145 lines
25 KiB
XML
145 lines
25 KiB
XML
<document id="EFBD5E410A8A3653CEA26CF00D8683DC" ID-DOI="10.1590/1806-9665-RBENT-2023-0088" ID-ISSN="1806-9665" ID-Zenodo-Dep="10665223" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1708000482662" checkinUser="felipe" docAuthor="Matson, Tanner Alex" docDate="2024" docId="03A1EF5ECB3FD829FFF09CF958F70619" docLanguage="en" docName="RevBrasEntomol.68.1.e20230088.pdf" docOrigin="Revista Brasileira de Entomologia (e 20230088) 68 (1)" docSource="http://dx.doi.org/10.1590/1806-9665-rbent-2023-0088" docStyle="DocumentStyle:0303D37C566E4E2A42C6591146119864.2:RevBrasEntomol.2015-.journal_article" docStyleId="0303D37C566E4E2A42C6591146119864" docStyleName="RevBrasEntomol.2015-.journal_article" docStyleVersion="2" docTitle="Cyphoedma Matson 2024, n. gen." docType="treatment" docVersion="2" lastPageNumber="2" masterDocId="FF989726CB3ED828FFA39D355C28035F" masterDocTitle="Cyphoedma n. gen. now described after a century of use, with the addition of a new species from Central America (Geometridae, Ennominae)" masterLastPageNumber="7" masterPageNumber="1" pageNumber="2" updateTime="1708000815060" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="959D3C4669A3DB5877686EA378A82F45">Cyphoedma n. gen. now described after a century of use, with the addition of a new species from Central America (Geometridae, Ennominae)</mods:title>
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<mods:namePart id="78EC4D3B3AA85F6E7509D53B38E09202">Matson, Tanner Alex</mods:namePart>
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<mods:affiliation id="8EDAF2A15BD3077670776C7694D3091E">Smithsonian National Museum of Natural History, Washington D. C., USA.</mods:affiliation>
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<mods:title id="3F4FCB27506B85A7267202432A786A9B">Revista Brasileira de Entomologia</mods:title>
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<mods:date id="2AC3A7A90CB6E9CC5CAF6E5B918BE7D2">2024</mods:date>
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<mods:number id="32F10483C20E34919DFC798A02987025">2024-01-22</mods:number>
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<mods:url id="B9717215EEF2ED4BE18385D8AAECD13A">http://dx.doi.org/10.1590/1806-9665-rbent-2023-0088</mods:url>
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<subSubSection id="C3120DC3CB3FD829FFF09CF95EDB0148" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph id="8BB75E48CB3FD829FFF09CF95D450280" blockId="1.[83,365,460,479]" box="[83,365,460,479]" pageId="1" pageNumber="2">
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<emphasis id="B97C825ACB3FD829FFF09CF95D450280" bold="true" box="[83,365,460,479]" pageId="1" pageNumber="2">
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<taxonomicName id="4C0825CBCB3FD829FFF09CF95D090280" ID-CoL="926HZ" authority="Matson" authorityName="Matson" authorityYear="2024" box="[83,289,460,479]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus" status="gen. nov.">Cyphoedma Matson</taxonomicName>
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,
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<taxonomicNameLabel id="A24F3F21CB3FD829FE899CF95D450280" box="[298,365,460,479]" pageId="1" pageNumber="2" rank="genus">n. gen.</taxonomicNameLabel>
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</emphasis>
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<paragraph id="8BB75E48CB3FD829FFDA9F315EDB0148" blockId="1.[82,757,516,1356]" box="[121,755,516,535]" pageId="1" pageNumber="2">
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urn:lsid:zoobank.org:act:
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<subSubSection id="C3120DC3CB3FD829FFDA9F175CE3010E" pageId="1" pageNumber="2" type="materials_examined">
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<paragraph id="8BB75E48CB3FD829FFDA9F175CE3010E" blockId="1.[82,757,516,1356]" pageId="1" pageNumber="2">
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<typeStatus id="54B3E0EACB3FD829FFDA9F175C80016A" box="[121,168,546,565]" pageId="1" pageNumber="2">Type</typeStatus>
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species:
