236 lines
22 KiB
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236 lines
22 KiB
XML
<document ID-DOI="10.11646/zootaxa.5244.1.2" ID-GBIF-Dataset="289c3a9e-6b37-4b9d-bcd9-028360cc1b7f" ID-ISSN="1175-5326" ID-Zenodo-Dep="7645709" ID-ZooBank="4DC67EBE-0C0A-4F12-9E34-75C5A9194CA8" checkinTime="1676539257576" checkinUser="plazi" docAuthor="Tasaku, Yuto & Maruyama, Munetoshi" docDate="2023" docId="8F5D742CFFC3FFF542D6A1D9760B2740" docLanguage="en" docName="zootaxa.5244.1.2.pdf" docOrigin="Zootaxa 5244 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5244.1.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudothinusa Tasaku & Maruyama 2023, gen. nov." docType="treatment" docVersion="2" lastPageNumber="37" masterDocId="73640C54FFC2FFF14241A1067211233D" masterDocTitle="Pseudothinusa sunahama, a new genus and species of the tribe Athetini (Coleoptera, Staphylinidae, Aleocharinae) from sandy beaches in Japan" masterLastPageNumber="40" masterPageNumber="33" pageNumber="34" updateTime="1676539908277" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Pseudothinusa sunahama, a new genus and species of the tribe Athetini (Coleoptera, Staphylinidae, Aleocharinae) from sandy beaches in Japan</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tasaku, Yuto</mods:namePart>
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<mods:affiliation>Department of Biology, School of Biological Sciences, Tokai University, Hokkaido, 005 - 8601, Japan.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Maruyama, Munetoshi</mods:namePart>
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<mods:affiliation>The Kyushu University Museum, Hakozaki 6 - 10 - 1, Higashi-ku, Fukuoka, 812 - 8581, Japan.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-16</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5244</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>33</mods:start>
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<mods:end>40</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5244.1.2</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5244.1.2</mods:identifier>
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<mods:identifier type="GBIF-Dataset">289c3a9e-6b37-4b9d-bcd9-028360cc1b7f</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7645709</mods:identifier>
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<mods:identifier type="ZooBank">4DC67EBE-0C0A-4F12-9E34-75C5A9194CA8</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7645711" ID-Zenodo-Dep="7645711" LSID="urn:lsid:plazi:treatment:8F5D742CFFC3FFF542D6A1D9760B2740" httpUri="http://treatment.plazi.org/id/8F5D742CFFC3FFF542D6A1D9760B2740" lastPageId="4" lastPageNumber="37" pageId="1" pageNumber="34">
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<subSubSection pageId="1" pageNumber="34" type="nomenclature">
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<paragraph blockId="1.[151,731,223,285]" box="[151,731,223,250]" pageId="1" pageNumber="34">
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<heading bold="true" box="[151,731,223,250]" fontSize="11" level="1" pageId="1" pageNumber="34" reason="1">
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<emphasis bold="true" box="[151,731,223,250]" pageId="1" pageNumber="34">
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<taxonomicName authority="Tasaku & Maruyama" authorityName="Tasaku & Maruyama" authorityYear="2023" box="[151,609,223,250]" class="Insecta" family="Staphylinidae" genus="Pseudothinusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis bold="true" box="[151,330,223,250]" italics="true" pageId="1" pageNumber="34">Pseudothinusa</emphasis>
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Tasaku & Maruyama
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</taxonomicName>
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,
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<taxonomicNameLabel box="[621,731,223,249]" pageId="1" pageNumber="34" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="1.[151,731,223,285]" box="[151,288,259,285]" pageId="1" pageNumber="34">(Figs. 1–18)</paragraph>
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</subSubSection>
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<subSubSection box="[151,608,331,357]" pageId="1" pageNumber="34" type="type_taxon">
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<paragraph blockId="1.[151,1436,331,826]" box="[151,608,331,357]" pageId="1" pageNumber="34">
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<emphasis bold="true" box="[151,306,331,357]" pageId="1" pageNumber="34">
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<typeStatus box="[151,210,331,357]" pageId="1" pageNumber="34">Type</typeStatus>
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species.
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</emphasis>
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<emphasis box="[313,608,331,357]" italics="true" pageId="1" pageNumber="34">
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<taxonomicName authorityName="Tasaku & Maruyama" authorityYear="2023" box="[313,604,331,357]" class="Insecta" family="Staphylinidae" genus="Pseudothinusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" species="sunahama">Pseudothinusa sunahama</taxonomicName>
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.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection box="[199,1313,367,394]" pageId="1" pageNumber="34" type="etymology">
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<paragraph blockId="1.[151,1436,331,826]" box="[199,1313,367,394]" pageId="1" pageNumber="34">
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<emphasis bold="true" box="[199,334,367,393]" pageId="1" pageNumber="34">Etymology.</emphasis>
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Meaning “false
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<taxonomicName authorityName="Casey" authorityYear="1984" box="[520,610,367,393]" class="Insecta" family="Staphylinidae" genus="Thinusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis box="[520,610,367,393]" italics="true" pageId="1" pageNumber="34">Thinusa</emphasis>
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</taxonomicName>
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” because of its morphological similarity to the genus
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<taxonomicName authorityName="Casey" authorityYear="1984" box="[1216,1306,367,393]" class="Insecta" family="Staphylinidae" genus="Thinusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis box="[1216,1306,367,393]" italics="true" pageId="1" pageNumber="34">Thinusa</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="34" type="diagnosis">
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<paragraph blockId="1.[151,1436,331,826]" pageId="1" pageNumber="34">
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<emphasis bold="true" box="[199,322,403,429]" pageId="1" pageNumber="34">Diagnosis.</emphasis>
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<taxonomicName authorityName="Tasaku & Maruyama" authorityYear="2023" box="[334,499,403,429]" class="Insecta" family="Staphylinidae" genus="Pseudothinusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis box="[334,499,403,429]" italics="true" pageId="1" pageNumber="34">Pseudothinusa</emphasis>
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</taxonomicName>
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may be distinguished from other athetine genera by combination of following character states: front and middle tibiae with spines; integument with strong microsculpture; mentum with short v-seta; mandible with slightly sharp apex; complete infraorbital carina.
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</paragraph>
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<paragraph blockId="1.[151,1436,331,826]" pageId="1" pageNumber="34">
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Among the coastal athetine species in
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<collectingCountry box="[635,743,511,538]" name="China" pageId="1" pageNumber="34">East Asia</collectingCountry>
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,
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<taxonomicName authority="Tasaku & Maruyama, 2023" authorityName="Tasaku & Maruyama" authorityYear="2023" box="[756,919,511,537]" class="Insecta" family="Staphylinidae" genus="Pseudothinusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis box="[756,919,511,537]" italics="true" pageId="1" pageNumber="34">Pseudothinusa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[928,1031,512,538]" pageId="1" pageNumber="34" rank="genus">gen. nov.</taxonomicNameLabel>
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is the most similar to
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<taxonomicName authority="Casey, 1910" authorityName="Casey" authorityYear="1910" class="Insecta" family="Staphylinidae" genus="Adota" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis box="[1288,1354,511,537]" italics="true" pageId="1" pageNumber="34">Adota</emphasis>
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Casey,
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<quantity box="[151,206,547,573]" metricMagnitude="1" metricUnit="m" metricValue="4.8514" pageId="1" pageNumber="34" unit="in" value="1910.0">1910</quantity>
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</taxonomicName>
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in general appearance. However,
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<taxonomicName authorityName="Tasaku & Maruyama" authorityYear="2023" box="[578,741,547,573]" class="Insecta" family="Staphylinidae" genus="Pseudothinusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis box="[578,741,547,573]" italics="true" pageId="1" pageNumber="34">Pseudothinusa</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName authorityName="Casey" authorityYear="1910" box="[885,951,547,573]" class="Insecta" family="Staphylinidae" genus="Adota" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis box="[885,951,547,573]" italics="true" pageId="1" pageNumber="34">Adota</emphasis>
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</taxonomicName>
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in having a complete infraorbital carina and spines on the front and middle tibiae.
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<taxonomicName authorityName="Tasaku & Maruyama" authorityYear="2023" box="[572,735,583,609]" class="Insecta" family="Staphylinidae" genus="Pseudothinusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis box="[572,735,583,609]" italics="true" pageId="1" pageNumber="34">Pseudothinusa</emphasis>
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</taxonomicName>
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also differs from
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<taxonomicName authority="Yosii & Sawada, 1976" authorityName="Yosii & Sawada" authorityYear="1976" box="[934,1342,583,609]" class="Insecta" family="Staphylinidae" genus="Psammostiba" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis box="[934,1083,583,609]" italics="true" pageId="1" pageNumber="34">Psammostiba</emphasis>
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<bibRefCitation author="Yosii, R. & Sawada, K." box="[1090,1342,583,609]" pageId="1" pageNumber="34" pagination="11 - 140" refId="ref4082" refString="Yosii, R. & Sawada, K. (1976) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae). II: Diagnostic characters of genera and subgenera with description of representative species. Contributions from the Biological Laboratory, Kyoto University, 25 (1), 11 - 140." type="journal article" year="1976">Yosii & Sawada, 1976</bibRefCitation>
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</taxonomicName>
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, another seashore-dwelling athetine genus, in having a smaller body size (2.0–2.8), tarsi with single empodial seta and the presence of spines on the front and middle tibiae. Compared to the genus
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<taxonomicName authority="Sawada, 1990" authorityName="Sawada" authorityYear="1990" box="[984,1260,655,681]" class="Insecta" family="Staphylinidae" genus="Eubadura" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis box="[984,1095,655,681]" italics="true" pageId="1" pageNumber="34">Eubadura</emphasis>
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<bibRefCitation author="Sawada, K." box="[1104,1260,655,681]" pageId="1" pageNumber="34" pagination="273 - 307" refId="ref3672" refString="Sawada, K. (1990) New Species of Aleocharinae from Japan II (Coleoptera: Staphylinidae). Contributions from the Biological Laboratory, Kyoto University, 27 (3), 273 - 307." type="journal article" year="1990">Sawada, 1990</bibRefCitation>
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</taxonomicName>
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and the coastal species of the genus
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<taxonomicName box="[377,449,691,717]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus">
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<emphasis box="[377,449,691,717]" italics="true" pageId="1" pageNumber="34">Atheta</emphasis>
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</taxonomicName>
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,
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<taxonomicName authority="Tasaku & Maruyama, 2023" authorityName="Tasaku & Maruyama" authorityYear="2023" box="[463,626,691,717]" class="Insecta" family="Staphylinidae" genus="Pseudothinusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis box="[463,626,691,717]" italics="true" pageId="1" pageNumber="34">Pseudothinusa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[632,732,692,718]" pageId="1" pageNumber="34" rank="genus">gen. nov.</taxonomicNameLabel>
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is clearly distinguishable by its slenderer body and the presence of spines on the front and middle tibiae. See also Discussion for comparisons with coastal species in other regions.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="37" pageId="1" pageNumber="34" type="description">
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<paragraph blockId="1.[151,1436,331,826]" pageId="1" pageNumber="34">
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<emphasis bold="true" box="[199,345,763,789]" pageId="1" pageNumber="34">Description.</emphasis>
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Body (Fig. 1) flat and subparallel; Microsculpture of head, pronotum, and abdominal segments strong (Figs. 1–2).
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7645714" ID-Zenodo-Dep="7645714" httpUri="https://zenodo.org/record/7645714/files/figure.png" pageId="1" pageNumber="34" startId="1.[151,264,1955,1979]" targetBox="[167,1436,864,1918]" targetPageId="1">
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<paragraph blockId="1.[151,1436,1955,2016]" pageId="1" pageNumber="34">
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<emphasis bold="true" box="[151,307,1955,1979]" pageId="1" pageNumber="34">FIGURES 1–3</emphasis>
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<taxonomicName authority="Tasaku & Maruyama, 2023" authorityName="Tasaku & Maruyama" authorityYear="2023" box="[313,571,1955,1979]" class="Insecta" family="Staphylinidae" genus="Pseudothinusa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="34" phylum="Arthropoda" rank="species" species="sunahama" status="sp. nov.">
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<emphasis box="[313,571,1955,1979]" italics="true" pageId="1" pageNumber="34">Pseudothinusa sunahama</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[577,657,1956,1980]" pageId="1" pageNumber="34">
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<taxonomicNameLabel box="[577,657,1956,1980]" pageId="1" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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: 1, habitus, dorsal view; 2, head, dorsolateral view, SEM photograph; 3, front tibia. Scale bars: 1.0 mm for 1; 0.1 mm for 2 & 3.
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</paragraph>
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</caption>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7645716" ID-Zenodo-Dep="7645716" httpUri="https://zenodo.org/record/7645716/files/figure.png" pageId="2" pageNumber="35" startId="2.[151,264,1943,1967]" targetBox="[229,1353,194,1898]" targetPageId="2">
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<paragraph blockId="2.[151,1437,1943,2040]" pageId="2" pageNumber="35">
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<emphasis bold="true" box="[151,323,1943,1967]" pageId="2" pageNumber="35">FIGURES 4–10</emphasis>
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<taxonomicName authority="Tasaku & Maruyama, 2023" authorityName="Tasaku & Maruyama" authorityYear="2023" box="[331,592,1944,1968]" class="Insecta" family="Staphylinidae" genus="Pseudothinusa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="35" phylum="Arthropoda" rank="species" species="sunahama" status="sp. nov.">
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<emphasis box="[331,592,1944,1968]" italics="true" pageId="2" pageNumber="35">Pseudothinusa sunahama</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[601,683,1944,1968]" pageId="2" pageNumber="35">
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<taxonomicNameLabel box="[601,683,1944,1968]" pageId="2" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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: mouthparts. 4, labrum, dorsal view; 5, epipharynx, ventral view; 6, left mandible, ventral view; 7, left mandible, ventral view; 8, prementum; 9, right maxilla, ventral view; 10, mentum. Scale bars: 0.1 mm.
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</paragraph>
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</caption>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7645721" ID-Zenodo-Dep="7645721" httpUri="https://zenodo.org/record/7645721/files/figure.png" pageId="3" pageNumber="36" startId="3.[151,264,1943,1967]" targetBox="[178,1410,196,1917]" targetPageId="3">
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<paragraph blockId="3.[151,1436,1943,2040]" pageId="3" pageNumber="36">
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<emphasis bold="true" box="[151,335,1943,1967]" pageId="3" pageNumber="36">FIGURES 11–18</emphasis>
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<taxonomicName authority="Tasaku & Maruyama, 2023" authorityName="Tasaku & Maruyama" authorityYear="2023" box="[344,607,1944,1968]" class="Insecta" family="Staphylinidae" genus="Pseudothinusa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="36" phylum="Arthropoda" rank="species" species="sunahama" status="sp. nov.">
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<emphasis box="[344,607,1944,1968]" italics="true" pageId="3" pageNumber="36">Pseudothinusa sunahama</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[616,700,1944,1968]" pageId="3" pageNumber="36">
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<taxonomicNameLabel box="[616,700,1944,1968]" pageId="3" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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: abdominal terminalia. 11, median lobe of aedeagus, lateral view; 12, median lobe of aedeagus, parameral view; 13, paramere, lateral view; 14, male tergite VIII; 15, male sternite VIII; 16, tergite VIII; 17, female sternite VIII; 18, spermatheca. Scale bars: 0.1 mm.
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</paragraph>
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</caption>
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<paragraph blockId="4.[151,1437,151,1149]" pageId="4" pageNumber="37">
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<emphasis box="[199,260,151,177]" italics="true" pageId="4" pageNumber="37">Head</emphasis>
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. Head approximately circular. Eyes with microsetae between facets (Fig. 2). Cervical carina and infraorbital carina complete. Antennae 11-segmented, moniliform.
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</paragraph>
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<paragraph blockId="4.[151,1437,151,1149]" pageId="4" pageNumber="37">
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<emphasis box="[199,338,223,249]" italics="true" pageId="4" pageNumber="37">Mouthparts.</emphasis>
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Labrum (Fig. 4) strongly transverse (width/length ratio 1.8), with 8 pairs of macrosetae. Epipharynx (Fig. 5) with normally developed sensillum
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<emphasis box="[661,675,259,284]" italics="true" pageId="4" pageNumber="37">a</emphasis>
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. Mandibles (Figs. 6–7) with slightly sharp apex, asymmetric; left mandible (Fig. 6) generalised, without teeth; Right mandible (Fig. 7) with a medium tooth at middle of inner margin, weakly serrate between tooth and apex. Labium as in Fig. 8. Prementum (Fig. 8) with 2 real pores and 1 setal pore latero-mesally; lateral lobe of apodeme long. Ligula (Fig. 8) bilobed, with each lobe rounded and covered with minute pores apically; apical setae absent; lateral margin weakly sclerotized. Labial palpus (Fig. 8) with segment I long and twice as long as II; segment III as long as I. Maxilla as in Fig. 9. Cardo as in Fig. 9. Stipes (Fig. 9) small, triangular, with 3 setae around base. Palpifer (Fig. 9) triangular in lateral view, with one large seta near apex, two small setae at middle and apex. Lacinia (Fig. 9) generalized; inner margin with a row of 8 spines in apical 1/3, dense setae in basal 2/3 and two setae in basal 1/3. Galea (Fig. 9) slender, about 3 times as long as wide, curved at middle, densely pubscent apically. Maxillary pulpus as in Fig. 9; segment I short; segment II with about 10 setae; segment III largest, 1.5 times longer than 2nd, with about 30 setae; IV segment without setae. Mentum (Fig. 10) trapeziform; v-seta short; surface moderately covered with pseudopores except for lateral area.
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</paragraph>
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<paragraph blockId="4.[151,1437,151,1149]" pageId="4" pageNumber="37">
|
||
<emphasis box="[199,288,691,717]" italics="true" pageId="4" pageNumber="37">Thorax.</emphasis>
|
||
Pronotum (Fig. 1) slightly transverse, with thick setae toward lateral side. Prosternum transverse. Scutellum generalized, covered with setae. Hypomera visible laterally. Mesoventral process pointed at apex. Metaventral process, pointed at apex, shorter than mesoventral process. Mesocoxal cavity unmargined, separated by isthmus. Hind wing present.
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,151,1149]" pageId="4" pageNumber="37">
|
||
<emphasis box="[199,260,835,860]" italics="true" pageId="4" pageNumber="37">Legs.</emphasis>
|
||
Tarsal formula 4-5-5; front and middle tibiae with spines (Fig. 3). Empodium simple; empodial seta as long as long as claw.
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,151,1149]" box="[199,1365,907,933]" pageId="4" pageNumber="37">
|
||
<emphasis box="[199,313,907,933]" italics="true" pageId="4" pageNumber="37">Abdomen.</emphasis>
|
||
Abdomen (Fig. 1) parallel side; abdominal chaetotaxy 02
|
||
<typeStatus box="[965,1013,907,933]" pageId="4" pageNumber="37">type</typeStatus>
|
||
(sensu
|
||
<bibRefCitation author="Yosii, R. & Sawada, K." box="[1098,1353,907,933]" pageId="4" pageNumber="37" pagination="11 - 140" refId="ref4082" refString="Yosii, R. & Sawada, K. (1976) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae). II: Diagnostic characters of genera and subgenera with description of representative species. Contributions from the Biological Laboratory, Kyoto University, 25 (1), 11 - 140." type="journal article" year="1976">Yosii & Sawada, 1976</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,151,1149]" pageId="4" pageNumber="37">
|
||
<emphasis box="[199,438,943,969]" italics="true" pageId="4" pageNumber="37">Male characteristics.</emphasis>
|
||
Tergite VIII (Fig. 14) posterior margin irregularly dentate; sternite VIII (Fig. 15) simple. Aedeagus: median lobe (Figs. 11–12) entirely oval, apical process broad. Apical lobe of paramere (Fig. 13) with 4 setae; a-seta as long as b-seta; c- and d-setae very short.
|
||
</paragraph>
|
||
<paragraph blockId="4.[151,1437,151,1149]" pageId="4" pageNumber="37">
|
||
<emphasis box="[199,465,1051,1077]" italics="true" pageId="4" pageNumber="37">Female characteristics.</emphasis>
|
||
Tergite VIII (Fig. 16) and sternite VIII (Fig. 17) simple, posterior margin not dentate. Spermatheca (Fig. 18) with basal part of stem twice curved around base.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,1050,1122,1149]" pageId="4" pageNumber="37" type="distribution">
|
||
<paragraph blockId="4.[151,1437,151,1149]" box="[199,1050,1122,1149]" pageId="4" pageNumber="37">
|
||
<emphasis bold="true" box="[199,353,1122,1148]" pageId="4" pageNumber="37">Distribution.</emphasis>
|
||
<collectingCountry box="[359,427,1123,1149]" name="Japan" pageId="4" pageNumber="37">Japan</collectingCountry>
|
||
:
|
||
<collectingRegion box="[438,551,1123,1149]" country="Japan" name="Hokkaido" pageId="4" pageNumber="37">Hokkaidô</collectingRegion>
|
||
, Honshû, Shikoku, Kyûshû (Nokono-shima)
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |