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<document id="985450B016C46E91955D09A467420A80" ID-CLB-Dataset="28041" ID-DOI="10.11646/zootaxa.4508.3.3" ID-GBIF-Dataset="9de070ed-9c35-4131-a350-72371ee93ddf" ID-ISSN="1175-5326" ID-Zenodo-Dep="2607216" ID-ZooBank="E53B4C53-554A-4913-8E3B-CE1C530DF004" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553586064605" checkinUser="plazi" docAuthor="Shavrin, Alexey V. &amp; Smetana, Aleš" docDate="2018" docId="DA218790FFBC2E22248FFAFE7B22FD71" docLanguage="en" docName="zootaxa.4508.3.3.pdf" docOrigin="Zootaxa 4508 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphichroum angustilobatum Shavrin &amp; Smetana 2018, sp.n." docType="treatment" docVersion="8" lastPageNumber="382" masterDocId="2618FFE8FFBD2E272418FFD37818FFBC" masterDocTitle="New species of the genus Amphichroum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) from China" masterLastPageNumber="402" masterPageNumber="377" pageNumber="378" updateTime="1698713048312" updateUser="ExternalLinkService">
<mods:mods id="10E50E1544CD39AB165775D44B6A4FAB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2A978A6EF9043BFF4FCA5B26175C8E3B">New species of the genus Amphichroum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) from China</mods:title>
</mods:titleInfo>
<mods:name id="B8C190172EC1088F7DD59FDFBDC81EC7" type="personal">
<mods:role id="86787C4E0B8A964D254CBC0EA0AA8446">
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</mods:role>
<mods:namePart id="2B3A5D1CDB15478EABC8DBE882120055">Shavrin, Alexey V.</mods:namePart>
</mods:name>
<mods:name id="99E862F51AB3ED9C77D676CB4251B5BD" type="personal">
<mods:role id="90F43FBF653F0FF40E4B228287778CB4">
<mods:roleTerm id="F153E13AFA30CD05EB99D1FDB7853E6D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FA7472F32AD382182741B0F6B9200183">Smetana, Aleš</mods:namePart>
</mods:name>
<mods:typeOfResource id="6168C5B24EC0F4809F69BC97871FE4A2">text</mods:typeOfResource>
<mods:relatedItem id="0A06AC0CFEAFED1C18592127E6C0FC07" type="host">
<mods:titleInfo id="17AF6165BC79DF4D762B262CD8CE1C30">
<mods:title id="56AE9B138C3268E2F63160E54F591446">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7243757AC795F498B4F05AF78ADF5B21">
<mods:date id="06DD094BA2498D3CDC9FD9B142F1E9E4">2018</mods:date>
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<mods:number id="B58CE31D90099D2700BF5629DB186140">2018-11-01</mods:number>
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<mods:detail id="78470855DCDCFFD035322F63F8C60A63" type="volume">
<mods:number id="26DACE621FDE2F0095B156162749A9DA">4508</mods:number>
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<mods:number id="91142E14FE529ACA9905C2196A19E700">3</mods:number>
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<mods:start id="7F37B4679960B771DBE1300A443993B2">377</mods:start>
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<mods:classification id="91DEE43FC564B00CDDDA28EBEA56FADB">journal article</mods:classification>
<mods:identifier id="2E818BA0D435A6ADC22E83BCB219209A" type="CLB-Dataset">28041</mods:identifier>
<mods:identifier id="A3B0230BB497FA2E4E802188C2BC5CBF" type="DOI">10.11646/zootaxa.4508.3.3</mods:identifier>
<mods:identifier id="6EA05501A26233391765ED5A8E129991" type="GBIF-Dataset">9de070ed-9c35-4131-a350-72371ee93ddf</mods:identifier>
<mods:identifier id="16373C0A0E28B6354A9D16BA07DF9ABA" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="DF0DDBB858DAE26184C279D2197DBEE9" type="Zenodo-Dep">2607216</mods:identifier>
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</mods:mods>
<treatment id="DA218790FFBC2E22248FFAFE7B22FD71" ID-DOI="http://doi.org/10.5281/zenodo.5962984" ID-GBIF-Taxon="154495877" ID-Zenodo-Dep="5962984" LSID="urn:lsid:plazi:treatment:DA218790FFBC2E22248FFAFE7B22FD71" httpUri="http://treatment.plazi.org/id/DA218790FFBC2E22248FFAFE7B22FD71" lastPageId="5" lastPageNumber="382" pageId="1" pageNumber="378">
<subSubSection id="1A92650DFFBC2E26248FFAFE7B4BFAFB" box="[151,851,1325,1351]" pageId="1" pageNumber="378" type="nomenclature">
<paragraph id="52373686FFBC2E26248FFAFE7B4BFAFB" blockId="1.[151,851,1325,1384]" box="[151,851,1325,1351]" pageId="1" pageNumber="378">
<heading id="097F81EAFFBC2E26248FFAFE7B4BFAFB" bold="true" box="[151,851,1325,1351]" fontSize="11" level="1" pageId="1" pageNumber="378" reason="1">
<emphasis id="60FCEA94FFBC2E26248FFAFE7B4BFAFB" bold="true" box="[151,851,1325,1351]" pageId="1" pageNumber="378">
<taxonomicName id="95884D05FFBC2E26248FFAFE7B14FAFB" ID-CoL="CZPL" authority="Shavrin &amp; Smetana" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[151,780,1325,1351]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="378" phylum="Arthropoda" rank="species" species="angustilobatum" status="sp. nov.">
<emphasis id="60FCEA94FFBC2E26248FFAFE7A14FAFB" bold="true" box="[151,524,1325,1351]" italics="true" pageId="1" pageNumber="378">Amphichroum angustilobatum</emphasis>
Shavrin &amp; Smetana
</taxonomicName>
,
<taxonomicNameLabel id="7BCF57EFFFBC2E262700FAFE7B4BFAFB" box="[792,851,1325,1351]" pageId="1" pageNumber="378" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFBC2E26248FFA9C7998FAD4" box="[151,384,1359,1384]" pageId="1" pageNumber="378" type="description">
<paragraph id="52373686FFBC2E26248FFA9C7998FAD4" blockId="1.[151,851,1325,1384]" box="[151,384,1359,1384]" pageId="1" pageNumber="378">
(
<figureCitation id="CAB32A03FFBC2E262487FA9C78F4FAD4" box="[159,236,1359,1384]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1954,1977]" captionTargetBox="[296,1236,546,1939]" captionTargetId="figure@2.[283,1307,539,1939]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus of Amphichroum (males): 1–A. angustilobatum sp.n. (holotype), 2–A. assingi sp.n. (holotype), 3–A. longilobatum sp.n. (paratype), 4–A. maculosum sp.n. (holotype). Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/2607218/files/figure.png" pageId="1" pageNumber="378">Figs. 1</figureCitation>
,
<figureCitation id="CAB32A03FFBC2E2624E2FA9C793EFAD4" box="[250,294,1359,1384]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1873,1896]" captionTargetBox="[231,1335,218,1829]" captionTargetId="figure@3.[210,1377,193,1853]" captionTargetPageId="3" captionText="FIGURES 5–10. Aedeagus of Amphichroum angustilobatum sp.n.(5–7: holotype) and A. assingi sp.n. (8–10: paratype): 5, 8– parameral view (outline of internal sac shown as point line), 6, 9–internal sac, 7, 10–lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607222/files/figure.png" pageId="1" pageNumber="378">5–7</figureCitation>
,
<figureCitation id="CAB32A03FFBC2E26252AFA837957FAD4" box="[306,335,1360,1384]" captionStart="FIGURE 11" captionStartId="4.[151,250,1055,1077]" captionTargetBox="[228,1360,185,1034]" captionTargetId="figure@4.[228,1360,185,1034]" captionTargetPageId="4" captionText="FIGURE 11. Type locality of Amphichroum angustilobatum sp.n.in the Meili Xue Shan range in Yunnan (photograph: M. Schülke (locality C2005-07A))." httpUri="https://zenodo.org/record/2607224/files/figure.png" pageId="1" pageNumber="378">11</figureCitation>
,
<figureCitation id="CAB32A03FFBC2E262543FA837960FAD4" box="[347,376,1360,1384]" captionStart="FIGURE 41" captionStartId="24.[151,250,1057,1079]" captionTargetBox="[179,1408,267,1035]" captionTargetId="figure@24.[179,1408,267,1035]" captionTargetPageId="24" captionText="FIGURE 41. Distribution of Amphichroum in China: A. angustilobatum sp.n. (black star), A. assingi sp.n. (white square with wide black margin), A. longilobatum sp.n. (white circle), A. maculosum sp.n. (white star), A. miaoershanum Watanabe, 1999 (circle with wide white margin), A. propinquum sp.n. (black square), A. rotundatum (white square), A. schuelkei sp.n. (black rhomb), A. squamosum sp.n. (black circle), A.subaequale sp.n. (white rhomb), A. subelongatum sp.n. (black triangle), A. tibetanum sp.n. (white triangle)." httpUri="https://zenodo.org/record/2607244/files/figure.png" pageId="1" pageNumber="378">41</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFBC2E26248FFA4478E7F96C" pageId="1" pageNumber="378" type="materials_examined">
<paragraph id="52373686FFBC2E26248FFA447CE9F9FC" blockId="1.[151,1437,1431,2033]" pageId="1" pageNumber="378">
<emphasis id="60FCEA94FFBC2E26248FFA4479DFFA0C" bold="true" box="[151,455,1431,1456]" pageId="1" pageNumber="378">Type material examined:</emphasis>
<materialsCitation id="E2E03CDBFFBC2E2625C9FA4478CEF9A1" ID-GBIF-Occurrence="2028139641" collectingDate="2005" collectingDateMax="2007" collectingDateMin="2005" collectionCode="VI" country="China" county="Deqin Co." elevation="2890" latitude="28.421667" location="Meili Xue Shan" longLatPrecision="12" longitude="98.80783" municipality="Diqing" pageId="1" pageNumber="378" specimenCount="1" specimenCount-male="1" stateProvince="Xizang" typeStatus="holotype">
<typeStatus id="8D338824FFBC2E2625C9FA447A21FA0C" box="[465,569,1431,1456]" pageId="1" pageNumber="378" type="holotype">Holotype</typeStatus>
<specimenCount id="448EFD0FFFBC2E26265BFA4B7A40FA13" box="[579,600,1432,1455]" pageId="1" pageNumber="378" type="male">♂</specimenCount>
[specimen dissected]: ‘
<collectingCountry id="2A9F7616FFBC2E26277EFA4B7BD0FA0C" box="[870,968,1432,1456]" name="China" pageId="1" pageNumber="378">CHINA</collectingCountry>
: N-Yunnan [C
<date id="26361046FFBC2E262074FA447CD5FA0C" box="[1132,1229,1431,1456]" pageId="1" pageNumber="378" value="2005" valueMax="2007">
<collectingDate id="3672E9AEFFBC2E262074FA447CD5FA0C" box="[1132,1229,1431,1456]" pageId="1" pageNumber="378" value="2005" valueMax="2007">2005-07</collectingDate>
</date>
A] |
<collectingMunicipality id="B253ACFCFFBC2E26211BFA447D49FA0C" box="[1283,1361,1431,1456]" pageId="1" pageNumber="378">Diqing</collectingMunicipality>
<collectingRegion id="904CF864FFBC2E262143FA447D81FA0C" box="[1371,1433,1431,1456]" country="China" name="Xizang" pageId="1" pageNumber="378">Tibet</collectingRegion>
. Aut. Pref., |
<collectingCounty id="BB564E0AFFBC2E26252CFA6F79A9FA68" box="[308,433,1468,1493]" pageId="1" pageNumber="378">Deqin Co.</collectingCounty>
,
<location id="5757605DFFBC2E2625DCFA6F7A9DFA69" LSID="urn:lsid:plazi:treatment:DA218790FFBC2E22248FFAFE7B22FD71:5757605DFFBC2E2625DCFA6F7A9DFA69" box="[452,645,1468,1493]" country="China" county="Deqin Co." latitude="28.421667" longLatPrecision="12" longitude="98.80783" municipality="Diqing" name="Meili Xue Shan" pageId="1" pageNumber="378" stateProvince="Xizang">Meili Xue Shan</location>
, E-side, |
<locationDeviation id="43852FA4FFBC2E262710FA6F7BFCFA69" box="[776,996,1468,1493]" location="Deqin" pageId="1" pageNumber="378">
<quantity id="95709B63FFBC2E262710FA6F7B4FFA69" box="[776,855,1468,1493]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="378" unit="km" value="12.0">12 km</quantity>
SW Deqin
</locationDeviation>
,
<quantity id="95709B63FFBC2E2627EEFA6F7C4CFA68" box="[1014,1108,1468,1492]" metricMagnitude="3" metricUnit="m" metricValue="2.8899999999999997" pageId="1" pageNumber="378" unit="m" value="2890.0">
<elevation id="D9A5D1B5FFBC2E2627EEFA6F7C4CFA68" box="[1014,1108,1468,1492]" metricMagnitude="3" metricUnit="m" metricValue="2.8899999999999997" pageId="1" pageNumber="378" unit="m" value="2890.0">2890 m</elevation>
</quantity>
, |
<geoCoordinate id="37BC5041FFBC2E262063FA6F7CE6FA69" box="[1147,1278,1468,1493]" direction="north" orientation="latitude" pageId="1" pageNumber="378" precision="9" value="28.421667">28°25.30'N</geoCoordinate>
,
<geoCoordinate id="37BC5041FFBC2E262108FA6F7D8BFA68" box="[1296,1427,1468,1492]" direction="east" orientation="longitude" pageId="1" pageNumber="378" precision="9" value="98.80783">98°48.47'E</geoCoordinate>
,’ &lt;rectangular label, printed&gt;,‘small creek valley, mixed forest | with bamboo, leaf litter, moss, | dead wood, sifted,
<date id="26361046FFBC2E26248FF9D778CEF9A1" box="[151,214,1540,1565]" pageId="1" pageNumber="378" value="2005-06-13">
13.
<collectionCode id="3499AE43FFBC2E2624ADF9D778CEF9A1" box="[181,214,1540,1565]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="378">VI</collectionCode>
</date>
</materialsCitation>
.2005, | leg. M. Schülke [C
<date id="26361046FFBC2E262636F9D77A8AF9A1" box="[558,658,1540,1565]" pageId="1" pageNumber="378" value="2005" valueMax="2007">2005-07</date>
A]’ &lt;rectangular label, printed&gt;, ‘
<materialsCitation id="E2E03CDBFFBC2E26203EF9D77CF5F9FC" ID-GBIF-Occurrence="2028139645" collectionCode="CSC" collectorName="Shavrin A. &amp; Smetana A." country="China" pageId="1" pageNumber="378" specimenCount="1" typeStatus="holotype">
<typeStatus id="8D338824FFBC2E26203EF9D77CA7F9A0" box="[1062,1215,1540,1564]" pageId="1" pageNumber="378" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="95884D05FFBC2E2620FCF9D77D90F9A1" authorityName="Kraatz" authorityYear="1857" box="[1252,1416,1540,1565]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="401" phylum="Arthropoda" rank="genus">Amphichroum</taxonomicName>
| angustilobatum
<emphasis id="60FCEA94FFBC2E262552F9F4799AF9FC" bold="true" box="[330,386,1575,1600]" pageId="1" pageNumber="378">
<taxonomicNameLabel id="7BCF57EFFFBC2E262552F9F4799AF9FC" box="[330,386,1575,1600]" pageId="1" pageNumber="378" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
|
<collectorName id="FF7D5350FFBC2E26258EF9F47A08F9FC" box="[406,528,1575,1600]" pageId="1" pageNumber="378">Shavrin A.</collectorName>
&amp;
<collectorName id="FF7D5350FFBC2E26262BF9FB7AAEF9FC" box="[563,694,1576,1600]" pageId="1" pageNumber="378">Smetana A.</collectorName>
des. 2018’ &lt;red rectangular label, printed&gt; (
<collectionCode id="3499AE43FFBC2E2620B3F9FB7CFDF9FC" box="[1195,1253,1576,1600]" country="Cuba" lsid="urn:lsid:biocol.org:col:12869" name="Colegio del Sagrado Corazón" pageId="1" pageNumber="378" type="Herbarium">CSC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="52373686FFBC2E2624DFF99E78E7F96C" blockId="1.[151,1437,1431,2033]" pageId="1" pageNumber="378">
<materialsCitation id="E2E03CDBFFBC2E2624DFF99E7A7BF934" ID-GBIF-Occurrence="2028139640" collectionCode="VI" collectorName="Aut. Pr. Deqin Co. Meili Xue" country="China" county="Aut. Pr. Deqin Co." elevation="2890" latitude="28.421667" location="Shan" longLatPrecision="12" longitude="98.80783" municipality="Diqing" pageId="1" pageNumber="378" specimenCount="1" specimenCount-female="1" stateProvince="Xizang" typeStatus="paratype">
<typeStatus id="8D338824FFBC2E2624DFF99E7931F9D9" box="[199,297,1613,1637]" pageId="1" pageNumber="378" type="paratype">Paratype</typeStatus>
<specimenCount id="448EFD0FFFBC2E262528F99E795FF9D8" box="[304,327,1613,1636]" pageId="1" pageNumber="378" type="female">♀</specimenCount>
: ‘
<collectingCountry id="2A9F7616FFBC2E262541F99F79A3F9D8" box="[345,443,1612,1636]" name="China" pageId="1" pageNumber="378">CHINA</collectingCountry>
: N-Yunnan
<collectingMunicipality id="B253ACFCFFBC2E262658F99F7A96F9D9" box="[576,654,1612,1637]" pageId="1" pageNumber="378">Diqing</collectingMunicipality>
<collectingRegion id="904CF864FFBC2E26268DF99F7ACBF9D9" box="[661,723,1612,1637]" country="China" name="Xizang" pageId="1" pageNumber="378">Tibet</collectingRegion>
. |
<collectorName id="FF7D5350FFBC2E2626F3F99F7C37F9D8" box="[747,1071,1612,1637]" pageId="1" pageNumber="378">
<collectingCounty id="BB564E0AFFBC2E2626F3F99F7BADF9D8" box="[747,949,1612,1637]" pageId="1" pageNumber="378">Aut.Pr. Deqin Co.</collectingCounty>
Meili Xue
</collectorName>
|
<location id="5757605DFFBC2E26205CF99F7C64F9D9" LSID="urn:lsid:plazi:treatment:DA218790FFBC2E22248FFAFE7B22FD71:5757605DFFBC2E26205CF99F7C64F9D9" box="[1092,1148,1612,1637]" country="China" county="Aut. Pr. Deqin Co." latitude="28.421667" longLatPrecision="12" longitude="98.80783" municipality="Diqing" name="Shan" pageId="1" pageNumber="378" stateProvince="Xizang">Shan</location>
E-side
<locationDeviation id="43852FA4FFBC2E2620CAF99F7D97F9D9" box="[1234,1423,1612,1637]" location="Deqin" pageId="1" pageNumber="378">
<quantity id="95709B63FFBC2E2620CAF99F7D0AF9D9" box="[1234,1298,1612,1637]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="378" unit="km" value="12.0">12km</quantity>
SW Deqin
</locationDeviation>
|
<geoCoordinate id="37BC5041FFBC2E26248FF9BC790EF934" box="[151,278,1647,1672]" direction="north" orientation="latitude" pageId="1" pageNumber="378" precision="9" value="28.421667">28°25.30'N</geoCoordinate>
<geoCoordinate id="37BC5041FFBC2E26253BF9A37987F934" box="[291,415,1648,1672]" direction="east" orientation="longitude" pageId="1" pageNumber="378" precision="9" value="98.80783">98°48.47'E</geoCoordinate>
<quantity id="95709B63FFBC2E2625B4F9A37A1EF93B" box="[428,518,1648,1672]" metricMagnitude="3" metricUnit="m" metricValue="2.8899999999999997" pageId="1" pageNumber="378" unit="m" value="2890.0">
<elevation id="D9A5D1B5FFBC2E2625B4F9A37A1EF93B" box="[428,518,1648,1672]" metricMagnitude="3" metricUnit="m" metricValue="2.8899999999999997" pageId="1" pageNumber="378" unit="m" value="2890.0">2890 m</elevation>
</quantity>
|
<date id="26361046FFBC2E26263DF9BC7A7BF934" box="[549,611,1647,1672]" pageId="1" pageNumber="378" value="2005-06-13">
13.
<collectionCode id="3499AE43FFBC2E26265BF9BC7A7BF934" box="[579,611,1647,1672]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="1" pageNumber="378">VI</collectionCode>
</date>
</materialsCitation>
.2005 A. Smetana [C160]’&lt;rectangular label, printed&gt;, ‘
<materialsCitation id="E2E03CDBFFBC2E2620E2F9A378E3F96C" ID-GBIF-Occurrence="2028139643" collectionCode="NSMT" collectorName="Shavrin A. &amp; Smetana A." country="China" pageId="1" pageNumber="378" specimenCount="1" typeStatus="paratype">
<typeStatus id="8D338824FFBC2E2620E2F9A37D92F934" box="[1274,1418,1648,1672]" pageId="1" pageNumber="378" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="95884D05FFBC2E26248FF9477923F911" authorityName="Kraatz" authorityYear="1857" box="[151,315,1684,1709]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="401" phylum="Arthropoda" rank="genus">Amphichroum</taxonomicName>
| angustilobatum
<emphasis id="60FCEA94FFBC2E262608F9477A50F911" bold="true" box="[528,584,1684,1709]" pageId="1" pageNumber="378">
<taxonomicNameLabel id="7BCF57EFFFBC2E262608F9477A50F911" box="[528,584,1684,1709]" pageId="1" pageNumber="378" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
|
<collectorName id="FF7D5350FFBC2E26267DF9477AFCF910" box="[613,740,1684,1709]" pageId="1" pageNumber="378">Shavrin A.</collectorName>
&amp;
<collectorName id="FF7D5350FFBC2E262709F9477B81F910" box="[785,921,1684,1708]" pageId="1" pageNumber="378">Smetana A.</collectorName>
des. 2018’ &lt;red rectangular label, printed&gt; (
<collectionCode id="3499AE43FFBC2E262487F96B78EBF96C" box="[159,243,1720,1744]" country="Japan" httpUri="http://grbio.org/cool/f678-ci92" name="National Science Museum (Natural History)" pageId="1" pageNumber="378">NSMT</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFBC2E2224DFF90F7A1EFE10" lastPageId="5" lastPageNumber="382" pageId="1" pageNumber="378" type="description">
<paragraph id="52373686FFBC2E2624DFF90F7BB6F881" blockId="1.[151,1437,1431,2033]" pageId="1" pageNumber="378">
<emphasis id="60FCEA94FFBC2E2624DFF90F7949F949" bold="true" box="[199,337,1756,1781]" pageId="1" pageNumber="378">Description</emphasis>
. Measurements (n=2): HW: 0.57–0.60; HL: 0.32–0.34; AL(
<typeStatus id="8D338824FFBC2E2627E3F90F7C7AF949" box="[1019,1122,1756,1781]" pageId="1" pageNumber="378" type="holotype">holotype</typeStatus>
): 1.13; OL: 0.17; PL: 0.57– 0.62; PW: 0.92; ESL: 1.12; EW: 1.14–1.31; AW: 1.12; MTbL(
<typeStatus id="8D338824FFBC2E262744F92C7BDAF8A4" box="[860,962,1791,1816]" pageId="1" pageNumber="378" type="holotype">holotype</typeStatus>
): 0.62; MTrL(
<typeStatus id="8D338824FFBC2E26207DF92C7CD2F8A4" box="[1125,1226,1791,1816]" pageId="1" pageNumber="378" type="holotype">holotype</typeStatus>
): 0.54 (MTrL 1–4: 0.32; MTrL 5: 0.22); AedL(
<typeStatus id="8D338824FFBC2E2625CAF8F77A20F881" box="[466,568,1828,1853]" pageId="1" pageNumber="378" type="holotype">holotype</typeStatus>
): 0.62; TL: 2.85–3.10 (
<typeStatus id="8D338824FFBC2E262722F8F77BBAF881" box="[826,930,1828,1853]" pageId="1" pageNumber="378" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="52373686FFBC2E2524DFF8947C03FE49" blockId="1.[151,1437,1431,2033]" lastBlockId="2.[151,1436,151,501]" lastPageId="2" lastPageNumber="379" pageId="1" pageNumber="378">Head, antennomeres 5–11 and disc of pronotum yellow-brown; mouthparts, antennomeres 1–4, lateral portions of pronotum, elytra and legs yellow; abdomen reddish-brown. Forebody with oily sheen, abdomen shiny; head with dense transverse microsculpture, with more or less rounded microsculpture on infraorbital ridges and neck; pronotum without microsculpture except for median third of disc with irregular and indistinct transverse microsculpture; scutellum with indistinct transverse microsculpture; abdominal tergites with very dense and distinct isodiametric microsculpture. Head without punctation except for posterior portion between grooves and infraorbital ridges with several smoothed large punctures; pronotum with dense, regular, small punctation, slightly deeper and larger on posterio-lateral portions; punctation of elytra dense, large and smoothed, distinctly denser, smaller and deeper on prescutellar area; abdominal tergites with irregular, indistinct and small punctures. Head with several very long setae on frontal portions, with two long yellow setae on median part of head between anterior parets of eyes and four pairs of long setae on posterior part of infraorbital ridge along occipital line; pronotum with dense, and very fine, regular yellow pubescence, and with two pairs of very long black setae near anterior margin and four pairs of similar setae on lateral margins; fine pubescence of elytra as that on pronotum, with additional five to six pairs of very long black setae on lateral portions; abdominal tergites with irregular setation and with long and erect black setae on paratergites and apical segments.</paragraph>
<caption id="06F7660EFFBF2E25248FF8717BF1F864" httpUri="https://zenodo.org/record/2607218/files/figure.png" pageId="2" pageNumber="379" startId="2.[151,264,1954,1977]" targetBox="[296,1236,546,1939]" targetPageId="2">
<paragraph id="52373686FFBF2E25248FF8717BF1F864" blockId="2.[151,1437,1954,2008]" pageId="2" pageNumber="379">
<emphasis id="60FCEA94FFBF2E25248FF8717924F804" bold="true" box="[151,316,1954,1977]" pageId="2" pageNumber="379">FIGURES 1–4.</emphasis>
Habitus of
<taxonomicName id="95884D05FFBF2E2525A0F8707A5FF805" authorityName="Kraatz" authorityYear="1857" box="[440,583,1955,1977]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="genus">
<emphasis id="60FCEA94FFBF2E2525A0F8707A5FF805" box="[440,583,1955,1977]" italics="true" pageId="2" pageNumber="379">Amphichroum</emphasis>
</taxonomicName>
(males): 1–
<taxonomicName id="95884D05FFBF2E2526DAF8777B66F805" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[706,894,1955,1977]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="angustilobatum" status="sp. nov.">
<emphasis id="60FCEA94FFBF2E2526DAF8777B66F805" box="[706,894,1955,1977]" italics="true" pageId="2" pageNumber="379">A. angustilobatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BCF57EFFFBF2E25279CF8777BADF805" box="[900,949,1956,1977]" pageId="2" pageNumber="379" rank="species">sp.n.</taxonomicNameLabel>
(holotype), 2–
<taxonomicName id="95884D05FFBF2E252057F8777CADF805" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[1103,1205,1956,1977]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="assingi" status="sp. nov.">
<emphasis id="60FCEA94FFBF2E252057F8777CADF805" box="[1103,1205,1956,1977]" italics="true" pageId="2" pageNumber="379">A. assingi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BCF57EFFFBF2E2520A4F8777CF4F805" box="[1212,1260,1956,1977]" pageId="2" pageNumber="379" rank="species">sp.n.</taxonomicNameLabel>
(holotype), 3–
<taxonomicName id="95884D05FFBF2E25219EF8777906F864" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="longilobatum" status="sp. nov.">
<emphasis id="60FCEA94FFBF2E25219EF8777906F864" italics="true" pageId="2" pageNumber="379">A. longilobatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BCF57EFFFBF2E25253CF810794CF864" box="[292,340,1987,2008]" pageId="2" pageNumber="379" rank="species">sp.n.</taxonomicNameLabel>
(paratype), 4–
<taxonomicName id="95884D05FFBF2E2525F0F8117A6FF864" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[488,631,1986,2008]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="379" phylum="Arthropoda" rank="species" species="maculosum" status="sp. nov.">
<emphasis id="60FCEA94FFBF2E2525F0F8117A6FF864" box="[488,631,1986,2008]" italics="true" pageId="2" pageNumber="379">A. maculosum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BCF57EFFFBF2E252666F8107AB6F864" box="[638,686,1987,2008]" pageId="2" pageNumber="379" rank="species">sp.n.</taxonomicNameLabel>
(holotype). Scale bar: 1.0 mm.
</paragraph>
</caption>
<caption id="06F7660EFFBE2E24248FF8827D29F83B" httpUri="https://zenodo.org/record/2607222/files/figure.png" pageId="3" pageNumber="380" startId="3.[151,264,1873,1896]" targetBox="[231,1335,218,1829]" targetPageId="3">
<paragraph id="52373686FFBE2E24248FF8827D29F83B" blockId="3.[151,1436,1873,1927]" pageId="3" pageNumber="380">
<emphasis id="60FCEA94FFBE2E24248FF8827950F8DB" bold="true" box="[151,328,1873,1896]" pageId="3" pageNumber="380">FIGURES 5–10.</emphasis>
Aedeagus of
<taxonomicName id="95884D05FFBE2E2425CEF8817B11F8D4" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[470,777,1874,1896]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="angustilobatum" status="sp. nov.">
<emphasis id="60FCEA94FFBE2E2425CEF8817B11F8D4" box="[470,777,1874,1896]" italics="true" pageId="3" pageNumber="380">Amphichroum angustilobatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BCF57EFFFBE2E242708F8817B58F8D4" box="[784,832,1874,1896]" pageId="3" pageNumber="380" rank="species">sp.n.</taxonomicNameLabel>
(5–7: holotype) and
<taxonomicName id="95884D05FFBE2E242017F8807C6CF8D4" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[1039,1140,1875,1896]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="380" phylum="Arthropoda" rank="species" species="assingi" status="sp. nov.">
<emphasis id="60FCEA94FFBE2E242017F8807C6CF8D4" box="[1039,1140,1875,1896]" italics="true" pageId="3" pageNumber="380">A. assingi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BCF57EFFFBE2E242062F8807CB2F8D4" box="[1146,1194,1875,1896]" pageId="3" pageNumber="380" rank="species">sp.n.</taxonomicNameLabel>
(8–10: paratype): 5, 8– parameral view (outline of internal sac shown as point line), 6, 9–internal sac, 7, 10–lateral view. Scale bar: 0.1 mm.
</paragraph>
</caption>
<caption id="06F7660EFFB92E23248FFBCC79C1FBE8" httpUri="https://zenodo.org/record/2607224/files/figure.png" pageId="4" pageNumber="381" startId="4.[151,250,1055,1077]" targetBox="[228,1360,185,1034]" targetPageId="4">
<paragraph id="52373686FFB92E23248FFBCC79C1FBE8" blockId="4.[151,1436,1055,1108]" pageId="4" pageNumber="381">
<emphasis id="60FCEA94FFB92E23248FFBCC7939FB88" bold="true" box="[151,289,1055,1077]" pageId="4" pageNumber="381">FIGURE 11.</emphasis>
Type locality of
<taxonomicName id="95884D05FFB92E2325C0FBCC7B16FB89" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[472,782,1055,1077]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="angustilobatum" status="sp. nov.">
<emphasis id="60FCEA94FFB92E2325C0FBCC7B16FB89" box="[472,782,1055,1077]" italics="true" pageId="4" pageNumber="381">Amphichroum angustilobatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="7BCF57EFFFB92E23270CFBCC7B5AFB89" box="[788,834,1055,1077]" pageId="4" pageNumber="381" rank="species">sp.n.</taxonomicNameLabel>
in the Meili Xue Shan range in Yunnan (photograph: M. Schülke (locality C2005-07A)).
</paragraph>
</caption>
<paragraph id="52373686FFB92E2324DFFBA87A05FA2C" blockId="4.[151,1437,1147,2036]" pageId="4" pageNumber="381">
Head subtrapezoidal, 1.7 times as wide as long, flattened in middle and slightly convex on posterior portions of infraorbital ridges; grooves in front of ocelli narrow, deep and long, diagonally stretching toward antennal bases to level of anterior margins of eyes. Ocelli very large, located at level of posterior margins of eyes near occipital furrow, distance between ocelli slightly longer than distance between ocellus and posterior margin of eye. Eyes very large, convex. Apical segment of maxillary palp 1.2 times as long as preceding segment, gradually narrowed apicad from basal third. Antenna moderately long, reaching one-fourth of elytral length when reclined; length × width of antennomeres (
<typeStatus id="8D338824FFB92E2325B0FA807A09FAD0" box="[424,529,1363,1388]" pageId="4" pageNumber="381" type="holotype">holotype</typeStatus>
): 1: 0.13 × 0.05; 2: 0.11 × 0.05; 3: 0.11 × 0.04; 4: 0.07 × 0.05; 5–8: 0.10 × 0.05; 9–10: 0.08 × 0.06; 11: 0.15 × 0.06.
</paragraph>
<paragraph id="52373686FFB92E2324DFFA4F7C7BFA40" blockId="4.[151,1437,1147,2036]" pageId="4" pageNumber="381">Pronotum 1.4–1.6 times as wide as long, 1.5–1.6 times as wide as head, widest at about middle, slightly more narrowed posteriad than anteriad; anterior angles rounded, indistinctly protruded apicad; posterior angles widely rounded; lateral portions flattened and slightly explanate, gradually widened posteriad.</paragraph>
<paragraph id="52373686FFB92E2324DFF9D47D1DF9FF" blockId="4.[151,1437,1147,2036]" pageId="4" pageNumber="381">Elytra slightly wider than long, distinctly widened apicad, reaching apical margin of abdominal tergite IV or V, slightly less than twice as long as pronotum, with widely rounded apical margins, truncated at suture.</paragraph>
<paragraph id="52373686FFB92E2324DFF9837A62F930" blockId="4.[151,1437,1147,2036]" pageId="4" pageNumber="381">Legs moderately long; metatibia about 1.1 times as long as metatarsus, with long strong thorns on inner and outer surface and long setae between them.</paragraph>
<paragraph id="52373686FFB92E2324DFF9447C18F96F" blockId="4.[151,1437,1147,2036]" pageId="4" pageNumber="381">Abdomen slightly or distinctly narrower than elytra, with a pair of very large transverse tomentose spots in middle of tergite IV, with wide palisade fringe on apical margin of tergite VII.</paragraph>
<paragraph id="52373686FFB92E2324DFF90C7A8AF84F" blockId="4.[151,1437,1147,2036]" pageId="4" pageNumber="381">
Male (
<figureCitation id="CAB32A03FFB92E232508F90C7949F94B" box="[272,337,1759,1784]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1954,1977]" captionTargetBox="[296,1236,546,1939]" captionTargetId="figure@2.[283,1307,539,1939]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus of Amphichroum (males): 1–A. angustilobatum sp.n. (holotype), 2–A. assingi sp.n. (holotype), 3–A. longilobatum sp.n. (paratype), 4–A. maculosum sp.n. (holotype). Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/2607218/files/figure.png" pageId="4" pageNumber="381">Fig. 1</figureCitation>
). Protarsomeres 1–4 very wide. Medial margin of apical part of protibia with two parallel rows of five short peg setae; mesotibia strongly curved mediad, with dense row of fourteen very short and wide thorns beginning from medial side of bend and stretching to apex of mesotibia; lateral margin of tibiae without long strong thorns, except for those around apex. Abdominal tergite VIII and sternite VIII with straight apical margin. Aedeagus (
<figureCitation id="CAB32A03FFB92E23250FF8BC7944F834" box="[279,348,1903,1928]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1873,1896]" captionTargetBox="[231,1335,218,1829]" captionTargetId="figure@3.[210,1377,193,1853]" captionTargetPageId="3" captionText="FIGURES 5–10. Aedeagus of Amphichroum angustilobatum sp.n.(5–7: holotype) and A. assingi sp.n. (8–10: paratype): 5, 8– parameral view (outline of internal sac shown as point line), 6, 9–internal sac, 7, 10–lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607222/files/figure.png" pageId="4" pageNumber="381">Fig. 5</figureCitation>
) with wide basal portion strongly narrowed apicad, median lobe narrow with elongated apical portion; parameres narrow, distinctly exceeding apex of median lobe, with two pairs of apical and preapical short setae; internal sac (
<figureCitation id="CAB32A03FFB92E23256AF86479AFF86C" box="[370,439,1975,2000]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1873,1896]" captionTargetBox="[231,1335,218,1829]" captionTargetId="figure@3.[210,1377,193,1853]" captionTargetPageId="3" captionText="FIGURES 5–10. Aedeagus of Amphichroum angustilobatum sp.n.(5–7: holotype) and A. assingi sp.n. (8–10: paratype): 5, 8– parameral view (outline of internal sac shown as point line), 6, 9–internal sac, 7, 10–lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607222/files/figure.png" pageId="4" pageNumber="381">Fig. 6</figureCitation>
) long, with two pairs of wide thorns in apical and complicated sclerotized structures in basal portion. Aedeagus laterally as in
<figureCitation id="CAB32A03FFB92E232651F8087A96F84F" box="[585,654,2011,2036]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1873,1896]" captionTargetBox="[231,1335,218,1829]" captionTargetId="figure@3.[210,1377,193,1853]" captionTargetPageId="3" captionText="FIGURES 5–10. Aedeagus of Amphichroum angustilobatum sp.n.(5–7: holotype) and A. assingi sp.n. (8–10: paratype): 5, 8– parameral view (outline of internal sac shown as point line), 6, 9–internal sac, 7, 10–lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607222/files/figure.png" pageId="4" pageNumber="381">Fig. 7</figureCitation>
.
</paragraph>
<paragraph id="52373686FFB82E2224DFFF447C26FF44" blockId="5.[151,1437,151,717]" pageId="5" pageNumber="382">Female. Protarsomeres 1–4 moderately narrow. Medial margin of protibia without peg setae; mesotibia without modifications, slender, gradually slightly widened apicad, with long strong thorns along entire length of lateral margin. Abdominal tergite VIII and sternite VIII with small rounded apical margin.</paragraph>
<paragraph id="52373686FFB82E2224DFFED77A1EFE10" blockId="5.[151,1437,151,717]" pageId="5" pageNumber="382">
<emphasis id="60FCEA94FFB82E2224DFFED779B7FEA1" bold="true" box="[199,431,260,285]" pageId="5" pageNumber="382">Comparative notes.</emphasis>
Based on the body size, coloration and general shape of the apical part of the aedeagus,
<taxonomicName id="95884D05FFB82E22219CFED7795DFEFC" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="angustilobatum" status="sp. nov.">
<emphasis id="60FCEA94FFB82E22219CFED7795DFEFC" italics="true" pageId="5" pageNumber="382">A. angustilobatum</emphasis>
</taxonomicName>
<emphasis id="60FCEA94FFB82E222555FEF4799DFEFC" bold="true" box="[333,389,295,320]" pageId="5" pageNumber="382">
<taxonomicNameLabel id="7BCF57EFFFB82E222555FEF4799DFEFC" box="[333,389,295,320]" pageId="5" pageNumber="382" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="95884D05FFB82E22264AFEFB7AF6FEFC" authorityName="Champion" authorityYear="1920" box="[594,750,296,320]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="pindarense">
<emphasis id="60FCEA94FFB82E22264AFEFB7AF6FEFC" box="[594,750,296,320]" italics="true" pageId="5" pageNumber="382">A. pindarense</emphasis>
</taxonomicName>
(
<collectingCountry id="2A9F7616FFB82E2226E6FEF47B24FEFC" box="[766,828,295,320]" name="India" pageId="5" pageNumber="382">India</collectingCountry>
,
<collectingCountry id="2A9F7616FFB82E222750FEF47B81FEFC" box="[840,921,295,320]" name="Bhutan" pageId="5" pageNumber="382">Bhutan</collectingCountry>
and Xizang) and
<taxonomicName id="95884D05FFB82E22207CFEFB7CF4FEFC" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[1124,1260,296,320]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="schuelkei" status="sp. nov.">
<emphasis id="60FCEA94FFB82E22207CFEFB7CF4FEFC" box="[1124,1260,296,320]" italics="true" pageId="5" pageNumber="382">A. schuelkei</emphasis>
</taxonomicName>
<emphasis id="60FCEA94FFB82E2220ECFEF47D34FEFC" bold="true" box="[1268,1324,295,320]" pageId="5" pageNumber="382">
<taxonomicNameLabel id="7BCF57EFFFB82E2220ECFEF47D34FEFC" box="[1268,1324,295,320]" pageId="5" pageNumber="382" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
(Yunnan), from which it differs by the wider basal part of the aedeagus and different structure of the internal sac. Additionally, from
<taxonomicName id="95884D05FFB82E2224CCFEA37943FE34" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[212,347,368,392]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="schuelkei" status="sp. nov.">
<emphasis id="60FCEA94FFB82E2224CCFEA37943FE34" box="[212,347,368,392]" italics="true" pageId="5" pageNumber="382">A. schuelkei</emphasis>
</taxonomicName>
<emphasis id="60FCEA94FFB82E22257AFEBC7982FE34" bold="true" box="[354,410,367,392]" pageId="5" pageNumber="382">
<taxonomicNameLabel id="7BCF57EFFFB82E22257AFEBC7982FE34" box="[354,410,367,392]" pageId="5" pageNumber="382" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
it differs by more distinct microsculpture of the head, longer antennomeres 4–10, and wider apical portions of the parameres.
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFB82E2224DFFE647B04FD39" pageId="5" pageNumber="382" type="distribution">
<paragraph id="52373686FFB82E2224DFFE6479B5FE48" blockId="5.[151,1437,151,717]" pageId="5" pageNumber="382">
<emphasis id="60FCEA94FFB82E2224DFFE647978FE6C" bold="true" box="[199,352,439,464]" pageId="5" pageNumber="382">Distribution.</emphasis>
The new species is at present known only from the
<typeStatus id="8D338824FFB82E2227B4FE6B7BC4FE6C" box="[940,988,440,464]" pageId="5" pageNumber="382">type</typeStatus>
locality in the Meili Xue Shan range in
<collectingRegion id="904CF864FFB82E22248FFE0E78E8FE48" box="[151,240,477,500]" country="China" name="Yunnan" pageId="5" pageNumber="382">Yunnan</collectingRegion>
,
<collectingCountry id="2A9F7616FFB82E2224E2FE0F7925FE49" box="[250,317,476,501]" name="China" pageId="5" pageNumber="382">China</collectingCountry>
(
<figureCitation id="CAB32A03FFB82E222555FE0F79B8FE48" box="[333,416,476,501]" captionStart="FIGURE 41" captionStartId="24.[151,250,1057,1079]" captionTargetBox="[179,1408,267,1035]" captionTargetId="figure@24.[179,1408,267,1035]" captionTargetPageId="24" captionText="FIGURE 41. Distribution of Amphichroum in China: A. angustilobatum sp.n. (black star), A. assingi sp.n. (white square with wide black margin), A. longilobatum sp.n. (white circle), A. maculosum sp.n. (white star), A. miaoershanum Watanabe, 1999 (circle with wide white margin), A. propinquum sp.n. (black square), A. rotundatum (white square), A. schuelkei sp.n. (black rhomb), A. squamosum sp.n. (black circle), A.subaequale sp.n. (white rhomb), A. subelongatum sp.n. (black triangle), A. tibetanum sp.n. (white triangle)." httpUri="https://zenodo.org/record/2607244/files/figure.png" pageId="5" pageNumber="382">Fig. 41</figureCitation>
).
</paragraph>
<paragraph id="52373686FFB82E2224DFFE2C7B04FD39" blockId="5.[151,1437,151,717]" pageId="5" pageNumber="382">
<emphasis id="60FCEA94FFB82E2224DFFE2C7952FDA4" bold="true" box="[199,330,511,536]" pageId="5" pageNumber="382">Bionomics.</emphasis>
Specimens were collected at elevation
<quantity id="95709B63FFB82E22271EFDD37B47FDAB" box="[774,863,512,536]" metricMagnitude="3" metricUnit="m" metricValue="2.8899999999999997" pageId="5" pageNumber="382" unit="m" value="2890.0">2890 m</quantity>
. The
<typeStatus id="8D338824FFB82E222785FE2C7BE7FDA4" box="[925,1023,511,536]" pageId="5" pageNumber="382" type="holotype">holotype</typeStatus>
was taken by sifting leaf litter, moss and dead wood in small creek valley in mixed forest with bamboo (locality C
<date id="26361046FFB82E2227E1FDF77C42FD81" box="[1017,1114,548,573]" pageId="5" pageNumber="382" value="2005" valueMax="2007">2005-07</date>
A:
<figureCitation id="CAB32A03FFB82E222062FDF77CD2FD80" box="[1146,1226,548,573]" captionStart="FIGURE 11" captionStartId="4.[151,250,1055,1077]" captionTargetBox="[228,1360,185,1034]" captionTargetId="figure@4.[228,1360,185,1034]" captionTargetPageId="4" captionText="FIGURE 11. Type locality of Amphichroum angustilobatum sp.n.in the Meili Xue Shan range in Yunnan (photograph: M. Schülke (locality C2005-07A))." httpUri="https://zenodo.org/record/2607224/files/figure.png" pageId="5" pageNumber="382">Fig. 11</figureCitation>
); the
<typeStatus id="8D338824FFB82E222111FDF67D71FD81" box="[1289,1385,549,573]" pageId="5" pageNumber="382" type="paratype">paratype</typeStatus>
was taken by sifting various debris along the creek and in a spray zone of a small waterfall in mixed broadleaved forest with broadleaved shrubs in undergrowth (locality: C160).
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFB82E2224DFFD5C7B22FD71" pageId="5" pageNumber="382" type="etymology">
<paragraph id="52373686FFB82E2224DFFD5C7B22FD71" blockId="5.[151,1437,151,717]" pageId="5" pageNumber="382">
<emphasis id="60FCEA94FFB82E2224DFFD5C7956FD14" bold="true" box="[199,334,655,680]" pageId="5" pageNumber="382">Etymology.</emphasis>
The specific epithet is a combination of Latin adjective
<emphasis id="60FCEA94FFB82E2227A7FD427C3BFD14" box="[959,1059,657,680]" italics="true" pageId="5" pageNumber="382">angustus</emphasis>
, -
<emphasis id="60FCEA94FFB82E222021FD427C5FFD14" box="[1081,1095,657,680]" italics="true" pageId="5" pageNumber="382">a</emphasis>
, -
<emphasis id="60FCEA94FFB82E222046FD427C98FD14" box="[1118,1152,657,680]" italics="true" pageId="5" pageNumber="382">um</emphasis>
(narrow) and
<emphasis id="60FCEA94FFB82E222106FD4378DAFD70" italics="true" pageId="5" pageNumber="382">lobatus, -a, -um</emphasis>
(lobed). It refers to narrow median lobe of the aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>
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