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<taxonomicName id="4C0825CBCB3FD829FF599F145EA0016B" authority="Walker, 1860: 40" authorityName="Walker" authorityPageNumber="40" authorityYear="1860" box="[250,648,545,564]" class="Insecta" family="Geometridae" genus="Cimicodes" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="transvolutata">
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Cimicodes transvolutata
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<bibRefCitation id="EF9923B9CB3FD829FE459F145EA0016B" author="Walker, F." box="[486,648,545,564]" pageId="1" pageNumber="2" refId="ref7114" refString="Walker, F., 1860. Catalogue of Lepidoptera Heterocera: List of Specimens of Lepidopterous Insects in the Collection of the British Museum. Trustees of the Natural History Museum, London, 278 pp." type="book" year="1860">Walker, 1860: 40</bibRefCitation>
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</taxonomicName>
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; by original designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3120DC3CB3FD829FFDA9F6958F70619" pageId="1" pageNumber="2" type="diagnosis">
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<paragraph id="8BB75E48CB3FD829FFDA9F695D140788" blockId="1.[82,757,516,1356]" pageId="1" pageNumber="2">
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<emphasis id="B97C825ACB3FD829FFDA9F695CCA0130" bold="true" box="[121,226,604,623]" pageId="1" pageNumber="2">Diagnosis.</emphasis>
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The wing shape and pattern of
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<taxonomicName id="4C0825CBCB3FD829FDB19F695EAF0130" authorityName="Matson" authorityYear="2024" box="[530,647,604,623]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Cyphoedma</taxonomicName>
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are similar to some members of
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<taxonomicName id="4C0825CBCB3FD829FE889F4C5DCC01D3" authority="Guenee" authorityName="Guenee" box="[299,484,633,652]" class="Insecta" family="Geometridae" genus="Cimicodes" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Cimicodes Guenée</taxonomicName>
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,
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<taxonomicName id="4C0825CBCB3FD829FE539F4C5EEC01D2" authority="Warren" authorityName="Warren" authorityYear="1895" box="[496,708,633,653]" class="Insecta" family="Geometridae" genus="Acrotomodes" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Acrotomodes Warren</taxonomicName>
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, and
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<taxonomicName id="4C0825CBCB3FD829FFF09FA35D1901F6" authority="Herrich-Schaffer" authorityName="Herrich-Schaffer" authorityYear="1855" box="[83,305,662,681]" class="Insecta" family="Geometridae" genus="Polla" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Polla Herrich-Schäffer</taxonomicName>
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(
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<bibRefCitation id="EF9923B9CB3FD829FEE19FA35D9501F5" author="Pitkin, L." box="[322,445,662,682]" pageId="1" pageNumber="2" pagination="121 - 401" refId="ref6703" refString="Pitkin, L., 2002. Neotropical Ennomine moths: a review of the genera (Lepidoptera: Geometridae). Zool. J. Linn. Soc. 135 (2 - 3), 121 - 401. https: // doi. org / 10.1046 / j. 1096 - 3642.2002.01200. x." type="journal article" year="2002">Pitkin, 2002</bibRefCitation>
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). However, it should be noted that while
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<taxonomicName id="4C0825CBCB3FD829FF669F815D150198" authorityName="Matson" authorityYear="2024" box="[197,317,692,711]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Cyphoedma</taxonomicName>
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and
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<taxonomicName id="4C0825CBCB3FD829FED09F815DF20198" box="[371,474,692,711]" class="Insecta" family="Geometridae" genus="Cimicodes" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Cimicodes</taxonomicName>
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appear to be sister genera,
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<taxonomicName id="4C0825CBCB3FD829FFF09FE45CAB01BB" authorityName="Herrich-Schaffer" authorityYear="1855" box="[83,131,721,740]" class="Insecta" family="Geometridae" genus="Polla" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Polla</taxonomicName>
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and
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<taxonomicName id="4C0825CBCB3FD829FF1B9FE45D1201BB" authorityName="Warren" authorityYear="1895" box="[184,314,721,740]" class="Insecta" family="Geometridae" genus="Acrotomodes" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Acrotomodes</taxonomicName>
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are not closely related (
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<bibRefCitation id="EF9923B9CB3FD829FD979FE45EC401BB" author="Brehm, G. & Murillo-Ramos, L. & Sihvonen, P. & Hausmann, A. & Schmidt, B. C. & Ounap, E. & Moser, A. & Mortter, R. & Bolt, D. & Bodner, F. & Lindt, A. & Parra, L. E. & Wahlberg, N." box="[564,748,721,741]" pageId="1" pageNumber="2" pagination="457 - 486" refId="ref6050" refString="Brehm, G., Murillo-Ramos, L., Sihvonen, P., Hausmann, A., Schmidt, B. C., Ounap, E., Moser, A., Mortter, R., Bolt, D., Bodner, F., Lindt, A., Parra, L. E., Wahlberg, N., 2019. New World geometrid moths (Lepidoptera: Geometridae): Molecular phylogeny, biogeography, taxonomic updates and description of 11 new tribes. Arthropod Syst. Phylogeny 77 (3), 457 - 486. https: // doi. org / 10.26049 / ASP 77 - 3 - 2019 - 5." type="journal article" year="2019">Brehm et al., 2019</bibRefCitation>
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) and the resemblance in wing shape is likely fortuitous or a result of convergent evolution on appearing to be a dead leaf to a foraging predator. One distinguishing feature of
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<taxonomicName id="4C0825CBCB3FD829FE739E1C5E6D0063" authorityName="Matson" authorityYear="2024" box="[464,581,809,828]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Cyphoedma</taxonomicName>
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is the presence of silvery-white, arcing, transverse medial line on the hindwing, which connects with the antemedial line of the forewing. In contrast, this line is either absent, straight, or does not traverse the entire wing in the other aforementioned genera. In
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<taxonomicName id="4C0825CBCB3FD829FE379EAB5E2100EE" authorityName="Matson" authorityYear="2024" box="[404,521,926,945]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Cyphoedma</taxonomicName>
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, the male valve appears unmodified, i.e., lacking significant processes or ridges. However, the valvae of
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<taxonomicName id="4C0825CBCB3FD829FF109EEC5CCB00B3" authorityName="Herrich-Schaffer" authorityYear="1855" box="[179,227,985,1004]" class="Insecta" family="Geometridae" genus="Polla" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Polla</taxonomicName>
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and
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<taxonomicName id="4C0825CBCB3FD829FEB79EEC5DBD00B3" authorityName="Warren" authorityYear="1895" box="[276,405,985,1004]" class="Insecta" family="Geometridae" genus="Acrotomodes" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Acrotomodes</taxonomicName>
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are divided and possess prominent processes, while the valve of
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<taxonomicName id="4C0825CBCB3FD829FED99EC35DF70756" box="[378,479,1014,1033]" class="Insecta" family="Geometridae" genus="Cimicodes" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Cimicodes</taxonomicName>
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features a large, sclerotized region with a spinose or setose ridge (
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<bibRefCitation id="EF9923B9CB3FD829FE7799215E640778" author="Pitkin, L." box="[468,588,1044,1063]" pageId="1" pageNumber="2" pagination="121 - 401" refId="ref6703" refString="Pitkin, L., 2002. Neotropical Ennomine moths: a review of the genera (Lepidoptera: Geometridae). Zool. J. Linn. Soc. 135 (2 - 3), 121 - 401. https: // doi. org / 10.1046 / j. 1096 - 3642.2002.01200. x." type="journal article" year="2002">Pitkin, 2002</bibRefCitation>
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). In females, the absence of a signum further separates
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<taxonomicName id="4C0825CBCB3FD829FE6D99045E6B071B" authorityName="Matson" authorityYear="2024" box="[462,579,1073,1092]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Cyphoedma</taxonomicName>
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from
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<taxonomicName id="4C0825CBCB3FD829FDDC99045E86071B" authorityName="Herrich-Schaffer" authorityYear="1855" box="[639,686,1073,1092]" class="Insecta" family="Geometridae" genus="Polla" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Polla</taxonomicName>
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. When at rest (
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<figureCitation id="133342CDCB3FD829FF07997B5CF3073D" box="[164,219,1102,1122]" captionStart="Figure 1" captionStartId="1.[84,138,1947,1964]" captionTargetBox="[85,754,1429,1931]" captionTargetId="figure-1001@1.[85,754,1429,1931]" captionTargetPageId="1" captionText="Figure 1 Natural resting posture of Cyphoedma. Image of female C. transvolutata (Walker) taken in Santa Catarina, Brazil.Photo credit: Douglas Meyer." figureDoi="http://doi.org/10.5281/zenodo.10665225" httpUri="https://zenodo.org/record/10665225/files/figure.png" pageId="1" pageNumber="2">Fig. 1</figureCitation>
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), the forewing costal margin of
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<taxonomicName id="4C0825CBCB3FD829FD8B997B5EB7073E" authorityName="Matson" authorityYear="2024" box="[552,671,1102,1121]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Cyphoedma</taxonomicName>
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is often perpendicular to the body axis. The recognition of
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<taxonomicName id="4C0825CBCB3FD829FDF399595EED0720" authorityName="Matson" authorityYear="2024" box="[592,709,1132,1151]" class="Insecta" family="Geometridae" genus="Cyphoedma" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">Cyphoedma</taxonomicName>
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as a distinct genus among related or visually similar genera is also supported by the mito-nuclear phylogenetic results of
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<bibRefCitation id="EF9923B9CB3FD829FE4C99935EDB07E5" author="Murillo-Ramos, L. C. & Brehm, G. & Sihvonen, P. & Hausmann, A. & Holm, S. & Ghanavi, H. R. & Ounap, E. & Truuverk, A. & Staude, H. & Friedrich, E. & Tammaru, T. & Wahlberg, N." box="[495,755,1190,1210]" pageId="1" pageNumber="2" pagination="7386" refId="ref6614" refString="Murillo-Ramos, L. C., Brehm, G., Sihvonen, P., Hausmann, A., Holm, S., Ghanavi, H. R., Ounap, E., Truuverk, A., Staude, H., Friedrich, E., Tammaru, T., Wahlberg, N., 2019. A comprehensive molecular phylogeny of Geometridae (Lepidoptera) with a focus on enigmatic small subfamilies. PeerJ 7, e 7386." type="journal article" year="2019">Murillo-Ramos et al. (2019)</bibRefCitation>
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||
and
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||
<bibRefCitation id="EF9923B9CB3FD829FFDE99F15D1F0788" author="Brehm, G. & Murillo-Ramos, L. & Sihvonen, P. & Hausmann, A. & Schmidt, B. C. & Ounap, E. & Moser, A. & Mortter, R. & Bolt, D. & Bodner, F. & Lindt, A. & Parra, L. E. & Wahlberg, N." box="[125,311,1220,1239]" pageId="1" pageNumber="2" pagination="457 - 486" refId="ref6050" refString="Brehm, G., Murillo-Ramos, L., Sihvonen, P., Hausmann, A., Schmidt, B. C., Ounap, E., Moser, A., Mortter, R., Bolt, D., Bodner, F., Lindt, A., Parra, L. E., Wahlberg, N., 2019. New World geometrid moths (Lepidoptera: Geometridae): Molecular phylogeny, biogeography, taxonomic updates and description of 11 new tribes. Arthropod Syst. Phylogeny 77 (3), 457 - 486. https: // doi. org / 10.26049 / ASP 77 - 3 - 2019 - 5." type="journal article" year="2019">Brehm et al. (2019)</bibRefCitation>
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.
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||
</paragraph>
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||
<paragraph id="8BB75E48CB3FD829FFDA99D458F70619" blockId="1.[82,757,516,1356]" lastBlockId="1.[800,1476,153,1350]" pageId="1" pageNumber="2">
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<emphasis id="B97C825ACB3FD829FFDA99D45D6C07AB" bold="true" box="[121,324,1249,1268]" pageId="1" pageNumber="2">Male description (</emphasis>
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<figureCitation id="133342CDCB3FD829FEE799D45DA007AB" box="[324,392,1249,1268]" captionStart="Figure 2" captionStartId="2.[112,165,970,987]" captionTargetBox="[135,1480,154,948]" captionTargetId="figure-185@2.[134,1483,151,953]" captionTargetPageId="2" captionText="Figure 2 Adult habitus of Cyphoedma. Arrows pointing to diagnostic wing differences. (a) C. ashleyorum n. sp., male holotype, USNMENT01771276, dorsal; (b) same, ventral; (c) C. ashleyorum n. sp., female paratype, USNM01771288, dorsal; (d) same, ventral; (e) C. mirafloresa (Dognin), male lectotype, USNMENT01771321, dorsal; (f) same, ventral; (g) C. mirafloresa (Dognin), female, USNMENT01771294, dorsal; (h) same, ventral; (i) C. transvolutata (Walker), male, USNMENT01771293, dorsal; (j) same, ventral; (k) C. transvolutata (Walker), female, USNMENT01771320, dorsal; (l) same, ventral. Scale bar = 2 cm." figureDoi="http://doi.org/10.5281/zenodo.10665229" httpUri="https://zenodo.org/record/10665229/files/figure.png" pageId="1" pageNumber="2">Figs. 2</figureCitation>
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a-b, 2e-f, 2i-j, 3). Forewing length
|
||
<quantity id="4CF0F3ADCB3FD829FFF099CB5CE9064D" box="[83,193,1278,1298]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="1" pageNumber="2" unit="mm" value="22.5" valueMax="25.0" valueMin="20.0">20–25 mm</quantity>
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. Head: Antenna filiform, brown scales above. Vertex dark brown with lighter brown scales near scape. Frons mostly dark brown, but with lighter brown scales medially; ventral margin white. Labial palpus porrect, slightly longer than diameter of eye, cream to white on inner surface, and variably brown and white on outer surface. Proboscis well developed. Chaetosemata well separated. Thorax: Patagium mostly gray. Tegula brown. Mesothorax brown above, white below. Foreleg with large epiphysis; tibial spur formula 0–2–4; hindtibia swollen, bearing plume of long androconia folded into longitudinal groove on inner surface (when not deployed). Legs often whiter on internal surface and variably banded or speckled with yellow and dark brown on external surface; hind leg generally less speckled. Forewing: Predominantly brown with slightly falcate apex speckled with white and yellow. Costal and subapical areas given toward mustard-brown; costa often with frosty highlights. Curved, transverse silvery-white antemedial and postmedial lines. Dark brown medial transverse line through frosted central part of wing. Small black discal spot. Irregular, central, light brown patch distad to postmedial line. Subterminal area sometimes with jagged silvery-white line, especially toward apex. Outer margin with silvery-white highlight. Underside pale brown to white in proximal half; transitioning to dark brown in distal half. Fringe brown.Hindwing: Concolorous with forewing, but more frosted in basal half; bearing arcing, medial, transverse silvery-white line that joins with antemedial line of forewing. Small black discal spot present. Subterminal area sometimes with small, lighter brown flecks. Outer margin with silvery-white highlight. Underside pale brown to white in proximal two-thirds; transitioning to dark brown in distal third. Fringe brown. Abdomen: Dark brown above, pale white to cream below. Third sternite with comb of setae. Male Genitalia: (
|
||
<figureCitation id="133342CDCB3FD829FB799E4C592600D2" box="[1242,1294,889,909]" captionStart="Figure 3" captionStartId="2.[113,165,1710,1727]" captionTargetBox="[262,1353,1132,1673]" captionTargetId="figure-239@2.[249,1370,1125,1696]" captionTargetPageId="2" captionText="Figure 3 Cyphoedma male genitalia.(a)C.ashleyorum n.sp.,TAM-2023-288(USNM 154251),USNMENT01771290,paratype,USNM,genital capsule;(b) same,phallus;(c)C.mirafloresa (Dognin),TAM-2023-279 (USNM154250),USNMENT01771323,genital capsule;(d) same,phallus;(e)C.transvolutata (Walker),TAM-2023-277 (USNM 154248),USNMENT01771293, genital capsule; (f) same, phallus.Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10665231" httpUri="https://zenodo.org/record/10665231/files/figure.png" pageId="1" pageNumber="2">Fig. 3</figureCitation>
|
||
): Uncus short and tapered toward apex. Valva without processes. Juxta plate-like and not strongly sclerotized. Vesica with two separate fields of small dentate cornuti on adjacent diverticula.
|
||
<emphasis id="B97C825ACB3FD829FBF49EE7593400BA" bold="true" box="[1111,1308,978,997]" pageId="1" pageNumber="2">Female description</emphasis>
|
||
(
|
||
<figureCitation id="133342CDCB3FD829FA8A9EE7594D00BA" box="[1321,1381,978,997]" captionStart="Figure 2" captionStartId="2.[112,165,970,987]" captionTargetBox="[135,1480,154,948]" captionTargetId="figure-185@2.[134,1483,151,953]" captionTargetPageId="2" captionText="Figure 2 Adult habitus of Cyphoedma. Arrows pointing to diagnostic wing differences. (a) C. ashleyorum n. sp., male holotype, USNMENT01771276, dorsal; (b) same, ventral; (c) C. ashleyorum n. sp., female paratype, USNM01771288, dorsal; (d) same, ventral; (e) C. mirafloresa (Dognin), male lectotype, USNMENT01771321, dorsal; (f) same, ventral; (g) C. mirafloresa (Dognin), female, USNMENT01771294, dorsal; (h) same, ventral; (i) C. transvolutata (Walker), male, USNMENT01771293, dorsal; (j) same, ventral; (k) C. transvolutata (Walker), female, USNMENT01771320, dorsal; (l) same, ventral. Scale bar = 2 cm." figureDoi="http://doi.org/10.5281/zenodo.10665229" httpUri="https://zenodo.org/record/10665229/files/figure.png" pageId="1" pageNumber="2">Figs. 2</figureCitation>
|
||
c-d, 2g-h, 2k-l, 4). Forewing length
|
||
<quantity id="4CF0F3ADCB3FD829FBB49EDA58AD075C" box="[1047,1157,1007,1027]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.6" metricValueMin="2.3" pageId="1" pageNumber="2" unit="mm" value="24.5" valueMax="26.0" valueMin="23.0">23–26 mm</quantity>
|
||
. Head: As in male. Thorax: As in male; though lacking hindtibial androconia.Forewing: As in male, but often with lighter brown ground color and less pronounced central light brown patch distad to postmedial line above. Underside brown to tan in proximal half; transitioning to darker brown in distal half. Hindwing: As in male, but underside brown to tan in proximal half and transitioning to darker brown in distal half (less contrasted than male). Abdomen: As in male but lacking third sternite comb of setae. Female Genitalia:(
|
||
<figureCitation id="133342CDCB3FD829FC7799EE582D07B1" box="[980,1029,1243,1262]" captionStart="Figure 4" captionStartId="3.[83,137,1049,1066]" captionTargetBox="[220,1328,160,1008]" captionTargetId="figure-47@3.[207,1350,151,1032]" captionTargetPageId="3" captionText="Figure 4 Cyphoedma female genitalia. (a) C. ashleyorum n. sp., TAM-2023-291 (USNM 154253), USNMENT01771290, paratype; (b) C. mirafloresa (Dognin), TAM-2023-290 (USNM 154252), USNMENT01771322; (c) C. transvolutata (Walker), TAM-2023-278 (USNM 154249), USNMENT01771320. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10665233" httpUri="https://zenodo.org/record/10665233/files/figure.png" pageId="1" pageNumber="2">Fig.4</figureCitation>
|
||
): Papillae anales rounded; posterior apophysis 2.5–3x longer than anterior apophysis. Ductus bursae more heavily sclerotized at junction with corpus bursae. Corpus bursae pyriform with lightly wrinkled surface; signum absent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |