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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.456.8465" ID-GBIF-Dataset="931d6fa7-7ffb-4039-b7ec-58c907d22f48" ID-PMC="PMC4329401" ID-Pensoft-Pub="1313-2970-456-85" ID-PubMed="25709523" ID-ZBK="71EC0955E0A648A888AE9C4F1E3BD53D" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-456-85" ModsDocOrigin="ZooKeys 456" ModsDocTitle="Description of six new species of Lycocerus Gorham (Coleoptera, Cantharidae), with taxonomic note and new distribution data of some other species" checkinTime="1451245090240" checkinUser="pensoft" docAuthor="Yang, Yuxia, Su, Junyan &amp; Yang, Xingke" docDate="2014" docId="09759F9F906E824472B90AD7C59DAEAB" docLanguage="en" docName="ZooKeys 456: 85-107" docOrigin="ZooKeys 456" docSource="http://dx.doi.org/10.3897/zookeys.456.8465" docTitle="Lycocerus quadrilineatus Y. Yang &amp; X. Yang, sp. n." docType="treatment" docUuid="EC8B55E5-A8DD-482B-8E79-BFE872697E21" docUuidSource="ZooBank" docVersion="4" lastPageNumber="97" masterDocId="7F5D2B6AFF9C1B1D961CC53389455A43" masterDocTitle="Description of six new species of Lycocerus Gorham (Coleoptera, Cantharidae), with taxonomic note and new distribution data of some other species" masterLastPageNumber="107" masterPageNumber="85" pageNumber="96" updateTime="1668159639272" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Description of six new species of Lycocerus Gorham (Coleoptera, Cantharidae), with taxonomic note and new distribution data of some other species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yang, Yuxia</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Su, Junyan</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yang, Xingke</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2014</mods:date>
<mods:detail type="volume">
<mods:number>456</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>85</mods:start>
<mods:end>107</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.456.8465</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.456.8465</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-456-85</mods:identifier>
<mods:identifier type="ZBK">71EC0955E0A648A888AE9C4F1E3BD53D</mods:identifier>
<mods:identifier type="ZooBank">71EC0955E0A648A888AE9C4F1E3BD53D</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152056683" LSID="urn:lsid:zoobank.org:act:EC8B55E5-A8DD-482B-8E79-BFE872697E21" httpUri="http://treatment.plazi.org/id/09759F9F906E824472B90AD7C59DAEAB" lastPageId="12" lastPageNumber="97" pageId="11" pageNumber="96">
<subSubSection pageId="11" pageNumber="96" type="multiple">
<paragraph pageId="11" pageNumber="96">
<pageBreakToken pageId="11" pageNumber="96" start="start">Taxon</pageBreakToken>
classification Animalia Coleoptera Cantharidae
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="96" type="nomenclature">
<paragraph pageId="11" pageNumber="96">
<taxonomicName LSID="http://zoobank.org/EC8B55E5-A8DD-482B-8E79-BFE872697E21" authority="Y. Yang &amp; X. Yang" class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lycocerus quadrilineatus" order="Coleoptera" pageId="11" pageNumber="96" phylum="Arthropoda" rank="species" species="quadrilineatus">Lycocerus quadrilineatus Y. Yang &amp; X. Yang</taxonomicName>
<taxonomicNameLabel pageId="11" pageNumber="96">sp. n.</taxonomicNameLabel>
Figs 6, 14-15, 31-33, 37, 41
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="97" type="type material">
<paragraph pageId="12" pageNumber="97">
<pageBreakToken pageId="12" pageNumber="97" start="start">Type</pageBreakToken>
material.
</paragraph>
<paragraph pageId="12" pageNumber="97">
Holotype ♂ (ZIN): &quot;N. Vietnam, Sa Pa env., 1200m, 20.V.1999, leg. Ozlov&quot;. Paratypes: 1♀ (ZIN): same data as holotype; 1♀ (ZIN):
<normalizedToken originalValue="“ВЬЕТНАМ">&quot;VETNAM</normalizedToken>
[Vietnam]
<normalizedToken originalValue="горы">gory</normalizedToken>
,
<normalizedToken originalValue="у">u</normalizedToken>
<normalizedToken originalValue="ША-ПА">SHA-PA</normalizedToken>
[Sa Pa], 1600-2000m, 5.6.1963
<normalizedToken originalValue="г">g</normalizedToken>
,
<normalizedToken originalValue="Каьакое">Kaakoe</normalizedToken>
[Kabakov]&quot;.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="97" type="distribution">
<paragraph pageId="12" pageNumber="97">Distribution.</paragraph>
<paragraph pageId="12" pageNumber="97">Vietnam (Sa Pa).</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="97" type="description">
<paragraph pageId="12" pageNumber="97">Description.</paragraph>
<paragraph pageId="12" pageNumber="97">Male (Fig. 6). Body black, mandibles dark brown, pronotum red, with a large dark brown marking, which almost extending to all margins of disc, elytra red, nearly black at the whole length of all elytral interstices.</paragraph>
<paragraph pageId="12" pageNumber="97">
Head evenly narrowed behind eyes, surface densely and finely punctate, semilustrous, covered with dense, fine, yellowish brown decumbent pubescence; eyes strongly protruding, head width across eyes distinctly wider than anterior margin of pronotum; terminal maxillary palpomere long-triangular, nearly truncate and sharp at apical one-third length of inner margin; antennae (Fig. 15) extending to apical one-fourth length of elytra, antennomeres II nearly as long as wide at apices,
<normalizedToken originalValue="IIIXI">III-XI</normalizedToken>
flattened,
<normalizedToken originalValue="IIIIX">III-IX</normalizedToken>
obliquely widened apically, nearly long-triangular, III about twice as long as wide at apices, IV slightly longer than III,
<normalizedToken originalValue="XXI">X-XI</normalizedToken>
nearly parallel-sided, XI slightly longer than X and pointed at apices,
<normalizedToken originalValue="IVXI">IV-XI</normalizedToken>
each with a round smooth impression at apical part of outer margin.
</paragraph>
<paragraph pageId="12" pageNumber="97">Pronotum slightly longer than wide, widest near posterior margin, anterior margin rounded, anterior angles rounded, lateral margins moderately diverging posteriorly, slightly sinuate at anterior portion, posterior angles rectangular, posterior margin slightly arcuate and narrowly bordered, disc moderately convex at posterolateral parts, surface punctate and pubescent like that of head, matt.</paragraph>
<paragraph pageId="12" pageNumber="97">Elytra about 4.3 times longer than pronotum, 3.5 times longer than humeral width, which about one-third wider than posterior margin of pronotum, lateral margins nearly parallel, elytral venations well-developed, moderately costate.</paragraph>
<paragraph pageId="12" pageNumber="97">All tarsal claws simple.</paragraph>
<paragraph pageId="12" pageNumber="97">Abdominal sternite IX long-triangular. Aedeagus (Figs 31-33): ventral process of each paramere normal and rounded at apex, even and nearly straight in lateral view; dorsal fig slightly shorter than ventral process, not narrowed apically, with inner angle rounded, outer angle obtuse-angled, inner margin nearly straight, apical margin rounded; laterophyse with apex pointed laterodorsally to outer angle of dorsal fig.</paragraph>
<paragraph pageId="12" pageNumber="97">
Female. Similar to the male, but eyes less protruding; antennae (Fig. 14) shorter, extending to basal one-third length of elytra, antennomeres III about 1.1 times longer than wide at apices,
<normalizedToken originalValue="IVXI">IV-XI</normalizedToken>
without impressions; pronotum nearly as long as wide, disc slightly convex on posterolateral parts; elytra with lateral margins slightly diverging posteriorly; pro- and meso-outer and inner tarsal claws each with a basal projection; abdominal sternite VIII (Fig. 37) very slightly emarginated in middle and largely emarginated on both sides of posterior margin, the portion between middle and lateral emarginations indistinctly angled at apex; internal reproductive organ of genitalia see Fig. 41.
</paragraph>
<paragraph pageId="12" pageNumber="97">Body length (both sexes): 8.5-10.0 mm; width: 2.0-2.2 mm.</paragraph>
<caption pageId="12" pageNumber="97">
<paragraph pageId="12" pageNumber="97">
Figures 34-38. Abdominal sternite VIII of female, ventral view: 34
<taxonomicName class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lycocerus hickeri" order="Coleoptera" pageId="12" pageNumber="97" phylum="Arthropoda" rank="species" species="hickeri">Lycocerus hickeri</taxonomicName>
(Pic, 1934) 35
<taxonomicName class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lycocerus sichuanus" order="Coleoptera" pageId="12" pageNumber="97" phylum="Arthropoda" rank="species" species="sichuanus">Lycocerus sichuanus</taxonomicName>
sp. n. 36
<taxonomicName class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lycocerus hubeiensis" order="Coleoptera" pageId="12" pageNumber="97" phylum="Arthropoda" rank="species" species="hubeiensis">Lycocerus hubeiensis</taxonomicName>
sp. n. 37
<taxonomicName class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lycocerus quadrilineatus" order="Coleoptera" pageId="12" pageNumber="97" phylum="Arthropoda" rank="species" species="quadrilineatus">Lycocerus quadrilineatus</taxonomicName>
sp. n. 38
<taxonomicName class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lycocerus obscurus" order="Coleoptera" pageId="12" pageNumber="97" phylum="Arthropoda" rank="species" species="obscurus">Lycocerus obscurus</taxonomicName>
Pic, 1916. Scale bars: 1.0 mm.
</paragraph>
</caption>
<caption pageId="12" pageNumber="97">
<paragraph pageId="12" pageNumber="97">
Figures 39-41. Female genitalia, lateral view: 39.
<taxonomicName class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lycocerus sichuanus" order="Coleoptera" pageId="12" pageNumber="97" phylum="Arthropoda" rank="species" species="sichuanus">Lycocerus sichuanus</taxonomicName>
sp. n.; 40
<taxonomicName class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lycocerus hubeiensis" order="Coleoptera" pageId="12" pageNumber="97" phylum="Arthropoda" rank="species" species="hubeiensis">Lycocerus hubeiensis</taxonomicName>
sp. n. 41
<taxonomicName class="Insecta" family="Cantharidae" genus="Lycocerus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lycocerus quadrilineatus" order="Coleoptera" pageId="12" pageNumber="97" phylum="Arthropoda" rank="species" species="quadrilineatus">Lycocerus quadrilineatus</taxonomicName>
sp. n. Scale bars: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="12" pageNumber="97" type="etymology">
<paragraph pageId="12" pageNumber="97">Etymology.</paragraph>
<paragraph pageId="12" pageNumber="97">The specific name is derived from Latin quadrus (four) and linea (stripe), referring to its elytra darkened at all of the four elytral interstices.</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="97" type="diagnosis">
<paragraph pageId="12" pageNumber="97">Diagnosis.</paragraph>
<paragraph pageId="12" pageNumber="97">This species can be easily distinguished from other species by the elytra darkened at all elytral interstices, elytral venation well-developed; all claws simple in male, pro- and meso-outer and inner tarsal claws each with a basal projection in female; aedeagus: dorsal fig of each paramere not narrowed apically.</paragraph>
</subSubSection>
</treatment>
</document>