308 lines
33 KiB
XML
308 lines
33 KiB
XML
<document id="A8AB2192A29ED07B919570A049204D69" ID-CLB-Dataset="3978" ID-DOI="10.11646/zootaxa.5052.4.3" ID-GBIF-Dataset="67302988-0889-4717-a9b9-52ca0cd5fa81" ID-ISSN="1175-5326" ID-Zenodo-Dep="5577237" ID-ZooBank="6B3BF392-AB36-47BB-B306-79161F76F805" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1634625773640" checkinUser="plazi" docAuthor="Parapar, Julio, Al-Kandari, Manal, Candás, María & Moreira, Juan" docDate="2021" docId="E81287982C4EF760FF518AC0FA4BFE8B" docLanguage="en" docName="zootaxa.5052.4.3.pdf" docOrigin="Zootaxa 5052 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Polyophthalmus Quatrefages 1850" docType="key" docVersion="8" lastPageNumber="523" masterDocId="142BFFE02C5BF776FFC68937FFD7FFA8" masterDocTitle="A new species of Polyophthalmus (Annelida, Opheliidae) from the Arabian Gulf with an insight on internal anatomy and diversity of the genus" masterLastPageNumber="528" masterPageNumber="501" pageNumber="522" updateTime="1699460166335" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="B31B8DCE47254C3121252FFE88173ACF" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="2CAF30C5FBF663B15759CE0EA7348167">
|
||
<mods:title id="66364D118F31C302CF947FB6B869402B">A new species of Polyophthalmus (Annelida, Opheliidae) from the Arabian Gulf with an insight on internal anatomy and diversity of the genus</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="2FF0664B83E61D6CDA9BDFF800D5E3C0" type="personal">
|
||
<mods:role id="D9CF3530E51DF613493E3647C190B2A2">
|
||
<mods:roleTerm id="4CCBB6001B23F8F084A9B2458F753DDA">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B70974CB47A6825A0844202A35A1CD0A">Parapar, Julio</mods:namePart>
|
||
<mods:affiliation id="FAD85470ED6ECE7C2B40064C22912BDE">Departamento de Bioloxía, Universidade da Coruña, Spain</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="4ECC0E7BB0C94D680A38F17DAC123622" type="personal">
|
||
<mods:role id="436D9326CE04C3BE4B83F3440A7C1F11">
|
||
<mods:roleTerm id="B14542307B6F32F06B22FA1AC7E2B7A4">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E84EAE4ECCD3FFBB355AB93E842AE6E3">Al-Kandari, Manal</mods:namePart>
|
||
<mods:affiliation id="D7FF3E5DD661704C19744ABE5B471E90">Kuwait Institute for Scientific Research, 22107 Salmiya, Kuwait.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="CD7C88ACBD6D672B082F93C0EEFCD5AC" type="personal">
|
||
<mods:role id="D3AAB5F71D8732F1A3D8B84811E2F9A0">
|
||
<mods:roleTerm id="B72A92E9A8C66647A21798E4559BD61F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2E7C384B325BEED761EA6A0C4A712AF9">Candás, María</mods:namePart>
|
||
<mods:affiliation id="ACD15B2C62AF0D0CE25AF21D12C60679">REBUSC-Estación de Bioloxía Mariña da Graña, Universidade de Santiago de Compostela, Spain.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="F43F67A62CB161DC78D6E039090B820F" type="personal">
|
||
<mods:role id="92728214BC1B02F5E05EFB7F9B36AE55">
|
||
<mods:roleTerm id="10FE6BD4FCBB690D7CCF9F502ADED2B8">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C8F111F90B241722F7FC16A8B57AA444">Moreira, Juan</mods:namePart>
|
||
<mods:affiliation id="F7543D6FD74B0CF1B079EDB5446E3E78">Departamento de Biología (Zoología) & Centro de Investigación en Biodiversidad y Cambio Global (CIBC-UAM), Facultad de Ciencias, Universidad Autónoma de Madrid, Spain.</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="E69E758A01DA71FBCC313E36A8D9780B">text</mods:typeOfResource>
|
||
<mods:relatedItem id="BF68A702856869C3232E504ED35CA502" type="host">
|
||
<mods:titleInfo id="D8D78ABD0CC9AD42A139A6D12C3EF69F">
|
||
<mods:title id="2F811907F019589A2F29B33CACB170E5">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="FFD88F0CB2535799AF0EFAD11E1DDCCF">
|
||
<mods:date id="5F4AADAE817DFE9018956BA1D0A83574">2021</mods:date>
|
||
<mods:detail id="15ADE212ABC6A8824D45FD684E6ACDA8" type="pubDate">
|
||
<mods:number id="E6ADF9823421A6D2CE7F7F820D03D2E5">2021-10-18</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="F140DB74B13269C81955A272E387E186" type="volume">
|
||
<mods:number id="C96085DD699D8CAE0EAEE0B516E11599">5052</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="7697311E310B5B657841E7CF233E62DB" type="issue">
|
||
<mods:number id="900C122EE1767A8F6D6187E9051205A0">4</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="452CAAD76C8C96DB14A5FD51A8BD8DB1" unit="page">
|
||
<mods:start id="4018F91B694062ED01E79CC053FB5B73">501</mods:start>
|
||
<mods:end id="F56F2028B1DAB6FE3A23883F509366D4">528</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="2322566A1F0B5832B92A17FBDBF5CED8">journal article</mods:classification>
|
||
<mods:identifier id="F66AFD97ACF3C9322F400F7263B7FFBC" type="CLB-Dataset">3978</mods:identifier>
|
||
<mods:identifier id="8F0ECAD15F0A88968F108CBF63303508" type="DOI">10.11646/zootaxa.5052.4.3</mods:identifier>
|
||
<mods:identifier id="5C89CE37085A4D20EE93DCFACA8DBC1E" type="GBIF-Dataset">67302988-0889-4717-a9b9-52ca0cd5fa81</mods:identifier>
|
||
<mods:identifier id="1FC6FA83B58822E92FCA05E5B423A2FA" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="32858686D68AC7DEE06DE26250F16E43" type="Zenodo-Dep">5577237</mods:identifier>
|
||
<mods:identifier id="139423A46E194957C200246B617308ED" type="ZooBank">6B3BF392-AB36-47BB-B306-79161F76F805</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="E81287982C4EF760FF518AC0FA4BFE8B" ID-DOI="http://doi.org/10.5281/zenodo.5579401" ID-GBIF-Taxon="189091505" ID-Zenodo-Dep="5579401" LSID="urn:lsid:plazi:treatment:E81287982C4EF760FF518AC0FA4BFE8B" httpUri="http://treatment.plazi.org/id/E81287982C4EF760FF518AC0FA4BFE8B" lastPageId="22" lastPageNumber="523" pageId="21" pageNumber="522">
|
||
<subSubSection id="28A165052C4EF763FF518AC0FB5BFBB8" box="[151,1164,1014,1041]" pageId="21" pageNumber="522" type="nomenclature">
|
||
<paragraph id="6004368E2C4EF763FF518AC0FB5BFBB8" blockId="21.[151,1164,1014,1041]" box="[151,1164,1014,1041]" pageId="21" pageNumber="522">
|
||
<heading id="3B4C81E22C4EF763FF518AC0FB5BFBB8" bold="true" box="[151,1164,1014,1041]" fontSize="11" level="1" pageId="21" pageNumber="522" reason="1">
|
||
<emphasis id="52CFEA9C2C4EF763FF518AC0FB5BFBB8" bold="true" box="[151,1164,1014,1041]" pageId="21" pageNumber="522">
|
||
Key to species of
|
||
<taxonomicName id="A7BB4D0D2C4EF763FEA88AC1FCCAFBB8" ID-CoL="84SK5" authority="Quatrefages, 1850" authorityName="Quatrefages" authorityYear="1850" box="[366,797,1014,1041]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="genus">
|
||
<emphasis id="52CFEA9C2C4EF763FEA88AC1FDE4FBB9" bold="true" box="[366,563,1014,1041]" italics="true" pageId="21" pageNumber="522">Polyophthalmus</emphasis>
|
||
<bibRefCitation id="042A4B7F2C4EF763FDFD8AC1FCCAFBB8" author="Quatrefages, A. de" box="[571,797,1014,1040]" pageId="21" pageNumber="522" pagination="5 - 46" refId="ref14480" refString="Quatrefages, A. de (1850) Etudes sur les types inferieurs de l'embranchement des Anneles. Memoire sur la famille des Polyphthalmiens (Polyphthalmea Nob.). Annales des Sciences Naturelles, Paris, Series 3, 13, 5 - 46." type="journal article" year="1850">Quatrefages, 1850</bibRefCitation>
|
||
</taxonomicName>
|
||
of Indian and Pacific Oceans
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="28A165052C4EF763FF518D08FD96F9A1" pageId="21" pageNumber="522" type="materials_examined">
|
||
<paragraph id="6004368E2C4EF763FF518D08FC7DFA99" blockId="21.[151,1437,1087,1545]" pageId="21" pageNumber="522">
|
||
The following tentative key comprises the seven
|
||
<taxonomicName id="A7BB4D0D2C4EF763FD0A8D08FC0BFBF1" box="[716,988,1087,1113]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="undetermined">
|
||
<emphasis id="52CFEA9C2C4EF763FD0A8D08FC55FBF1" box="[716,898,1087,1113]" italics="true" pageId="21" pageNumber="522">Polyophthalmus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
previously described in the Indian and Pacific oceans by
|
||
<bibRefCitation id="042A4B7F2C4EF763FEA78D54FE2DFBD6" author="Grube, A. E." box="[353,506,1123,1150]" pageId="21" pageNumber="522" pagination="59 - 68" refId="ref12791" refString="Grube, A. E. (1869) Familie der Opheliaceen. Schlesische gesellschaft fur vaterlandische kultur Breslau Jahresbericht, 46, 59 - 68." type="journal article" year="1869">Grube (1869)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="042A4B7F2C4EF763FDC08D54FD19FBD5" author="Kukenthal, W." box="[518,718,1123,1149]" pageId="21" pageNumber="522" pagination="361 - 373" refId="ref13441" refString="Kukenthal, W. (1887) Die Opheliaceen der expedition der Vettore Pisani. Jenaische Zeitschrift fur Naturwissenschaft, 21, 361 - 373." type="journal article" year="1887">Kükenthal (1887)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="042A4B7F2C4EF763FD1F8D54FC79FBD5" author="Michaelsen, W." box="[729,942,1123,1149]" pageId="21" pageNumber="522" pagination="1 - 23" refId="ref13643" refString="Michaelsen, W. (1892) Polychaeten von Ceylon. Jahrbuch der Hamburgischen wissenschaftlichen Anstalten, 9 (2), 1 - 23." type="journal article" year="1892">Michaelsen (1892)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="042A4B7F2C4EF763FC7F8D54FB78FBD5" author="Purschke, G. & Ding, Z. & Muller, M. C." box="[953,1199,1122,1149]" pageId="21" pageNumber="522" pagination="229 - 241" refId="ref14413" refString="Purschke, G., Ding, Z. & Muller, M. C. (1995) Ultrastructural differences as a taxonomic marker: the segmental ocelli of Polyophthalmus pictus and Polyophthalmus qingdaoensis sp. n. (Polychaeta, Opheliidae). Zoomorphology, 115 (4), 229 - 241. https: // doi. org / 10.1007 / BF 00393803" type="journal article" year="1995">
|
||
Purschke
|
||
<emphasis id="52CFEA9C2C4EF763FBE08D54FB89FBD5" box="[1062,1118,1123,1149]" italics="true" pageId="21" pageNumber="522">et al.</emphasis>
|
||
(1995)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="042A4B7F2C4EF763FB238D54FF33FB09" author="Magalhaes, W. F. & Rizzo, A. E. & Bailey-Brock, J. H." pageId="21" pageNumber="522" pagination="209 - 235" refId="ref13534" refString="Magalhaes, W. F., Rizzo, A. E. & Bailey-Brock, J. H. (2019) Opheliidae (Annelida: Polychaeta) from the western Pacific islands, including five new species. Zootaxa, 4555 (2), 209 - 235. https: // doi. org / 10.11646 / zootaxa. 4555.2.3" type="journal article" year="2019">
|
||
Magalhães
|
||
<emphasis id="52CFEA9C2C4EF763FAA38D54FA4AFBD5" box="[1381,1437,1123,1149]" italics="true" pageId="21" pageNumber="522">et al.</emphasis>
|
||
(2019)
|
||
</bibRefCitation>
|
||
, as well as the new species described here,
|
||
<taxonomicName id="A7BB4D0D2C4EF763FD078DB0FC91FB09" authority="Parapar & Al-Kandari & Candás & Moreira, 2021" authorityName="Parapar & Al-Kandari & Candás & Moreira" authorityYear="2021" box="[705,838,1159,1185]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="zhadanae" status="sp. nov.">
|
||
<emphasis id="52CFEA9C2C4EF763FD078DB0FC91FB09" box="[705,838,1159,1185]" italics="true" pageId="21" pageNumber="522">P. zhadanae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="52CFEA9C2C4EF763FC8B8DB0FC72FB09" bold="true" box="[845,933,1159,1185]" pageId="21" pageNumber="522">
|
||
<taxonomicNameLabel id="49FC57E72C4EF763FC8B8DB0FC72FB09" box="[845,933,1159,1185]" pageId="21" pageNumber="522" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Some of these species had been synonimized with the NE Atlantic
|
||
<taxonomicName id="A7BB4D0D2C4EF763FE558D9CFE22FB6C" baseAuthorityName="Dujardin" baseAuthorityYear="1839" box="[403,501,1195,1220]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="pictus">
|
||
<emphasis id="52CFEA9C2C4EF763FE558D9CFE22FB6C" box="[403,501,1195,1220]" italics="true" pageId="21" pageNumber="522">P. pictus</emphasis>
|
||
</taxonomicName>
|
||
and therefore the latter is also included in the key for comparisons. Material of previously described species has not been examined here; original descriptions and
|
||
<bibRefCitation id="042A4B7F2C4EF763FB8D8DF8FA84FB41" author="Magalhaes, W. F. & Rizzo, A. E. & Bailey-Brock, J. H." box="[1099,1363,1231,1257]" pageId="21" pageNumber="522" pagination="209 - 235" refId="ref13534" refString="Magalhaes, W. F., Rizzo, A. E. & Bailey-Brock, J. H. (2019) Opheliidae (Annelida: Polychaeta) from the western Pacific islands, including five new species. Zootaxa, 4555 (2), 209 - 235. https: // doi. org / 10.11646 / zootaxa. 4555.2.3" type="journal article" year="2019">
|
||
Magalhães
|
||
<emphasis id="52CFEA9C2C4EF763FB0B8DF8FB28FB41" box="[1229,1279,1231,1257]" italics="true" pageId="21" pageNumber="522">et al</emphasis>
|
||
. (2019
|
||
</bibRefCitation>
|
||
;
|
||
<tableCitation id="2D3903352C4EF763FA988DF8FF74FAA5" captionStart="TABLE 1" captionStartId="4.[151,239,471,497]" captionTargetPageId="4" captionText="TABLE 1. Locality and collection data, museum registration numbers for type material of new species (except for KISR) and reference to figures, of examined specimens of Polyophthalmus zhadanae sp. nov. Abbreviations: H—high intertidal; L—low intertidal; M—middle intertidal; n.d. = no data; ST—subtidal. Specimens deposited at the KISR do not have collection numbers. (*) SEM stub. Geographical coordinates in decimal degrees (degree symbols omitted); latitude is North (N) and longitude is East (E) unless noted (West: W)." httpUri="http://table.plazi.org/id/34C466062C5FF772FF5188E0FEDCFDF5" pageId="21" pageNumber="522" tableUuid="34C466062C5FF772FF5188E0FEDCFDF5">Table 1</tableCitation>
|
||
) were consulted to build the key. Some characters not explicitly mentioned in original descriptions were extracted from original illustrations.
|
||
<typeStatus id="BF00882C2C4EF763FE028C20FE2BFA99" box="[452,508,1303,1329]" pageId="21" pageNumber="522">Type</typeStatus>
|
||
locality of each species is in brackets.
|
||
</paragraph>
|
||
<paragraph id="6004368E2C4EF763FF018C0CFD96F9A1" blockId="21.[151,1437,1087,1545]" pageId="21" pageNumber="522">
|
||
Characters used in
|
||
<taxonomicName id="A7BB4D0D2C4EF763FE558C0CFD9EFAFD" authorityName="Quatrefages" authorityYear="1850" box="[403,585,1339,1365]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="genus">
|
||
<emphasis id="52CFEA9C2C4EF763FE558C0CFD9EFAFD" box="[403,585,1339,1365]" italics="true" pageId="21" pageNumber="522">Polyophthalmus</emphasis>
|
||
</taxonomicName>
|
||
keys are traditionally focused on body pigmentation, distribution of segmental eyes and chaetal length. We avoided diagnostic characters such as presence of pygidial cirri and parapodial cirri because they can only be visualized properly with SEM and therefore have been only described in recent works. In the same way, the presence and distribution of interparapodial organs is unknown for most of species. Study of
|
||
<typeStatus id="BF00882C2C4EF763FAAA8C90FA4BFA69" box="[1388,1436,1447,1473]" pageId="21" pageNumber="522">type</typeStatus>
|
||
material and newly collected specimens in the Indo-Pacific will help to better characterize these taxa regarding the aforementioned diagnostic characters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="28A165052C4EF760FF518F03FA4BFE8B" lastPageId="22" lastPageNumber="523" pageId="21" pageNumber="522" type="key">
|
||
<keyStep id="DB4F2E2B2C4EF763FF518F03FA4BF9CE" pageId="21" pageNumber="522">
|
||
<paragraph id="6004368E2C4EF763FF518F03FA4BF9E2" blockId="21.[151,1437,1588,2020]" box="[151,1436,1588,1610]" pageId="21" pageNumber="522">
|
||
<keyLead id="DB4A95BB2C4EF763FF518F03FA4BF9E2" box="[151,1436,1588,1610]" pageId="21" pageNumber="522">
|
||
1. Segmental eyes in almost all body chaetigers....................................
|
||
<taxonomicName id="A7BB4D0D2C4EF763FB8B8F03FA4BF9E2" ID-CoL="4LCTF" authority="(Dujardin, 1839) (France)" baseAuthorityName="Dujardin" baseAuthorityYear="1839" box="[1101,1436,1588,1610]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="pictus">
|
||
<emphasis id="52CFEA9C2C4EF763FB8B8F03FB4AF9E1" box="[1101,1181,1588,1609]" italics="true" pageId="21" pageNumber="522">P. pictus</emphasis>
|
||
(
|
||
<bibRefCitation id="042A4B7F2C4EF763FB6C8F03FAE9F9E2" author="Dujardin, F." box="[1194,1342,1588,1610]" pageId="21" pageNumber="522" pagination="287 - 294" refId="ref12092" refString="Dujardin, F. (1839) Observations sur quelques Annelides marines. Annales des Sciences Naturelles, Paris, 11, 287 - 294." type="journal article" year="1839">Dujardin, 1839</bibRefCitation>
|
||
) (
|
||
<collectingCountry id="18AC761E2C4EF763FA978F03FA42F9E2" box="[1361,1429,1588,1610]" name="France" pageId="21" pageNumber="522">France</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6004368E2C4EF763FF518F66FA4BF9CE" blockId="21.[151,1437,1588,2020]" box="[151,1436,1617,1639]" pageId="21" pageNumber="522">
|
||
<keyLead id="DB4A95BB2C4EF763FF518F66FA4BF9CE" box="[151,1436,1617,1639]" pageId="21" pageNumber="522">- Segmental eyes absent in anterior most and posterior most chaetigers............................................ 2</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="DB4F2E2B2C4EF763FF518F59FA4BF909" pageId="21" pageNumber="522">
|
||
<paragraph id="6004368E2C4EF763FF518F59FA4BF92B" blockId="21.[151,1437,1588,2020]" box="[151,1436,1646,1668]" pageId="21" pageNumber="522">
|
||
<keyLead id="DB4A95BB2C4EF763FF518F59FA4BF92B" box="[151,1436,1646,1668]" pageId="21" pageNumber="522">2. Chaetae of last 5–7 chaetigers longer than previous ones and surpassing pygidium................................. 3</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6004368E2C4EF763FF518FBBFA4BF909" blockId="21.[151,1437,1588,2020]" box="[151,1436,1675,1698]" pageId="21" pageNumber="522">
|
||
<keyLead id="DB4A95BB2C4EF763FF518FBBFA4BF909" box="[151,1436,1675,1698]" pageId="21" pageNumber="522">- Chaetae of last 5–7 chaetigers same length as previous ones................................................... 4</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="DB4F2E2B2C4EF763FF518F9EFA4AF974" pageId="21" pageNumber="522">
|
||
<paragraph id="6004368E2C4EF763FF518F9EFA4BF917" blockId="21.[151,1437,1588,2020]" box="[151,1436,1705,1727]" pageId="21" pageNumber="522">
|
||
<keyLead id="DB4A95BB2C4EF763FF518F9EFA4BF917" box="[151,1436,1705,1727]" pageId="21" pageNumber="522">
|
||
3. Anal marginal papillae elongate......................................
|
||
<taxonomicName id="A7BB4D0D2C4EF763FC2F8F9DFAFFF916" ID-CoL="8RDG8" authority="Michaelsen, 1892" authorityName="Michaelsen" authorityYear="1892" box="[1001,1320,1705,1727]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="longisetosus">
|
||
<emphasis id="52CFEA9C2C4EF763FC2F8F9DFBAFF916" box="[1001,1144,1705,1727]" italics="true" pageId="21" pageNumber="522">P. longisetosus</emphasis>
|
||
<bibRefCitation id="042A4B7F2C4EF763FBB88F9EFAFFF916" author="Michaelsen, W." box="[1150,1320,1705,1727]" pageId="21" pageNumber="522" pagination="1 - 23" refId="ref13643" refString="Michaelsen, W. (1892) Polychaeten von Ceylon. Jahrbuch der Hamburgischen wissenschaftlichen Anstalten, 9 (2), 1 - 23." type="journal article" year="1892">Michaelsen, 1892</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="18AC761E2C4EF763FAF38F9EFA41F917" box="[1333,1430,1705,1727]" name="Sri Lanka" pageId="21" pageNumber="522">Sri Lanka</collectingCountry>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6004368E2C4EF763FF518FF0FA4AF974" blockId="21.[151,1437,1588,2020]" box="[151,1437,1734,1756]" pageId="21" pageNumber="522">
|
||
<keyLead id="DB4A95BB2C4EF763FF518FF0FA4AF974" box="[151,1437,1734,1756]" pageId="21" pageNumber="522">
|
||
- Anal marginal papillae digitate, short...................
|
||
<taxonomicName id="A7BB4D0D2C4EF763FCF08FF0FA4AF974" ID-CoL="4LCTB" authority="Magalhaes, Rizzo and Bailey-Brock, 2019" authorityName="Magalhaes, Rizzo and Bailey-Brock" authorityYear="2019" box="[822,1437,1734,1756]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="mauliola">
|
||
<emphasis id="52CFEA9C2C4EF763FCF08FF0FC74F973" box="[822,931,1734,1756]" italics="true" pageId="21" pageNumber="522">P. mauliola</emphasis>
|
||
<bibRefCitation id="042A4B7F2C4EF763FC6F8FF1FA96F973" author="Magalhaes, W. F. & Rizzo, A. E. & Bailey-Brock, J. H." box="[937,1345,1734,1756]" pageId="21" pageNumber="522" pagination="209 - 235" refId="ref13534" refString="Magalhaes, W. F., Rizzo, A. E. & Bailey-Brock, J. H. (2019) Opheliidae (Annelida: Polychaeta) from the western Pacific islands, including five new species. Zootaxa, 4555 (2), 209 - 235. https: // doi. org / 10.11646 / zootaxa. 4555.2.3" type="journal article" year="2019">Magalhães, Rizzo and Bailey-Brock, 2019</bibRefCitation>
|
||
(Hawaii)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="DB4F2E2B2C4EF763FF518FD3FA4BF8BF" pageId="21" pageNumber="522">
|
||
<paragraph id="6004368E2C4EF763FF518FD3FA4BF951" blockId="21.[151,1437,1588,2020]" box="[151,1436,1763,1786]" pageId="21" pageNumber="522">
|
||
<keyLead id="DB4A95BB2C4EF763FF518FD3FA4BF951" box="[151,1436,1763,1786]" pageId="21" pageNumber="522">4. Prostomium with distinctive pigmentation.................................................................. 5</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6004368E2C4EF763FF518E36FA4BF8BF" blockId="21.[151,1437,1588,2020]" box="[151,1436,1793,1815]" pageId="21" pageNumber="522">
|
||
<keyLead id="DB4A95BB2C4EF763FF518E36FA4BF8BF" box="[151,1436,1793,1815]" pageId="21" pageNumber="522">- Prostomium lacking distinctive pigmentation............................................................... 6</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="DB4F2E2B2C4EF763FF518E29FA4AF8C7" pageId="21" pageNumber="522">
|
||
<paragraph id="6004368E2C4EF763FF518E29FA4BF89C" blockId="21.[151,1437,1588,2020]" box="[151,1436,1822,1844]" pageId="21" pageNumber="522">
|
||
<keyLead id="DB4A95BB2C4EF763FF518E29FA4BF89C" box="[151,1436,1822,1844]" pageId="21" pageNumber="522">
|
||
5. Pigmentation present in dorsal region of prostomium and peristomium...........
|
||
<taxonomicName id="A7BB4D0D2C4EF763FBD18E28FAFFF89B" ID-CoL="8RDG7" authority="Michaelsen, 1892" authorityName="Michaelsen" authorityYear="1892" box="[1047,1320,1822,1844]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="collaris">
|
||
<emphasis id="52CFEA9C2C4EF763FBD18E28FBA0F89B" box="[1047,1143,1822,1844]" italics="true" pageId="21" pageNumber="522">P. collaris</emphasis>
|
||
<bibRefCitation id="042A4B7F2C4EF763FBB88E29FAFFF89B" author="Michaelsen, W." box="[1150,1320,1822,1844]" pageId="21" pageNumber="522" pagination="1 - 23" refId="ref13643" refString="Michaelsen, W. (1892) Polychaeten von Ceylon. Jahrbuch der Hamburgischen wissenschaftlichen Anstalten, 9 (2), 1 - 23." type="journal article" year="1892">Michaelsen, 1892</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="18AC761E2C4EF763FAF38E29FA41F89C" box="[1333,1430,1822,1844]" name="Sri Lanka" pageId="21" pageNumber="522">Sri Lanka</collectingCountry>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6004368E2C4EF763FF518E0BFA4AF8C7" blockId="21.[151,1437,1588,2020]" pageId="21" pageNumber="522">
|
||
<keyLead id="DB4A95BB2C4EF763FF518E0BFA4AF8C7" pageId="21" pageNumber="522">
|
||
- About 10 dark pigmented structures inside brain including ocelli; peristomium not pigmented.........................................................................................
|
||
<taxonomicName id="A7BB4D0D2C4EF763FC218E6DFA4AF8C7" ID-CoL="4LCTG" authority="Purschke et al., 1995" authorityName="Purschke" authorityYear="1995" box="[999,1437,1881,1903]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="qingdaoensis">
|
||
<emphasis id="52CFEA9C2C4EF763FC218E6DFBAAF8C6" box="[999,1149,1881,1903]" italics="true" pageId="21" pageNumber="522">P. qingdaoensis</emphasis>
|
||
<bibRefCitation id="042A4B7F2C4EF763FB458E6EFA9AF8C7" author="Purschke, G. & Ding, Z. & Muller, M. C." box="[1155,1357,1881,1903]" pageId="21" pageNumber="522" pagination="229 - 241" refId="ref14413" refString="Purschke, G., Ding, Z. & Muller, M. C. (1995) Ultrastructural differences as a taxonomic marker: the segmental ocelli of Polyophthalmus pictus and Polyophthalmus qingdaoensis sp. n. (Polychaeta, Opheliidae). Zoomorphology, 115 (4), 229 - 241. https: // doi. org / 10.1007 / BF 00393803" type="journal article" year="1995">
|
||
Purschke
|
||
<emphasis id="52CFEA9C2C4EF763FB278E6DFADCF8C6" box="[1249,1291,1881,1903]" italics="true" pageId="21" pageNumber="522">et al</emphasis>
|
||
., 1995
|
||
</bibRefCitation>
|
||
(
|
||
<collectingCountry id="18AC761E2C4EF763FA9C8E6EFA41F8C7" box="[1370,1430,1881,1903]" name="China" pageId="21" pageNumber="522">China</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="DB4F2E2B2C4EF763FF518E41FA4BF801" pageId="21" pageNumber="522">
|
||
<paragraph id="6004368E2C4EF763FF518E41FA4BF824" blockId="21.[151,1437,1588,2020]" box="[151,1436,1910,1932]" pageId="21" pageNumber="522">
|
||
<keyLead id="DB4A95BB2C4EF763FF518E41FA4BF824" box="[151,1436,1910,1932]" pageId="21" pageNumber="522">6. Transverse bands of brown pigmentation similar in almost all segments.......................................... 7</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6004368E2C4EF763FF518EA3FA4BF801" blockId="21.[151,1437,1588,2020]" box="[151,1436,1940,1962]" pageId="21" pageNumber="522">
|
||
<keyLead id="DB4A95BB2C4EF763FF518EA3FA4BF801" box="[151,1436,1940,1962]" pageId="21" pageNumber="522">- Transverse bands of brown pigmentation in two different patterns............................................... 8</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="DB4F2E2B2C4EF763FF518E86FA4AF84C" pageId="21" pageNumber="522">
|
||
<paragraph id="6004368E2C4EF763FF518E86FA4BF86F" blockId="21.[151,1437,1588,2020]" box="[151,1436,1969,1991]" pageId="21" pageNumber="522">
|
||
<keyLead id="DB4A95BB2C4EF763FF518E86FA4BF86F" box="[151,1436,1969,1991]" pageId="21" pageNumber="522">
|
||
7. Segmental eyes from
|
||
<collectingCountry id="18AC761E2C4EF763FE748E86FE03F86E" box="[434,468,1969,1990]" name="Switzerland" pageId="21" pageNumber="522">CH</collectingCountry>
|
||
7 to
|
||
<collectingCountry id="18AC761E2C4EF763FE388E86FDF6F86E" box="[510,545,1969,1990]" name="Switzerland" pageId="21" pageNumber="522">CH</collectingCountry>
|
||
17........................................
|
||
<taxonomicName id="A7BB4D0D2C4EF763FBE68E85FADDF86F" ID-CoL="8RDG5" authority="Grube, 1869" authorityName="Grube" authorityYear="1869" box="[1056,1290,1969,1991]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="australis">
|
||
<emphasis id="52CFEA9C2C4EF763FBE68E85FB5BF86E" box="[1056,1164,1969,1991]" italics="true" pageId="21" pageNumber="522">P. australis</emphasis>
|
||
<bibRefCitation id="042A4B7F2C4EF763FB548E86FADDF86F" author="Grube, A. E." box="[1170,1290,1969,1991]" pageId="21" pageNumber="522" pagination="59 - 68" refId="ref12791" refString="Grube, A. E. (1869) Familie der Opheliaceen. Schlesische gesellschaft fur vaterlandische kultur Breslau Jahresbericht, 46, 59 - 68." type="journal article" year="1869">Grube, 1869</bibRefCitation>
|
||
</taxonomicName>
|
||
(NE
|
||
<collectingCountry id="18AC761E2C4EF763FAFA8E86FA41F86F" box="[1340,1430,1969,1991]" name="Australia" pageId="21" pageNumber="522">Australia</collectingCountry>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6004368E2C4EF763FF518EF8FA4AF84C" blockId="21.[151,1437,1588,2020]" box="[151,1437,1998,2020]" pageId="21" pageNumber="522">
|
||
<keyLead id="DB4A95BB2C4EF763FF518EF8FA4AF84C" box="[151,1437,1998,2020]" pageId="21" pageNumber="522">
|
||
- Segmental eyes from
|
||
<collectingCountry id="18AC761E2C4EF763FE748EF9FE02F84C" box="[434,469,1998,2020]" name="Switzerland" pageId="21" pageNumber="522">CH</collectingCountry>
|
||
5 to
|
||
<collectingCountry id="18AC761E2C4EF763FE388EF9FDF6F84C" box="[510,545,1998,2020]" name="Switzerland" pageId="21" pageNumber="522">CH</collectingCountry>
|
||
16.......................................
|
||
<taxonomicName id="A7BB4D0D2C4EF763FBD68EF8FAC1F84B" ID-CoL="8RDG9" authority="Kukenthal, 1887" authorityName="Kukenthal" authorityYear="1887" box="[1040,1302,1998,2020]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="21" pageNumber="522" phylum="Annelida" rank="species" species="striatus">
|
||
<emphasis id="52CFEA9C2C4EF763FBD68EF8FBA7F84C" box="[1040,1136,1999,2020]" italics="true" pageId="21" pageNumber="522">P. striatus</emphasis>
|
||
<bibRefCitation id="042A4B7F2C4EF763FBB08EF9FAC1F84B" author="Kukenthal, W." box="[1142,1302,1998,2020]" pageId="21" pageNumber="522" pagination="361 - 373" refId="ref13441" refString="Kukenthal, W. (1887) Die Opheliaceen der expedition der Vettore Pisani. Jenaische Zeitschrift fur Naturwissenschaft, 21, 361 - 373." type="journal article" year="1887">Kükenthal, 1887</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="18AC761E2C4EF763FAE48EF9FA41F84C" box="[1314,1430,1998,2020]" name="Hong Kong" pageId="21" pageNumber="522">Hong Kong</collectingCountry>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="DB4F2E2B2C4DF760FF5189AFFA4BFE8B" pageId="22" pageNumber="523">
|
||
<paragraph id="6004368E2C4DF760FF5189AFFA4BFF63" blockId="22.[151,1437,151,291]" pageId="22" pageNumber="523">
|
||
<keyLead id="DB4A95BB2C4DF760FF5189AFFA4BFF63" pageId="22" pageNumber="523">
|
||
8. Nine pairs of lateral eyespots (
|
||
<collectingCountry id="18AC761E2C4DF760FDCC89A0FDF8FF05" box="[522,559,151,173]" name="Switzerland" pageId="22" pageNumber="523">CH</collectingCountry>
|
||
7–15), all same size. Anterior body region (
|
||
<collectingCountry id="18AC761E2C4DF760FC1489A0FC20FF05" box="[978,1015,151,173]" name="Switzerland" pageId="22" pageNumber="523">CH</collectingCountry>
|
||
1–5) with two light brown bands; following segments with dorsal dark brown patches................................
|
||
<taxonomicName id="A7BB4D0D2C4DF760FC3A8982FAFFFF62" ID-CoL="8RDG6" authority="Kukenthal, 1887" authorityName="Kukenthal" authorityYear="1887" box="[1020,1320,181,203]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="22" pageNumber="523" phylum="Annelida" rank="species" species="ceylonensis">
|
||
<emphasis id="52CFEA9C2C4DF760FC3A8982FB54FF62" box="[1020,1155,181,202]" italics="true" pageId="22" pageNumber="523">P. ceylonensis</emphasis>
|
||
<bibRefCitation id="042A4B7F2C4DF760FB4E8982FAFFFF62" author="Kukenthal, W." box="[1160,1320,181,203]" pageId="22" pageNumber="523" pagination="361 - 373" refId="ref13441" refString="Kukenthal, W. (1887) Die Opheliaceen der expedition der Vettore Pisani. Jenaische Zeitschrift fur Naturwissenschaft, 21, 361 - 373." type="journal article" year="1887">Kükenthal, 1887</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="18AC761E2C4DF760FAF38982FA41FF63" box="[1333,1430,181,203]" name="Sri Lanka" pageId="22" pageNumber="523">Sri Lanka</collectingCountry>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="6004368E2C4DF760FF5189E4FA4BFE8B" blockId="22.[151,1437,151,291]" pageId="22" pageNumber="523">
|
||
<keyLead id="DB4A95BB2C4DF760FF5189E4FA4BFE8B" pageId="22" pageNumber="523">
|
||
- Twelve pairs of lateral eyespots (
|
||
<collectingCountry id="18AC761E2C4DF760FDE589E5FD9FFF40" box="[547,584,210,232]" name="Switzerland" pageId="22" pageNumber="523">CH</collectingCountry>
|
||
7–18); anterior and posterior ones smaller. Anterior body region (
|
||
<collectingCountry id="18AC761E2C4DF760FB0789E5FB31FF40" box="[1217,1254,210,232]" name="Switzerland" pageId="22" pageNumber="523">CH</collectingCountry>
|
||
1–10) with a dense aggregation of dark brown spots and stripes (see Description), following segments with a combination of one anterior dorsal transverse band and two posterior dark brown spots........................
|
||
<emphasis id="52CFEA9C2C4DF760FC35883AFB6BFE8A" bold="true" box="[1011,1212,268,290]" pageId="22" pageNumber="523">
|
||
<taxonomicName id="A7BB4D0D2C4DF760FC35883AFBBDFE8A" ID-CoL="8RW9V" authority="Parapar & Al-Kandari & Candás & Moreira, 2021" authorityName="Parapar & Al-Kandari & Candás & Moreira" authorityYear="2021" box="[1011,1130,268,290]" class="Polychaeta" family="Opheliidae" genus="Polyophthalmus" kingdom="Animalia" pageId="22" pageNumber="523" phylum="Annelida" rank="species" species="zhadanae" status="sp. nov.">
|
||
<emphasis id="52CFEA9C2C4DF760FC35883AFBBDFE8A" bold="true" box="[1011,1130,268,290]" italics="true" pageId="22" pageNumber="523">P. zhadanae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="49FC57E72C4DF760FBB6883AFB6BFE8A" box="[1136,1212,269,290]" pageId="22" pageNumber="523" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<collectingCountry id="18AC761E2C4DF760FB0F883AFAC6FE8B" box="[1225,1297,269,291]" name="Kuwait" pageId="22" pageNumber="523">Kuwait</collectingCountry>
|
||
/
|
||
<collectingCountry id="18AC761E2C4DF760FADE883AFA41FE8B" box="[1304,1430,269,291]" name="Saudi Arabia" pageId="22" pageNumber="523">Saudi Arabia</collectingCountry>
|
||
)
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |