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<document id="B8BA062548D7883FFFBDD4EB336B3464" ID-CLB-Dataset="28041" ID-DOI="10.11646/zootaxa.4508.3.3" ID-GBIF-Dataset="9de070ed-9c35-4131-a350-72371ee93ddf" ID-ISSN="1175-5326" ID-Zenodo-Dep="2607216" ID-ZooBank="E53B4C53-554A-4913-8E3B-CE1C530DF004" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553586064605" checkinUser="plazi" docAuthor="Shavrin, Alexey V. &amp; Smetana, Aleš" docDate="2018" docId="DA218790FFB82E21248FFCCC7D8EFCA8" docLanguage="en" docName="zootaxa.4508.3.3.pdf" docOrigin="Zootaxa 4508 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphichroum assingi Shavrin &amp; Smetana 2018, sp.n." docType="treatment" docVersion="8" lastPageNumber="383" masterDocId="2618FFE8FFBD2E272418FFD37818FFBC" masterDocTitle="New species of the genus Amphichroum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) from China" masterLastPageNumber="402" masterPageNumber="377" pageNumber="382" updateTime="1698713048312" updateUser="ExternalLinkService">
<mods:mods id="E9B308E38E7B53A525BCF15C77C7D9DC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4EF84D65D8AAC1C39B20C8AC887EBC3E">
<mods:title id="8491CD479ED79C8D2356239938FD9A18">New species of the genus Amphichroum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) from China</mods:title>
</mods:titleInfo>
<mods:name id="D90291A0222C5D2BDC409B029E597D00" type="personal">
<mods:role id="2E9AC56CB894DF4527ADD73F2A619426">
<mods:roleTerm id="48B88BE14D0DE209B0D136FDB34249BE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B8FDF1588424334BD657BC558C5684EF">Shavrin, Alexey V.</mods:namePart>
</mods:name>
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<mods:role id="0CE4794C93B755C11C8F71B10B3C92F8">
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</mods:role>
<mods:namePart id="DC32F7653131271A3058ED9DA6D08D7B">Smetana, Aleš</mods:namePart>
</mods:name>
<mods:typeOfResource id="F4DF85767AD8C5319C9F9CFDA4B61191">text</mods:typeOfResource>
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<mods:title id="4FABFB4374A9112A913664C851D326DF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6E070F425FB43FCEA51E24A5AEB036A8">
<mods:date id="E41EDCB1909396CF662B067FFB894357">2018</mods:date>
<mods:detail id="01D19E0FADD180821714566AF5318836" type="pubDate">
<mods:number id="292A108E37333EB2909F7E70FF07A98A">2018-11-01</mods:number>
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<mods:detail id="0D4241D33083F3F97B10276EE6531135" type="volume">
<mods:number id="251B4961A16F33C034F6E70A8A821FFE">4508</mods:number>
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<mods:classification id="E4916CE5BE8ECC69FEB008DEFA103D04">journal article</mods:classification>
<mods:identifier id="C5BBE3E960B2F24904186173229D5677" type="CLB-Dataset">28041</mods:identifier>
<mods:identifier id="75DEFB457EA6B8B74766D8E927811DE3" type="DOI">10.11646/zootaxa.4508.3.3</mods:identifier>
<mods:identifier id="B603A777CD3F1B2FD18BDA9FEFE3FBE1" type="GBIF-Dataset">9de070ed-9c35-4131-a350-72371ee93ddf</mods:identifier>
<mods:identifier id="33CB8DE1E8C3F5757DF208275D2D45AD" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="DE925F05B226A06C1491B7CA58A6B04C" type="Zenodo-Dep">2607216</mods:identifier>
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</mods:mods>
<treatment id="DA218790FFB82E21248FFCCC7D8EFCA8" ID-DOI="http://doi.org/10.5281/zenodo.5962988" ID-GBIF-Taxon="154495879" ID-Zenodo-Dep="5962988" LSID="urn:lsid:plazi:treatment:DA218790FFB82E21248FFCCC7D8EFCA8" httpUri="http://treatment.plazi.org/id/DA218790FFB82E21248FFCCC7D8EFCA8" lastPageId="6" lastPageNumber="383" pageId="5" pageNumber="382">
<subSubSection id="1A92650DFFB82E22248FFCCC7AF3FC86" box="[151,747,799,826]" pageId="5" pageNumber="382" type="nomenclature">
<paragraph id="52373686FFB82E22248FFCCC7AF3FC86" blockId="5.[151,747,799,859]" box="[151,747,799,826]" pageId="5" pageNumber="382">
<heading id="097F81EAFFB82E22248FFCCC7AF3FC86" bold="true" box="[151,747,799,826]" fontSize="11" level="1" pageId="5" pageNumber="382" reason="1">
<emphasis id="60FCEA94FFB82E22248FFCCC7AF3FC86" bold="true" box="[151,747,799,826]" pageId="5" pageNumber="382">
<taxonomicName id="95884D05FFB82E22248FFCCC7ABCFC86" ID-CoL="CZPN" authority="Shavrin &amp; Smetana" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[151,676,799,826]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="assingi" status="sp. nov.">
<emphasis id="60FCEA94FFB82E22248FFCCC79BBFC85" bold="true" box="[151,419,799,825]" italics="true" pageId="5" pageNumber="382">Amphichroum assingi</emphasis>
Shavrin &amp; Smetana
</taxonomicName>
,
<taxonomicNameLabel id="7BCF57EFFFB82E2226A8FCF37AF3FC86" box="[688,747,800,826]" pageId="5" pageNumber="382" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFB82E22248FFC91797DFCE7" box="[151,357,834,859]" pageId="5" pageNumber="382" type="description">
<paragraph id="52373686FFB82E22248FFC91797DFCE7" blockId="5.[151,747,799,859]" box="[151,357,834,859]" pageId="5" pageNumber="382">
(
<figureCitation id="CAB32A03FFB82E222487FC9178F6FCE7" box="[159,238,834,859]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1954,1977]" captionTargetBox="[296,1236,546,1939]" captionTargetId="figure@2.[283,1307,539,1939]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus of Amphichroum (males): 1–A. angustilobatum sp.n. (holotype), 2–A. assingi sp.n. (holotype), 3–A. longilobatum sp.n. (paratype), 4–A. maculosum sp.n. (holotype). Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/2607218/files/figure.png" pageId="5" pageNumber="382">Figs. 2</figureCitation>
,
<figureCitation id="CAB32A03FFB82E2224E2FC90792DFCE7" box="[250,309,835,859]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1873,1896]" captionTargetBox="[231,1335,218,1829]" captionTargetId="figure@3.[210,1377,193,1853]" captionTargetPageId="3" captionText="FIGURES 5–10. Aedeagus of Amphichroum angustilobatum sp.n.(5–7: holotype) and A. assingi sp.n. (8–10: paratype): 5, 8– parameral view (outline of internal sac shown as point line), 6, 9–internal sac, 7, 10–lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607222/files/figure.png" pageId="5" pageNumber="382">8–10</figureCitation>
,
