476 lines
56 KiB
XML
476 lines
56 KiB
XML
<document id="B1B959DC6F95B69F31D4300839279A2C" ID-DOI="10.11646/zootaxa.3637.2.7" ID-GBIF-Dataset="594e95c0-4ac3-450b-815c-c6683b7b9e01" ID-ISSN="1175-5326" ID-Zenodo-Dep="223729" ID-ZooBank="844B7B6C-9468-4A1D-9A53-32C646880252" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460591024090" checkinUser="plazi" docAuthor="Greenfield, David W. & Suzuki, Toshiyuki" docDate="2013" docId="03C987C906229F5EADC9A72CFD3AFE6D" docLanguage="en" docName="zt03637p175.pdf" docOrigin="Zootaxa 3637 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eviota nigramembrana Greenfield & Suzuki, 2013, n. sp." docType="treatment" docVersion="7" lastPageNumber="174" masterDocId="FFF0FFB106229F5BAD5EA06EFF9FFFEA" masterDocTitle="Eviota nigramembrana, a new dwarfgoby from the Western Pacific (Teleostei: Gobiidae)" masterLastPageNumber="175" masterPageNumber="169" pageNumber="169" updateTime="1698325953227" updateUser="plazi">
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<mods:mods id="23C63C61A4FE7311DA783A0BAAB96F85" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="45286808E7D76C16EB5F8D293CC3D020">
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<mods:title id="226D73BF998DB4AC23C0CFC203640341">Eviota nigramembrana, a new dwarfgoby from the Western Pacific (Teleostei: Gobiidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="6DF0A519622D9DF4D96FC4E0670657AC" type="personal">
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<mods:role id="7F1BF82F13EF4550C79FD15DC9F4DCD3">
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<mods:roleTerm id="17AACA38A8CA2FD7C41C7A5ED087FB5A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="934BFD2A46F80EC1AB0D86A7DD67E0C6">Greenfield, David W.</mods:namePart>
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</mods:name>
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<mods:name id="972223052C94DABB166368663AC1A857" type="personal">
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<mods:role id="B02ABF847DB78E2F237DD979F85FA940">
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<mods:roleTerm id="906CF75733CEE8F59EB6204BBE0458D2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C47C142FD7F8D20369D647D5B08DBD4F">Suzuki, Toshiyuki</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="CBCF1D3745B71E09D7A2B1610F988471">text</mods:typeOfResource>
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<mods:titleInfo id="567E4C26F81B8B6EE5886202D510E7F5">
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<mods:title id="B7D9CB9AA228690AF21EFCA812FE5024">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="8C300AAB83D6532DB66CE30DB9D5B4FE">
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<mods:date id="F5B35E31A059DEAEAC1A02F7BE71721F">2013</mods:date>
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<mods:detail id="E67B4F17C54F996CF0D8A6E73BE87AF1" type="volume">
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<mods:number id="8EA3BAA258C760E93D1D0CCC7037DC28">3637</mods:number>
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</mods:detail>
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<mods:detail id="AF152C9D8DE625D49ECF450D64CF5A06" type="issue">
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<mods:number id="1ED2755F727F0FAE095766E03206EE98">2</mods:number>
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</mods:detail>
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<mods:extent id="F1DC25A06B5475E6C1D7131AE3C05178" unit="page">
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<mods:start id="BA180A81FFE6AE120B7872F3C4D4A0DB">169</mods:start>
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<mods:end id="4BBBFC524A0141293D185C5E3DF7B9D8">175</mods:end>
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<mods:classification id="482A52E5F866B3D296BB0A048435CC40">journal article</mods:classification>
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<mods:identifier id="735A1A8C974CB043C291E0F00A4A3C83" type="DOI">10.11646/zootaxa.3637.2.7</mods:identifier>
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<mods:identifier id="8112B505BC910A39EA7C1285075CF45B" type="GBIF-Dataset">594e95c0-4ac3-450b-815c-c6683b7b9e01</mods:identifier>
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<mods:identifier id="98A9E9F0C8CCC7F7B05685496CE5D945" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="470B8AA029B71571246D539954D73E16" type="Zenodo-Dep">223729</mods:identifier>
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<mods:identifier id="F83EB54F6D7DC8FD0AF25B614843CB12" type="ZooBank">844B7B6C-9468-4A1D-9A53-32C646880252</mods:identifier>
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</mods:mods>
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<treatment id="03C987C906229F5EADC9A72CFD3AFE6D" ID-DOI="http://doi.org/10.5281/zenodo.6152136" ID-GBIF-Taxon="119565672" ID-Zenodo-Dep="6152136" LSID="urn:lsid:plazi:treatment:03C987C906229F5EADC9A72CFD3AFE6D" httpUri="http://treatment.plazi.org/id/03C987C906229F5EADC9A72CFD3AFE6D" lastPageId="5" lastPageNumber="174" pageId="0" pageNumber="169">
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<subSubSection id="C37A655406229F5AADC9A72CFDBEFF26" lastPageId="1" lastPageNumber="170" pageId="0" pageNumber="169" type="nomenclature">
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<paragraph id="8BDF36DF06229F5BADC9A72CFE66F8B6" blockId="0.[151,557,1858,1989]" box="[151,505,1858,1884]" pageId="0" pageNumber="169">
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<heading id="D09781B306229F5BADC9A72CFE66F8B6" bold="true" box="[151,505,1858,1884]" fontSize="11" level="1" pageId="0" pageNumber="169" reason="1">
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<emphasis id="B914EACD06229F5BADC9A72CFE66F8B6" bold="true" box="[151,505,1858,1884]" pageId="0" pageNumber="169">
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||
<taxonomicName id="4C604D5C06229F5BADC9A72CFE30F8B6" ID-CoL="3DHMD" box="[151,431,1858,1884]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="0" pageNumber="169" phylum="Chordata" rank="species" species="nigramembrana" status="sp. nov.">
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<emphasis id="B914EACD06229F5BADC9A72CFE30F8B6" bold="true" box="[151,431,1858,1884]" italics="true" pageId="0" pageNumber="169">Eviota nigramembrana</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22757B606229F5BACE8A72CFE66F8B6" box="[438,505,1858,1884]" pageId="0" pageNumber="169" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BDF36DF06229F5BADC9A70BFEE3F897" blockId="0.[151,557,1858,1989]" box="[151,380,1893,1917]" pageId="0" pageNumber="169">Blackbar dwarfgoby</paragraph>
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<paragraph id="8BDF36DF06229F5BADC9A7E4FE99F82F" blockId="0.[151,557,1858,1989]" pageId="0" pageNumber="169">
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Japanese name—Kometubu-isohaze
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<figureCitation id="135B2A5A06229F5BADC9A7C3FE9DF82F" box="[151,258,1965,1989]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStartId-0="2.[151,249,977,1000]" captionStartId-1="2.[151,249,1930,1953]" captionStartId-2="3.[151,249,1537,1560]" captionStartId-3="4.[151,249,1061,1084]" captionStartId-4="4.[151,249,1957,1980]" captionStartId-5="5.[151,250,1045,1068]" captionTargetBox-0="[156,1436,517,945]" captionTargetBox-1="[283,1303,1049,1909]" captionTargetBox-2="[221,1407,841,1503]" captionTargetBox-3="[155,1436,441,1032]" captionTargetBox-4="[158,1418,1266,1933]" captionTargetBox-5="[404,1183,445,1023]" captionTargetId-0="figure@2.[156,1441,517,947]" captionTargetId-1="figure@2.[283,1304,1049,1909]" captionTargetId-2="figure@3.[154,1439,841,1510]" captionTargetId-3="figure@4.[155,1440,441,1032]" captionTargetId-4="figure@4.[151,1436,1215,1936]" captionTargetId-5="figure@5.[404,1184,445,1024]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionTargetPageId-4="4" captionTargetPageId-5="5" captionText-0="FIGURE 1. Color in preservative of holotype of Eviota nigramembrana, OMNH-P. 35642. Photograph by D. W. Greenfield." captionText-1="FIGURE 2. Dark pigment on opercular membrane of Eviota nigramembrana, OMNH-P. 35644. Photograph by D. W. Greenfield." captionText-2="FIGURE 3. Color of fresh holotype of Eviota nigramembrana, OMNH-P. 35642. Photograph by T. Suzuki." captionText-3="FIGURE 4. Underwater photograph of Eviota nigramembrana, Kasari-wan Inlet, Amami-oshima Island, the Ryukyu Islands, Japan, 6 m depth, 29 Aug. 2005, Photograph by Hiroyuki Kanehara." captionText-4="FIGURE 5. Color of fresh paratype of Eviota nigramembrana, OMNH-P. 34238. Photograph by T. Suzuki." captionText-5="FIGURE 6. Pigmented opercular membrane of Philippine Island specimen of Eviota nigramembrana, USNM 220062, 17.2 mm male. Photograph by D. W. Greenfield." httpUri-0="https://zenodo.org/record/223730/files/figure.png" httpUri-1="https://zenodo.org/record/223731/files/figure.png" httpUri-2="https://zenodo.org/record/223732/files/figure.png" httpUri-3="https://zenodo.org/record/223733/files/figure.png" httpUri-4="https://zenodo.org/record/223734/files/figure.png" httpUri-5="https://zenodo.org/record/223735/files/figure.png" pageId="0" pageNumber="169">Figs. 1–6</figureCitation>
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.
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</paragraph>
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<paragraph id="8BDF36DF06239F5AADC9A0F7FDBEFF26" blockId="1.[151,791,152,205]" pageId="1" pageNumber="170">
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<taxonomicName id="4C604D5C06239F5AADC9A0F7FF46FF44" box="[151,217,153,174]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="170" phylum="Chordata" rank="genus">
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<emphasis id="B914EACD06239F5AADC9A0F7FF46FF44" box="[151,217,153,174]" italics="true" pageId="1" pageNumber="170">Eviota</emphasis>
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</taxonomicName>
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sp. C. Hayashi
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<emphasis id="B914EACD06239F5AAC27A0F7FE33FF44" box="[377,428,152,174]" italics="true" pageId="1" pageNumber="170">et al.</emphasis>
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1990: 131; Senou
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<emphasis id="B914EACD06239F5AAF35A0F7FD01FF44" box="[619,670,152,174]" italics="true" pageId="1" pageNumber="170">et al.</emphasis>
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, 2004: 122.
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<taxonomicName id="4C604D5C06239F5AADC9A0D6FF46FF27" box="[151,217,184,205]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="170" phylum="Chordata" rank="genus">
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<emphasis id="B914EACD06239F5AADC9A0D6FF46FF27" box="[151,217,184,205]" italics="true" pageId="1" pageNumber="170">Eviota</emphasis>
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</taxonomicName>
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sp. 3. Akihito
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<emphasis id="B914EACD06239F5AAC31A0D6FE3CFF27" box="[367,419,183,205]" italics="true" pageId="1" pageNumber="170">et al.</emphasis>
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2002: 1178.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37A655406239F5AADC9A097FD81FCE3" pageId="1" pageNumber="170" type="materials_examined">
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||
<paragraph id="8BDF36DF06239F5AADC9A097FF49FEB3" blockId="1.[151,1437,249,2007]" pageId="1" pageNumber="170">
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||
<emphasis id="B914EACD06239F5AADC9A097FE94FEFB" bold="true" box="[151,267,249,273]" pageId="1" pageNumber="170">
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<typeStatus id="54DB887D06239F5AADC9A097FE99FEFB" box="[151,262,249,273]" pageId="1" pageNumber="170" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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OMNH-P 35642,
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<quantity id="4C989B3A06239F5AACB9A097FDD0FEFB" box="[487,591,249,273]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" pageId="1" pageNumber="170" unit="mm" value="13.9">13.9 mm</quantity>
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female, Okinawa Prefecture, the Ryukyu Islands, Uehara, Iriomote-jima
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<collectingCountry id="F377764F06239F5AADC9A173FF41FEDF" box="[151,222,285,309]" name="Iceland" pageId="1" pageNumber="170">Island</collectingCountry>
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||
,
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<geoCoordinate id="EE54501806239F5AADB1A173FEFEFEDF" box="[239,353,285,309]" direction="north" orientation="latitude" pageId="1" pageNumber="170" precision="5" value="24.4325">24.4325N</geoCoordinate>
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,
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<geoCoordinate id="EE54501806239F5AAC32A173FE77FEDF" box="[364,488,285,309]" direction="east" orientation="longitude" pageId="1" pageNumber="170" precision="5" value="123.8148">123.8148E</geoCoordinate>
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||
,
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||
<quantity id="4C989B3A06239F5AACADA173FDA6FEDF" box="[499,569,285,309]" metricMagnitude="0" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="1" pageNumber="170" unit="m" value="4.0" valueMax="5.0" valueMin="3.0">3–5 m</quantity>
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depth, field number S-5671, Toshiyuki Suzuki & Masatomi Suzuki,
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<date id="FFDE101F06239F5AA869A173FF4CFEB3" pageId="1" pageNumber="170" value="1996-08-08">8 August 1996</date>
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||
.
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</paragraph>
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||
<paragraph id="8BDF36DF06239F5AAD97A10BFD81FCE3" blockId="1.[151,1437,249,2007]" pageId="1" pageNumber="170">
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<emphasis id="B914EACD06239F5AAD97A10BFED5FE97" bold="true" box="[201,330,357,381]" pageId="1" pageNumber="170">
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<typeStatus id="54DB887D06239F5AAD97A10BFEDAFE97" box="[201,325,357,381]" pageId="1" pageNumber="170" type="paratype">Paratypes</typeStatus>
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:
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</emphasis>
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All from the Okinawa Prefecture, the Ryukyu Islands—USNM 406763 (ex OMNH-P 4285),
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<quantity id="4C989B3A06239F5AA835A10BFF5CFE4B" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="1" pageNumber="170" unit="mm" value="14.8">14.8 mm</quantity>
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female, Aragusuku, Gusukube-cho, Miyako-gun, Miyako-jima
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<collectingCountry id="F377764F06239F5AAE86A1E7FB80FE4B" box="[984,1055,393,417]" name="Iceland" pageId="1" pageNumber="170">Island</collectingCountry>
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, Aragusuku Beach,
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<geoCoordinate id="EE54501806239F5AA878A1E7FA07FE4B" box="[1318,1432,393,417]" direction="north" orientation="latitude" pageId="1" pageNumber="170" precision="5" value="24.7572">24.7572N</geoCoordinate>
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,
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<geoCoordinate id="EE54501806239F5AADC9A1C3FE8BFE2F" box="[151,276,429,453]" direction="east" orientation="longitude" pageId="1" pageNumber="170" precision="5" value="125.4263">125.4263E</geoCoordinate>
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,
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<quantity id="4C989B3A06239F5AAC7BA1C3FEC9FE2F" box="[293,342,429,453]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="170" unit="m" value="3.0">3 m</quantity>
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depth, field number S-3718, Toshiyuki Suzuki & Masatomi Hosokawa,
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<date id="FFDE101F06239F5AA99FA1C3FA08FE2F" box="[1217,1431,429,453]" pageId="1" pageNumber="170" value="1994-02-13">13 February 1994</date>
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||
; OMNH-P 34238,
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||
<quantity id="4C989B3A06239F5AAC3FA1BFFE5AFE03" box="[353,453,465,489]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="1" pageNumber="170" unit="mm" value="11.4">11.4 mm</quantity>
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male, Uehara, Iriomote-jima
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||
<collectingCountry id="F377764F06239F5AAE4DA1BFFCC5FE03" box="[787,858,465,489]" name="Iceland" pageId="1" pageNumber="170">Island</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE54501806239F5AAE38A1BFFC46FE03" box="[870,985,465,489]" direction="north" orientation="latitude" pageId="1" pageNumber="170" precision="5" value="24.4298">24.4298N</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE54501806239F5AAEBAA1BFFBFFFE03" box="[996,1120,465,489]" direction="east" orientation="longitude" pageId="1" pageNumber="170" precision="5" value="123.8262">123.8262E</geoCoordinate>
|
||
,
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||
<quantity id="4C989B3A06239F5AA932A1BCFB08FE03" box="[1132,1175,466,489]" metricMagnitude="0" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="170" unit="m" value="7.0">7 m</quantity>
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||
depth, field number S- 17076, Toshiyuki Suzuki, Masatomi Suzuki & Akira Kawai,
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||
<date id="FFDE101F06239F5AAE3BA19BFC6BFDE7" box="[869,1012,501,525]" pageId="1" pageNumber="170" value="2008-07-08">8 July 2008</date>
|
||
; OMNH-P 35639,
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||
<quantity id="4C989B3A06239F5AA988A19BFAA1FDE7" box="[1238,1342,501,525]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="1" pageNumber="170" unit="mm" value="13.8">13.8 mm</quantity>
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||
female, Uehara, Iriomote-jima
|
||
<collectingCountry id="F377764F06239F5AACFFA277FE77FDDB" box="[417,488,537,561]" name="Iceland" pageId="1" pageNumber="170">Island</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE54501806239F5AACA7A277FDF4FDDB" box="[505,619,537,561]" direction="north" orientation="latitude" pageId="1" pageNumber="170" precision="5" value="24.4325">24.4325N</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE54501806239F5AAF24A277FD67FDDB" box="[634,760,537,561]" direction="east" orientation="longitude" pageId="1" pageNumber="170" precision="5" value="123.8148">123.8148E</geoCoordinate>
|
||
,
|
||
<quantity id="4C989B3A06239F5AAE59A277FCCDFDDB" box="[775,850,537,561]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="1" pageNumber="170" unit="m" value="5.5" valueMax="6.0" valueMin="5.0">5–6 m</quantity>
|
||
depth, field number S-3906, Toshiyuki Suzuki & Masatomi Suzuki,
|
||
<date id="FFDE101F06239F5AAC2FA253FDB1FDBF" box="[369,558,573,598]" pageId="1" pageNumber="170" value="1994-08-22">22 August 1994</date>
|
||
; OMNH-P 35641,
|
||
<quantity id="4C989B3A06239F5AAE53A253FCEAFDBF" box="[781,885,573,597]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" pageId="1" pageNumber="170" unit="mm" value="14.1">14.1 mm</quantity>
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||
male, Uehara, Iriomote-jima
|
||
<collectingCountry id="F377764F06239F5AA98EA253FA88FDBF" box="[1232,1303,573,597]" name="Iceland" pageId="1" pageNumber="170">Island</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE54501806239F5AA878A253FA07FDBF" box="[1318,1432,573,597]" direction="north" orientation="latitude" pageId="1" pageNumber="170" precision="5" value="24.4325">24.4325N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE54501806239F5AADC9A20FFE8BFD93" box="[151,276,609,633]" direction="east" orientation="longitude" pageId="1" pageNumber="170" precision="5" value="123.8148">123.8148E</geoCoordinate>
|
||
, field number S-3987, 7–
|
||
<quantity id="4C989B3A06239F5AAF1CA20FFDF0FD93" box="[578,623,609,633]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="170" unit="m" value="8.0">8 m</quantity>
|
||
depth, Toshiyuki Suzuki & Masatomi Suzuki,
|
||
<date id="FFDE101F06239F5AA9C6A20FFAC9FD93" box="[1176,1366,609,634]" pageId="1" pageNumber="170" value="1994-08-27">27 August 1994</date>
|
||
; CAS 234537-(ex OMNH-P 35643), 14.0 mm male, Aakabanare, Iriomote-jima
|
||
<collectingCountry id="F377764F06239F5AA95DA2EBFBD5FD77" box="[1027,1098,645,669]" name="Iceland" pageId="1" pageNumber="170">Island</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE54501806239F5AA905A2EBFB52FD77" box="[1115,1229,645,669]" direction="north" orientation="latitude" pageId="1" pageNumber="170" precision="5" value="24.4004">24.4004N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE54501806239F5AA980A2EBFAC4FD77" box="[1246,1371,645,669]" direction="east" orientation="longitude" pageId="1" pageNumber="170" precision="5" value="123.8715">123.8715E</geoCoordinate>
|
||
,
|
||
<quantity id="4C989B3A06239F5AA832A2EBFA03FD77" box="[1388,1436,645,669]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="170" unit="m" value="3.0">3 m</quantity>
|
||
depth, field number S-5983, Toshiyuki Suzuki & Masatomi Suzuki,
|
||
<date id="FFDE101F06239F5AAEE9A2C7FBCCFD2B" box="[951,1107,681,705]" pageId="1" pageNumber="170" value="1997-07-24">24 July 1997</date>
|
||
; OMNH-P 35644, 14.0 mm female, Uehara, Iriomote-jima
|
||
<collectingCountry id="F377764F06239F5AACACA2A3FDA6FD0F" box="[498,569,717,741]" name="Iceland" pageId="1" pageNumber="170">Island</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE54501806239F5AAF1AA2A3FD28FD0F" box="[580,695,717,741]" direction="north" orientation="latitude" pageId="1" pageNumber="170" precision="5" value="24.4325">24.4325N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE54501806239F5AAF9FA2A3FCA0FD0F" box="[705,831,717,741]" direction="east" orientation="longitude" pageId="1" pageNumber="170" precision="5" value="123.8148">123.8148E</geoCoordinate>
|
||
,
|
||
<quantity id="4C989B3A06239F5AAE17A2A3FC0FFD0F" box="[841,912,717,741]" metricMagnitude="0" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="1" pageNumber="170" unit="m" value="5.5" valueMax="6.0" valueMin="5.0">5–6 m</quantity>
|
||
depth, field number S-6178, Toshiyuki Suzuki & Masatomi Suzuki,
|
||
<date id="FFDE101F06239F5AACD8A29FFD85FCE3" box="[390,538,753,777]" pageId="1" pageNumber="170" value="1997-07-29">29 July 1997</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A655406239F5AAD97A37BFDB1FBEF" pageId="1" pageNumber="170" type="description">
|
||
<paragraph id="8BDF36DF06239F5AAD97A37BFDB1FBEF" blockId="1.[151,1437,249,2007]" pageId="1" pageNumber="170">
|
||
<emphasis id="B914EACD06239F5AAD97A37BFDB1FCC7" bold="true" box="[201,558,789,814]" pageId="1" pageNumber="170">
|
||
Non-types:
|
||
<collectingCountry id="F377764F06239F5AAC0BA37BFDB1FCC7" box="[341,558,789,814]" name="Philippines" pageId="1" pageNumber="170">Philippine Islands</collectingCountry>
|
||
</emphasis>
|
||
-Negros Oriental, Siquijor Id.- USNM 223062 (3)
|
||
<quantity id="4C989B3A06239F5AA92AA37BFA84FCC7" box="[1140,1307,789,813]" metricMagnitude="-2" metricUnit="m" metricValue="1.4849999999999999" metricValueMax="1.72" metricValueMin="1.25" pageId="1" pageNumber="170" unit="mm" value="14.85" valueMax="17.2" valueMin="12.5">12.5–17.2 mm</quantity>
|
||
,
|
||
<quantity id="4C989B3A06239F5AA877A37BFA08FCC7" box="[1321,1431,789,813]" metricMagnitude="0" metricUnit="m" metricValue="5.35" metricValueMax="10.7" metricValueMin="0.0" pageId="1" pageNumber="170" unit="m" value="5.35" valueMax="10.7" valueMin="0.0">0–10.7 m</quantity>
|
||
, SP78-7; USNM 223063 (19) 13.0–
|
||
<quantity id="4C989B3A06239F5AAF76A357FDE6FCBB" box="[552,633,825,849]" metricMagnitude="1" metricUnit="m" metricValue="1.8199999999999998" pageId="1" pageNumber="170" unit="m" value="18.2">18.2 m</quantity>
|
||
,
|
||
<quantity id="4C989B3A06239F5AAFD9A357FD78FCBB" box="[647,743,825,849]" metricMagnitude="-1" metricUnit="m" metricValue="6.0" metricValueMax="12.0" metricValueMin="0.0" pageId="1" pageNumber="170" unit="m" value="0.6" valueMax="1.2" valueMin="0.0">0–1.2 m</quantity>
