FBA3" box="[322,352,1017,1043]" italics="true" pageId="2" pageNumber="441">
Z
<date id="FFD08891FFDBFF95DB7EC0CBFEB6FBA3" box="[341,352,1017,1043]" pageId="2" pageNumber="441" value="1933-01-30">1</date>
</emphasis>
3033,
<emphasis id="B91A7243FFDBFF95DB91C0CBFE01FBA3" box="[442,471,1017,1043]" italics="true" pageId="2" pageNumber="441">
Z
<date id="FFD08891FFDBFF95DBE1C0CBFE01FBA3" box="[458,471,1017,1043]" pageId="2" pageNumber="441" value="1926-04-21">4</date>
</emphasis>
2126. Peritrematal groove 248258 long, expanding to beyond coxa I. Cuticle shows many regularly distributed pits.
</paragraph>
<paragraph id="8BD1AE51FFDBFF95DAECC773FB34FAE7" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="441">
<emphasis id="B91A7243FFDBFF95DAECC773FE5EFBEB" box="[199,392,1089,1115]" italics="true" pageId="2" pageNumber="441">Ventral idiosoma</emphasis>
(
<figureCitation id="1355B2D4FFDBFF95DBB3C773FDDDFBEB" box="[408,523,1088,1115]" captionStart="FIGURES 1519" captionStartId="7.[151,264,1924,1949]" captionTargetBox="[164,1387,178,1897]" captionTargetId="figure@7.[151,1399,166,1904]" captionTargetPageId="7" captionText="FIGURES 1519. Trachygamasus karuni sp. nov., male. 15. Ventral side of idiosoma, abbreviations as in Figure 2. 16. Anterior margin of sternum with genital lamina flanked by presternal plates. 17. Hypostome in ventral perspective showing corniculi, hypostomatic setae, palpcoxa seta and palptrochanter. 18. Gnathotectum, two aspects. 19. Chelicera adaxially." figureDoi="http://doi.org/10.5281/zenodo.3568908" httpUri="https://zenodo.org/record/3568908/files/figure.png" pageId="2" pageNumber="441">Figure 15</figureCitation>
). Tritosternum absent, presternal plates large (
<figureCitation id="1355B2D4FFDBFF95DE36C773FB27FBEC" box="[1053,1265,1088,1116]" captionStart="FIGURES 1519" captionStartId="7.[151,264,1924,1949]" captionTargetBox="[164,1387,178,1897]" captionTargetId="figure@7.[151,1399,166,1904]" captionTargetPageId="7" captionText="FIGURES 1519. Trachygamasus karuni sp. nov., male. 15. Ventral side of idiosoma, abbreviations as in Figure 2. 16. Anterior margin of sternum with genital lamina flanked by presternal plates. 17. Hypostome in ventral perspective showing corniculi, hypostomatic setae, palpcoxa seta and palptrochanter. 18. Gnathotectum, two aspects. 19. Chelicera adaxially." figureDoi="http://doi.org/10.5281/zenodo.3568908" httpUri="https://zenodo.org/record/3568908/files/figure.png" pageId="2" pageNumber="441">Figures 15 and 16</figureCitation>
). Sternogenital shield anterior margin (
<figureCitation id="1355B2D4FFDBFF95DBB0C757FDB3FB30" box="[411,613,1124,1152]" captionStart="FIGURES 1519" captionStartId="7.[151,264,1924,1949]" captionTargetBox="[164,1387,178,1897]" captionTargetId="figure@7.[151,1399,166,1904]" captionTargetPageId="7" captionText="FIGURES 1519. Trachygamasus karuni sp. nov., male. 15. Ventral side of idiosoma, abbreviations as in Figure 2. 16. Anterior margin of sternum with genital lamina flanked by presternal plates. 17. Hypostome in ventral perspective showing corniculi, hypostomatic setae, palpcoxa seta and palptrochanter. 18. Gnathotectum, two aspects. 19. Chelicera adaxially." figureDoi="http://doi.org/10.5281/zenodo.3568908" httpUri="https://zenodo.org/record/3568908/files/figure.png" pageId="2" pageNumber="441">Figures 15 and 16</figureCitation>
) with a concavity behind the genital lamina. Genital lamina shows a short posterior part covered with sternum; in its anterior part there are two lateral thickenings and a bar-shaped anterior thickening. Poorly visible, anteriorly protruded, apically barbed hyaline extension is present (
<figureCitation id="1355B2D4FFDBFF95DEE8C79FFAE0FB78" box="[1219,1334,1197,1224]" captionStart="FIGURES 1519" captionStartId="7.[151,264,1924,1949]" captionTargetBox="[164,1387,178,1897]" captionTargetId="figure@7.[151,1399,166,1904]" captionTargetPageId="7" captionText="FIGURES 1519. Trachygamasus karuni sp. nov., male. 15. Ventral side of idiosoma, abbreviations as in Figure 2. 16. Anterior margin of sternum with genital lamina flanked by presternal plates. 17. Hypostome in ventral perspective showing corniculi, hypostomatic setae, palpcoxa seta and palptrochanter. 18. Gnathotectum, two aspects. 19. Chelicera adaxially." figureDoi="http://doi.org/10.5281/zenodo.3568908" httpUri="https://zenodo.org/record/3568908/files/figure.png" pageId="2" pageNumber="441">Figure 16</figureCitation>
). Sternal setae simple and similar, their length: 1721 (
<emphasis id="B91A7243FFDBFF95D8B2C7E3FD3EFB5B" box="[665,744,1233,1259]" italics="true" pageId="2" pageNumber="441">st1st3</emphasis>
), 1617 (
<emphasis id="B91A7243FFDBFF95D97DC7E3FCA1FB5B" box="[854,887,1233,1259]" italics="true" pageId="2" pageNumber="441">st4</emphasis>
), 1213 (
<emphasis id="B91A7243FFDBFF95D9CEC7E3FBD3FB5B" box="[997,1029,1233,1259]" italics="true" pageId="2" pageNumber="441">st5</emphasis>
); opisthogastric setae simple except for
<emphasis id="B91A7243FFDBFF95DAEBC7C7FF3AFABF" box="[192,236,1269,1295]" italics="true" pageId="2" pageNumber="441">JV4</emphasis>
, as well as
<emphasis id="B91A7243FFDBFF95DB59C7C7FE48FABF" box="[370,414,1269,1295]" italics="true" pageId="2" pageNumber="441">JV5</emphasis>
and
<emphasis id="B91A7243FFDBFF95DBFDC7C7FDD0FABF" box="[470,518,1269,1295]" italics="true" pageId="2" pageNumber="441">ZV4</emphasis>
which are larger and terminally pilose and located marginally, or setae
<emphasis id="B91A7243FFDBFF95DF1FC7C7FAB6FABF" box="[1332,1376,1269,1295]" italics="true" pageId="2" pageNumber="441">JV5</emphasis>
even somewhat dorsally. Their length: 2021 (
<emphasis id="B91A7243FFDBFF95D840C62BFD40FA83" box="[619,662,1305,1331]" italics="true" pageId="2" pageNumber="441">JV1</emphasis>
), 2527 (
<emphasis id="B91A7243FFDBFF95D92CC62BFCE5FA83" box="[775,819,1305,1331]" italics="true" pageId="2" pageNumber="441">JV2</emphasis>
and
<emphasis id="B91A7243FFDBFF95D947C62BFC41FA83" box="[876,919,1305,1331]" italics="true" pageId="2" pageNumber="441">JV3</emphasis>
), 2933 (
<emphasis id="B91A7243FFDBFF95DE23C62BFBE2FA83" box="[1032,1076,1305,1331]" italics="true" pageId="2" pageNumber="441">JV4</emphasis>
,
<emphasis id="B91A7243FFDBFF95DE68C62BFBB9FA83" box="[1091,1135,1305,1331]" italics="true" pageId="2" pageNumber="441">JV5</emphasis>
,
<emphasis id="B91A7243FFDBFF95DE55C62BFB7BFA83" box="[1150,1197,1305,1331]" italics="true" pageId="2" pageNumber="441">ZV4</emphasis>
), 910 (
<emphasis id="B91A7243FFDBFF95DF24C62BFAEBFA83" box="[1295,1341,1305,1331]" italics="true" pageId="2" pageNumber="441">ZV1</emphasis>
), 2427 (
<emphasis id="B91A7243FFDBFF95DA8AC60FFF19FAE7" box="[161,207,1341,1367]" italics="true" pageId="2" pageNumber="441">ZV2</emphasis>
,
<emphasis id="B91A7243FFDBFF95DAF6C60FFEDAFAE7" box="[221,268,1341,1367]" italics="true" pageId="2" pageNumber="441">ZV3</emphasis>
), 1214 (
<emphasis id="B91A7243FFDBFF95DB51C60FFE4FFAE7" box="[378,409,1341,1367]" italics="true" pageId="2" pageNumber="441">Ad</emphasis>
). Sternal poroids
<emphasis id="B91A7243FFDBFF95D84AC60FFD52FAE7" box="[609,644,1341,1367]" italics="true" pageId="2" pageNumber="441">
<date id="FFD08891FFDBFF95D84AC60FFD52FAE7" box="[609,644,1341,1367]" pageId="2" pageNumber="441">iv3</date>
</emphasis>
circular. Idiosomal margin behind the anus is convex.
</paragraph>
<paragraph id="8BD1AE51FFDBFF95DAECC653FAA9F9E3" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="441">
<taxonomicName id="4C6ED5D2FFDBFF95DAECC653FE80FACB" authorityName="Krefft" authorityYear="1871" box="[199,342,1377,1403]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="2" pageNumber="441" phylum="Nematoda" rank="genus">
<emphasis id="B91A7243FFDBFF95DAECC653FE80FACB" box="[199,342,1377,1403]" italics="true" pageId="2" pageNumber="441">Gnathosoma</emphasis>
</taxonomicName>
(
<figureCitation id="1355B2D4FFDBFF95DB4FC653FE12FACB" box="[356,452,1376,1403]" captionStart="FIGURES 35" captionStartId="5.[151,264,577,602]" captionTargetBox="[155,1430,169,554]" captionTargetId="figure@5.[151,1436,166,558]" captionTargetPageId="5" captionText="FIGURES 35. Trachygamasus karuni sp. nov. 3. Anterior part of podonotum in female with anterior ends of peritremes, podonotal setae, poroids idj1 and gland pores gdj2, the last-named with double openings. 4 &amp; 5. Ventral view of gnathosoma in female (Figure 4) and male (Figure 5). Holotype: Figure 4; paratypes: Figures 3 and 5." figureDoi="http://doi.org/10.5281/zenodo.3568902" httpUri="https://zenodo.org/record/3568902/files/figure.png" pageId="2" pageNumber="441">Figure 5</figureCitation>
,
<figureCitation id="1355B2D4FFDBFF95DBE4C653FDCFFACB" box="[463,537,1377,1403]" captionStart="FIGURES 1519" captionStartId="7.[151,264,1924,1949]" captionTargetBox="[164,1387,178,1897]" captionTargetId="figure@7.[151,1399,166,1904]" captionTargetPageId="7" captionText="FIGURES 1519. Trachygamasus karuni sp. nov., male. 15. Ventral side of idiosoma, abbreviations as in Figure 2. 16. Anterior margin of sternum with genital lamina flanked by presternal plates. 17. Hypostome in ventral perspective showing corniculi, hypostomatic setae, palpcoxa seta and palptrochanter. 18. Gnathotectum, two aspects. 19. Chelicera adaxially." figureDoi="http://doi.org/10.5281/zenodo.3568908" httpUri="https://zenodo.org/record/3568908/files/figure.png" pageId="2" pageNumber="441">1719</figureCitation>
). Gnathotectum trispinate (
<figureCitation id="1355B2D4FFDBFF95D96EC653FC65FACB" box="[837,947,1377,1403]" captionStart="FIGURES 1519" captionStartId="7.[151,264,1924,1949]" captionTargetBox="[164,1387,178,1897]" captionTargetId="figure@7.[151,1399,166,1904]" captionTargetPageId="7" captionText="FIGURES 1519. Trachygamasus karuni sp. nov., male. 15. Ventral side of idiosoma, abbreviations as in Figure 2. 16. Anterior margin of sternum with genital lamina flanked by presternal plates. 17. Hypostome in ventral perspective showing corniculi, hypostomatic setae, palpcoxa seta and palptrochanter. 18. Gnathotectum, two aspects. 19. Chelicera adaxially." figureDoi="http://doi.org/10.5281/zenodo.3568908" httpUri="https://zenodo.org/record/3568908/files/figure.png" pageId="2" pageNumber="441">Figure 18</figureCitation>
), central prong larger than the lateral prongs and rounded terminally, ending with an additional pointed apex. Lateral prongs acuminate as in female. Corniculi (
<figureCitation id="1355B2D4FFDBFF95DAB4C69BFEDBFA73" box="[159,269,1448,1475]" captionStart="FIGURES 35" captionStartId="5.[151,264,577,602]" captionTargetBox="[155,1430,169,554]" captionTargetId="figure@5.[151,1436,166,558]" captionTargetPageId="5" captionText="FIGURES 35. Trachygamasus karuni sp. nov. 3. Anterior part of podonotum in female with anterior ends of peritremes, podonotal setae, poroids idj1 and gland pores gdj2, the last-named with double openings. 4 &amp; 5. Ventral view of gnathosoma in female (Figure 4) and male (Figure 5). Holotype: Figure 4; paratypes: Figures 3 and 5." figureDoi="http://doi.org/10.5281/zenodo.3568902" httpUri="https://zenodo.org/record/3568902/files/figure.png" pageId="2" pageNumber="441">Figures 5</figureCitation>
and
<figureCitation id="1355B2D4FFDBFF95DB61C69BFEB1FA73" box="[330,359,1449,1475]" captionStart="FIGURES 1519" captionStartId="7.[151,264,1924,1949]" captionTargetBox="[164,1387,178,1897]" captionTargetId="figure@7.[151,1399,166,1904]" captionTargetPageId="7" captionText="FIGURES 1519. Trachygamasus karuni sp. nov., male. 15. Ventral side of idiosoma, abbreviations as in Figure 2. 16. Anterior margin of sternum with genital lamina flanked by presternal plates. 17. Hypostome in ventral perspective showing corniculi, hypostomatic setae, palpcoxa seta and palptrochanter. 18. Gnathotectum, two aspects. 19. Chelicera adaxially." figureDoi="http://doi.org/10.5281/zenodo.3568908" httpUri="https://zenodo.org/record/3568908/files/figure.png" pageId="2" pageNumber="441">17</figureCitation>
) short and conical; hypognathal groove with 56 posterior rows of denticles (
<figureCitation id="1355B2D4FFDBFF95DED8C69BFA8EFA73" box="[1267,1368,1448,1475]" captionStart="FIGURES 35" captionStartId="5.[151,264,577,602]" captionTargetBox="[155,1430,169,554]" captionTargetId="figure@5.[151,1436,166,558]" captionTargetPageId="5" captionText="FIGURES 35. Trachygamasus karuni sp. nov. 3. Anterior part of podonotum in female with anterior ends of peritremes, podonotal setae, poroids idj1 and gland pores gdj2, the last-named with double openings. 4 &amp; 5. Ventral view of gnathosoma in female (Figure 4) and male (Figure 5). Holotype: Figure 4; paratypes: Figures 3 and 5." figureDoi="http://doi.org/10.5281/zenodo.3568902" httpUri="https://zenodo.org/record/3568902/files/figure.png" pageId="2" pageNumber="441">Figure 5</figureCitation>
). Hypostomatic and palpcoxal setae simple; internal posterior hypostomatics more than double length of external ones. Chelicera (
<figureCitation id="1355B2D4FFDBFF95DB3FC6C3FE51F9BB" box="[276,391,1521,1547]" captionStart="FIGURES 1519" captionStartId="7.[151,264,1924,1949]" captionTargetBox="[164,1387,178,1897]" captionTargetId="figure@7.[151,1399,166,1904]" captionTargetPageId="7" captionText="FIGURES 1519. Trachygamasus karuni sp. nov., male. 15. Ventral side of idiosoma, abbreviations as in Figure 2. 16. Anterior margin of sternum with genital lamina flanked by presternal plates. 17. Hypostome in ventral perspective showing corniculi, hypostomatic setae, palpcoxa seta and palptrochanter. 18. Gnathotectum, two aspects. 19. Chelicera adaxially." figureDoi="http://doi.org/10.5281/zenodo.3568908" httpUri="https://zenodo.org/record/3568908/files/figure.png" pageId="2" pageNumber="441">Figure 19</figureCitation>
): fixed digit with two minute denticles in front of
<emphasis id="B91A7243FFDBFF95D9FFC6C3FBB0F9BB" box="[980,1126,1521,1547]" italics="true" pageId="2" pageNumber="441">pilus dentilis</emphasis>
and dentate lamellar ridge behind
<emphasis id="B91A7243FFDBFF95DAC0C527FEACF99F" box="[235,378,1557,1583]" italics="true" pageId="2" pageNumber="441">pilus dentilis</emphasis>
. Movable digit with one tooth followed by a sinuous ridge. Palp trochanter (
<figureCitation id="1355B2D4FFDBFF95DEF3C527FA91F99F" box="[1240,1351,1557,1583]" captionStart="FIGURES 1519" captionStartId="7.[151,264,1924,1949]" captionTargetBox="[164,1387,178,1897]" captionTargetId="figure@7.[151,1399,166,1904]" captionTargetPageId="7" captionText="FIGURES 1519. Trachygamasus karuni sp. nov., male. 15. Ventral side of idiosoma, abbreviations as in Figure 2. 16. Anterior margin of sternum with genital lamina flanked by presternal plates. 17. Hypostome in ventral perspective showing corniculi, hypostomatic setae, palpcoxa seta and palptrochanter. 18. Gnathotectum, two aspects. 19. Chelicera adaxially." figureDoi="http://doi.org/10.5281/zenodo.3568908" httpUri="https://zenodo.org/record/3568908/files/figure.png" pageId="2" pageNumber="441">Figure 17</figureCitation>
) with a thickened ridge anteroventrally; seta
<date id="FFD08891FFDBFF95D81DC50BFD74F9E3" box="[566,674,1593,1619]" pageId="2" pageNumber="441">
<emphasis id="B91A7243FFDBFF95D81DC50BFD86F9E3" box="[566,592,1593,1619]" italics="true" pageId="2" pageNumber="441">v1</emphasis>
and
<emphasis id="B91A7243FFDBFF95D8ACC50BFD74F9E3" box="[647,674,1593,1619]" italics="true" pageId="2" pageNumber="441">v2</emphasis>
</date>
simple. Anterolateral setae on the palp femur and genu spatulate.
