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<mods:title id="4C91AE36A94F57654D39CB51F842C350">The milliped family Trichopetalidae, Part 1: Introduction and Genera Trigenotyla Causey, Nannopetalum n. gen., and Causeyella n. gen. (Diplopoda: Chordeumatida, Cleidogonoidea)</mods:title>
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<paragraph id="7F67B855FF873D0D987817CAFD19F9B2" blockId="11.[264,772,1364,1545]" box="[264,745,1519,1545]" pageId="11" pageNumber="12">
<heading id="242F0F39FF873D0D987817CAFD19F9B2" bold="true" box="[264,745,1519,1545]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<emphasis id="4DAC6447FF873D0D987817CAFD19F9B2" bold="true" box="[264,745,1519,1545]" pageId="11" pageNumber="12">
Family
<taxonomicName id="B8D8C3D6FF873D0D981717CAFD19F9B2" ID-CoL="7NLDS" authority="Verhoeff, 1914" authorityName="Verhoeff" authorityYear="1914" box="[359,745,1519,1545]" class="Diplopoda" family="Trichopetalidae" kingdom="Animalia" order="Chordeumatida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">
Trichopetalidae
<bibRefCitation id="1B49C5A4FF873D0D9B4317CAFD19F9B2" author="Verhoeff" box="[563,745,1519,1545]" pageId="11" pageNumber="36" refString="Verhoeff, K. W. (1914) Ascospermophoren aus Japan. Zoologischer Anzeiger, 43, 125 - 143." type="journal article" year="1914">Verhoeff, 1914</bibRefCitation>
</taxonomicName>
</emphasis>
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</paragraph>
<paragraph id="7F67B855FF873D0D98781469FD7BF9D8" blockId="11.[264,1232,1612,1667]" box="[264,651,1612,1635]" pageId="11" pageNumber="12">
Trichopetalinae
<bibRefCitation id="1B49C5A4FF873D0D98C51469FDA8F9D8" author="Verhoeff" box="[437,600,1612,1635]" pageId="11" pageNumber="36" refString="Verhoeff, K. W. (1914) Ascospermophoren aus Japan. Zoologischer Anzeiger, 43, 125 - 143." type="journal article" year="1914">Verhoeff, 1914</bibRefCitation>
:347.
</paragraph>
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<paragraph id="7F67B855FF873D0D98781449FB20F938" blockId="11.[264,1232,1612,1667]" box="[264,1232,1644,1667]" pageId="11" pageNumber="12">
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<taxonomicName id="B8D8C3D6FF873D0D98781449FD76F938" ID-CoL="7NLDS" authority="Verhoeff, 1932:485" authorityName="Verhoeff" authorityPageNumber="485" authorityYear="1932" box="[264,646,1644,1667]" class="Diplopoda" family="Trichopetalidae" kingdom="Animalia" order="Chordeumatida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="family">
Trichopetalidae
<treatmentCitation id="FE799E44FF873D0D98C51449FD76F938" author="Verhoeff" box="[437,646,1644,1667]" page="485" pageId="11" pageNumber="12" year="1932">
<bibRefCitation id="1B49C5A4FF873D0D98C51449FDA8F938" author="Verhoeff" box="[437,600,1644,1667]" pageId="11" pageNumber="36" refString="Verhoeff, K. W. (1932) Diplopoden-Beitrage. (124. Diplopoden-Aufsatz.) Zoologische Jahrbucher, Abteilung fur Sytematik, 62, 459 - 524." type="journal article" year="1932">Verhoeff, 1932</bibRefCitation>
:485
</treatmentCitation>
</taxonomicName>
.
<treatmentCitation id="FE799E44FF873D0D9BE11449FC96F938" author="Hoffman" box="[657,870,1644,1667]" page="262" pageId="11" pageNumber="12" year="1961">
<bibRefCitation id="1B49C5A4FF873D0D9BE11449FCC5F938" author="Hoffman" box="[657,821,1644,1667]" pageId="11" pageNumber="36" refString="Hoffman, R. L. (1961) Systematic and morphological notes on North American conotylid Diplopoda. Transactions of the American Entomological Society, 87, 259 - 272." type="journal article" year="1961">Hoffman, 1961</bibRefCitation>
:262
</treatmentCitation>
; 1999:233 (list).
<treatmentCitation id="FE799E44FF873D0D9D6B1449FB3DF938" author="Shear" box="[1051,1229,1644,1667]" page="274" pageId="11" pageNumber="12" year="1972">
<bibRefCitation id="1B49C5A4FF873D0D9D6B1449FB6DF938" author="Shear" box="[1051,1181,1644,1667]" pageId="11" pageNumber="36" refString="Shear, W. A. (1972) Studies in the milliped Order Chordeumida (Diplopoda): a revision of the Family Cleidogonidae and a reclassification of the Order Chordeumida in the New World. Bulletin of the Museum of Comparative Zoology, 144, 151 - 352." type="journal article" year="1972">Shear, 1972</bibRefCitation>
:274
</treatmentCitation>
.
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</paragraph>
</subSubSection>
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<paragraph id="7F67B855FF873D0D98781491FD17F975" blockId="11.[264,1323,1716,1902]" box="[264,743,1716,1742]" pageId="11" pageNumber="12">
<emphasis id="4DAC6447FF873D0D98781491FE7AF976" box="[264,394,1716,1741]" italics="true" pageId="11" pageNumber="12">
<typeStatus id="A06306F7FF873D0D98781491FECFF976" box="[264,319,1716,1741]" pageId="11" pageNumber="12">Type</typeStatus>
genus
</emphasis>
:
<taxonomicName id="B8D8C3D6FF873D0D98EA1491FD13F975" authority="Harger 1872" authorityName="Harger" authorityYear="1872" box="[410,739,1716,1742]" class="Diplopoda" family="Trichopetalidae" genus="Trichopetalum" kingdom="Animalia" order="Chordeumatida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="4DAC6447FF873D0D98EA1491FDB5F976" box="[410,581,1716,1741]" italics="true" pageId="11" pageNumber="12">Trichopetalum</emphasis>
<bibRefCitation id="1B49C5A4FF873D0D9B3D1491FD13F975" author="Harger" box="[589,739,1716,1742]" pageId="11" pageNumber="35" refString="Harger, O. (1872) Descriptions of new North American myriapods. American Journal of Sciences and Arts, 4, 117 - 121." type="journal article" year="1872">Harger 1872</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="7F67B855FF873D0A984814F9FECCFEE5" blockId="11.[264,1323,1716,1902]" lastBlockId="12.[264,1323,284,910]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="4DAC6447FF873D0D984814F9FE07F94E" box="[312,503,1756,1781]" italics="true" pageId="11" pageNumber="12">Included genera</emphasis>
:
<taxonomicName id="B8D8C3D6FF873D0D9B7714F9FCBEF94D" authority="Harger 1872" authorityName="Harger" authorityYear="1872" box="[519,846,1756,1782]" class="Diplopoda" family="Trichopetalidae" genus="Trichopetalum" kingdom="Animalia" order="Chordeumatida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="4DAC6447FF873D0D9B7714F9FD42F94E" box="[519,690,1756,1781]" italics="true" pageId="11" pageNumber="12">Trichopetalum</emphasis>
<bibRefCitation id="1B49C5A4FF873D0D9BC914F9FCBEF94D" author="Harger" box="[697,846,1756,1782]" pageId="11" pageNumber="35" refString="Harger, O. (1872) Descriptions of new North American myriapods. American Journal of Sciences and Arts, 4, 117 - 121." type="journal article" year="1872">Harger 1872</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B8D8C3D6FF873D0D9A2B14F9FBA6F94D" authority="Cope 1872" authorityName="Cope" authorityYear="1872" box="[859,1110,1756,1782]" class="Diplopoda" family="Trichopetalidae" genus="Scoterpes" kingdom="Animalia" order="Chordeumatida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="4DAC6447FF873D0D9A2B14F9FC3DF94E" box="[859,973,1756,1781]" italics="true" pageId="11" pageNumber="12">Scoterpes</emphasis>
<bibRefCitation id="1B49C5A4FF873D0D9AA414F9FBA6F94D" author="Cope" box="[980,1110,1756,1782]" pageId="11" pageNumber="35" refString="Cope, E. D. (1872) On the Wyandotte Cave and its fauna. American Naturalist, 6, 406 - 422." type="journal article" year="1872">Cope 1872</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B8D8C3D6FF873D0D9D1314F9FE83F8A5" authority="Causey 1951" authorityName="Causey" authorityYear="1951" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="4DAC6447FF873D0D9D1314F9FB19F94E" box="[1123,1257,1756,1781]" italics="true" pageId="11" pageNumber="12">Trigenotyla</emphasis>
Causey 1951
</taxonomicName>
,
<taxonomicName id="B8D8C3D6FF873D0D98F01521FD55F8A5" authority="Causey 1963" authorityName="Causey" authorityYear="1963" box="[384,677,1796,1822]" class="Diplopoda" family="Trichopetalidae" genus="Mexiterpes" kingdom="Animalia" order="Chordeumatida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="4DAC6447FF873D0D98F01521FDF3F8A6" box="[384,515,1796,1821]" italics="true" pageId="11" pageNumber="12">Mexiterpes</emphasis>
<bibRefCitation id="1B49C5A4FF873D0D9B791521FD55F8A5" author="Causey" box="[521,677,1796,1822]" pageId="11" pageNumber="35" refString="Causey, Nell B. (1963) Mexiterpes sabinus, new genus and new species, a Mexican troglobite (Diplopoda: Trichopetalidae). Psyche, 70, 235 - 239." type="journal article" year="1963">Causey 1963</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B8D8C3D6FF873D0D9BC01521FCC2F8A6" box="[688,818,1796,1821]" class="Diplopoda" family="Trichopetalidae" genus="Causeyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="4DAC6447FF873D0D9BC01521FCC2F8A6" box="[688,818,1796,1821]" italics="true" pageId="11" pageNumber="12">Causeyella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="569FD93CFF873D0D9A4A1521FC7BF8A5" box="[826,907,1796,1822]" pageId="11" pageNumber="12" rank="genus">n. gen.</taxonomicNameLabel>
,
<taxonomicName id="B8D8C3D6FF873D0D9AE71521FBB3F8A6" box="[919,1091,1796,1821]" class="Diplopoda" family="Trichopetalidae" genus="Nannopetalum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Chordeumatida" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="4DAC6447FF873D0D9AE71521FBB3F8A6" box="[919,1091,1796,1821]" italics="true" pageId="11" pageNumber="12">Nannopetalum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="569FD93CFF873D0D9D3B1521FB6AF8A5" box="[1099,1178,1796,1822]" pageId="11" pageNumber="12" rank="genus">n. gen.</taxonomicNameLabel>
<emphasis id="4DAC6447FF873D0D98481509FE5FF8FE" box="[312,431,1836,1861]" italics="true" pageId="11" pageNumber="12">Diagnosis</emphasis>
: A family of Cleidogonoidea distinct in the modification of the ninth legs of males­­reduction of the telopodite to a single fused article; great development of the segmental setae; and in the (usual) presence of a strong fimbriate branch on the anterior gonopod.
</paragraph>
</subSubSection>
<subSubSection id="37C2EBDEFF803D0A98481349FE99FE45" pageId="12" pageNumber="13" type="distribution">
<paragraph id="7F67B855FF803D0A98481349FE99FE45" blockId="12.[264,1323,284,910]" pageId="12" pageNumber="13">
<emphasis id="4DAC6447FF803D0A98481349FE21FE3E" box="[312,465,364,389]" italics="true" pageId="12" pageNumber="13">Distribution:</emphasis>
North
<collectingCountry id="07CFF8C5FF803D0A9B5E1349FD66FE3D" box="[558,662,364,390]" name="United States of America" pageId="12" pageNumber="13">America</collectingCountry>
, from Newfoundland west to Wisconsin and south to
<collectingCountry id="07CFF8C5FF803D0A987813B1FE99FE15" box="[264,361,404,430]" name="Georgia" pageId="12" pageNumber="13">Georgia</collectingCountry>
, Alabama, Mississippi, Arkansas and Oklahoma (not present on the coastal plain from Virginia to
<collectingCountry id="07CFF8C5FF803D0A98BF1399FDDDFE6D" box="[463,557,444,470]" name="Georgia" pageId="12" pageNumber="13">Georgia</collectingCountry>
or the Gulf Coastal Plain), highlands of northeastern and central
<collectingCountry id="07CFF8C5FF803D0A987813C1FE95FE45" box="[264,357,484,510]" name="Mexico" pageId="12" pageNumber="13">Mexico</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="37C2EBDEFF803D0A98481029FC06FC35" pageId="12" pageNumber="13" type="discussion">
<paragraph id="7F67B855FF803D0A98481029FC06FC35" blockId="12.[264,1323,284,910]" pageId="12" pageNumber="13">
<emphasis id="4DAC6447FF803D0A98481029FE33FD9E" box="[312,451,524,549]" italics="true" pageId="12" pageNumber="13">Discussion:</emphasis>
<bibRefCitation id="1B49C5A4FF803D0A98A21029FD75FD9D" author="Causey" box="[466,645,524,550]" pageId="12" pageNumber="35" refString="Causey, N. B. (1959) A second species of the milliped genus Trigenotyla (Chordeumidea: Conotylidae: Trichopetalinae: Scoterpini). Proceedings of the Biological Society of Washington, 72, 75 - 78." type="journal article" year="1959">Causey (1959)</bibRefCitation>
established a tribe Scoterpini (
<emphasis id="4DAC6447FF803D0A9D6A1029FBAEFD9E" box="[1050,1118,524,549]" italics="true" pageId="12" pageNumber="13">recte:</emphasis>
Scoterpetini) to include the genera
<taxonomicName id="B8D8C3D6FF803D0A989C1011FDAEFDF6" box="[492,606,564,589]" class="Diplopoda" family="Trichopetalidae" genus="Scoterpes" kingdom="Animalia" order="Chordeumatida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="4DAC6447FF803D0A989C1011FDAEFDF6" box="[492,606,564,589]" italics="true" pageId="12" pageNumber="13">Scoterpes</emphasis>
</taxonomicName>
and
<emphasis id="4DAC6447FF803D0A9BEB1011FCDAFDF6" box="[667,810,564,589]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="B8D8C3D6FF803D0A9BEB1011FCD3FDF6" box="[667,803,564,589]" class="Diplopoda" family="Trichopetalidae" genus="Trigenotyla" kingdom="Animalia" order="Chordeumatida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">Trigenotyla</taxonomicName>
;
</emphasis>
this was missed by Hoffman in his monumental
<emphasis id="4DAC6447FF803D0A982F1079FC1EFDCE" box="[351,1006,604,629]" italics="true" pageId="12" pageNumber="13">
Checklist of the Millipeds of North and Middle
<collectingCountry id="07CFF8C5FF803D0A9AFB1079FC1EFDCE" box="[907,1006,604,629]" name="United States of America" pageId="12" pageNumber="13">America</collectingCountry>
</emphasis>
(1999), although Causeys paper appears in his bibliography. Otherwise no division of the family into subordinate family­level taxa has been made. Causey considered trichopetalids to be a subfamily of
<taxonomicName id="B8D8C3D6FF803D0A987810F1FE67FD55" box="[264,407,724,750]" class="Diplopoda" family="Conotylidae" kingdom="Animalia" order="Chordeumatida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Conotylidae</taxonomicName>
(see Introduction, above); had she recognized the family
<taxonomicName id="B8D8C3D6FF803D0A9D2110F1FAF8FD55" box="[1105,1288,724,750]" class="Diplopoda" family="Trichopetalidae" kingdom="Animalia" order="Chordeumatida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="family">Trichopetalidae</taxonomicName>
as raised to that status by Verhoeff in 1932, she undoubtedly would have considered her tribe to be a subfamily. However, before deciding the validity of any subfamilial or tribal grouping of the trichopetalids, I would prefer to complete my work on the large genera
<taxonomicName id="B8D8C3D6FF803D0A9D8E1169FE61FC36" class="Diplopoda" family="Trichopetalidae" genus="Trichopetalum" kingdom="Animalia" order="Chordeumatida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="4DAC6447FF803D0A9D8E1169FE61FC36" italics="true" pageId="12" pageNumber="13">Trichopetalum</emphasis>
</taxonomicName>
and
<emphasis id="4DAC6447FF803D0A98BA1151FDB4FC36" box="[458,580,884,909]" italics="true" pageId="12" pageNumber="13">
<taxonomicName id="B8D8C3D6FF803D0A98BA1151FDCFFC36" box="[458,575,884,909]" class="Diplopoda" family="Trichopetalidae" genus="Scoterpes" kingdom="Animalia" order="Chordeumatida" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">Scoterpes</taxonomicName>
,
</emphasis>
to be published in a subsequent part.
</paragraph>
</subSubSection>
</treatment>
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