|
||
, LK 79-15; USNM 223067 (1)
|
||
<quantity id="4C989B3A06239F5AA908A357FB20FCBB" box="[1110,1215,825,849]" metricMagnitude="-2" metricUnit="m" metricValue="1.33" pageId="1" pageNumber="170" unit="mm" value="13.3">13.3 mm</quantity>
|
||
,
|
||
<quantity id="4C989B3A06239F5AA995A357FAB4FCBB" box="[1227,1323,825,849]" metricMagnitude="0" metricUnit="m" metricValue="3.05" metricValueMax="6.1" metricValueMin="0.0" pageId="1" pageNumber="170" unit="m" value="3.05" valueMax="6.1" valueMin="0.0">0–6.1 m</quantity>
|
||
, SP 78-4; USNM 223068 (1)
|
||
<quantity id="4C989B3A06239F5AAC2DA333FE45FC9F" box="[371,474,861,885]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="1" pageNumber="170" unit="mm" value="16.3">16.3 mm</quantity>
|
||
,
|
||
<quantity id="4C989B3A06239F5AACB8A333FDDBFC9F" box="[486,580,861,885]" metricMagnitude="-1" metricUnit="m" metricValue="4.5" metricValueMax="9.0" metricValueMin="0.0" pageId="1" pageNumber="170" unit="m" value="0.45" valueMax="0.9" valueMin="0.0">0–0.9 m</quantity>
|
||
, LK 79-14. Apo Id.—USNM 223071 (2) 12.7 &
|
||
<quantity id="4C989B3A06239F5AA92CA333FB46FC9F" box="[1138,1241,861,885]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="1" pageNumber="170" unit="mm" value="16.8">16.8 mm</quantity>
|
||
,
|
||
<quantity id="4C989B3A06239F5AA9BBA333FADBFC9F" box="[1253,1348,861,885]" metricMagnitude="0" metricUnit="m" metricValue="3.05" metricValueMax="6.1" metricValueMin="0.0" pageId="1" pageNumber="170" unit="m" value="3.05" valueMax="6.1" valueMin="0.0">0–6.1 m</quantity>
|
||
, SP 78- 35; CAS 47914 (2) 16.6 &
|
||
<quantity id="4C989B3A06239F5AAC8BA3EFFDB7FC73" box="[469,552,897,921]" metricMagnitude="1" metricUnit="m" metricValue="1.68" pageId="1" pageNumber="170" unit="m" value="16.8">16.8 m</quantity>
|
||
,
|
||
<quantity id="4C989B3A06239F5AAF68A3EFFD09FC73" box="[566,662,897,921]" metricMagnitude="0" metricUnit="m" metricValue="1.2" metricValueMax="2.4" metricValueMin="0.0" pageId="1" pageNumber="170" unit="m" value="1.2" valueMax="2.4" valueMin="0.0">0–2.4 m</quantity>
|
||
, LK 79-20. Zamboanga del Norte, Allibay Id.—USNM 223066 (1)
|
||
<quantity id="4C989B3A06239F5AADC9A3CBFF60FC57" box="[151,255,933,957]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="1" pageNumber="170" unit="mm" value="13.6">13.6 mm</quantity>
|
||
,
|
||
<quantity id="4C989B3A06239F5AAC55A3CBFEF5FC57" box="[267,362,933,957]" metricMagnitude="0" metricUnit="m" metricValue="1.55" metricValueMax="3.1" metricValueMin="0.0" pageId="1" pageNumber="170" unit="m" value="1.55" valueMax="3.1" valueMin="0.0">0–3.1 m</quantity>
|
||
, LK 79-5. Solino (Selinog) Id.—AMS I.22206-001 (3)
|
||
<quantity id="4C989B3A06239F5AAEBDA3CBFB15FC57" box="[995,1162,933,957]" metricMagnitude="-2" metricUnit="m" metricValue="1.4449999999999998" metricValueMax="1.5699999999999998" metricValueMin="1.3199999999999998" pageId="1" pageNumber="170" unit="mm" value="14.45" valueMax="15.7" valueMin="13.2">13.2–15.7 mm</quantity>
|
||
,
|
||
<quantity id="4C989B3A06239F5AA9C8A3CBFAB9FC57" box="[1174,1318,933,957]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="2.13" metricValueMin="1.3699999999999999" pageId="1" pageNumber="170" unit="m" value="17.5" valueMax="21.3" valueMin="13.7">13.7–21.3 m</quantity>
|
||
, LK 79-6; USNM 223065 (1)
|
||
<quantity id="4C989B3A06239F5AACDCA3A7FE71FC0B" box="[386,494,969,993]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="1" pageNumber="170" unit="mm" value="12.6">12.6 mm</quantity>
|
||
,
|
||
<quantity id="4C989B3A06239F5AACA0A3A7FD0DFC0B" box="[510,658,969,993]" metricMagnitude="1" metricUnit="m" metricValue="1.525" metricValueMax="1.83" metricValueMin="1.22" pageId="1" pageNumber="170" unit="m" value="15.25" valueMax="18.3" valueMin="12.2">12.2–18.3 m</quantity>
|
||
, LK 79-9. Palawan Province, Tagauayan Id.—USNM 223064 (4)
|
||
<quantity id="4C989B3A06239F5AADC9A383FEA3FBEF" box="[151,316,1005,1029]" metricMagnitude="-2" metricUnit="m" metricValue="1.395" metricValueMax="1.42" metricValueMin="1.3699999999999999" pageId="1" pageNumber="170" unit="mm" value="13.95" valueMax="14.2" valueMin="13.7">13.7–14.2 mm</quantity>
|
||
,
|
||
<quantity id="4C989B3A06239F5AAC19A383FE2CFBEF" box="[327,435,1005,1029]" metricMagnitude="0" metricUnit="m" metricValue="6.85" metricValueMax="13.7" metricValueMin="0.0" pageId="1" pageNumber="170" unit="m" value="6.85" valueMax="13.7" valueMin="0.0">0–13.7 m</quantity>
|
||
, SP 78-24.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A655406239F5AAD99A47FFD18FB0A" pageId="1" pageNumber="170" type="diagnosis">
|
||
<paragraph id="8BDF36DF06239F5AAD99A47FFD18FB0A" blockId="1.[151,1437,249,2007]" pageId="1" pageNumber="170">
|
||
<emphasis id="B914EACD06239F5AAD99A47FFEA5FBC0" bold="true" box="[199,314,1041,1066]" pageId="1" pageNumber="170">Diagnosis</emphasis>
|
||
. The following combination of characters distinguishes
|
||
<taxonomicName id="4C604D5C06239F5AAEAEA47CFB53FBC3" box="[1008,1228,1041,1065]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="170" phylum="Chordata" rank="species" species="nigramembrana">
|
||
<emphasis id="B914EACD06239F5AAEAEA47CFB53FBC3" box="[1008,1228,1041,1065]" italics="true" pageId="1" pageNumber="170">E. nigramembrana</emphasis>
|
||
</taxonomicName>
|
||
from congeners: cephalic sensory-pore system pattern group I (complete); dorsal/anal fin-ray formula 8/8; 5th pelvic-fin ray absent; five internal bands between anal-fin origin and caudal-fin base; base of pectoral fin pale; a dark rectangular caudalpeduncle spot above midline; spinous dorsal fin pale, without broad dark and light bands; posterior edge of opercular membrane with a concentration of dark chromatophores; in life red-orange spots along bases of both dorsal fins, and belly with distinct red spots.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A655406239F58AD99A485FD3BFCFE" lastPageId="3" lastPageNumber="172" pageId="1" pageNumber="170" type="description">
|
||
<paragraph id="8BDF36DF06239F5AAD99A485FF67F95D" blockId="1.[151,1437,249,2007]" pageId="1" pageNumber="170">
|
||
<emphasis id="B914EACD06239F5AAD99A485FEC6FAEE" bold="true" box="[199,345,1259,1284]" pageId="1" pageNumber="170">Description.</emphasis>
|
||
Dorsal-fin rays VI-I,8 [I,8 (6), I,9 (1-OMNH P. 34238)]; anal-fin rays I,8; dorsal and anal soft rays branched except first, the last ray branched to base; pectoral-fin rays 16 (16–17, usually 16), lower rays
|
||
<date id="FFDE101F06239F5AA808A57EFF58FAA7" pageId="1" pageNumber="170" value="2010-05-16">10–16 may</date>
|
||
be branched (usually 10–14); pelvic fins joined by membrane only at extreme base; no pelvic frenum; pelvicfin rays I,4, 4th pelvic-fin ray with 8 (6–8, usually 7 or 8) branches, 1–3 segments between branches; 5th pelvic-fin ray absent; 11 branched caudal-fin rays; segmented caudal-fin rays 17; lateral scale rows 23 (23–24); transverse scale rows 7; no scales on head, nape, breast, or pectoral-fin base; vertebrae 26; dorsal-fin spines filamentous in males and females, extending back to 3rd or 4th element of second dorsal fin; cephalic sensory-pore system compete (Group I); male genital papilla non-fimbriate, not reaching anal-fin base, sides smooth and curved with a blunt tip with a few short skin flaps on the end; mouth oblique, forming an angle of about 45° to horizontal axis of body, lower jaw not projecting; maxilla extending to center of pupil; nostrils short, not extending to upper lip and light in color; gill opening extending forward to midway between edge of preoperculum and posterior end of opercular membrane; gill membranes attached anteriorly to isthmus, without a free fold; scales on body finely ctenoid,.
|
||
</paragraph>
|
||
<paragraph id="8BDF36DF06239F5AAD99A6ADFE08F865" blockId="1.[151,1437,249,2007]" pageId="1" pageNumber="170">
|
||
<emphasis id="B914EACD06239F5AAD99A6ADFEE9F931" bold="true" box="[199,374,1731,1755]" pageId="1" pageNumber="170">Measurements</emphasis>
|
||
(based on
|
||
<typeStatus id="54DB887D06239F5AACA0A6ADFDFFF931" box="[510,608,1731,1755]" pageId="1" pageNumber="170" type="holotype">holotype</typeStatus>
|
||
and 6
|
||
<typeStatus id="54DB887D06239F5AAFE7A6AAFCB8F936" box="[697,807,1732,1756]" pageId="1" pageNumber="170" type="paratype">paratypes</typeStatus>
|
||
,
|
||
<quantity id="4C989B3A06239F5AAE69A6ADFC7EF931" box="[823,993,1731,1755]" metricMagnitude="-2" metricUnit="m" metricValue="1.31" metricValueMax="1.48" metricValueMin="1.14" pageId="1" pageNumber="170" unit="mm" value="13.1" valueMax="14.8" valueMin="11.4">11.4–14.8 mm</quantity>
|
||
). Head length 31.6 (30.5–34.2, 31.8); origin of first dorsal fin 36.0 (33.6–37.4, 35.7), above posterior end of pectoral-fin base; origin of second dorsal fin 56.1 (55.4–59.6, 57.2), slightly in advance of anal-fin origin; origin of anal fin 63.3 (58.5–63.3, 61.6); caudalpeduncle length 25.2 (22.9–28.5, 25.5); caudal-peduncle relatively deep, 15.8 (13.9–15.8, 15.0); body of moderate depth 23.7 (21.8–25.2, 23.6); eye diameter 10.1 (9.8–11.2, 10.4); snout length 6.1 (4.2–6.1, 5.1); upper-jaw length 10.8 (10.3–12.7, 11.2).
|
||
</paragraph>
|
||
<paragraph id="8BDF36DF06239F59AD99A7F4FC3CFE25" blockId="1.[151,1437,249,2007]" lastBlockId="2.[151,1436,151,463]" lastPageId="2" lastPageNumber="171" pageId="1" pageNumber="170">
|
||
<emphasis id="B914EACD06239F5AAD99A7F4FDCEF859" bold="true" box="[199,593,1946,1971]" pageId="1" pageNumber="170">
|
||
Color in preservative of
|
||
<typeStatus id="54DB887D06239F5AACB4A7F5FDCEF859" box="[490,593,1947,1971]" pageId="1" pageNumber="170" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
: (
|
||
<figureCitation id="135B2A5A06239F5AAF36A7F5FD35F859" box="[616,682,1947,1971]" captionStart="FIGURE 1" captionStartId="2.[151,249,977,1000]" captionTargetBox="[156,1436,517,945]" captionTargetId="figure@2.[156,1441,517,947]" captionTargetPageId="2" captionText="FIGURE 1. Color in preservative of holotype of Eviota nigramembrana, OMNH-P. 35642. Photograph by D. W. Greenfield." httpUri="https://zenodo.org/record/223730/files/figure.png" pageId="1" pageNumber="170">Fig. 1</figureCitation>
|
||
). Background color of head and body pale yellowish. Head with a diffuse occipital spot about the size of the pupil (absent or faint in most
|
||
<typeStatus id="54DB887D06239F5AAE97A7D1FBA6F83D" box="[969,1081,1983,2007]" pageId="1" pageNumber="170" type="paratype">paratypes</typeStatus>
|
||
), a few scattered melanophores on side of head and nape (more intense in some
|
||
<typeStatus id="54DB887D06209F59AFEAA0F9FCBAFF45" box="[692,805,151,175]" pageId="2" pageNumber="171" type="paratype">paratypes</typeStatus>
|
||
). Opercular membrane with an obvious concentration of melanophores contrasting with a lighter operculum and pectoral-fin base (
|
||
<figureCitation id="135B2A5A06209F59A94CA0D2FBC1FF3E" box="[1042,1118,188,212]" captionStart="FIGURE 2" captionStartId="2.[151,249,1930,1953]" captionTargetBox="[283,1303,1049,1909]" captionTargetId="figure@2.[283,1304,1049,1909]" captionTargetPageId="2" captionText="FIGURE 2. Dark pigment on opercular membrane of Eviota nigramembrana, OMNH-P. 35644. Photograph by D. W. Greenfield." httpUri="https://zenodo.org/record/223731/files/figure.png" pageId="2" pageNumber="171">Fig. 2</figureCitation>
|
||
). Melanophores also more concentrated on membrane covering branchiostegals below operculum. Iris of eye black and pupil translucent yellow. Body without distinctive markings except for a subcutaneous rectangular dark spot above the midline over the ural centrum and five subcutaneous dark marks along the ventral surface between the anal-fin origin and caudal-fin base. Pectoral and pelvic-fin rays edged with tiny melanophores, membranes clear. Caudal-fin membranes peppered with light brown chromatophores, distal margin edged with brown. Anal fin covered with light brown chromatophores. Basal one-quarter of first and second dorsal fins heavily peppered with melanophores, remainder lightly peppered with distal margins darker.
|
||
</paragraph>
|
||
<caption id="DF1F665706209F59ADC9A3BFFA1AFC0D" box="[151,1413,977,1000]" httpUri="https://zenodo.org/record/223730/files/figure.png" pageId="2" pageNumber="171" targetBox="[156,1436,517,945]" targetPageId="2">
|
||
<paragraph id="8BDF36DF06209F59ADC9A3BFFA1AFC0D" blockId="2.[151,1413,977,1000]" box="[151,1413,977,1000]" pageId="2" pageNumber="171">
|
||
<emphasis id="B914EACD06209F59ADC9A3BFFE8DFC0D" bold="true" box="[151,274,977,1000]" pageId="2" pageNumber="171">FIGURE 1.</emphasis>
|
||
Color in preservative of holotype of
|
||
<taxonomicName id="4C604D5C06209F59AFD4A3BCFCE9FC0D" box="[650,886,977,999]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="171" phylum="Chordata" rank="species" species="nigramembrana">
|
||
<emphasis id="B914EACD06209F59AFD4A3BCFCE9FC0D" box="[650,886,977,999]" italics="true" pageId="2" pageNumber="171">Eviota nigramembrana</emphasis>
|
||
</taxonomicName>
|
||
, OMNH-P. 35642. Photograph by D.W. Greenfield.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1F665706209F59ADC9A7E4FE95F82A" httpUri="https://zenodo.org/record/223731/files/figure.png" pageId="2" pageNumber="171" targetBox="[283,1303,1049,1909]" targetPageId="2">
|
||
<paragraph id="8BDF36DF06209F59ADC9A7E4FE95F82A" blockId="2.[151,1437,1930,1984]" pageId="2" pageNumber="171">
|
||
<emphasis id="B914EACD06209F59ADC9A7E4FE86F84A" bold="true" box="[151,281,1930,1953]" pageId="2" pageNumber="171">FIGURE 2.</emphasis>
|
||
Dark pigment on opercular membrane of
|
||
<taxonomicName id="4C604D5C06209F59AFADA7E2FC7AF84B" box="[755,997,1931,1953]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="171" phylum="Chordata" rank="species" species="nigramembrana">
|
||
<emphasis id="B914EACD06209F59AFADA7E2FC7AF84B" box="[755,997,1931,1953]" italics="true" pageId="2" pageNumber="171">Eviota nigramembrana</emphasis>
|
||
</taxonomicName>
|
||
, OMNH-P. 35644. Photograph by D.W. Greenfield.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDF36DF06219F58AD99A0F9FD3BFCFE" blockId="3.[151,1436,151,789]" pageId="3" pageNumber="172">
|
||
<emphasis id="B914EACD06219F58AD99A0F9FE7FFF5A" bold="true" box="[199,480,151,176]" pageId="3" pageNumber="172">
|
||
Color of fresh
|
||
<typeStatus id="54DB887D06219F58AC27A0F6FE7FFF5A" box="[377,480,152,176]" pageId="3" pageNumber="172" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
: (
|
||
<figureCitation id="135B2A5A06219F58ACA7A0F9FDA0FF45" box="[505,575,151,175]" captionStart="FIGURE 3" captionStartId="3.[151,249,1537,1560]" captionTargetBox="[221,1407,841,1503]" captionTargetId="figure@3.[154,1439,841,1510]" captionTargetPageId="3" captionText="FIGURE 3. Color of fresh holotype of Eviota nigramembrana, OMNH-P. 35642. Photograph by T. Suzuki." httpUri="https://zenodo.org/record/223732/files/figure.png" pageId="3" pageNumber="172">Fig. 3</figureCitation>
|
||
). Background color of head translucent sky blue as is the lower half of the body. Background color of upper half of body and nape translucent cream. Head overlaid by a number of narrow irregular red-orange lines: four under the eye, the anteriormost from the anteroventral portion of the eye down across the upper lip; the second posterior to the first from the eye across the posterior part of the upper jaw; the third from under the center of the eye down posterior to the jaws continuing down under the head; the fourth extends from the posteroventral margin of the eye across the cheek onto the branchiostegal membrane. Cheek, operculum and nape also covered with narrow irregular red-orange lines. A black occipital spot present above preopercle. Scale pockets with bold red-orange pigment, the pigment on the belly scale pockets smaller, looking like spots on a white background. Five dark subcutaneous marks between anal-fin origin and caudal-fin base, each mark extending up onto the body forming lighter subcutaneous bars: the first above the first two soft rays of the anal fin; the second over rays six and seven; the next three evenly spaced along the caudal peduncle. A black rectangular spot over the ural centrum above its midpoint. Pectoral and pelvic fins sky blue. Anal fin heavily peppered with melanophores. Caudal fin with a red-orange crescent across its base, followed by a clear sky blue area. Remainder of caudal fin heavily peppered with melanophores, with larger black spots on basal half. First and second dorsal fins with red-orange spots along basal one-quarter with dense melanophores between them. Distal margins of both fins with a dark peppering of melanophores, the space in the center relatively clear. Color in life agrees with color of fresh
|
||
<typeStatus id="54DB887D06219F58ACE4A2B9FD83FD05" box="[442,540,727,751]" pageId="3" pageNumber="172" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="135B2A5A06219F58AF72A2B9FDEFFD05" box="[556,624,727,751]" captionStart="FIGURE 4" captionStartId="4.[151,249,1061,1084]" captionTargetBox="[155,1436,441,1032]" captionTargetId="figure@4.[155,1440,441,1032]" captionTargetPageId="4" captionText="FIGURE 4. Underwater photograph of Eviota nigramembrana, Kasari-wan Inlet, Amami-oshima Island, the Ryukyu Islands, Japan, 6 m depth, 29 Aug. 2005, Photograph by Hiroyuki Kanehara." httpUri="https://zenodo.org/record/223733/files/figure.png" pageId="3" pageNumber="172">Fig. 4</figureCitation>
|
||
). One
|
||
<typeStatus id="54DB887D06219F58AFE4A2B6FC85FD1A" box="[698,794,728,752]" pageId="3" pageNumber="172" type="paratype">paratype</typeStatus>
|
||
(OMNH-P. 34238) has more red-orange pigmentation on the fins and body than other
|
||
<typeStatus id="54DB887D06219F58AC89A293FDDCFCFF" box="[471,579,765,789]" pageId="3" pageNumber="172" type="paratype">paratypes</typeStatus>
|
||
(
|
||
<figureCitation id="135B2A5A06219F58AF0CA292FD09FCFE" box="[594,662,764,788]" captionStart="FIGURE 5" captionStartId="4.[151,249,1957,1980]" captionTargetBox="[158,1418,1266,1933]" captionTargetId="figure@4.[151,1436,1215,1936]" captionTargetPageId="4" captionText="FIGURE 5. Color of fresh paratype of Eviota nigramembrana, OMNH-P. 34238. Photograph by T. Suzuki." httpUri="https://zenodo.org/record/223734/files/figure.png" pageId="3" pageNumber="172">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF1F665706219F58ADC9A66FFB44F9F2" box="[151,1243,1537,1560]" httpUri="https://zenodo.org/record/223732/files/figure.png" pageId="3" pageNumber="172" targetBox="[221,1407,841,1503]" targetPageId="3">
|
||
<paragraph id="8BDF36DF06219F58ADC9A66FFB44F9F2" blockId="3.[151,1243,1537,1560]" box="[151,1243,1537,1560]" pageId="3" pageNumber="172">
|
||
<emphasis id="B914EACD06219F58ADC9A66FFE8DF9FD" bold="true" box="[151,274,1537,1560]" pageId="3" pageNumber="172">FIGURE 3.</emphasis>
|
||
Color of fresh holotype of
|
||
<taxonomicName id="4C604D5C06219F58AF79A66DFC8DF9F2" box="[551,786,1538,1560]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="172" phylum="Chordata" rank="species" species="nigramembrana">
|
||
<emphasis id="B914EACD06219F58AF79A66DFC8DF9F2" box="[551,786,1538,1560]" italics="true" pageId="3" pageNumber="172">Eviota nigramembrana</emphasis>
|
||
</taxonomicName>
|
||
, OMNH-P. 35642. Photograph by T. Suzuki.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37A655406219F58AD99A651FF7EF929" pageId="3" pageNumber="172" type="distribution">
|
||
<paragraph id="8BDF36DF06219F58AD99A651FF7EF929" blockId="3.[151,1436,1599,2022]" pageId="3" pageNumber="172">
|
||
<emphasis id="B914EACD06219F58AD99A651FEFFF9B2" bold="true" box="[199,352,1599,1624]" pageId="3" pageNumber="172">Distribution.</emphasis>
|
||
Amami-oshima
|
||
<collectingCountry id="F377764F06219F58AF78A651FDF2F9BD" box="[550,621,1599,1623]" name="Iceland" pageId="3" pageNumber="172">Island</collectingCountry>
|
||
, Miyako-jima
|
||
<collectingCountry id="F377764F06219F58AE45A651FCC0F9BD" box="[795,863,1599,1623]" name="Iceland" pageId="3" pageNumber="172">Island</collectingCountry>
|
||
and Yayama Islands, the Ryukyu Islands,
|
||
<collectingCountry id="F377764F06219F58A802A62EFA03F9BD" box="[1372,1436,1600,1623]" name="Japan" pageId="3" pageNumber="172">Japan</collectingCountry>
|
||
(Akihito
|
||
<emphasis id="B914EACD06219F58ADA1A60AFEA6F991" box="[255,313,1635,1659]" italics="true" pageId="3" pageNumber="172">et al.</emphasis>
|
||
2002; Hayashi
|
||
<emphasis id="B914EACD06219F58ACB1A60AFDB6F991" box="[495,553,1635,1659]" italics="true" pageId="3" pageNumber="172">et al.</emphasis>
|
||
1990; Senou
|
||
<emphasis id="B914EACD06219F58AF94A60AFC9BF991" box="[714,772,1635,1659]" italics="true" pageId="3" pageNumber="172">et al.</emphasis>
|
||
, 2004; in this study).
|
||
<collectingCountry id="F377764F06219F58A95DA60DFB4CF991" box="[1027,1235,1635,1659]" name="Philippines" pageId="3" pageNumber="172">Philippine Islands</collectingCountry>
|
||
: Negros Oriental, Siquijor
|
||
<collectingCountry id="F377764F06219F58ADA4A6E9FEA1F975" box="[250,318,1671,1695]" name="Iceland" pageId="3" pageNumber="172">Island</collectingCountry>
|
||
and Apo
|
||
<collectingCountry id="F377764F06219F58ACEEA6E9FE67F975" box="[432,504,1671,1695]" name="Iceland" pageId="3" pageNumber="172">Island</collectingCountry>
|
||
; Zamboanga del Norte, Allibay and Solino Islands; Palawan province, Tagauayan island.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A655406219F5FAD99A6BEFB24FE8F" lastPageId="4" lastPageNumber="173" pageId="3" pageNumber="172" type="etymology">
|
||
<paragraph id="8BDF36DF06219F58AD99A6BEFC3CF8E1" blockId="3.[151,1436,1599,2022]" pageId="3" pageNumber="172">
|
||
<emphasis id="B914EACD06219F58AD99A6BEFED2F902" bold="true" box="[199,333,1744,1768]" pageId="3" pageNumber="172">Etymology.</emphasis>
|
||
The specific epithet is an adjective combining the Latin
|
||
<emphasis id="B914EACD06219F58AE9BA6BEFB9DF90D" box="[965,1026,1744,1767]" italics="true" pageId="3" pageNumber="172">nigra</emphasis>
|
||
(black) and
|
||
<emphasis id="B914EACD06219F58A9D0A6A1FA99F90D" box="[1166,1286,1743,1767]" italics="true" pageId="3" pageNumber="172">membrana</emphasis>
|
||
(membrane), referring to the distinctive black pigment on the opercular membrane.
|
||
</paragraph>
|
||
<paragraph id="8BDF36DF06219F5FAD99A779FB24FE8F" blockId="3.[151,1436,1599,2022]" lastBlockId="4.[151,1436,151,357]" lastPageId="4" lastPageNumber="173" pageId="3" pageNumber="172">
|
||
<emphasis id="B914EACD06219F58AD99A779FEF9F8DA" bold="true" box="[199,358,1815,1840]" pageId="3" pageNumber="172">Comparisons</emphasis>
|
||
. Of the 27 described species of
|
||
<taxonomicName id="4C604D5C06219F58AF82A776FCBAF8C5" box="[732,805,1816,1839]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="172" phylum="Chordata" rank="genus">
|
||
<emphasis id="B914EACD06219F58AF82A776FCBAF8C5" box="[732,805,1816,1839]" italics="true" pageId="3" pageNumber="172">Eviota</emphasis>
|
||
</taxonomicName>
|
||
belonging to the cephalic sensory-pore pattern group I (Greenfield & Suzuki, 2012), only three (
|
||
<taxonomicName id="4C604D5C06219F58AFC8A752FC3DF8B9" authority="Smith, E." authorityName="Smith, E." box="[662,930,1851,1875]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="172" phylum="Chordata" rank="species" species="nebulosa">
|
||
<emphasis id="B914EACD06219F58AFC8A752FCBDF8B9" box="[662,802,1851,1875]" italics="true" pageId="3" pageNumber="172">E. nebulosa</emphasis>
|
||
Smith,
|
||
<emphasis id="B914EACD06219F58AED7A752FC3DF8B9" box="[905,930,1852,1875]" italics="true" pageId="3" pageNumber="172">E.</emphasis>
|
||
</taxonomicName>
|
||
nigripinna Lachner and Karnella, and
|
||
<taxonomicName id="4C604D5C06219F58A8DAA752FDDDF89D" authority="Greenfield and Randall" authorityName="Greenfield and Randall" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="172" phylum="Chordata" rank="species" species="winterbottomi">
|
||
<emphasis id="B914EACD06219F58A8DAA752FEABF89D" italics="true" pageId="3" pageNumber="172">E. winterbottomi</emphasis>
|
||
Greenfield and Randall
|
||
</taxonomicName>
|
||
) have the following characters shared with
|
||
<taxonomicName id="4C604D5C06219F58A975A70EFB61F89D" box="[1067,1278,1887,1911]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="172" phylum="Chordata" rank="species" species="nigramembrana">
|
||
<emphasis id="B914EACD06219F58A975A70EFB61F89D" box="[1067,1278,1887,1911]" italics="true" pageId="3" pageNumber="172">E. nigramembrana</emphasis>
|
||
</taxonomicName>
|
||
: a dorsal/anal formula almost always 8/8; 5th pelvic-fin ray absent or rudimentary; five or fewer dark internal spots or bands between anal-fin origin and caudal fin; no distinct markings on pectoral-fin base; and similar moderately deep body.
|
||
<taxonomicName id="4C604D5C06219F58AD82A7A1FE7DF80C" box="[220,482,1998,2022]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="172" phylum="Chordata" rank="species" species="nigramembrana">
|
||
<emphasis id="B914EACD06219F58AD82A7A1FE7DF80C" box="[220,482,1998,2022]" italics="true" pageId="3" pageNumber="172">Eviota nigramembrana</emphasis>
|
||
</taxonomicName>
|
||
has a dark internal mark above the midline of the ural centrum whereas it is on the middle of the centrum in
|
||
<taxonomicName id="4C604D5C06269F5FACE7A0F7FDD0FF5A" box="[441,591,153,176]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="nigripinna">
|
||
<emphasis id="B914EACD06269F5FACE7A0F7FDD0FF5A" box="[441,591,153,176]" italics="true" pageId="4" pageNumber="173">E. nigripinna</emphasis>
|
||
</taxonomicName>
|
||
and over the entire centrum in
|
||
<taxonomicName id="4C604D5C06269F5FAEE9A0F7FBEBFF5A" box="[951,1140,152,176]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="winterbottomi">
|
||
<emphasis id="B914EACD06269F5FAEE9A0F7FBEBFF5A" box="[951,1140,152,176]" italics="true" pageId="4" pageNumber="173">E. winterbottomi</emphasis>
|
||
</taxonomicName>
|
||
. The spinous dorsal fin is light in
|
||
<taxonomicName id="4C604D5C06269F5FADAAA0D3FE53FF3E" box="[244,460,188,212]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="nigramembrana">
|
||
<emphasis id="B914EACD06269F5FADAAA0D3FE53FF3E" box="[244,460,188,212]" italics="true" pageId="4" pageNumber="173">E. nigramembrana</emphasis>
|
||
</taxonomicName>
|
||
whereas it is black in
|
||
<taxonomicName id="4C604D5C06269F5FAF84A0D3FCCFFF3E" box="[730,848,189,212]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="nigripinna">
|
||
<emphasis id="B914EACD06269F5FAF84A0D3FCCFFF3E" box="[730,848,189,212]" italics="true" pageId="4" pageNumber="173">nigripinna</emphasis>
|
||
</taxonomicName>
|
||
and has broad dark and light bars in
|
||
<taxonomicName id="4C604D5C06269F5FA853A0D3FA0BFF3E" box="[1293,1428,188,212]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="nebulosa">
|
||
<emphasis id="B914EACD06269F5FA853A0D3FA0BFF3E" box="[1293,1428,188,212]" italics="true" pageId="4" pageNumber="173">E. nebulosa</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C604D5C06269F5FADC9A08FFE02FF12" box="[151,413,224,248]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="nigramembrana">
|
||
<emphasis id="B914EACD06269F5FADC9A08FFE02FF12" box="[151,413,224,248]" italics="true" pageId="4" pageNumber="173">Eviota nigramembrana</emphasis>
|
||
</taxonomicName>
|
||
also differs from all three in live coloration:
|
||
<taxonomicName id="4C604D5C06269F5FAEC5A08FFBAEFF12" box="[923,1073,225,248]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="nigripinna">
|
||
<emphasis id="B914EACD06269F5FAEC5A08FFBAEFF12" box="[923,1073,225,248]" italics="true" pageId="4" pageNumber="173">E. nigripinna</emphasis>
|
||
</taxonomicName>
|
||
has four narrow red bars across the nape that are lacking in
|
||
<taxonomicName id="4C604D5C06269F5FAC92A16BFD3FFEF6" box="[460,672,260,284]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="nigramembrana">
|
||
<emphasis id="B914EACD06269F5FAC92A16BFE42FEF6" box="[460,477,261,284]" italics="true" pageId="4" pageNumber="173">E</emphasis>
|
||
.
|
||
<emphasis id="B914EACD06269F5FACB2A16AFD3FFEF6" box="[492,672,260,284]" italics="true" pageId="4" pageNumber="173">nigramembrana</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C604D5C06269F5FAFF1A16BFCF4FEF6" box="[687,875,260,284]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="winterbottomi">
|
||
<emphasis id="B914EACD06269F5FAFF1A16BFCF4FEF6" box="[687,875,260,284]" italics="true" pageId="4" pageNumber="173">E. winterbottomi</emphasis>
|
||
</taxonomicName>
|
||
has large red spots on the side of the head and six broad dark bars on the side of the body that are lacking in
|
||
<taxonomicName id="4C604D5C06269F5FAE7AA147FC67FEAA" box="[804,1016,296,320]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="nigramembrana">
|
||
<emphasis id="B914EACD06269F5FAE7AA147FC67FEAA" box="[804,1016,296,320]" italics="true" pageId="4" pageNumber="173">E. nigramembrana</emphasis>
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="4C604D5C06269F5FA968A147FB25FEAA" box="[1078,1210,296,320]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="nebulosa">
|
||
<emphasis id="B914EACD06269F5FA968A147FB25FEAA" box="[1078,1210,296,320]" italics="true" pageId="4" pageNumber="173">E. nebulosa</emphasis>
|
||
</taxonomicName>
|
||
has two red bars on the cheek under the eye whereas
|
||
<taxonomicName id="4C604D5C06269F5FAF57A123FD43FE8E" box="[521,732,332,356]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="nigramembrana">
|
||
<emphasis id="B914EACD06269F5FAF57A123FD43FE8E" box="[521,732,332,356]" italics="true" pageId="4" pageNumber="173">E. nigramembrana</emphasis>
|
||
</taxonomicName>
|
||
has several narrow red bars under the eye.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF1F665706269F5FADC9A44BFCD5FBB0" httpUri="https://zenodo.org/record/223733/files/figure.png" pageId="4" pageNumber="173" targetBox="[155,1436,441,1032]" targetPageId="4">
|
||
<paragraph id="8BDF36DF06269F5FADC9A44BFCD5FBB0" blockId="4.[151,1436,1061,1114]" pageId="4" pageNumber="173">
|
||
<emphasis id="B914EACD06269F5FADC9A44BFE93FBD1" bold="true" box="[151,268,1061,1084]" pageId="4" pageNumber="173">FIGURE 4</emphasis>
|
||
. Underwater photograph of
|
||
<taxonomicName id="4C604D5C06269F5FAF6EA449FC83FBD6" box="[560,796,1062,1084]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="nigramembrana">
|
||
<emphasis id="B914EACD06269F5FAF6EA449FC83FBD6" box="[560,796,1062,1084]" italics="true" pageId="4" pageNumber="173">Eviota nigramembrana</emphasis>
|
||
</taxonomicName>
|
||
, Kasari-wan Inlet, Amami-oshima Island, the Ryukyu Islands, Japan, 6 m depth, 29 Aug. 2005, Photograph by Hiroyuki Kanehara.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1F665706269F5FADC9A7CBFB45F851" box="[151,1242,1957,1980]" httpUri="https://zenodo.org/record/223734/files/figure.png" pageId="4" pageNumber="173" targetBox="[158,1418,1266,1933]" targetPageId="4">
|
||
<paragraph id="8BDF36DF06269F5FADC9A7CBFB45F851" blockId="4.[151,1242,1957,1980]" box="[151,1242,1957,1980]" pageId="4" pageNumber="173">
|
||
<emphasis id="B914EACD06269F5FADC9A7CBFE93F851" bold="true" box="[151,268,1957,1980]" pageId="4" pageNumber="173">FIGURE 5</emphasis>
|
||
. Color of fresh paratype of
|
||
<taxonomicName id="4C604D5C06269F5FAF7BA7C9FC8EF856" box="[549,785,1958,1980]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="173" phylum="Chordata" rank="species" species="nigramembrana">
|
||
<emphasis id="B914EACD06269F5FAF7BA7C9FC8EF856" box="[549,785,1958,1980]" italics="true" pageId="4" pageNumber="173">Eviota nigramembrana</emphasis>
|
||
</taxonomicName>
|
||
, OMNH-P. 34238. Photograph by T. Suzuki.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37A655406279F5EAD99A0F6FD3AFE6D" pageId="5" pageNumber="174" type="discussion">
|
||
<paragraph id="8BDF36DF06279F5EAD99A0F6FD3AFE6D" blockId="5.[151,1436,151,392]" pageId="5" pageNumber="174">
|
||
<emphasis id="B914EACD06279F5EAD99A0F6FEA4FF5A" bold="true" box="[199,315,152,176]" pageId="5" pageNumber="174">Remarks.</emphasis>
|
||
We have restricted our
|
||
<typeStatus id="54DB887D06279F5EAF16A0F6FDE7FF5A" box="[584,632,152,176]" pageId="5" pageNumber="174">type</typeStatus>
|
||
material to the Ryukyu Islands,
|
||
<collectingCountry id="F377764F06279F5EAEB6A0F6FBB7FF45" box="[1000,1064,152,175]" name="Japan" pageId="5" pageNumber="174">Japan</collectingCountry>
|
||
because we have information on live and fresh color from there. The preserved material from the
|
||
<collectingCountry id="F377764F06279F5EAE2FA0D2FBA4FF3E" box="[881,1083,188,212]" name="Philippines" pageId="5" pageNumber="174">Philippine Islands</collectingCountry>
|
||
agrees with preserved material from
|
||
<collectingCountry id="F377764F06279F5EAD8BA08EFE8AFF1D" box="[213,277,224,247]" name="Japan" pageId="5" pageNumber="174">Japan</collectingCountry>
|
||
in distinguishing characters, including the distinctive dark opercular membrane (
|
||
<figureCitation id="135B2A5A06279F5EA9EEA0B1FB6AFF1D" box="[1200,1269,223,247]" captionStart="FIGURE 6" captionStartId="5.[151,250,1045,1068]" captionTargetBox="[404,1183,445,1023]" captionTargetId="figure@5.[404,1184,445,1024]" captionTargetPageId="5" captionText="FIGURE 6. Pigmented opercular membrane of Philippine Island specimen of Eviota nigramembrana, USNM 220062, 17.2 mm male. Photograph by D. W. Greenfield." httpUri="https://zenodo.org/record/223735/files/figure.png" pageId="5" pageNumber="174">Fig. 6</figureCitation>
|
||
); however, we do not have information on live coloration. There are differences in the coloration of preserved specimens from the
|
||
<typeStatus id="54DB887D06279F5EADC9A146FF58FEAA" box="[151,199,296,320]" pageId="5" pageNumber="174">type</typeStatus>
|
||
series with the
|
||
<collectingCountry id="F377764F06279F5EAC23A149FE6FFED5" box="[381,496,295,319]" name="Philippines" pageId="5" pageNumber="174">Philippine</collectingCountry>
|
||
specimens having more small melanophores on the cheeks and nape and the scale pockets more heavily pigmented (
|
||
<figureCitation id="135B2A5A06279F5EAF6EA122FDE2FE8E" box="[560,637,332,356]" captionStart="FIGURE 7" captionStartId="5.[151,249,1576,1599]" captionTargetBox="[195,1414,1203,1514]" captionTargetId="figure@5.[164,1449,1148,1550]" captionTargetPageId="5" captionText="FIGURE 7. Eviota nigramembrana from the Philippine Islands, USNM 223063, 17.6 mm male. Photograph by K. Sandved." httpUri="https://zenodo.org/record/223736/files/figure.png" pageId="5" pageNumber="174">Fig. 7</figureCitation>
|
||
). Until future DNA studies determine otherwise, we consider the
|
||
<collectingCountry id="F377764F06279F5EADC9A101FE95FE6D" box="[151,266,367,391]" name="Philippines" pageId="5" pageNumber="174">Philippine</collectingCountry>
|
||
specimens to be
|
||
<taxonomicName id="4C604D5C06279F5EAC95A11FFD01FE62" box="[459,670,368,392]" class="Actinopterygii" family="Gobiidae" genus="Eviota" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="174" phylum="Chordata" rank="species" species="nigramembrana">
|
||
<emphasis id="B914EACD06279F5EAC95A11FFD01FE62" box="[459,670,368,392]" italics="true" pageId="5" pageNumber="174">E. nigramembrana</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |