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<document id="EBDA8A83D55CAB19E3F589BAAC74BC82" ID-CLB-Dataset="276306" ID-DOI="10.1111/j.1096-3642.2011.00810.x" ID-GBIF-Dataset="f3f27409-856f-44d3-b62b-ea61d6668d7f" ID-ISSN="0024-4082" ID-Zenodo-Dep="5407725" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630347746520" checkinUser="marcus" docAuthor="Köhler, Frank &amp; Johnson, Michael S." docDate="2012" docId="D2543F71FC23FF99BB83FCFB5D5AC46E" docLanguage="en" docName="j.1096-3642.2011.00810.x.pdf" docOrigin="Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 165 (2)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2011.00810.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Amplirhagada tealei Köhler &amp; Johnson 2012, SP. NOV." docType="treatment" docVersion="7" lastPageNumber="360" masterDocId="2E6D4709FC35FF8EB814FF915E50C301" masterDocTitle="Species limits in molecular phylogenies: a cautionary tale from Australian land snails (Camaenidae: Amplirhagada Iredale, 1933)" masterLastPageNumber="362" masterPageNumber="337" pageNumber="359" updateTime="1699239322058" updateUser="plazi">
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<mods:title id="6B51B37A1BB0C4014C8E6568F23FD0D0">Species limits in molecular phylogenies: a cautionary tale from Australian land snails (Camaenidae: Amplirhagada Iredale, 1933)</mods:title>
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<mods:namePart id="AF45E98BA2ED6345FB175DFF4C5D6A1B">Köhler, Frank</mods:namePart>
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<mods:namePart id="08468E8149528004C2FBE46A52DA9C5F">Johnson, Michael S.</mods:namePart>
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<subSubSection id="12E7DDECFC23FF98BB83FCFB5B4CC07E" box="[919,1308,874,895]" pageId="22" pageNumber="359" type="nomenclature">
<paragraph id="5A428E67FC23FF98BB83FCFB5B4CC07E" blockId="22.[919,1308,874,928]" box="[919,1308,874,895]" pageId="22" pageNumber="359">
<heading id="010A390BFC23FF98BB83FCFB5B4CC07E" box="[919,1308,874,895]" centered="true" fontSize="9" level="2" pageId="22" pageNumber="359" reason="4">
<taxonomicName id="9DFDF5E4FC23FF98BB83FCFB5AFDC07E" ID-CoL="D7S5" authority="Köhler &amp; Johnson, 2012" authorityName="Köhler &amp; Johnson" authorityYear="2012" box="[919,1197,874,895]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="359" phylum="Mollusca" rank="species" species="tealei" status="sp. nov.">AMPLIRHAGADA TEALEI</taxonomicName>
<emphasis id="68895275FC23FF98BCA3FCFF5B4CC07E" bold="true" box="[1207,1308,874,895]" pageId="22" pageNumber="359">
<taxonomicNameLabel id="73BAEF0EFC23FF98BCA3FCFF5B4CC07E" box="[1207,1308,874,895]" pageId="22" pageNumber="359" rank="species">SP. NOV.</taxonomicNameLabel>
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<subSubSection id="12E7DDECFC23FF99BBF5FC1A5D5AC46E" lastPageId="23" lastPageNumber="360" pageId="22" pageNumber="359" type="description">
<paragraph id="5A428E67FC23FF98BBF5FC1A5A82C0A1" blockId="22.[919,1308,874,928]" box="[993,1234,906,928]" pageId="22" pageNumber="359">
<figureCitation id="C2C692E2FC23FF98BBF5FC1A5A2AC09E" box="[993,1146,906,928]" captionStart="Figure 21" captionStartId="21.[826,905,786,805]" captionTargetBox="[833,1441,200,756]" captionTargetId="figure-384@21.[826,1442,196,756]" captionTargetPageId="21" captionText="Figure 21. Shells of CLADES 4 and 5. A, Amplirhagada imperialis sp. nov. paratype WAM S42949 (Boongaree Island). B, A. boongareensis Köhler, 2010a holotype WAM S34616 (Boongaree Island). C, Amplirhagada tealei sp. nov. paratype WAM 42951 (Prince Regent Reserve). Note that foot tissue protrudes from some of the shells. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5407761" httpUri="https://zenodo.org/record/5407761/files/figure.png" pageId="22" pageNumber="359">FIGURES 21C</figureCitation>