<figureCitation id="CAB32A03FFB82E222558FC907945FCE7" box="[320,349,835,859]" captionStart="FIGURE 41" captionStartId="24.[151,250,1057,1079]" captionTargetBox="[179,1408,267,1035]" captionTargetId="figure@24.[179,1408,267,1035]" captionTargetPageId="24" captionText="FIGURE 41. Distribution of Amphichroum in China: A. angustilobatum sp.n. (black star), A. assingi sp.n. (white square with wide black margin), A. longilobatum sp.n. (white circle), A. maculosum sp.n. (white star), A. miaoershanum Watanabe, 1999 (circle with wide white margin), A. propinquum sp.n. (black square), A. rotundatum (white square), A. schuelkei sp.n. (black rhomb), A. squamosum sp.n. (black circle), A.subaequale sp.n. (white rhomb), A. subelongatum sp.n. (black triangle), A. tibetanum sp.n. (white triangle)." httpUri="https://zenodo.org/record/2607244/files/figure.png" pageId="5" pageNumber="382">41</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFB82E22248FFC597C21FB8F" pageId="5" pageNumber="382" type="materials_examined">
<paragraph id="52373686FFB82E22248FFC597BFCFC57" blockId="5.[151,1437,906,2011]" pageId="5" pageNumber="382">
<emphasis id="60FCEA94FFB82E22248FFC5979D9FC1F" bold="true" box="[151,449,906,931]" pageId="5" pageNumber="382">Type material examined:</emphasis>
<materialsCitation id="E2E03CDBFFB82E2225DFFC597B7DFC7B" ID-GBIF-Occurrence="2028139628" collectingDate="2005-06-13" country="China" location="Judian" municipality="Pass" pageId="5" pageNumber="382" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="8D338824FFB82E2225DFFC597A37FC1F" box="[455,559,906,931]" pageId="5" pageNumber="382" type="holotype">Holotype</typeStatus>
<specimenCount id="448EFD0FFFB82E22262EFC587A53FC1E" box="[566,587,907,930]" pageId="5" pageNumber="382" type="male">♂</specimenCount>
[specimen without left metatarsus]: ‘
<collectingCountry id="2A9F7616FFB82E2227F1FC587C53FC1F" box="[1001,1099,907,931]" name="China" pageId="5" pageNumber="382">CHINA</collectingCountry>
:
<collectingRegion id="904CF864FFB82E22204DFC587CB2FC1E" box="[1109,1194,907,930]" country="China" name="Yunnan" pageId="5" pageNumber="382">Yunnan</collectingRegion>
|
<collectingMunicipality id="B253ACFCFFB82E2220A6FC587CE8FC1E" box="[1214,1264,907,930]" pageId="5" pageNumber="382">Pass</collectingMunicipality>
<quantity id="95709B63FFB82E2220EFFC597D26FC1F" box="[1271,1342,906,931]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="382" unit="km" value="50.0">50 km</quantity>
W from |
<location id="5757605DFFB82E2224BFFC7D78EBFC7B" LSID="urn:lsid:plazi:treatment:DA218790FFB82E21248FFCCC7D8EFCA8:5757605DFFB82E2224BFFC7D78EBFC7B" box="[167,243,942,967]" country="China" municipality="Pass" name="Judian" pageId="5" pageNumber="382" stateProvince="Yunnan">Judian</location>
, 10-
<date id="26361046FFB82E22253CFC7D7980FC7B" box="[292,408,942,967]" pageId="5" pageNumber="382" value="2005-06-13">
<collectingDate id="3672E9AEFFB82E22253CFC7D7980FC7B" box="[292,408,942,967]" pageId="5" pageNumber="382" value="2005-06-13">13.6.2005</collectingDate>
</date>
| Ivo Jeniš’ &lt;rectangular label, printed
</materialsCitation>
&gt;, ‘
<materialsCitation id="E2E03CDBFFB82E22279FFC7C7BF8FC57" ID-GBIF-Occurrence="2028139632" collectionCode="CA" collectorName="Shavrin A. &amp; Smetana A." country="China" pageId="5" pageNumber="382" specimenCount="1" typeStatus="holotype">
<typeStatus id="8D338824FFB82E22279FFC7C7C38FC7B" box="[903,1056,943,967]" pageId="5" pageNumber="382" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="95884D05FFB82E222024FC7D7CC7FC7B" authorityName="Kraatz" authorityYear="1857" box="[1084,1247,942,967]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="401" phylum="Arthropoda" rank="genus">Amphichroum</taxonomicName>
| assingi
<emphasis id="60FCEA94FFB82E22214BFC7D7D93FC7B" bold="true" box="[1363,1419,942,967]" pageId="5" pageNumber="382">
<taxonomicNameLabel id="7BCF57EFFFB82E22214BFC7D7D93FC7B" box="[1363,1419,942,967]" pageId="5" pageNumber="382" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
|
<collectorName id="FF7D5350FFB82E22248FFC017908FC57" box="[151,272,978,1003]" pageId="5" pageNumber="382">Shavrin A.</collectorName>
&amp;
<collectorName id="FF7D5350FFB82E22252CFC0079AFFC57" box="[308,439,979,1003]" pageId="5" pageNumber="382">Smetana A.</collectorName>
des. 2018’ &lt;red rectangular label, printed&gt; (
<collectionCode id="3499AE43FFB82E2227B4FC007BC0FC57" box="[940,984,979,1003]" country="USA" name="Chicago Academy of Sciences" pageId="5" pageNumber="382" type="Museum">CA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="52373686FFB82E2224DFFC247C21FB8F" blockId="5.[151,1437,906,2011]" pageId="5" pageNumber="382">
<materialsCitation id="E2E03CDBFFB82E2224DFFC247D84FBB3" ID-GBIF-Occurrence="2028139631" box="[199,1436,1014,1039]" collectionCode="CAS, CA" collectorName="Shavrin A. &amp; Smetana A." country="China" pageId="5" pageNumber="382" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="8D338824FFB82E2224DFFC247921FBB3" box="[199,313,1015,1039]" pageId="5" pageNumber="382" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="448EFD0FFFB82E22255BFC247977FBB2" box="[323,367,1015,1039]" pageId="5" pageNumber="382" type="male">1 ♂</specimenCount>
,
<specimenCount id="448EFD0FFFB82E222564FC2479B0FBB2" box="[380,424,1015,1039]" pageId="5" pageNumber="382" type="female">1 ♀</specimenCount>
: same data as the holotype (
<specimenCount id="448EFD0FFFB82E2226F1FC247B0CFBB2" box="[745,788,1015,1039]" pageId="5" pageNumber="382" type="male">1 ♂</specimenCount>
:
<collectionCode id="3499AE43FFB82E222738FC247B42FBB3" box="[800,858,1015,1039]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="5" pageNumber="382" type="Museum">CAS</collectionCode>
;
<specimenCount id="448EFD0FFFB82E22277EFC247B8BFBB2" box="[870,915,1015,1039]" pageId="5" pageNumber="382" type="female">1 ♀</specimenCount>
:
<collectionCode id="3499AE43FFB82E2227B8FC247BD3FBB3" box="[928,971,1015,1039]" country="USA" name="Chicago Academy of Sciences" pageId="5" pageNumber="382" type="Museum">CA</collectionCode>
). All
<typeStatus id="8D338824FFB82E222011FC247C6CFBB3" box="[1033,1140,1015,1039]" pageId="5" pageNumber="382" type="paratype">paratypes</typeStatus>
with red rectangular label
</materialsCitation>
‘
<typeStatus id="8D338824FFB82E222483FBC87934FB8F" box="[155,300,1051,1075]" pageId="5" pageNumber="382" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="95884D05FFB82E222527FBC979FAFB8F" authorityName="Kraatz" authorityYear="1857" box="[319,482,1050,1075]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="401" phylum="Arthropoda" rank="genus">Amphichroum</taxonomicName>
| assingi
<emphasis id="60FCEA94FFB82E222652FBC97A9AFB8F" bold="true" box="[586,642,1050,1075]" pageId="5" pageNumber="382">
<taxonomicNameLabel id="7BCF57EFFFB82E222652FBC97A9AFB8F" box="[586,642,1050,1075]" pageId="5" pageNumber="382" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
| Shavrin A. &amp; Smetana A. des. 2018’.
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFB82E2124DFFBED7B1DFD38" lastPageId="6" lastPageNumber="383" pageId="5" pageNumber="382" type="description">
<paragraph id="52373686FFB82E2224DFFBED78C0FB7F" blockId="5.[151,1437,906,2011]" pageId="5" pageNumber="382">
<emphasis id="60FCEA94FFB82E2224DFFBED7949FBEB" bold="true" box="[199,337,1086,1111]" pageId="5" pageNumber="382">Description</emphasis>
. Measurements (n=3): HW: 0.70–0.82; HL: 0.37–0.47; AL (
<typeStatus id="8D338824FFB82E222019FBED7C72FBEB" box="[1025,1130,1086,1111]" pageId="5" pageNumber="382" type="holotype">holotype</typeStatus>
): 1.32; OL: 0.18–0.20; PL: 0.67–0.79; PW: 1.07–1.33; ESL: 1.25–1.55; EW: 1.22–1.54; AW: 1.20–1.55; MTbL (
<typeStatus id="8D338824FFB82E222089FBB17CE2FBC7" box="[1169,1274,1122,1147]" pageId="5" pageNumber="382" type="holotype">holotype</typeStatus>
): 0.77; MTrL (
<typeStatus id="8D338824FFB82E222487FB557910FB23" box="[159,264,1158,1183]" pageId="5" pageNumber="382" type="holotype">holotype</typeStatus>
): 0.60 (MTrL 1–4: 0.35; MTrL 5: 0.25); AedL (
<typeStatus id="8D338824FFB82E222722FB557BBBFB23" box="[826,931,1158,1183]" pageId="5" pageNumber="382" type="holotype">holotype</typeStatus>
): 0.85; TL: 3.55 (
<typeStatus id="8D338824FFB82E22206BFB557CC0FB23" box="[1139,1240,1158,1183]" pageId="5" pageNumber="382" type="holotype">holotype</typeStatus>
)–4.95 (
<typeStatus id="8D338824FFB82E222129FB557D81FB23" box="[1329,1433,1158,1183]" pageId="5" pageNumber="382" type="holotype">holotype</typeStatus>
: 3.55).
</paragraph>
<paragraph id="52373686FFB82E2224DFFB1D79DEF9F3" blockId="5.[151,1437,906,2011]" pageId="5" pageNumber="382">
Head, disc of pronotum and apical segment of maxillary palpomere reddish-brown; mouthparts, antennomeres 1–3, lateral portions of pronotum, legs, elytra and paratergites yellow to yellowish-brown; antennomeres 4–11 brown; abdomen reddish-brown to brown (female
<typeStatus id="8D338824FFB82E2226C2FAC47B24FA93" box="[730,828,1303,1327]" pageId="5" pageNumber="382" type="paratype">paratype</typeStatus>
) immature, yellow-brown with slightly darker head). Head with dense transverse microsculpture, somewhat diagonal on posterior portions of infraorbital ridges; neck with coarse isodiametric microsculpture; pronotum with dense, and coarse isodiametric microsculpture, sometimes sparser on medio-basal third (
<typeStatus id="8D338824FFB82E222616FA517A6EFA27" box="[526,630,1410,1435]" pageId="5" pageNumber="382" type="holotype">holotype</typeStatus>
); scutellum with transverse coarse meshes; elytra with indistinct isodiametric microsculpture, more distinct and coarser near scutellum. Head with indistinct to distinct punctation, denser and deeper on posterior portion between grooves and on infraorbital ridges (one
<typeStatus id="8D338824FFB82E2220CFFA187D2FFA5F" box="[1239,1335,1483,1507]" pageId="5" pageNumber="382" type="paratype">paratype</typeStatus>
without punctation except for several very indistinct punctures on infraorbital ridges); pronotum with indistinct to distinct small punctation, sparser on medio-basal third; scutellum without punctures; punctation of elytra dense, moderately large and deep.
</paragraph>
<paragraph id="52373686FFB82E2224DFF9897B1FF8BF" blockId="5.[151,1437,906,2011]" pageId="5" pageNumber="382">
Head 1.7–1.8 times as wide as long, slightly convex posteriorly; grooves in front of ocelli very deep and wide, narrowing apicad. Ocelli located lower of level of posterior margin of eyes, distance between ocelli distinctly shorter than distance between ocellus and posterior margin of eye. Eyes large and strongly convex. Length × width of antennomeres (
<typeStatus id="8D338824FFB82E22257AF91579D3F963" box="[354,459,1734,1759]" pageId="5" pageNumber="382" type="holotype">holotype</typeStatus>
): length × width of antennomeres (
<typeStatus id="8D338824FFB82E222740F9157BD9F963" box="[856,961,1734,1759]" pageId="5" pageNumber="382" type="holotype">holotype</typeStatus>
): 1: 0.16 × 0.05; 2: 0.12 × 0.05; 3: 0.12 × 0.04; 4: 0.10 × 0.05; 5–10: 0.11 × 0.06; 11: 0.16 × 0.06.
</paragraph>
<paragraph id="52373686FFB82E2224DFF8DC795EF8D3" blockId="5.[151,1437,906,2011]" pageId="5" pageNumber="382">
Pronotum 1.5–1.6 times as wide as long, evenly narrowed both anteriad and posteriad (
<typeStatus id="8D338824FFB82E2220AAF8DD7D00F89B" box="[1202,1304,1806,1831]" pageId="5" pageNumber="382" type="holotype">holotype</typeStatus>
) or slightly more narrowed anteriad than posteriad; anterior angles rounded, distinctly protruded anteriad; lateral portions markedly wide.
</paragraph>
<paragraph id="52373686FFB82E2224DFF8A97A50F80A" blockId="5.[151,1437,906,2011]" pageId="5" pageNumber="382">Elytra slightly longer than wide, indistinctly widened from middle, reaching basal margin of abdominal tergite IV, 1.8–1.9 times as long as pronotum.</paragraph>
<paragraph id="52373686FFB82E2224DFF8117A84F866" blockId="5.[151,1437,906,2011]" box="[199,668,1986,2011]" pageId="5" pageNumber="382">Metatibia 1.2 times as long as metatarsus.</paragraph>
<paragraph id="52373686FFBB2E2124DFFF447AFFFF0C" blockId="6.[151,1436,151,789]" box="[199,743,151,176]" pageId="6" pageNumber="383">Abdomen slightly narrower or wider than elytra.</paragraph>
<paragraph id="52373686FFBB2E2124DFFF6F7930FE10" blockId="6.[151,1436,151,789]" pageId="6" pageNumber="383">
Male (
<figureCitation id="CAB32A03FFBB2E212517FF6F794CFF68" box="[271,340,188,213]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1954,1977]" captionTargetBox="[296,1236,546,1939]" captionTargetId="figure@2.[283,1307,539,1939]" captionTargetPageId="2" captionText="FIGURES 1–4. Habitus of Amphichroum (males): 1–A. angustilobatum sp.n. (holotype), 2–A. assingi sp.n. (holotype), 3–A. longilobatum sp.n. (paratype), 4–A. maculosum sp.n. (holotype). Scale bar: 1.0 mm." httpUri="https://zenodo.org/record/2607218/files/figure.png" pageId="6" pageNumber="383">Fig. 2</figureCitation>
). Protarsomeres 1–4 wide. Medial margin of apical part of protibia with two parallel rows of six to eight short peg setae; mesotibia moderately curved mediad, with dense row consisiting of sixteen to eighteen short thorns. Apical margin of abdominal tergite VIII straight. Apical margin of sternite VIII slightly sinuate. Aedeagus (
<figureCitation id="CAB32A03FFBB2E212487FEF478F2FEFC" box="[159,234,295,320]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1873,1896]" captionTargetBox="[231,1335,218,1829]" captionTargetId="figure@3.[210,1377,193,1853]" captionTargetPageId="3" captionText="FIGURES 5–10. Aedeagus of Amphichroum angustilobatum sp.n.(5–7: holotype) and A. assingi sp.n. (8–10: paratype): 5, 8– parameral view (outline of internal sac shown as point line), 6, 9–internal sac, 7, 10–lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607222/files/figure.png" pageId="6" pageNumber="383">Fig. 8</figureCitation>
) with narrow median lobe, gradually narrowing to subacute apex; parameres narrow, significantly exceeding apex of median lobe, with two pairs of small apical and preapical setae; internal sac (
<figureCitation id="CAB32A03FFBB2E2120CDFE9F7D02FED8" box="[1237,1306,332,357]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1873,1896]" captionTargetBox="[231,1335,218,1829]" captionTargetId="figure@3.[210,1377,193,1853]" captionTargetPageId="3" captionText="FIGURES 5–10. Aedeagus of Amphichroum angustilobatum sp.n.(5–7: holotype) and A. assingi sp.n. (8–10: paratype): 5, 8– parameral view (outline of internal sac shown as point line), 6, 9–internal sac, 7, 10–lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607222/files/figure.png" pageId="6" pageNumber="383">Fig. 9</figureCitation>
) very long, with fields of large thorns in apical and basal portions and with ring-like structure in basal part. Aedeagus laterally as in
<figureCitation id="CAB32A03FFBB2E2124CAFE47793CFE10" box="[210,292,404,429]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1873,1896]" captionTargetBox="[231,1335,218,1829]" captionTargetId="figure@3.[210,1377,193,1853]" captionTargetPageId="3" captionText="FIGURES 5–10. Aedeagus of Amphichroum angustilobatum sp.n.(5–7: holotype) and A. assingi sp.n. (8–10: paratype): 5, 8– parameral view (outline of internal sac shown as point line), 6, 9–internal sac, 7, 10–lateral view. Scale bar: 0.1 mm." httpUri="https://zenodo.org/record/2607222/files/figure.png" pageId="6" pageNumber="383">Fig. 10</figureCitation>
.
</paragraph>
<paragraph id="52373686FFBB2E2124DFFE647C91FE49" blockId="6.[151,1436,151,789]" pageId="6" pageNumber="383">Female. Protarsomeres 1–4 moderately narrow. Medial margin of protibia and mesotibia without modifications. Abdominal tergite VIII and sternite VIII with small rounded apical margin.</paragraph>
<paragraph id="52373686FFBB2E2124DFFE2C7B1DFD38" blockId="6.[151,1436,151,789]" pageId="6" pageNumber="383">
<emphasis id="60FCEA94FFBB2E2124DFFE2C79AAFDA4" bold="true" box="[199,434,511,536]" pageId="6" pageNumber="383">Comparative notes.</emphasis>
Based on the body size, the elongated parameres and on the presence of fields of large thorns in the internal sac,
<taxonomicName id="95884D05FFBB2E2125DBFDF77A2BFD80" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[451,563,548,572]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="383" phylum="Arthropoda" rank="species" species="assingi" status="sp. nov.">
<emphasis id="60FCEA94FFBB2E2125DBFDF77A2BFD80" box="[451,563,548,572]" italics="true" pageId="6" pageNumber="383">A. assingi</emphasis>
</taxonomicName>
<emphasis id="60FCEA94FFBB2E212625FDF77A6CFD81" bold="true" box="[573,628,548,573]" pageId="6" pageNumber="383">
<taxonomicNameLabel id="7BCF57EFFFBB2E212625FDF77A6CFD81" box="[573,628,548,573]" pageId="6" pageNumber="383" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="95884D05FFBB2E21271FFDF77BB2FD80" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[775,938,548,572]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="383" phylum="Arthropoda" rank="species" species="rotundatum" status="sp. nov.">
<emphasis id="60FCEA94FFBB2E21271FFDF77BB2FD80" box="[775,938,548,572]" italics="true" pageId="6" pageNumber="383">A. rotundatum</emphasis>
</taxonomicName>
<emphasis id="60FCEA94FFBB2E2127ABFDF77BF2FD81" bold="true" box="[947,1002,548,573]" pageId="6" pageNumber="383">
<taxonomicNameLabel id="7BCF57EFFFBB2E2127ABFDF77BF2FD81" box="[947,1002,548,573]" pageId="6" pageNumber="383" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="95884D05FFBB2E212004FDF77CB6FD80" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[1052,1198,548,572]" class="Insecta" family="Staphylinidae" genus="Amphichroum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="383" phylum="Arthropoda" rank="species" species="tibetanum" status="sp. nov.">
<emphasis id="60FCEA94FFBB2E212004FDF77CB6FD80" box="[1052,1198,548,572]" italics="true" pageId="6" pageNumber="383">A. tibetanum</emphasis>
</taxonomicName>
<emphasis id="60FCEA94FFBB2E2120AFFDF77CF6FD81" bold="true" box="[1207,1262,548,573]" pageId="6" pageNumber="383">
<taxonomicNameLabel id="7BCF57EFFFBB2E2120AFFDF77CF6FD81" box="[1207,1262,548,573]" pageId="6" pageNumber="383" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
, from which it differs by the shape of anterior angles of the pronotum distinctly protruded anteriad, by the narrowed apical portion of the median lobe and by structures of the internal sac.
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFBB2E2124DFFD5C7B34FD4C" pageId="6" pageNumber="383" type="distribution">
<paragraph id="52373686FFBB2E2124DFFD5C79BEFD70" blockId="6.[151,1436,151,789]" pageId="6" pageNumber="383">
<emphasis id="60FCEA94FFBB2E2124DFFD5C7978FD14" bold="true" box="[199,352,655,680]" pageId="6" pageNumber="383">Distribution.</emphasis>
The new species is at present known only from the
<typeStatus id="8D338824FFBB2E2127A7FD437BF7FD14" box="[959,1007,656,680]" pageId="6" pageNumber="383">type</typeStatus>
locality in the Hengduan Mountains,
<collectingRegion id="904CF864FFBB2E21248FFD6678E8FD70" box="[151,240,693,716]" country="China" name="Yunnan" pageId="6" pageNumber="383">Yunnan</collectingRegion>
,
<collectingCountry id="2A9F7616FFBB2E2124E2FD677959FD71" box="[250,321,692,717]" name="China" pageId="6" pageNumber="383">China</collectingCountry>
(
<figureCitation id="CAB32A03FFBB2E212551FD677981FD70" box="[329,409,692,717]" captionStart="FIGURE 41" captionStartId="24.[151,250,1057,1079]" captionTargetBox="[179,1408,267,1035]" captionTargetId="figure@24.[179,1408,267,1035]" captionTargetPageId="24" captionText="FIGURE 41. Distribution of Amphichroum in China: A. angustilobatum sp.n. (black star), A. assingi sp.n. (white square with wide black margin), A. longilobatum sp.n. (white circle), A. maculosum sp.n. (white star), A. miaoershanum Watanabe, 1999 (circle with wide white margin), A. propinquum sp.n. (black square), A. rotundatum (white square), A. schuelkei sp.n. (black rhomb), A. squamosum sp.n. (black circle), A.subaequale sp.n. (white rhomb), A. subelongatum sp.n. (black triangle), A. tibetanum sp.n. (white triangle)." httpUri="https://zenodo.org/record/2607244/files/figure.png" pageId="6" pageNumber="383">Fig. 41</figureCitation>
).
</paragraph>
<paragraph id="52373686FFBB2E2124DFFD047B34FD4C" blockId="6.[151,1436,151,789]" box="[199,812,727,752]" pageId="6" pageNumber="383">
<emphasis id="60FCEA94FFBB2E2124DFFD047952FD4C" bold="true" box="[199,330,727,752]" pageId="6" pageNumber="383">Bionomics.</emphasis>
The detailed ecological data are unknown.
</paragraph>
</subSubSection>
<subSubSection id="1A92650DFFBB2E2124DFFD2F7D8EFCA8" box="[199,1430,764,789]" pageId="6" pageNumber="383" type="etymology">
<paragraph id="52373686FFBB2E2124DFFD2F7D8EFCA8" blockId="6.[151,1436,151,789]" box="[199,1430,764,789]" pageId="6" pageNumber="383">
<emphasis id="60FCEA94FFBB2E2124DFFD2F7956FCA9" bold="true" box="[199,334,764,789]" pageId="6" pageNumber="383">Etymology.</emphasis>
Patronymic, the species is named to honour our colleague, Volker Assing of Hannover,
<collectingCountry id="2A9F7616FFBB2E212130FD2F7D8AFCA8" box="[1320,1426,764,788]" name="Germany" pageId="6" pageNumber="383">Germany</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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