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<document id="F14801C8887ABEBD433CF89D83E40DBB" ID-CLB-Dataset="9003" ID-DOI="10.11646/zootaxa.4903.3.6" ID-GBIF-Dataset="b1578e9d-8724-4bfe-8e55-7094e750f94d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4428032" ID-ZooBank="0F610083-8759-444F-96A0-F8FAD3D3A097" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610099967254" checkinUser="plazi" docAuthor="Jiang, Xuan-Kong, Zhang, Zu-Xu, Chen, Hui-Ming &amp; Xie, Zhi-Cai" docDate="2021" docId="03A74D388F6AFFC7A7DEF9BDFF6AFE80" docLanguage="en" docName="zootaxa.4903.3.6.pdf" docOrigin="Zootaxa 4903 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glyphiulus polytrichus Jiang &amp; Zhang &amp; Chen &amp; Xie 2021, new combination" docType="treatment" docVersion="7" lastPageNumber="414" masterDocId="FF9E35408F62FFCDA749FF8BFF9AFFE5" masterDocTitle="Description of Hypocambala zizhongi sp. nov. and the new combination, Glyphiulus polytrichus (Golovatch et al., 2011) comb. nov., based on morphological and molecular data (Spirostreptida: Cambalidea: Cambalopsidae)" masterLastPageNumber="418" masterPageNumber="405" pageNumber="413" updateTime="1698924094466" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4B8F8CAE7934CBAD8D5DD1830996C4FA">Description of Hypocambala zizhongi sp. nov. and the new combination, Glyphiulus polytrichus (Golovatch et al., 2011) comb. nov., based on morphological and molecular data (Spirostreptida: Cambalidea: Cambalopsidae)</mods:title>
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<mods:namePart id="0A12E2D5F0C536F24F8F57E79F07747A">Jiang, Xuan-Kong</mods:namePart>
<mods:affiliation id="351EB70FB10AF3379ED8F92EF55D1C1D">Institute of Biology, Guizhou Academy of Sciences, Guiyang 550009, China &amp; CAS Key Laboratory of Aquatic Biodiversity and Conservation, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan 430072, China &amp; University of Chinese Academy of Sciences, Beijing 100049, China</mods:affiliation>
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<mods:namePart id="378B508151D9FFF654E119AA16635C01">Zhang, Zu-Xu</mods:namePart>
<mods:nameIdentifier id="3C01690FE3C67947D41FC5BC6C058390" type="ORCID">0000-0002-8439-0231</mods:nameIdentifier>
<mods:affiliation id="0B98AA41BFB2C0EF17FD494E81841845">Institute of Biology, Guizhou Academy of Sciences, Guiyang 550009, China &amp; 826011851 @ qq. com; https: // orcid. org / 0000 - 0002 - 8439 - 0231</mods:affiliation>
<mods:nameIdentifier id="E66905009DF0319D602CBF17A02A7A03" type="email">826011851@qq.com</mods:nameIdentifier>
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<mods:namePart id="5150CF329E7986CCFD169E66929C4B80">Chen, Hui-Ming</mods:namePart>
<mods:nameIdentifier id="B4D8AF59031712D8C7718175F9FB263A" type="ORCID">0000-0002-2449-3036</mods:nameIdentifier>
<mods:affiliation id="750821B2BD0D2E3A0835EB7EB5E63A0E">Institute of Biology, Guizhou Academy of Sciences, Guiyang 550009, China &amp; mei 0601 @ 126. com; https: // orcid. org / 0000 - 0002 - 2449 - 3036</mods:affiliation>
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<mods:namePart id="9DA865C7B6C664113A5AC930782D04C2">Xie, Zhi-Cai</mods:namePart>
<mods:nameIdentifier id="66D0610A07B475F98DF036340FC9F36B" type="ORCID">0000-0001-9658-4888</mods:nameIdentifier>
<mods:affiliation id="EF418B3317FD615424BD1A3058F72762">CAS Key Laboratory of Aquatic Biodiversity and Conservation, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan 430072, China &amp; zhcxie @ ihb. ac. cn; https: // orcid. org / 0000 - 0001 - 9658 - 4888</mods:affiliation>
<mods:nameIdentifier id="8F80E1DDA567767019EEB9E13BA91CF8" type="email">zhcxie@ihb.ac.cn</mods:nameIdentifier>
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<mods:title id="2FFEC070FBC875E737D26BE8F72E3DDB">Zootaxa</mods:title>
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<mods:date id="DFC821403CA6230769AC33951D7A115D">2021</mods:date>
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<mods:identifier id="9475C2F49257B36BEEB0E1301D99991A" type="DOI">10.11646/zootaxa.4903.3.6</mods:identifier>
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<treatment id="03A74D388F6AFFC7A7DEF9BDFF6AFE80" ID-DOI="http://doi.org/10.5281/zenodo.4428049" ID-GBIF-Taxon="176566830" ID-Zenodo-Dep="4428049" LSID="urn:lsid:plazi:treatment:03A74D388F6AFFC7A7DEF9BDFF6AFE80" httpUri="http://treatment.plazi.org/id/03A74D388F6AFFC7A7DEF9BDFF6AFE80" lastPageId="10" lastPageNumber="414" pageId="8" pageNumber="413">
<subSubSection id="C314AFA58F6AFFC5A7DEF9BDFAC7F9B5" box="[151,1373,1590,1617]" pageId="8" pageNumber="413" type="nomenclature">
<paragraph id="8BB1FC2E8F6AFFC5A7DEF9BDFAC7F9B5" blockId="8.[151,1373,1590,1653]" box="[151,1373,1590,1617]" pageId="8" pageNumber="413">
<heading id="D0F94B428F6AFFC5A7DEF9BDFAC7F9B5" bold="true" box="[151,1373,1590,1617]" fontSize="11" level="1" pageId="8" pageNumber="413" reason="1">
<emphasis id="B97A203C8F6AFFC5A7DEF9BDFAC7F9B5" bold="true" box="[151,1373,1590,1617]" pageId="8" pageNumber="413">
<taxonomicName id="4C0E87AD8F6AFFC5A7DEF9BDFBE0F9B5" authority="(Golovatch, Geoffroy, Mauries and VandenSpiegel, 2011)" authorityName="Jiang &amp; Zhang &amp; Chen &amp; Xie" authorityYear="2021" baseAuthorityName="Golovatch, Geoffroy, Mauries and VandenSpiegel" baseAuthorityYear="2011" box="[151,1146,1590,1617]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="413" phylum="Arthropoda" rank="species" species="polytrichus" status="comb. nov.">
<emphasis id="B97A203C8F6AFFC5A7DEF9BDFE31F9B4" bold="true" box="[151,427,1590,1617]" italics="true" pageId="8" pageNumber="413">Glyphiulus polytrichus</emphasis>
(Golovatch, Geoffroy, Mauriès and VandenSpiegel, 2011)
</taxonomicName>
,
<taxonomicNameLabel id="A2499D478F6AFFC5A3CEF9BCFAC7F9B5" box="[1159,1373,1590,1617]" pageId="8" pageNumber="413" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C314AFA58F6AFFC5A7DEF9D0FEABF990" box="[151,305,1627,1653]" pageId="8" pageNumber="413" type="description">
<paragraph id="8BB1FC2E8F6AFFC5A7DEF9D0FEABF990" blockId="8.[151,1373,1590,1653]" box="[151,305,1627,1653]" pageId="8" pageNumber="413">
<figureCitation id="1335E0AB8F6AFFC5A7DEF9D0FF44F990" box="[151,222,1627,1653]" captionStart="FIGURES 1–2" captionStartId="1.[151,264,1355,1379]" captionTargetBox="[151,1435,918,1329]" captionTargetId="figure-275@1.[151,1435,918,1329]" captionTargetPageId="1" captionText="FIGURES 1–2. Habitus of Hypocambala zizhongi sp. nov. and Glyphiulus polytrichus (Golovatch et al., 2011) comb. nov. 1. Hypocambala zizhongi sp. nov.; 2. Glyphiulus polytrichus (Golovatch et al., 2011) comb. nov." figureDoi="http://doi.org/10.5281/zenodo.4428034" httpUri="https://zenodo.org/record/4428034/files/figure.png" pageId="8" pageNumber="413">Figs 2</figureCitation>
,
<figureCitation id="1335E0AB8F6AFFC5A7A3F9D0FEABF990" box="[234,305,1627,1653]" captionStart="FIGURES 23–34" captionStartId="9.[151,264,1634,1658]" captionTargetBox="[151,1435,381,1611]" captionTargetId="figure-135@9.[151,1435,381,1611]" captionTargetPageId="9" captionText="FIGURES 23–34. Male of Glyphiulus polytrichus (Golovatch et al., 2011) comb. nov. 23. anterior part of body, dorsal view; 24. same, lateral view; 25. midbody rings, dorsal view; 26. same, lateral view; 27. posterior part of body, dorsal view; 28. same, lateral view; 29. legs I, posterior view; 30. coxosternal processes of legs I, posterior view; 31. anterior gonopods, posterior view; 32. coxite process of anterior gonopod, posterior view; 33. posterior gonopods, anterior view; 34. flagellum of posterior gonopod, anterior view.Abbreviations: csp, coxosternal process; cx, coxite; cxp, coxite process; f, flagellum; s, sternite; t, telopodite. Scale bars: 23–26 = 0.5 mm; 27–29, 31 = 0.2 mm; 30 = 0.05 mm; 32 = 0.4 mm; 33 = 0.1 mm; 34 = 0.04 mm." figureDoi="http://doi.org/10.5281/zenodo.4428045" httpUri="https://zenodo.org/record/4428045/files/figure.png" pageId="8" pageNumber="413">23–34</figureCitation>
</paragraph>
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<paragraph id="8BB1FC2E8F6AFFC5A7DEF914FB83F952" blockId="8.[151,1049,1695,1719]" box="[151,1049,1695,1719]" pageId="8" pageNumber="413">
<treatmentCitationGroup id="AB1EDB008F6AFFC5A7DEF914FB83F952" box="[151,1049,1695,1719]" pageId="8" pageNumber="413">
<treatmentCitation id="0AAFDA3F8F6AFFC5A7DEF914FB8FF952" author="Golovatch, S. I. &amp; Geoffroy, J. - J. &amp; Mauries, J. - P. &amp; VandenSpiegel, D." box="[151,1045,1695,1719]" page="169" pageId="8" pageNumber="413" year="2011">
<taxonomicName id="4C0E87AD8F6AFFC5A7DEF914FB8FF952" authority="Golovatch, Geoffroy, Mauries and VandenSpiegel, 2011 b: 169" authorityName="Golovatch, Geoffroy, Mauries and VandenSpiegel" authorityPageNumber="169" authorityYear="2011" box="[151,1045,1695,1719]" class="Diplopoda" family="Glyphiulidae" genus="Hypocambala" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="413" phylum="Arthropoda" rank="species" species="polytricha">
<emphasis id="B97A203C8F6AFFC5A7DEF914FE0EF952" box="[151,404,1695,1719]" italics="true" pageId="8" pageNumber="413">Hypocambala polytricha</emphasis>
<bibRefCitation id="EF9F81DF8F6AFFC5A6D3F914FB8FF952" author="Golovatch, S. I. &amp; Geoffroy, J. - J. &amp; Mauries, J. - P. &amp; VandenSpiegel, D." box="[410,1045,1695,1719]" pageId="8" pageNumber="413" pagination="167 - 174" refId="ref7307" refString="Golovatch, S. I., Geoffroy, J. - J., Mauries, J. - P. &amp; VandenSpiegel, D. (2011 b) Two new species of the millipede genus Hypocambala Silvestri, 1895 from China and Vietnam (Diplopoda: Spirostreptida: Cambalopsidae). Arthropoda Selecta, 20 (3), 167 - 174. https: // doi. org / 10.15298 / arthsel. 20.3.03" type="journal article" year="2011">Golovatch, Geoffroy, Mauriès and VandenSpiegel, 2011b: 169</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C314AFA58F6AFFC5A7DEF96EFD5DF8A2" pageId="8" pageNumber="413" type="materials_examined">
<paragraph id="8BB1FC2E8F6AFFC5A7DEF96EFD5DF8A2" blockId="8.[151,1437,1765,2008]" pageId="8" pageNumber="413">
<emphasis id="B97A203C8F6AFFC5A7DEF96EFEE2F91A" bold="true" box="[151,376,1765,1791]" pageId="8" pageNumber="413">Material examined</emphasis>
:
<materialsCitation id="3B66F6738F6AFFC5A6CFF96EFD59F8A2" ID-GBIF-Occurrence="3014733303" collectingDate="2018-10-18" collectionCode="IBGAS" collectorName="X. K. Jiang &amp; H. M. Chen" country="China" county="Longzhou County" elevation="359" latitude="22.672297" location="Jinlong Town" longLatPrecision="1" longitude="106.769005" municipality="Autonomous Region" pageId="8" pageNumber="413" specimenCount="30" specimenCount-female="10" specimenCount-juvenile="6" specimenCount-male="14" stateProvince="Guangxi">
<specimenCount id="9D0837A78F6AFFC5A6CFF96EFE71F91A" box="[390,491,1765,1791]" pageId="8" pageNumber="413" type="male">14 males</specimenCount>
,
<specimenCount id="9D0837A78F6AFFC5A6BCF96EFDF4F91A" box="[501,622,1765,1791]" pageId="8" pageNumber="413" type="female">10 females</specimenCount>
and
<specimenCount id="9D0837A78F6AFFC5A5EBF96EFC84F91A" box="[674,798,1765,1791]" pageId="8" pageNumber="413" type="juvenile">6 juveniles</specimenCount>
,
<collectingCountry id="F319BCBE8F6AFFC5A46EF96EFCF7F91A" box="[807,877,1765,1791]" name="China" pageId="8" pageNumber="413">China</collectingCountry>
,
<collectingRegion id="49CA32CC8F6AFFC5A43EF96EFBA9F91A" box="[887,1075,1765,1791]" country="China" name="Guangxi" pageId="8" pageNumber="413">Guangxi Zhuang</collectingRegion>
<collectingMunicipality id="6BD566548F6AFFC5A37EF96DFABBF91A" box="[1079,1313,1765,1792]" pageId="8" pageNumber="413">Autonomous Region</collectingMunicipality>
,
<collectingCounty id="62D084A28F6AFFC5A262F96EFF71F8C6" pageId="8" pageNumber="413">Longzhou County</collectingCounty>
,
<location id="8ED1AAF58F6AFFC5A7BCF882FE0AF8C1" LSID="urn:lsid:plazi:treatment:03A74D388F6AFFC7A7DEF9BDFF6AFE80:8ED1AAF58F6AFFC5A7BCF882FE0AF8C1" box="[245,400,1801,1828]" country="China" county="Longzhou County" latitude="22.672297" longLatPrecision="1" longitude="106.769005" municipality="Autonomous Region" name="Jinlong Town" pageId="8" pageNumber="413" stateProvince="Guangxi">Jinlong Town</location>
,
<location id="8ED1AAF58F6AFFC5A6D2F882FDCFF8C6" LSID="urn:lsid:plazi:treatment:03A74D388F6AFFC7A7DEF9BDFF6AFE80:8ED1AAF58F6AFFC5A6D2F882FDCFF8C6" box="[411,597,1801,1827]" country="China" county="Longzhou County" latitude="22.672297" longLatPrecision="1" longitude="106.769005" municipality="Autonomous Region" name="Hengluo Village" pageId="8" pageNumber="413" stateProvince="Guangxi">Hengluo Village</location>
,
<location id="8ED1AAF58F6AFFC5A516F882FD40F8C6" LSID="urn:lsid:plazi:treatment:03A74D388F6AFFC7A7DEF9BDFF6AFE80:8ED1AAF58F6AFFC5A516F882FD40F8C6" box="[607,730,1801,1827]" country="China" county="Longzhou County" latitude="22.672297" longLatPrecision="1" longitude="106.769005" municipality="Autonomous Region" name="Zhawaitun" pageId="8" pageNumber="413" stateProvince="Guangxi">Zhawaitun</location>
,
<location id="8ED1AAF58F6AFFC5A5ADF881FCC8F8C6" LSID="urn:lsid:plazi:treatment:03A74D388F6AFFC7A7DEF9BDFF6AFE80:8ED1AAF58F6AFFC5A5ADF881FCC8F8C6" box="[740,850,1801,1828]" country="China" county="Longzhou County" latitude="22.672297" longLatPrecision="1" longitude="106.769005" municipality="Autonomous Region" name="Yan Cave" pageId="8" pageNumber="413" stateProvince="Guangxi">Yan Cave</location>
,
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,
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, alt.
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<elevation id="00231B1D8F6AFFC5A3AFF882FAB5F8C1" box="[1254,1327,1801,1828]" metricMagnitude="2" metricUnit="m" metricValue="3.59" pageId="8" pageNumber="413" unit="m" value="359.0">359 m</elevation>
</quantity>
,
<date id="FFB0DAEE8F6AFFC5A273F882FF67F8A2" pageId="8" pageNumber="413" value="2018-10-18">
<collectingDate id="EFF423068F6AFFC5A273F882FF67F8A2" pageId="8" pageNumber="413" value="2018-10-18">18 October 2018</collectingDate>
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,
<collectorName id="26FB99F88F6AFFC5A641F8A5FE1AF8A2" box="[264,384,1837,1864]" pageId="8" pageNumber="413">X.K. Jiang</collectorName>
&amp;
<collectorName id="26FB99F88F6AFFC5A6EDF8A5FDBBF8A2" box="[420,545,1837,1864]" pageId="8" pageNumber="413">H.M. Chen</collectorName>
leg. (
<collectionCode id="ED1F64EB8F6AFFC5A528F8A6FD21F8A2" box="[609,699,1837,1863]" pageId="8" pageNumber="413">IBGAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C314AFA58F6AFFC4A78EF8DAFC14FE80" lastPageId="9" lastPageNumber="414" pageId="8" pageNumber="413" type="diagnosis">
<paragraph id="8BB1FC2E8F6AFFC4A78EF8DAFC14FE80" blockId="8.[151,1437,1765,2008]" lastBlockId="9.[151,1437,151,357]" lastPageId="9" lastPageNumber="414" pageId="8" pageNumber="413">
<emphasis id="B97A203C8F6AFFC5A78EF8DAFEA0F88E" bold="true" box="[199,314,1873,1899]" pageId="8" pageNumber="413">Diagnosis</emphasis>
: This species can be distinguished from the
<taxonomicName id="4C0E87AD8F6AFFC5A47EF8D9FC49F88E" baseAuthorityName="Gervais" baseAuthorityYear="1847" box="[823,979,1873,1900]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="413" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B97A203C8F6AFFC5A47EF8D9FC49F88E" box="[823,979,1873,1900]" italics="true" pageId="8" pageNumber="413">G. granulatus</emphasis>
</taxonomicName>
-group by the male legs I with complete telopodites and the coxosternal processes situated close to each other (
<figureCitation id="1335E0AB8F6AFFC5A4F7F8FEFBDFF86A" box="[958,1093,1909,1935]" captionStart="FIGURES 23–34" captionStartId="9.[151,264,1634,1658]" captionTargetBox="[151,1435,381,1611]" captionTargetId="figure-135@9.[151,1435,381,1611]" captionTargetPageId="9" captionText="FIGURES 23–34. Male of Glyphiulus polytrichus (Golovatch et al., 2011) comb. nov. 23. anterior part of body, dorsal view; 24. same, lateral view; 25. midbody rings, dorsal view; 26. same, lateral view; 27. posterior part of body, dorsal view; 28. same, lateral view; 29. legs I, posterior view; 30. coxosternal processes of legs I, posterior view; 31. anterior gonopods, posterior view; 32. coxite process of anterior gonopod, posterior view; 33. posterior gonopods, anterior view; 34. flagellum of posterior gonopod, anterior view.Abbreviations: csp, coxosternal process; cx, coxite; cxp, coxite process; f, flagellum; s, sternite; t, telopodite. Scale bars: 23–26 = 0.5 mm; 27–29, 31 = 0.2 mm; 30 = 0.05 mm; 32 = 0.4 mm; 33 = 0.1 mm; 34 = 0.04 mm." figureDoi="http://doi.org/10.5281/zenodo.4428045" httpUri="https://zenodo.org/record/4428045/files/figure.png" pageId="8" pageNumber="413">Figs 29, 30</figureCitation>
). It can be distinguished from other species of the
<taxonomicName id="4C0E87AD8F6AFFC5A6C9F811FD8AF856" authorityName="Carl" authorityYear="1911" box="[384,528,1946,1972]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="413" phylum="Arthropoda" rank="species" species="javanicus">
<emphasis id="B97A203C8F6AFFC5A6C9F811FD8AF856" box="[384,528,1946,1972]" italics="true" pageId="8" pageNumber="413">G. javanicus</emphasis>
</taxonomicName>
-group except
<taxonomicName id="4C0E87AD8F6AFFC5A5FEF812FB35F856" authority="Golovatch et al., 2007" authorityName="Golovatch" authorityYear="2007" box="[695,1199,1945,1972]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="413" phylum="Arthropoda" rank="species" species="costulifer">
<emphasis id="B97A203C8F6AFFC5A5FEF812FC3FF856" box="[695,933,1945,1971]" italics="true" pageId="8" pageNumber="413">Glyphiulus costulifer</emphasis>
Golovatch
<emphasis id="B97A203C8F6AFFC5A365F811FBFDF856" box="[1068,1127,1945,1971]" italics="true" pageId="8" pageNumber="413">et al.</emphasis>
, 2007
</taxonomicName>
and
<taxonomicName id="4C0E87AD8F6AFFC5A3A3F811FE0BF832" authority="Golovatch et al., 2007" authorityName="Golovatch" authorityYear="2007" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="8" pageNumber="413" phylum="Arthropoda" rank="species" species="percostulifer">
<emphasis id="B97A203C8F6AFFC5A3A3F811FA06F856" box="[1258,1436,1945,1972]" italics="true" pageId="8" pageNumber="413">G. percostulifer</emphasis>
Golovatch
<emphasis id="B97A203C8F6AFFC5A65AF835FED1F832" box="[275,331,1981,2007]" italics="true" pageId="8" pageNumber="413">et al.</emphasis>
, 2007
</taxonomicName>
by the male legs I with complete telopodites and the relatively more strongly separated coxosternal processes (
<figureCitation id="1335E0AB8F6BFFC4A63CFF1CFE66FF54" box="[373,508,151,177]" captionStart="FIGURES 23–34" captionStartId="9.[151,264,1634,1658]" captionTargetBox="[151,1435,381,1611]" captionTargetId="figure-135@9.[151,1435,381,1611]" captionTargetPageId="9" captionText="FIGURES 23–34. Male of Glyphiulus polytrichus (Golovatch et al., 2011) comb. nov. 23. anterior part of body, dorsal view; 24. same, lateral view; 25. midbody rings, dorsal view; 26. same, lateral view; 27. posterior part of body, dorsal view; 28. same, lateral view; 29. legs I, posterior view; 30. coxosternal processes of legs I, posterior view; 31. anterior gonopods, posterior view; 32. coxite process of anterior gonopod, posterior view; 33. posterior gonopods, anterior view; 34. flagellum of posterior gonopod, anterior view.Abbreviations: csp, coxosternal process; cx, coxite; cxp, coxite process; f, flagellum; s, sternite; t, telopodite. Scale bars: 23–26 = 0.5 mm; 27–29, 31 = 0.2 mm; 30 = 0.05 mm; 32 = 0.4 mm; 33 = 0.1 mm; 34 = 0.04 mm." figureDoi="http://doi.org/10.5281/zenodo.4428045" httpUri="https://zenodo.org/record/4428045/files/figure.png" pageId="9" pageNumber="414">Figs 29, 30</figureCitation>
); crests on medial parts of collum slightly degenerated with a carinotaxic pattern of I–IV+5a+pc+ma (
<figureCitation id="1335E0AB8F6BFFC4A6CAFF30FD9CFF30" box="[387,518,187,213]" captionStart="FIGURES 23–34" captionStartId="9.[151,264,1634,1658]" captionTargetBox="[151,1435,381,1611]" captionTargetId="figure-135@9.[151,1435,381,1611]" captionTargetPageId="9" captionText="FIGURES 23–34. Male of Glyphiulus polytrichus (Golovatch et al., 2011) comb. nov. 23. anterior part of body, dorsal view; 24. same, lateral view; 25. midbody rings, dorsal view; 26. same, lateral view; 27. posterior part of body, dorsal view; 28. same, lateral view; 29. legs I, posterior view; 30. coxosternal processes of legs I, posterior view; 31. anterior gonopods, posterior view; 32. coxite process of anterior gonopod, posterior view; 33. posterior gonopods, anterior view; 34. flagellum of posterior gonopod, anterior view.Abbreviations: csp, coxosternal process; cx, coxite; cxp, coxite process; f, flagellum; s, sternite; t, telopodite. Scale bars: 23–26 = 0.5 mm; 27–29, 31 = 0.2 mm; 30 = 0.05 mm; 32 = 0.4 mm; 33 = 0.1 mm; 34 = 0.04 mm." figureDoi="http://doi.org/10.5281/zenodo.4428045" httpUri="https://zenodo.org/record/4428045/files/figure.png" pageId="9" pageNumber="414">Figs 23, 24</figureCitation>
) (see
<bibRefCitation id="EF9F81DF8F6BFFC4A505FF30FCC8FF30" author="Golovatch, S. I. &amp; Geoffroy, J. - J. &amp; Mauries, J. - P. &amp; VandenSpiegel, D." box="[588,850,187,213]" pageId="9" pageNumber="414" pagination="7 - 49" refId="ref7077" refString="Golovatch, S. I., Geoffroy, J. - J., Mauries, J. - P. &amp; VandenSpiegel, D. (2007 a) Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1. The granulatus - group. Zoosystema, 29 (1), 7 - 49." type="journal article" year="2007">
Golovatch
<emphasis id="B97A203C8F6BFFC4A583FF30FC99FF30" box="[714,771,187,213]" italics="true" pageId="9" pageNumber="414">et al.</emphasis>
2007a
</bibRefCitation>
, b for an explanation of this formula); flagellum of posterior gonopod short (
<figureCitation id="1335E0AB8F6BFFC4A6E6FF54FDAAFF1C" box="[431,560,223,249]" captionStart="FIGURES 23–34" captionStartId="9.[151,264,1634,1658]" captionTargetBox="[151,1435,381,1611]" captionTargetId="figure-135@9.[151,1435,381,1611]" captionTargetPageId="9" captionText="FIGURES 23–34. Male of Glyphiulus polytrichus (Golovatch et al., 2011) comb. nov. 23. anterior part of body, dorsal view; 24. same, lateral view; 25. midbody rings, dorsal view; 26. same, lateral view; 27. posterior part of body, dorsal view; 28. same, lateral view; 29. legs I, posterior view; 30. coxosternal processes of legs I, posterior view; 31. anterior gonopods, posterior view; 32. coxite process of anterior gonopod, posterior view; 33. posterior gonopods, anterior view; 34. flagellum of posterior gonopod, anterior view.Abbreviations: csp, coxosternal process; cx, coxite; cxp, coxite process; f, flagellum; s, sternite; t, telopodite. Scale bars: 23–26 = 0.5 mm; 27–29, 31 = 0.2 mm; 30 = 0.05 mm; 32 = 0.4 mm; 33 = 0.1 mm; 34 = 0.04 mm." figureDoi="http://doi.org/10.5281/zenodo.4428045" httpUri="https://zenodo.org/record/4428045/files/figure.png" pageId="9" pageNumber="414">Figs 33, 34</figureCitation>
). The new species can be separated from
<taxonomicName id="4C0E87AD8F6BFFC4A4B3FF54FB1FFF1C" box="[1018,1157,223,249]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="414" phylum="Arthropoda" rank="species" species="costulifer">
<emphasis id="B97A203C8F6BFFC4A4B3FF54FB1FFF1C" box="[1018,1157,223,249]" italics="true" pageId="9" pageNumber="414">G. costulifer</emphasis>
</taxonomicName>
by crest 5a being as long as crest ma on collum (
<figureCitation id="1335E0AB8F6BFFC4A6D1FE88FE71FEF8" box="[408,491,259,285]" captionStart="FIGURES 23–34" captionStartId="9.[151,264,1634,1658]" captionTargetBox="[151,1435,381,1611]" captionTargetId="figure-135@9.[151,1435,381,1611]" captionTargetPageId="9" captionText="FIGURES 23–34. Male of Glyphiulus polytrichus (Golovatch et al., 2011) comb. nov. 23. anterior part of body, dorsal view; 24. same, lateral view; 25. midbody rings, dorsal view; 26. same, lateral view; 27. posterior part of body, dorsal view; 28. same, lateral view; 29. legs I, posterior view; 30. coxosternal processes of legs I, posterior view; 31. anterior gonopods, posterior view; 32. coxite process of anterior gonopod, posterior view; 33. posterior gonopods, anterior view; 34. flagellum of posterior gonopod, anterior view.Abbreviations: csp, coxosternal process; cx, coxite; cxp, coxite process; f, flagellum; s, sternite; t, telopodite. Scale bars: 23–26 = 0.5 mm; 27–29, 31 = 0.2 mm; 30 = 0.05 mm; 32 = 0.4 mm; 33 = 0.1 mm; 34 = 0.04 mm." figureDoi="http://doi.org/10.5281/zenodo.4428045" httpUri="https://zenodo.org/record/4428045/files/figure.png" pageId="9" pageNumber="414">Fig. 23</figureCitation>
), whereas crest 5a is three times as long as ma in
<taxonomicName id="4C0E87AD8F6BFFC4A359FE88FB01FEF8" box="[1040,1179,259,285]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="414" phylum="Arthropoda" rank="species" species="costulifer">
<emphasis id="B97A203C8F6BFFC4A359FE88FB01FEF8" box="[1040,1179,259,285]" italics="true" pageId="9" pageNumber="414">G. costulifer</emphasis>
</taxonomicName>
; flagellum of posterior gonopod pilose (
<figureCitation id="1335E0AB8F6BFFC4A619FEACFE39FEA4" box="[336,419,295,321]" captionStart="FIGURES 23–34" captionStartId="9.[151,264,1634,1658]" captionTargetBox="[151,1435,381,1611]" captionTargetId="figure-135@9.[151,1435,381,1611]" captionTargetPageId="9" captionText="FIGURES 23–34. Male of Glyphiulus polytrichus (Golovatch et al., 2011) comb. nov. 23. anterior part of body, dorsal view; 24. same, lateral view; 25. midbody rings, dorsal view; 26. same, lateral view; 27. posterior part of body, dorsal view; 28. same, lateral view; 29. legs I, posterior view; 30. coxosternal processes of legs I, posterior view; 31. anterior gonopods, posterior view; 32. coxite process of anterior gonopod, posterior view; 33. posterior gonopods, anterior view; 34. flagellum of posterior gonopod, anterior view.Abbreviations: csp, coxosternal process; cx, coxite; cxp, coxite process; f, flagellum; s, sternite; t, telopodite. Scale bars: 23–26 = 0.5 mm; 27–29, 31 = 0.2 mm; 30 = 0.05 mm; 32 = 0.4 mm; 33 = 0.1 mm; 34 = 0.04 mm." figureDoi="http://doi.org/10.5281/zenodo.4428045" httpUri="https://zenodo.org/record/4428045/files/figure.png" pageId="9" pageNumber="414">Fig. 34</figureCitation>
),
<emphasis id="B97A203C8F6BFFC4A6FFFEACFE4EFEA5" box="[438,468,295,320]" italics="true" pageId="9" pageNumber="414">vs.</emphasis>
smooth in
<taxonomicName id="4C0E87AD8F6BFFC4A507FEACFD40FEA4" box="[590,730,295,321]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="414" phylum="Arthropoda" rank="species" species="costulifer">
<emphasis id="B97A203C8F6BFFC4A507FEACFD40FEA4" box="[590,730,295,321]" italics="true" pageId="9" pageNumber="414">G. costulifer</emphasis>
</taxonomicName>
. It can be separated from
<taxonomicName id="4C0E87AD8F6BFFC4A4BEFEACFB3CFEA4" box="[1015,1190,295,321]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="414" phylum="Arthropoda" rank="species" species="percostulifer">
<emphasis id="B97A203C8F6BFFC4A4BEFEACFB3CFEA4" box="[1015,1190,295,321]" italics="true" pageId="9" pageNumber="414">G. percostulifer</emphasis>
</taxonomicName>
by the slender telopodites of the anterior gonopod,
<emphasis id="B97A203C8F6BFFC4A6A0FEC0FD9DFE81" box="[489,519,331,356]" italics="true" pageId="9" pageNumber="414">vs.</emphasis>
short and stout in
<taxonomicName id="4C0E87AD8F6BFFC4A59EFEC0FC1DFE80" box="[727,903,331,357]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="414" phylum="Arthropoda" rank="species" species="percostulifer">
<emphasis id="B97A203C8F6BFFC4A59EFEC0FC1DFE80" box="[727,903,331,357]" italics="true" pageId="9" pageNumber="414">G. percostulifer</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF71ACA68F6BFFC4A7DEF9E9FB76F8CB" ID-DOI="http://doi.org/10.5281/zenodo.4428045" ID-Zenodo-Dep="4428045" httpUri="https://zenodo.org/record/4428045/files/figure.png" pageId="9" pageNumber="414" startId="9.[151,264,1634,1658]" targetBox="[151,1435,381,1611]" targetPageId="9">
<paragraph id="8BB1FC2E8F6BFFC4A7DEF9E9FB76F8CB" blockId="9.[151,1437,1634,1839]" pageId="9" pageNumber="414">
<emphasis id="B97A203C8F6BFFC4A7DEF9E9FECFF99F" bold="true" box="[151,341,1634,1658]" pageId="9" pageNumber="414">FIGURES 23–34.</emphasis>
Male of
<taxonomicName id="4C0E87AD8F6BFFC4A6FAF9E9FC0CF99E" authority="(Golovatch et al., 2011)" authorityName="Jiang &amp; Zhang &amp; Chen &amp; Xie" authorityYear="2021" baseAuthorityName="Jiang &amp; Zhang &amp; Chen &amp; Xie" baseAuthorityYear="2011" box="[435,918,1634,1659]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="414" phylum="Arthropoda" rank="species" species="polytrichus" status="comb. nov.">
<emphasis id="B97A203C8F6BFFC4A6FAF9E9FD01F99F" box="[435,667,1634,1658]" italics="true" pageId="9" pageNumber="414">Glyphiulus polytrichus</emphasis>
(Golovatch
<emphasis id="B97A203C8F6BFFC4A452F9E8FCD5F99F" box="[795,847,1634,1658]" italics="true" pageId="9" pageNumber="414">et al.</emphasis>
, 2011)
</taxonomicName>
<emphasis id="B97A203C8F6BFFC4A4D4F9E8FB88F99E" bold="true" box="[925,1042,1635,1659]" pageId="9" pageNumber="414">
<taxonomicNameLabel id="A2499D478F6BFFC4A4D4F9E8FB88F99E" box="[925,1042,1635,1659]" pageId="9" pageNumber="414" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
23. anterior part of body, dorsal view; 24. same, lateral view; 25. midbody rings, dorsal view; 26. same, lateral view; 27. posterior part of body, dorsal view; 28. same, lateral view; 29. legs I, posterior view; 30. coxosternal processes of legs I, posterior view; 31. anterior gonopods, posterior view; 32. coxite process of anterior gonopod, posterior view; 33. posterior gonopods, anterior view; 34. flagellum of posterior gonopod, anterior view. Abbreviations: csp, coxosternal process; cx, coxite; cxp, coxite process; f, flagellum; s, sternite; t, telopodite. Scale bars: 23–26 = 0.5 mm; 27–29, 31 = 0.2 mm; 30
<emphasis id="B97A203C8F6BFFC4A5F5F89CFD50F8CA" bold="true" box="[700,714,1815,1839]" pageId="9" pageNumber="414">=</emphasis>
0.05 mm; 32 = 0.4 mm; 33 = 0.1 mm; 34
<emphasis id="B97A203C8F6BFFC4A33FF89CFB1EF8CA" bold="true" box="[1142,1156,1815,1839]" pageId="9" pageNumber="414">=</emphasis>
0.04 mm.
</paragraph>
</caption>
<subSubSection id="C314AFA58F6BFFC4A78EF8D7FD68F892" box="[199,754,1884,1911]" pageId="9" pageNumber="414" type="distribution">
<paragraph id="8BB1FC2E8F6BFFC4A78EF8D7FD68F892" blockId="9.[151,1437,1884,2019]" box="[199,754,1884,1911]" pageId="9" pageNumber="414">
<emphasis id="B97A203C8F6BFFC4A78EF8D7FEC3F893" bold="true" box="[199,345,1884,1910]" pageId="9" pageNumber="414">Distribution</emphasis>
: Known only from the
<typeStatus id="54B5428C8F6BFFC4A528F8D6FD0BF892" box="[609,657,1885,1911]" pageId="9" pageNumber="414">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C314AFA58F6BFFC7A78EF80AFF6AFE80" lastPageId="10" lastPageNumber="415" pageId="9" pageNumber="414" type="discussion">
<paragraph id="8BB1FC2E8F6BFFC7A78EF80AFF6AFE80" blockId="9.[151,1437,1884,2019]" lastBlockId="10.[151,1437,151,357]" lastPageId="10" lastPageNumber="415" pageId="9" pageNumber="414">
<emphasis id="B97A203C8F6BFFC4A78EF80AFEAEF87E" bold="true" box="[199,308,1921,1947]" pageId="9" pageNumber="414">Remarks</emphasis>
: According to the review of
<taxonomicName id="4C0E87AD8F6BFFC4A539F80AFD71F87E" authorityName="Gervais" authorityYear="1847" box="[624,747,1921,1947]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="414" phylum="Arthropoda" rank="genus">
<emphasis id="B97A203C8F6BFFC4A539F80AFD71F87E" box="[624,747,1921,1947]" italics="true" pageId="9" pageNumber="414">Glyphiulus</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF9F81DF8F6BFFC4A5B0F80AFC63F87E" author="Golovatch, S. I. &amp; Geoffroy, J. - J. &amp; Mauries, J. - P. &amp; VandenSpiegel, D." box="[761,1017,1921,1947]" pageId="9" pageNumber="414" pagination="7 - 49" refId="ref7077" refString="Golovatch, S. I., Geoffroy, J. - J., Mauries, J. - P. &amp; VandenSpiegel, D. (2007 a) Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1. The granulatus - group. Zoosystema, 29 (1), 7 - 49." type="journal article" year="2007">
Golovatch
<emphasis id="B97A203C8F6BFFC4A43CF809FC37F87E" box="[885,941,1921,1947]" italics="true" pageId="9" pageNumber="414">et al.</emphasis>
2007a
</bibRefCitation>
, b), no character can exclude
<taxonomicName id="4C0E87AD8F6BFFC4A208F80AFF7FF85A" authorityName="Jiang &amp; Zhang &amp; Chen &amp; Xie" authorityYear="2021" baseAuthorityName="Jiang &amp; Zhang &amp; Chen &amp; Xie" baseAuthorityYear="2011" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="414" phylum="Arthropoda" rank="species" species="polytrichus" status="comb. nov.">
<emphasis id="B97A203C8F6BFFC4A208F80AFF7FF85A" italics="true" pageId="9" pageNumber="414">G. polytrichus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2499D478F6BFFC4A7A4F82EFEF0F85A" box="[237,362,1957,1983]" pageId="9" pageNumber="414" rank="species">comb. nov.</taxonomicNameLabel>
from this genus, and the reassignment is also supported by the phylogenetic analysis. The two species groups of
<taxonomicName id="4C0E87AD8F6BFFC4A62CF842FE7AF806" authorityName="Gervais" authorityYear="1847" box="[357,480,1993,2019]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="9" pageNumber="414" phylum="Arthropoda" rank="genus">
<emphasis id="B97A203C8F6BFFC4A62CF842FE7AF806" box="[357,480,1993,2019]" italics="true" pageId="9" pageNumber="414">Glyphiulus</emphasis>
</taxonomicName>
can be separated by the characters of the first leg-pair of the male (
<bibRefCitation id="EF9F81DF8F6BFFC7A3ADF842FF44FF54" author="Golovatch, S. I. &amp; Geoffroy, J. - J. &amp; Mauries, J. - P. &amp; VandenSpiegel, D." lastPageId="10" lastPageNumber="415" pageId="9" pageNumber="414" pagination="7 - 49" refId="ref7077" refString="Golovatch, S. I., Geoffroy, J. - J., Mauries, J. - P. &amp; VandenSpiegel, D. (2007 a) Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1. The granulatus - group. Zoosystema, 29 (1), 7 - 49." type="journal article" year="2007">
Golovatch
<emphasis id="B97A203C8F6BFFC4A22AF841FA07F806" box="[1379,1437,1993,2019]" italics="true" pageId="9" pageNumber="414">et al.</emphasis>
2007a
</bibRefCitation>
, b, 2011a, b, 2012;
<bibRefCitation id="EF9F81DF8F68FFC7A6F2FF1CFDE0FF54" author="Jiang, X. K. &amp; Guo, X. &amp; Yu, Z. G. &amp; Chen, H. M." box="[443,634,151,177]" pageId="10" pageNumber="415" pagination="197 - 208" refId="ref7609" refString="Jiang, X. K., Lv, J. C., Guo, X., Yu, Z. G. &amp; Chen, H. M. (2017) Two new species of the millipede genus Glyphiulus Gervais, 1847 from Southwest China (Diplopoda: Spirostreptida: Cambalopsidae). Zootaxa, 4323 (2), 197 - 208. https: // doi. org / 10.11646 / zootaxa. 4323.2.3" type="journal article" year="2017">
Jiang
<emphasis id="B97A203C8F68FFC7A6B7FF1CFDACFF54" box="[510,566,151,177]" italics="true" pageId="10" pageNumber="415">et al.</emphasis>
2017
</bibRefCitation>
). On the one hand, based on morphological traits,
<taxonomicName id="4C0E87AD8F68FFC7A3E6FF1CFACAFF54" authorityName="Jiang &amp; Zhang &amp; Chen &amp; Xie" authorityYear="2021" baseAuthorityName="Jiang &amp; Zhang &amp; Chen &amp; Xie" baseAuthorityYear="2011" box="[1199,1360,151,177]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="415" phylum="Arthropoda" rank="species" species="polytrichus" status="comb. nov.">
<emphasis id="B97A203C8F68FFC7A3E6FF1CFACAFF54" box="[1199,1360,151,177]" italics="true" pageId="10" pageNumber="415">G. polytrichus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2499D478F68FFC7A21EFF1CFF5DFF30" pageId="10" pageNumber="415" rank="species">comb. nov.</taxonomicNameLabel>
should be placed in the
<taxonomicName id="4C0E87AD8F68FFC7A69CFF30FDF8FF31" authorityName="Carl" authorityYear="1911" box="[469,610,187,213]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="415" phylum="Arthropoda" rank="species" species="javanicus">
<emphasis id="B97A203C8F68FFC7A69CFF30FDF8FF31" box="[469,610,187,213]" italics="true" pageId="10" pageNumber="415">G. javanicus</emphasis>
</taxonomicName>
-group by the presence of contiguous processes on the coxosternum of the male first leg-pair, and possibly close to those species which show complete telopodites, e.g.
<taxonomicName id="4C0E87AD8F68FFC7A3EAFF54FAF7FF1D" box="[1187,1389,223,249]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="415" phylum="Arthropoda" rank="species" species="guangnanensis">
<emphasis id="B97A203C8F68FFC7A3EAFF54FAF7FF1D" box="[1187,1389,223,249]" italics="true" pageId="10" pageNumber="415">G. guangnanensis</emphasis>
</taxonomicName>
. On the other hand, the phylogenetic analysis undoubtedly assigns this species close to the
<taxonomicName id="4C0E87AD8F68FFC7A331FE88FA8FFEF8" baseAuthorityName="Gervais" baseAuthorityYear="1847" box="[1144,1301,259,285]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="415" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B97A203C8F68FFC7A331FE88FA8FFEF8" box="[1144,1301,259,285]" italics="true" pageId="10" pageNumber="415">G. granulatus</emphasis>
</taxonomicName>
-group. It is not possible to resolve this disagreement for now, and here we reassign this species to
<taxonomicName id="4C0E87AD8F68FFC7A32BFEACFB47FEA4" authorityName="Gervais" authorityYear="1847" box="[1122,1245,295,321]" class="Diplopoda" family="Glyphiulidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="10" pageNumber="415" phylum="Arthropoda" rank="genus">
<emphasis id="B97A203C8F68FFC7A32BFEACFB47FEA4" box="[1122,1245,295,321]" italics="true" pageId="10" pageNumber="415">Glyphiulus</emphasis>
</taxonomicName>
as an ungrouped species.
</paragraph>
</subSubSection>
</treatment>
</document>
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