,
<figureCitation id="C2C692E2FC23FF98BC9DFC1B5AF8C09E" box="[1161,1192,906,927]" captionStart="Figure 24" captionStartId="22.[806,885,573,592]" captionTargetBox="[806,1420,198,541]" captionTargetId="figure-447@22.[806,1422,197,544]" captionTargetPageId="22" captionText="Figure 24. Genital anatomy of Amplirhagada tealei sp. nov. holotype WAM S42950 (Prince Regent Reserve). Abbreviations: ag, albumen gland; at, atrium; bc, bursa copulatrix; hd, hermaphroditic duct; p, penis; rm, penial retractor muscle; so, spermoviduct; vd, vas deferens. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5407765" httpUri="https://zenodo.org/record/5407765/files/figure.png" pageId="22" pageNumber="359">24</figureCitation>
,
<figureCitation id="C2C692E2FC23FF98BCA2FC1B5A82C0A1" box="[1206,1234,906,928]" captionStart="Figure 25" captionStartId="23.[164,243,1401,1420]" captionTargetBox="[176,769,204,1363]" captionTargetId="figure-444@23.[164,781,196,1371]" captionTargetPageId="23" captionText="Figure 25. Penial anatomy of Amplirhagada tealei sp. nov. holotype WAM S42950 (Prince Regent Reserve). Abbreviations: at, atrium; iwp, inner penial wall; mp, main pilaster of inner penial wall; pv, penial verge; rm, penial retractor muscle; sh, penial sheath; vd, vas deferens; vg, vagina. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5407769" httpUri="https://zenodo.org/record/5407769/files/figure.png" pageId="22" pageNumber="359">25</figureCitation>
</paragraph>
<paragraph id="5A428E67FC23FF98BB32FC235A3EC705" blockId="22.[806,1421,945,1028]" pageId="22" pageNumber="359">
<materialsCitation id="EA95843AFC23FF98BB32FC235A3AC705" ID-GBIF-Occurrence="4435312336" latitude="-15.116501" location="Cape Brewster" longLatPrecision="1" longitude="124.924164" municipality="Kimberley" pageId="22" pageNumber="359" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
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<typeStatus id="854630C5FC23FF98BB32FC235D0AC0C6" box="[806,858,946,967]" pageId="22" pageNumber="359">Type</typeStatus>
locality:
</emphasis>
<collectingRegion id="98394085FC23FF98BBDEFC205AF1C0C6" box="[970,1185,945,967]" country="Australia" name="Western Australia" pageId="22" pageNumber="359">Western Australia</collectingRegion>
, near coast of
<collectingMunicipality id="BA26141DFC23FF98BD40FC205D3CC0E4" pageId="22" pageNumber="359">Kimberley</collectingMunicipality>
mainland, about
<quantity id="9D052382FC23FF98BC23FC5E5AD4C0E4" box="[1079,1156,975,997]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="22" pageNumber="359" unit="km" value="1.6">1.6 km</quantity>
east of
<location id="5F22D8BCFC23FF98BCCAFC415BD9C0E4" LSID="urn:lsid:plazi:treatment:D2543F71FC23FF99BB83FCFB5D5AC46E:5F22D8BCFC23FF98BCCAFC415BD9C0E4" box="[1246,1417,976,997]" latitude="-15.116501" longLatPrecision="1" longitude="124.924164" municipality="Kimberley" name="Cape Brewster" pageId="22" pageNumber="359" stateProvince="Western Australia">Cape Brewster</location>
,
<geoCoordinate id="3FC9E8A0FC23FF98BB32FC7F5DE8C702" box="[806,952,1005,1028]" degrees="15" direction="south" minutes="06" orientation="latitude" pageId="22" pageNumber="359" precision="1" seconds="59.4" value="-15.116501">15°0659.4″S</geoCoordinate>
,
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.
</paragraph>
<paragraph id="5A428E67FC23FF98BB32FBA05A33C7A2" blockId="22.[806,1421,1073,1187]" pageId="22" pageNumber="359">
<emphasis id="68895275FC23FF98BB32FBA05DAFC747" box="[806,1023,1073,1094]" italics="true" pageId="22" pageNumber="359">Material examined</emphasis>
(
<tableCitation id="177FBBDCFC23FF98BC06FBA05A34C746" box="[1042,1124,1073,1095]" captionStart="Table 1" captionStartId="3.[144,208,429,450]" captionText="Table 1. Details of the material studied" pageId="22" pageNumber="359">Table 1</tableCitation>
):
<materialsCitation id="EA95843AFC23FF98BC6BFBA05A46C767" ID-GBIF-Occurrence="4435312306" accessionNumber="S42950" collectionCode="WAM" pageId="22" pageNumber="359" specimenCount="1" typeStatus="holotype">
<typeStatus id="854630C5FC23FF98BC6BFBA05AB5C746" box="[1151,1253,1073,1095]" pageId="22" pageNumber="359" type="holotype">Holotype</typeStatus>
<collectionCode id="3CEC16A2FC23FF98BCE4FBA05B7FC747" box="[1264,1327,1073,1094]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="22" pageNumber="359" type="Museum">WAM</collectionCode>
<accessionNumber id="45AE1384FC23FF98BD2DFBA05BDDC746" box="[1337,1421,1073,1095]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S42950" pageId="22" pageNumber="359" type="EnaNcbi">S42950</accessionNumber>
(dissected specimen)
</materialsCitation>
.
<materialsCitation id="EA95843AFC23FF98BC30FBC15A0EC7A2" ID-GBIF-Occurrence="4435312333" accessionNumber="S42951" collectionCode="WAM, AM" pageId="22" pageNumber="359" specimenCount="1" typeStatus="paratype">
<typeStatus id="854630C5FC23FF98BC30FBC15AC7C764" box="[1060,1175,1104,1125]" pageId="22" pageNumber="359" type="paratype">Paratypes</typeStatus>
<collectionCode id="3CEC16A2FC23FF98BCB5FBC15AB0C764" box="[1185,1248,1104,1125]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="22" pageNumber="359" type="Museum">WAM</collectionCode>
<accessionNumber id="45AE1384FC23FF98BCFEFBC15B6FC767" box="[1258,1343,1104,1126]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S42951" pageId="22" pageNumber="359" type="EnaNcbi">S42951</accessionNumber>
(three preserved specimens;
<figureCitation id="C2C692E2FC23FF98BC3BFBFF5AC2C784" box="[1071,1170,1134,1157]" captionStart="Figure 21" captionStartId="21.[826,905,786,805]" captionTargetBox="[833,1441,200,756]" captionTargetId="figure-384@21.[826,1442,196,756]" captionTargetPageId="21" captionText="Figure 21. Shells of CLADES 4 and 5. A, Amplirhagada imperialis sp. nov. paratype WAM S42949 (Boongaree Island). B, A. boongareensis Köhler, 2010a holotype WAM S34616 (Boongaree Island). C, Amplirhagada tealei sp. nov. paratype WAM 42951 (Prince Regent Reserve). Note that foot tissue protrudes from some of the shells. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5407761" httpUri="https://zenodo.org/record/5407761/files/figure.png" pageId="22" pageNumber="359">Fig. 21C</figureCitation>
),
<collectionCode id="3CEC16A2FC23FF98BCBFFBFE5A84C785" box="[1195,1236,1135,1156]" country="Australia" name="Australian Museum" pageId="22" pageNumber="359" type="Museum">AM</collectionCode>
C.470994 (two preserved bodies) (
<tableCitation id="177FBBDCFC23FF98BC15FB1C5A04C7A2" box="[1025,1108,1165,1187]" captionStart="Table 1" captionStartId="3.[144,208,429,450]" captionText="Table 1. Details of the material studied" pageId="22" pageNumber="359">Table 1</tableCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="5A428E67FC23FF98BB32FB415D0EC640" blockId="22.[806,1422,1232,1345]" pageId="22" pageNumber="359">
<emphasis id="68895275FC23FF98BB32FB415DF4C7E4" box="[806,932,1232,1253]" italics="true" pageId="22" pageNumber="359">Etymology:</emphasis>
For Roy Teale, who collected the material and has contributed substantially to the collection of Western Australian camaenids; noun in the genitive case.
</paragraph>
<paragraph id="5A428E67FC23FF99BB32FAE15ADBC274" blockId="22.[806,1421,1391,1903]" lastBlockId="23.[826,1441,198,373]" lastPageId="23" lastPageNumber="360" pageId="22" pageNumber="359">
<emphasis id="68895275FC23FF98BB32FAE15DFEC684" box="[806,942,1392,1413]" italics="true" pageId="22" pageNumber="359">Description:</emphasis>
Shell (
<figureCitation id="C2C692E2FC23FF98BC03FAFE5A28C684" box="[1047,1144,1391,1413]" captionStart="Figure 21" captionStartId="21.[826,905,786,805]" captionTargetBox="[833,1441,200,756]" captionTargetId="figure-384@21.[826,1442,196,756]" captionTargetPageId="21" captionText="Figure 21. Shells of CLADES 4 and 5. A, Amplirhagada imperialis sp. nov. paratype WAM S42949 (Boongaree Island). B, A. boongareensis Köhler, 2010a holotype WAM S34616 (Boongaree Island). C, Amplirhagada tealei sp. nov. paratype WAM 42951 (Prince Regent Reserve). Note that foot tissue protrudes from some of the shells. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5407761" httpUri="https://zenodo.org/record/5407761/files/figure.png" pageId="22" pageNumber="359">Fig. 21C</figureCitation>
) dome shaped, broad with comparatively low spire, angulated periphery, flattened whorls, shallow suture (H = 15.1 ± 0.8, D = 22.4 ± 0.5, W = 4.7 ± 0.1, average ± SD of three shells); colour brownish, diffuse brown, sub-sutural and mid-whorl bands may be present, darker near lip; outer lip light purplish; teleoconch with regular axial growth lines. Umbilicus completely concealed by columellar reflection. Aperture simple, well expanded, slightly reflected, with weak basal node. Genital anatomy (
<figureCitation id="C2C692E2FC23FF98BB89F9305DA3C5B7" box="[925,1011,1697,1719]" captionStart="Figure 24" captionStartId="22.[806,885,573,592]" captionTargetBox="[806,1420,198,541]" captionTargetId="figure-447@22.[806,1422,197,544]" captionTargetPageId="22" captionText="Figure 24. Genital anatomy of Amplirhagada tealei sp. nov. holotype WAM S42950 (Prince Regent Reserve). Abbreviations: ag, albumen gland; at, atrium; bc, bursa copulatrix; hd, hermaphroditic duct; p, penis; rm, penial retractor muscle; so, spermoviduct; vd, vas deferens. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5407765" httpUri="https://zenodo.org/record/5407765/files/figure.png" pageId="22" pageNumber="359">Figs 24</figureCitation>
,
<figureCitation id="C2C692E2FC23FF98BC16F9305A70C5B6" box="[1026,1056,1697,1719]" captionStart="Figure 25" captionStartId="23.[164,243,1401,1420]" captionTargetBox="[176,769,204,1363]" captionTargetId="figure-444@23.[164,781,196,1371]" captionTargetPageId="23" captionText="Figure 25. Penial anatomy of Amplirhagada tealei sp. nov. holotype WAM S42950 (Prince Regent Reserve). Abbreviations: at, atrium; iwp, inner penial wall; mp, main pilaster of inner penial wall; pv, penial verge; rm, penial retractor muscle; sh, penial sheath; vd, vas deferens; vg, vagina. Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5407769" httpUri="https://zenodo.org/record/5407769/files/figure.png" pageId="22" pageNumber="359">25</figureCitation>
). Penis long, tubular, straight; vas deferens entering penial sheath near apical end, penial retractor muscle short; penial verge long (&gt; 1/3 length of penial chamber), elongated; inner penial wall with large, conical, corrugated main pilaster, comprising entire length of penial wall, broadest near anterior end, tapering in thickness and development described elsewhere (
<bibRefCitation id="3E6CF396FC22FF99BC2EFF575AB1C3DA" author="Kohler F" box="[1082,1249,198,220]" pageId="23" pageNumber="360" pagination="217 - 284" refId="ref17211" refString="Kohler F. 2010 a. Uncovering local endemism in the Kimberley, Western Australia: description of new species of the genus Amplirhagada Iredale, 1933 (Pulmonata, Camaenidae). Records of the Australian Museum 62: 217 - 284." type="journal article" year="2010">Köhler, 2010a</bibRefCitation>
,
<bibRefCitation id="3E6CF396FC22FF99BCE7FF575B69C3DA" author="Kohler F" box="[1267,1337,198,219]" pageId="23" pageNumber="360" pagination="167 - 202" refId="ref17348" refString="Kohler F. 2011 a. Descriptions of new species of the diverse and endemic land snail Amplirhagada Iredale, 1933 from rainforest patches across the Kimberley, Western Australia (Pulmonata, Camaenidae). Records of the Australian Museum 63: 167 - 202." type="journal article" year="2011">2011a</bibRefCitation>
, b). The penial wall sculpture is typical, consisting of dense longitudinal pilasters that cover the entire surface of the inner penial wall and a long corrugated main pilaster extending the entire whorl and being broadest close to the anterior end.
</paragraph>
<paragraph id="5A428E67FC22FF99BB2EFE3B5BF1C2DE" blockId="23.[826,1441,426,570]" pageId="23" pageNumber="360">
<emphasis id="68895275FC22FF99BB2EFE3B5D9CC2BE" box="[826,972,426,447]" italics="true" pageId="23" pageNumber="360">Distribution:</emphasis>
Known from the
<typeStatus id="854630C5FC22FF99BCDEFE3A5AABC2C1" box="[1226,1275,427,448]" pageId="23" pageNumber="360">type</typeStatus>
locality, the coastal portion of Prince Regent Reserve only (
<figureCitation id="C2C692E2FC22FF99BD45FE585BC4C2DE" box="[1361,1428,457,479]" captionStart="Figure 6" captionStartId="11.[826,905,885,904]" captionTargetBox="[826,1442,198,855]" captionTargetId="graphics-452@11.[604,1929,165,1235]" captionTargetPageId="11" captionText="Figure 6. Map of the study area with known occurrences of Amplirhagada species as documented by Solem (1981) and Köhler (2010a, 2011a, c). Triangles (in italics): Localities of known species: and, A. anderdonensis; bas, A. basilica; ber, A. berthierana; boo, A. boongareensis; buf, A. buffonensis; cam, A. camdenensis; cof, A. coffea; dec, A. decora; dis, A. discoidea; epi; A. epiphallica; for, A. forrestiana; gem, A. gemina; gib, A. gibsoni; glo, A. globosa; int, A. intermedia; inu, A. inusitata; kes, A. kessneri; kim, A. kimberleyana; lam, A. lamarckiana; lin, A. lindsayae; mon, A. montalivetensis; pue, A. puescheli; reg, A. regia; sph, A. sphaeroidea; tri, A. tricenaria; uwi, A. uwinsensis; via, A. vialae; yor, A. yorkensis. Circles (in bold): material studied herein: alb, A. albertiana sp. nov. (Albert Island); emo, A. ambulator sp. nov. (East Montalivet Island group); lul-wip, A. gemina; nma, A. intermedia (North Maret Island); prr, A. tealei sp. nov. (Mainland View, Prince Regent Reserve); sb, A. imperialis sp. nov. (Boongaree Island); sma, A. intermedia (South Maret and Natfi Island); tur, A. turbinensis sp. nov. (Turbin island); wal, A. montalivetensis (Walker Island); wmo, A. fitzpatricki sp. nov. (West Montalivet Island)." figureDoi="http://doi.org/10.5281/zenodo.5407731" httpUri="https://zenodo.org/record/5407731/files/figure.png" pageId="23" pageNumber="360">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="5A428E67FC22FF99BBDFFDB35B41C13B" blockId="23.[826,1441,426,570]" box="[971,1297,546,570]" pageId="23" pageNumber="360">ACKNOWLEDGEMENTS</paragraph>
<paragraph id="5A428E67FC22FF99BB2EFDC05A11C03F" blockId="23.[826,1441,593,830]" pageId="23" pageNumber="360">
Special thanks are due to Martin Püschel for producing, as usual, excellent drawings of genital organs. The study was made possible through grants from the Department of Environment and Conservation, Western Australia, and from the Australian Government (ABRS
<collectingCountry id="22EACEF7FC22FF99BBC4FD7A5DA4C000" box="[976,1012,747,769]" name="Russia" pageId="23" pageNumber="360">RF</collectingCountry>
211-10) to F.K. Thanks are also due to two anonymous reviewers for carefully reading the submitted manuscript.
</paragraph>
<caption id="0E82DEEFFC22FF99B8B0FAE85C43C51E" ID-DOI="http://doi.org/10.5281/zenodo.5407769" ID-Zenodo-Dep="5407769" httpUri="https://zenodo.org/record/5407769/files/figure.png" pageId="23" pageNumber="360" startId="23.[164,243,1401,1420]" targetBox="[176,769,204,1363]" targetPageId="23">
<paragraph id="5A428E67FC22FF99B8B0FAE85C43C51E" blockId="23.[164,779,1401,1567]" pageId="23" pageNumber="360">
<emphasis id="68895275FC22FF99B8B0FAE85F49C68D" bold="true" box="[164,281,1401,1420]" pageId="23" pageNumber="360">Figure 25.</emphasis>
Penial anatomy of
<emphasis id="68895275FC22FF99B9FAFAE85E83C6A8" bold="true" pageId="23" pageNumber="360">
<taxonomicName id="9DFDF5E4FC22FF99B9FAFAE85CB2C68D" authority="Köhler &amp; Johnson, 2012" authorityName="Köhler &amp; Johnson" authorityYear="2012" box="[494,738,1401,1420]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="23" pageNumber="360" phylum="Mollusca" rank="species" species="tealei" status="sp. nov.">
<emphasis id="68895275FC22FF99B9FAFAE85CB2C68D" bold="true" box="[494,738,1401,1420]" italics="true" pageId="23" pageNumber="360">Amplirhagada tealei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73BAEF0EFC22FF99BAFEFAE85E83C6A8" pageId="23" pageNumber="360" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype WAM
<accessionNumber id="45AE1384FC22FF99B983FA075FB4C6A8" box="[407,484,1430,1449]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S42950" pageId="23" pageNumber="360" type="EnaNcbi">S42950</accessionNumber>
(Prince Regent Reserve). Abbreviations: at, atrium; iwp, inner penial wall; mp, main pilaster of inner penial wall; pv, penial verge; rm, penial retractor muscle; sh, penial sheath; vd, vas deferens; vg, vagina. Scale bar = 5 mm.
</paragraph>
</caption>
<paragraph id="5A428E67FC22FF99B8B0F9D85CB3C5F9" blockId="23.[164,779,1609,1784]" pageId="23" pageNumber="360">posteriorly; entire inner penial wall covered with narrow, regular longitudinal pilasters, slightly corrugated near anterior end. No pustulation. Vagina moderate in length (shorter than penis), tubular; bursa copulatrix moderately long, tubular, extending to anterior end of spermoviduct, with inflated head.</paragraph>
<paragraph id="5A428E67FC22FF99B8B0F88D5D5AC46E" blockId="23.[164,778,1820,1903]" pageId="23" pageNumber="360">
<emphasis id="68895275FC22FF99B8B0F88D5FE4C430" box="[164,436,1820,1841]" italics="true" pageId="23" pageNumber="360">Comparative remarks:</emphasis>
Differs from all species described herein by its dome-shaped, globular shell, which is similar that of a number of other species
</paragraph>
</subSubSection>
</treatment>
</document>