</paragraph>
<paragraph id="8BD1AE51FFDBFF95DAECC56CFBA0F8B7" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="441">
<emphasis id="B91A7243FFDBFF95DAECC56CFF2AF9C7" box="[199,252,1630,1655]" italics="true" pageId="2" pageNumber="441">Legs</emphasis>
. Characteristic ambulacra of legs IIIV as in the female. Leg II (
<figureCitation id="1355B2D4FFDBFF95D9C8C56FFB5FF9C7" box="[995,1161,1629,1655]" captionStart="FIGURES 2024" captionStartId="8.[151,264,1924,1949]" captionTargetBox="[166,1364,181,1889]" captionTargetId="figure@8.[151,1376,166,1904]" captionTargetPageId="8" captionText="FIGURES 2024. Trachygamasus karuni sp. nov., male, leg II. 20. Femur, genu and tibia in ventral perspective. 21. Femur, genu and tibia posterolaterally. 22. Spur on genu anterolaterally. 23 &amp; 24. Spurs on femur II and genu II, enlarged. Setae anterolateral (al), posteroventral (pv), posterolateral (pl), and posterodorsal (pd) are marked." figureDoi="http://doi.org/10.5281/zenodo.3568910" httpUri="https://zenodo.org/record/3568910/files/figure.png" pageId="2" pageNumber="441">Figures 2024</figureCitation>
) spurred as follows: the femur with one minute spur ventrally, genual spur located anteroventrally. Both spurs minute and conical. Posterolateral seta
<emphasis id="B91A7243FFDBFF95DB32C597FEEBF90F" box="[281,317,1701,1727]" italics="true" pageId="2" pageNumber="441">pl1</emphasis>
on the femur, posterolateral setae
<emphasis id="B91A7243FFDBFF95D8EDC597FD3CF90F" box="[710,746,1701,1727]" italics="true" pageId="2" pageNumber="441">pl1</emphasis>
and
<emphasis id="B91A7243FFDBFF95D909C597FC91F90F" box="[802,839,1701,1727]" italics="true" pageId="2" pageNumber="441">pl2</emphasis>
on the genu and tibia, as well as posterodorsal setae
<emphasis id="B91A7243FFDBFF95DABCC5FBFF14F953" box="[151,194,1737,1763]" italics="true" pageId="2" pageNumber="441">pd1</emphasis>
on the genu and the tibia thickened and terminally pilose. The other setae on leg II simple. Some anterodorsal setae on femur IV and genu IV terminally blunt. Other aspects of the legs unremarkable.
</paragraph>
</subSubSection>
<subSubSection id="C374FDDAFFDBFF95DAECC423FE21F8FF" pageId="2" pageNumber="441" type="etymology">
<paragraph id="8BD1AE51FFDBFF95DAECC423FE21F8FF" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="441">
<emphasis id="B91A7243FFDBFF95DAECC423FE98F89B" bold="true" box="[199,334,1809,1835]" pageId="2" pageNumber="441">Etymology.</emphasis>
The specific name
<taxonomicName id="4C6ED5D2FFDBFF95D807C423FDA3F89B" authorityName="Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński" authorityYear="2019" box="[556,629,1809,1835]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="441" phylum="Arthropoda" rank="species" species="karuni">
<emphasis id="B91A7243FFDBFF95D807C423FDA3F89B" box="[556,629,1809,1835]" italics="true" pageId="2" pageNumber="441">karuni</emphasis>
</taxonomicName>
refers to the Karun, Irans only navigable river, running along the sam- pling areas for the new species.
</paragraph>
</subSubSection>
<subSubSection id="C374FDDAFFDBFF9EDAECC46BFD4DFEF3" lastPageId="9" lastPageNumber="448" pageId="2" pageNumber="441" type="discussion">
<paragraph id="8BD1AE51FFDBFF94DAECC46BFAA9FF67" blockId="2.[151,1437,153,2015]" lastBlockId="3.[151,1437,153,215]" lastPageId="3" lastPageNumber="442" pageId="2" pageNumber="441">
<emphasis id="B91A7243FFDBFF95DAECC46BFEEDF8C3" bold="true" box="[199,315,1881,1907]" pageId="2" pageNumber="441">Remarks.</emphasis>
As shown in the key to
<taxonomicName id="4C6ED5D2FFDBFF95D84BC46BFCC6F8C3" box="[608,784,1881,1907]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="441" phylum="Arthropoda" rank="genus">
<emphasis id="B91A7243FFDBFF95D84BC46BFCC6F8C3" box="[608,784,1881,1907]" italics="true" pageId="2" pageNumber="441">Trachygamasus</emphasis>
</taxonomicName>
species, the newly described species is morphologically similar to
<taxonomicName id="4C6ED5D2FFDBFF95DB26C44CFDBFF827" authority="Ma &amp; Wang, 1996" authorityName="Ma &amp; Wang" authorityYear="1996" box="[269,617,1917,1943]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="441" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B91A7243FFDBFF95DB26C44CFECBF827" box="[269,285,1918,1943]" italics="true" pageId="2" pageNumber="441">T</emphasis>
.
<emphasis id="B91A7243FFDBFF95DB06C44FFE5EF827" box="[301,392,1917,1943]" italics="true" pageId="2" pageNumber="441">borealis</emphasis>
<bibRefCitation id="EFFFD3A0FFDBFF95DBBBC44FFDBFF827" author="Ma, L. - M. &amp; Wang, S." box="[400,617,1917,1943]" pageId="2" pageNumber="441" pagination="81 - 88" refId="ref7836" refString="Ma, L. - M. &amp; Wang, S. (1996) Four new species of the family Parasitidae and discovery of the genus Trachygamasus in China. Acta Arachnologica Sinica, 5, 81 - 88. [in Chinese, with English abstract]" type="journal article" year="1996">Ma &amp; Wang, 1996</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C6ED5D2FFDBFF95D85DC44CFC42F827" authority="Witalinski, 2017" authorityName="Witalinski" authorityYear="2017" box="[630,916,1917,1943]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="441" phylum="Arthropoda" rank="species" species="gerdi">
<emphasis id="B91A7243FFDBFF95D85DC44CFD50F827" box="[630,646,1918,1943]" italics="true" pageId="2" pageNumber="441">T</emphasis>
.
<emphasis id="B91A7243FFDBFF95D8BDC44FFD06F827" box="[662,720,1917,1943]" italics="true" pageId="2" pageNumber="441">gerdi</emphasis>
<bibRefCitation id="EFFFD3A0FFDBFF95D8F2C44FFC42F827" author="Witalinski, W." box="[729,916,1917,1943]" pageId="2" pageNumber="441" pagination="407 - 416" refId="ref8173" refString="Witalinski, W. (2017) A new species of Trachygamasus from Poland, a new definition of the genus, and a key to the world species (Parasitiformes: Parasitidae). Zootaxa, 4303 (3), 407 - 416. https: // doi. org / 10.11646 / zootaxa. 4303.3.6" type="journal article" year="2017">Witaliński, 2017</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C6ED5D2FFDBFF95D9E4C44CFA4FF827" authority="Yao, Jin &amp; Zhang, 2019" authorityName="Yao, Jin &amp; Zhang" authorityYear="2019" box="[975,1433,1917,1943]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="441" phylum="Arthropoda" rank="species" species="pseudogerdi">
<emphasis id="B91A7243FFDBFF95D9E4C44CFC09F827" box="[975,991,1918,1943]" italics="true" pageId="2" pageNumber="441">T</emphasis>
.
<emphasis id="B91A7243FFDBFF95D9C4C44FFBAEF827" box="[1007,1144,1917,1943]" italics="true" pageId="2" pageNumber="441">pseudogerdi</emphasis>
<bibRefCitation id="EFFFD3A0FFDBFF95DEAAC44FFA4FF827" author="Yao, M. &amp; Yi, T. - C. &amp; Guo, J. - J. &amp; Jin, D. - C. &amp; Zhang, R. - Z." box="[1153,1433,1917,1943]" pageId="2" pageNumber="441" pagination="1465 - 1489" refId="ref8231" refString="Yao, M., Yi, T. - C., Guo, J. - J., Jin, D. - C. &amp; Zhang, R. - Z. (2019) Three new species of Trachygamasus (Mesostigmata: Parasitidae) from China, with a key to world species of the genus. Systematic and Applied Acarology, 24, 1465 - 1489. https: // doi. org / 10.11158 / saa. 24.8.9" type="journal article" year="2019">Yao, Jin &amp; Zhang, 2019</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C6ED5D2FFDBFF95DABCC493FE4EF80B" authorityName="Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński" authorityYear="2019" box="[151,408,1953,1979]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="441" phylum="Arthropoda" rank="species" species="karuni">
<emphasis id="B91A7243FFDBFF95DABCC493FE4EF80B" box="[151,408,1953,1979]" italics="true" pageId="2" pageNumber="441">Trachygamasus karuni</emphasis>
</taxonomicName>
can be easily distinguished from these species due to the characteristic sternal (paragynal) poroids
<emphasis id="B91A7243FFDBFF95DAD9C4F7FEC2F86F" box="[242,276,1989,2015]" italics="true" pageId="2" pageNumber="441">
<date id="FFD08891FFDBFF95DAD9C4F7FEC2F86F" box="[242,276,1989,2015]" pageId="2" pageNumber="441">iv3</date>
</emphasis>
, which are roundish rather than elongate, as well as the shape of the saccular endogynium in the female. The males of the newly described species are distinguishable due to leg II spurs, one on femur II and one on genu II, but not on tibia II. To date,
<taxonomicName id="4C6ED5D2FFDAFF94DBE2C38CFDE6FF67" authority="Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński, 2019" authorityName="Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński" authorityYear="2019" box="[457,560,189,215]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="karuni" status="sp. nov.">
<emphasis id="B91A7243FFDAFF94DBE2C38CFE0FFF67" box="[457,473,190,215]" italics="true" pageId="3" pageNumber="442">T</emphasis>
.
<emphasis id="B91A7243FFDAFF94DBCCC38FFDE6FF67" box="[487,560,189,215]" italics="true" pageId="3" pageNumber="442">karuni</emphasis>
</taxonomicName>
<emphasis id="B91A7243FFDAFF94D81CC38FFD47FF67" bold="true" box="[567,657,189,215]" pageId="3" pageNumber="442">
<taxonomicNameLabel id="A229CF38FFDAFF94D81CC38FFD47FF67" box="[567,657,189,215]" pageId="3" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the only
<taxonomicName id="4C6ED5D2FFDAFF94D938C38FFC15FF67" box="[787,963,189,215]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="442" phylum="Arthropoda" rank="genus">
<emphasis id="B91A7243FFDAFF94D938C38FFC15FF67" box="[787,963,189,215]" italics="true" pageId="3" pageNumber="442">Trachygamasus</emphasis>
</taxonomicName>
this
<typeStatus id="54D510F3FFDAFF94D9D1C38FFBFCFF67" box="[1018,1066,189,215]" pageId="3" pageNumber="442">type</typeStatus>
of spur arrangement on leg II.
</paragraph>
<caption id="DF11FED9FFDAFF94DABCC5C7FC8EF8E6" ID-DOI="http://doi.org/10.5281/zenodo.3568898" ID-Zenodo-Dep="3568898" httpUri="https://zenodo.org/record/3568898/files/figure.png" pageId="3" pageNumber="442" startId="3.[151,250,1781,1806]" targetBox="[251,1340,262,1686]" targetPageId="3">
<paragraph id="8BD1AE51FFDAFF94DABCC5C7FC8EF8E6" blockId="3.[151,1437,1781,1878]" pageId="3" pageNumber="442">
<emphasis id="B91A7243FFDAFF94DABCC5C7FDA2F8BE" bold="true" box="[151,628,1781,1806]" pageId="3" pageNumber="442">
FIGURE 1.
<taxonomicName id="4C6ED5D2FFDAFF94DB0BC5C7FDC3F8BE" authority="Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński, 2019" authorityName="Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński" authorityYear="2019" box="[288,533,1781,1806]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="442" phylum="Arthropoda" rank="species" species="karuni" status="sp. nov.">
<emphasis id="B91A7243FFDAFF94DB0BC5C7FDC3F8BE" bold="true" box="[288,533,1781,1806]" italics="true" pageId="3" pageNumber="442">Trachygamasus karuni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229CF38FFDAFF94D80BC5C4FDA2F8BE" box="[544,628,1782,1806]" pageId="3" pageNumber="442" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. A. Dorsal side of idiosoma, paratype. Dorsal setae and setae series (left-hand side), as well as the dorsal poroids and gland pores (right-hand side) are marked and specifically acknowledged. Pseudoperforated cuticle is marked in the right lower part. B. Seta
<emphasis id="B91A7243FFDAFF94D910C40CFC87F8E6" box="[827,849,1854,1878]" italics="true" pageId="3" pageNumber="442">r3</emphasis>
.
</paragraph>
</caption>
<caption id="DF11FED9FFDDFF93DABCC42CFBF5F887" ID-DOI="http://doi.org/10.5281/zenodo.3568900" ID-Zenodo-Dep="3568900" box="[151,1059,1822,1847]" httpUri="https://zenodo.org/record/3568900/files/figure.png" pageId="4" pageNumber="443" startId="4.[151,250,1822,1847]" targetBox="[248,1346,216,1731]" targetPageId="4">
<paragraph id="8BD1AE51FFDDFF93DABCC42CFBF5F887" blockId="4.[151,1059,1822,1847]" box="[151,1059,1822,1847]" pageId="4" pageNumber="443">
<emphasis id="B91A7243FFDDFF93DABCC42CFDB4F887" bold="true" box="[151,610,1822,1847]" pageId="4" pageNumber="443">
FIGURE 2.
<taxonomicName id="4C6ED5D2FFDDFF93DB32C42CFDDDF887" authority="Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński, 2019" authorityName="Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński" authorityYear="2019" box="[281,523,1822,1847]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="443" phylum="Arthropoda" rank="species" species="karuni" status="sp. nov.">
<emphasis id="B91A7243FFDDFF93DB32C42CFDDDF887" bold="true" box="[281,523,1822,1847]" italics="true" pageId="4" pageNumber="443">Trachygamasus karuni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229CF38FFDDFF93D839C42DFDB4F887" box="[530,610,1823,1847]" pageId="4" pageNumber="443" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. Ventral side of idiosoma, paratype.
</paragraph>
</caption>
<caption id="DF11FED9FFDCFF92DABCC173FBD2FD12" ID-DOI="http://doi.org/10.5281/zenodo.3568902" ID-Zenodo-Dep="3568902" httpUri="https://zenodo.org/record/3568902/files/figure.png" pageId="5" pageNumber="444" startId="5.[151,264,577,602]" targetBox="[155,1430,169,554]" targetPageId="5">
<paragraph id="8BD1AE51FFDCFF92DABCC173FBD2FD12" blockId="5.[151,1437,577,674]" pageId="5" pageNumber="444">
<emphasis id="B91A7243FFDCFF92DABCC173FD43FDEA" bold="true" box="[151,661,577,602]" pageId="5" pageNumber="444">
FIGURES 35.
<taxonomicName id="4C6ED5D2FFDCFF92DB6EC173FDEFFDEA" authority="Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński, 2019" authorityName="Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński" authorityYear="2019" box="[325,569,577,602]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="karuni" status="sp. nov.">
<emphasis id="B91A7243FFDCFF92DB6EC173FDEFFDEA" bold="true" box="[325,569,577,602]" italics="true" pageId="5" pageNumber="444">Trachygamasus karuni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229CF38FFDCFF92D869C170FD43FDEA" box="[578,661,578,602]" pageId="5" pageNumber="444" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
3. Anterior part of podonotum in female with anterior ends of peritremes, podonotal setae, poroids
<emphasis id="B91A7243FFDCFF92DBBFC154FE6DFDCE" box="[404,443,614,638]" italics="true" pageId="5" pageNumber="444">idj1</emphasis>
and gland pores
<emphasis id="B91A7243FFDCFF92D84CC154FD42FDCE" box="[615,660,614,638]" italics="true" pageId="5" pageNumber="444">gdj2</emphasis>
, the last-named with double openings. 4 &amp; 5. Ventral view of
<taxonomicName id="4C6ED5D2FFDCFF92DF23C157FA54FDCE" authorityName="Krefft" authorityYear="1871" box="[1288,1410,613,638]" class="Chromadorea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="5" pageNumber="441" phylum="Nematoda" rank="genus">gnathosoma</taxonomicName>
in female (Figure 4) and male (Figure 5). Holotype: Figure 4; paratypes: Figures 3 and 5.
</paragraph>
</caption>
<caption id="DF11FED9FFDCFF92DABCC458FCA3F85F" ID-DOI="http://doi.org/10.5281/zenodo.3568904" ID-Zenodo-Dep="3568904" httpUri="https://zenodo.org/record/3568904/files/figure.png" pageId="5" pageNumber="444" startId="5.[151,264,1898,1923]" targetBox="[189,1398,721,1869]" targetPageId="5">
<paragraph id="8BD1AE51FFDCFF92DABCC458FCA3F85F" blockId="5.[151,1436,1898,2031]" pageId="5" pageNumber="444">
<emphasis id="B91A7243FFDCFF92DABCC458FD48F833" bold="true" box="[151,670,1898,1923]" pageId="5" pageNumber="444">
FIGURES 613.
<taxonomicName id="4C6ED5D2FFDCFF92DB7BC458FD95F833" authority="Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński, 2019" authorityName="Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński" authorityYear="2019" box="[336,579,1898,1923]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="444" phylum="Arthropoda" rank="species" species="karuni" status="sp. nov.">
<emphasis id="B91A7243FFDCFF92DB7BC458FD95F833" bold="true" box="[336,579,1898,1923]" italics="true" pageId="5" pageNumber="444">Trachygamasus karuni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229CF38FFDCFF92D867C459FD48F833" box="[588,670,1899,1923]" pageId="5" pageNumber="444" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female. 6. Presternal plates. 7. Genital region showing epigynium and an outline of the endogynium
<emphasis id="B91A7243FFDCFF92DB81C4BDFE3FF816" box="[426,489,1935,1958]" italics="true" pageId="5" pageNumber="444">in situ</emphasis>
. Setae and pores as in Figure 2. 8. Endogynium, two aspects. 9. Gnathotectum. 10. Chelicera antiaxially. 11. Coxa I ventrally. 12. Ambulacra on leg II (Figure 12 left) and leg III (Figure 12 right). 13. Tibia II, posterolateral view. Holotype: Figures 7 and 8 right; paratypes: Figures 6, 8 left, 913.
</paragraph>
</caption>
<caption id="DF11FED9FFDFFF91DABCC5C5FB01F8A0" ID-DOI="http://doi.org/10.5281/zenodo.3568906" ID-Zenodo-Dep="3568906" box="[151,1239,1783,1808]" httpUri="https://zenodo.org/record/3568906/files/figure.png" pageId="6" pageNumber="445" startId="6.[151,250,1783,1808]" targetBox="[293,1195,224,1660]" targetPageId="6">
<paragraph id="8BD1AE51FFDFFF91DABCC5C5FB01F8A0" blockId="6.[151,1239,1783,1808]" box="[151,1239,1783,1808]" pageId="6" pageNumber="445">
<emphasis id="B91A7243FFDFFF91DABCC5C5FDB9F8A0" bold="true" box="[151,623,1783,1808]" pageId="6" pageNumber="445">
FIGURE 14.
<taxonomicName id="4C6ED5D2FFDFFF91DB0DC5C5FDCEF8A0" authority="Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński, 2019" authorityName="Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński" authorityYear="2019" box="[294,536,1783,1808]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="445" phylum="Arthropoda" rank="species" species="karuni" status="sp. nov.">
<emphasis id="B91A7243FFDFFF91DB0DC5C5FDCEF8A0" bold="true" box="[294,536,1783,1808]" italics="true" pageId="6" pageNumber="445">Trachygamasus karuni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229CF38FFDFFF91D835C5CAFDB9F8A0" box="[542,623,1784,1808]" pageId="6" pageNumber="445" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. Dorsal side of idiosoma, abbreviations as in Figure 1.
</paragraph>
</caption>
<caption id="DF11FED9FFDEFF90DABCC4B6FB24F855" ID-DOI="http://doi.org/10.5281/zenodo.3568908" ID-Zenodo-Dep="3568908" httpUri="https://zenodo.org/record/3568908/files/figure.png" pageId="7" pageNumber="446" startId="7.[151,264,1924,1949]" targetBox="[164,1387,178,1897]" targetPageId="7">
<paragraph id="8BD1AE51FFDEFF90DABCC4B6FB24F855" blockId="7.[151,1437,1924,2021]" pageId="7" pageNumber="446">
<emphasis id="B91A7243FFDEFF90DABCC4B6FD75F82D" bold="true" box="[151,675,1924,1949]" pageId="7" pageNumber="446">
FIGURES 1519.
<taxonomicName id="4C6ED5D2FFDEFF90DB71C4B6FD9AF82D" authority="Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński, 2019" authorityName="Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński" authorityYear="2019" box="[346,588,1924,1949]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="446" phylum="Arthropoda" rank="species" species="karuni" status="sp. nov.">
<emphasis id="B91A7243FFDEFF90DB71C4B6FD9AF82D" bold="true" box="[346,588,1924,1949]" italics="true" pageId="7" pageNumber="446">Trachygamasus karuni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229CF38FFDEFF90D878C4B7FD75F82D" box="[595,675,1925,1949]" pageId="7" pageNumber="446" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male. 15. Ventral side of idiosoma, abbreviations as in Figure 2. 16. Anterior margin of sternum with genital lamina flanked by presternal plates. 17. Hypostome in ventral perspective showing corniculi, hypostomatic setae, palpcoxa seta and palptrochanter. 18. Gnathotectum, two aspects. 19. Chelicera adaxially.
</paragraph>
</caption>
<caption id="DF11FED9FFD1FF9FDABCC4B6FBC2F855" ID-DOI="http://doi.org/10.5281/zenodo.3568910" ID-Zenodo-Dep="3568910" httpUri="https://zenodo.org/record/3568910/files/figure.png" pageId="8" pageNumber="447" startId="8.[151,264,1924,1949]" targetBox="[166,1364,181,1889]" targetPageId="8">
<paragraph id="8BD1AE51FFD1FF9FDABCC4B6FBC2F855" blockId="8.[151,1437,1924,2021]" pageId="8" pageNumber="447">
<emphasis id="B91A7243FFD1FF9FDABCC4B6FD71F82D" bold="true" box="[151,679,1924,1949]" pageId="8" pageNumber="447">
FIGURES 2024.
<taxonomicName id="4C6ED5D2FFD1FF9FDB77C4B6FD98F82D" authority="Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński, 2019" authorityName="Farahi &amp; Shishehbor &amp; Nemati &amp; Witaliński" authorityYear="2019" box="[348,590,1924,1949]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="447" phylum="Arthropoda" rank="species" species="karuni" status="sp. nov.">
<emphasis id="B91A7243FFD1FF9FDB77C4B6FD98F82D" bold="true" box="[348,590,1924,1949]" italics="true" pageId="8" pageNumber="447">Trachygamasus karuni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A229CF38FFD1FF9FD87EC4B7FD71F82D" box="[597,679,1925,1949]" pageId="8" pageNumber="447" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, leg II. 20. Femur, genu and tibia in ventral perspective. 21. Femur, genu and tibia posterolaterally. 22. Spur on genu anterolaterally. 23 &amp; 24. Spurs on femur II and genu II, enlarged. Setae anterolateral (
<emphasis id="B91A7243FFD1FF9FDAD1C4FFFED8F855" box="[250,270,1997,2021]" italics="true" pageId="8" pageNumber="447">al</emphasis>
), posteroventral (
<emphasis id="B91A7243FFD1FF9FDBE8C4FCFE0DF855" box="[451,475,1998,2021]" italics="true" pageId="8" pageNumber="447">pv</emphasis>
), posterolateral (
<emphasis id="B91A7243FFD1FF9FD8A3C4FFFD4AF855" box="[648,668,1997,2021]" italics="true" pageId="8" pageNumber="447">pl</emphasis>
), and posterodorsal (
<emphasis id="B91A7243FFD1FF9FD958C4FFFC5AF855" box="[883,908,1997,2021]" italics="true" pageId="8" pageNumber="447">pd</emphasis>
) are marked.
</paragraph>
</caption>
<paragraph id="8BD1AE51FFD0FF9EDAECC3ABFD4DFEF3" blockId="9.[151,1436,153,323]" pageId="9" pageNumber="448">
The new species is the fifth species of
<taxonomicName id="4C6ED5D2FFD0FF9ED8AAC3ABFCE7FF03" box="[641,817,153,179]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="genus">
<emphasis id="B91A7243FFD0FF9ED8AAC3ABFCE7FF03" box="[641,817,153,179]" italics="true" pageId="9" pageNumber="448">Trachygamasus</emphasis>
</taxonomicName>
showing spurs on leg II in the male. The others are
<taxonomicName id="4C6ED5D2FFD0FF9EDFADC3A8FE5AFF67" authority="Ma &amp; Wang" authorityName="Ma &amp; Wang" authorityYear="1996" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B91A7243FFD0FF9EDFADC3A8FA40FF03" box="[1414,1430,154,179]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9EDABCC38FFF24FF67" box="[151,242,189,215]" italics="true" pageId="9" pageNumber="448">borealis</emphasis>
Ma &amp; Wang
</taxonomicName>
,
<taxonomicName id="4C6ED5D2FFD0FF9EDBB2C38CFD67FF67" authority="Tichomirov" authorityName="Tichomirov" box="[409,689,189,215]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="medianus">
<emphasis id="B91A7243FFD0FF9EDBB2C38CFE7FFF67" box="[409,425,190,215]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9EDB93C38FFDF5FF67" box="[440,547,189,215]" italics="true" pageId="9" pageNumber="448">medianus</emphasis>
Tichomirov
</taxonomicName>
,
<taxonomicName id="4C6ED5D2FFD0FF9ED895C38CFC0AFF67" authority="Witalinski, 2017" authorityName="Witalinski" authorityYear="2017" box="[702,988,189,215]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="gerdi">
<emphasis id="B91A7243FFD0FF9ED895C38CFD18FF67" box="[702,718,190,215]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9ED8F5C38FFCCEFF67" box="[734,792,189,215]" italics="true" pageId="9" pageNumber="448">gerdi</emphasis>
<bibRefCitation id="EFFFD3A0FFD0FF9ED90AC38FFC0AFF67" author="Witalinski, W." box="[801,988,189,215]" pageId="9" pageNumber="448" pagination="407 - 416" refId="ref8173" refString="Witalinski, W. (2017) A new species of Trachygamasus from Poland, a new definition of the genus, and a key to the world species (Parasitiformes: Parasitidae). Zootaxa, 4303 (3), 407 - 416. https: // doi. org / 10.11646 / zootaxa. 4303.3.6" type="journal article" year="2017">Witaliński, 2017</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C6ED5D2FFD0FF9EDE3CC38CFF05FF4B" authority="Yao, Jin &amp; Zhang, 2019" authorityName="Yao, Jin &amp; Zhang" authorityYear="2019" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="species" species="pseudogerdi">
<emphasis id="B91A7243FFD0FF9EDE3CC38CFBF1FF67" box="[1047,1063,190,215]" italics="true" pageId="9" pageNumber="448">T</emphasis>
.
<emphasis id="B91A7243FFD0FF9EDE1CC38FFB16FF67" box="[1079,1216,189,215]" italics="true" pageId="9" pageNumber="448">pseudogerdi</emphasis>
<bibRefCitation id="EFFFD3A0FFD0FF9EDEE2C38FFF05FF4B" author="Yao, M. &amp; Yi, T. - C. &amp; Guo, J. - J. &amp; Jin, D. - C. &amp; Zhang, R. - Z." pageId="9" pageNumber="448" pagination="1465 - 1489" refId="ref8231" refString="Yao, M., Yi, T. - C., Guo, J. - J., Jin, D. - C. &amp; Zhang, R. - Z. (2019) Three new species of Trachygamasus (Mesostigmata: Parasitidae) from China, with a key to world species of the genus. Systematic and Applied Acarology, 24, 1465 - 1489. https: // doi. org / 10.11158 / saa. 24.8.9" type="journal article" year="2019">Yao, Jin &amp; Zhang, 2019</bibRefCitation>
</taxonomicName>
. It should be emphasised that the spurs in
<taxonomicName id="4C6ED5D2FFD0FF9ED88BC3D3FC86FF4B" box="[672,848,225,251]" class="Arachnida" family="Parasitidae" genus="Trachygamasus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="448" phylum="Arthropoda" rank="genus">
<emphasis id="B91A7243FFD0FF9ED88BC3D3FC86FF4B" box="[672,848,225,251]" italics="true" pageId="9" pageNumber="448">Trachygamasus</emphasis>
</taxonomicName>
males are minute, even if present, and thus may have been overlooked, especially in earlier descriptions. It is the fourth species with a characteristic, hyaline uniramous anterior protrusion of the male genital lamina.
</paragraph>
</subSubSection>
</treatment>
</document>