treatments-xml/data/CB/9B/D8/CB9BD80813515D1897E6B3A267CEE553.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1132.86022" ID-Pensoft-Pub="1313-2970-1132-127" ID-Pensoft-UUID="E55644A3C1EC5872B08C63B5E2F3912F" ID-ZooBank="46E68453A2FA4D22A0D74509DFFB7C1B" ModsDocID="1313-2970-1132-127" checkinTime="1669812760590" checkinUser="pensoft" docAuthor="Kato, Daichi, Watanabe, Kozo &amp; Kolcsar, Levente-Peter" docDate="2022" docId="CB9BD80813515D1897E6B3A267CEE553" docLanguage="en" docName="ZooKeys 1132: 127-162" docOrigin="ZooKeys 1132" docPubDate="2022-11-29" docSource="http://dx.doi.org/10.3897/zookeys.1132.86022" docTitle="Ormosia (Parormosia) phalara Kato &amp; Kolcsar 2022, sp. nov." docType="treatment" docUuid="65BBAF0A-C395-4E8F-8CF5-D674396F44CA" docUuidSource="ZooBank" docVersion="1" id="E55644A3C1EC5872B08C63B5E2F3912F" lastPageNumber="127" masterDocId="E55644A3C1EC5872B08C63B5E2F3912F" masterDocTitle="Japanese species of Ormosia Rondani (Diptera, Limoniidae): revision of the subgenera Oreophila Lackschewitz and Parormosia Alexander" masterLastPageNumber="162" masterPageNumber="127" pageNumber="127" updateTime="1669812760590" updateUser="pensoft">
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<mods:title>Japanese species of Ormosia Rondani (Diptera, Limoniidae): revision of the subgenera Oreophila Lackschewitz and Parormosia Alexander</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kato, Daichi</mods:namePart>
<mods:affiliation>Echigo-Matsunoyama Museum of Natural Science, ' Kyororo', 1712 - 2 Matsunoyama-Matsuguchi, Tokamachi, 942 - 1411, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Watanabe, Kozo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7062-595X</mods:nameIdentifier>
<mods:affiliation>Center for Marine Environmental Studies (CMES), Ehime University, Matsuyama, Ehime 790 - 8577, Japan</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kolcsar, Levente-Peter</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7784-2386</mods:nameIdentifier>
<mods:affiliation>Center for Marine Environmental Studies (CMES), Ehime University, Matsuyama, Ehime 790 - 8577, Japan</mods:affiliation>
<mods:nameIdentifier type="email">kolcsar.peter@gmail.com</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2022</mods:date>
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<mods:number>2022-11-29</mods:number>
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<mods:detail type="volume">
<mods:number>1132</mods:number>
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<mods:start>127</mods:start>
<mods:end>162</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1132.86022</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1132.86022</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:65BBAF0A-C395-4E8F-8CF5-D674396F44CA" httpUri="http://treatment.plazi.org/id/CB9BD80813515D1897E6B3A267CEE553" lastPageNumber="127" pageId="0" pageNumber="127">
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<paragraph pageId="0" pageNumber="127">
<taxonomicName LSID="https://zoobank.org/65BBAF0A-C395-4E8F-8CF5-D674396F44CA" authority="Kato &amp; Kolcsár" authorityName="Kato &amp; Kolcsar" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Parormosia) phalara" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="phalara" status="sp. nov." subGenus="Parormosia">
Ormosia (Parormosia) phalara Kato &amp; 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="127">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="description">
<paragraph pageId="0" pageNumber="127">
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution map of Japanese species of Ormosia (Oreophila) and Ormosia (Parormosia). A Ormosia (Oreophila) confluenta Alexander, 1922 (red), Ormosia (Oreophila) komazawai Kato &amp; Kolcsar, sp. nov. (blue) B Ormosia (Oreophila) sootryeni Lackschewitz, 1935 C Ormosia (Parormosia) diversipes Alexander, 1919 D Ormosia (Parormosia) nippoalpina Alexander, 1941 (red), Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. (blue)." figureDoi="10.3897/zookeys.1132.86022.figure5" httpUri="https://binary.pensoft.net/fig/775347" pageId="0" pageNumber="127">Figs 5D</figureCitation>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Habitus of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A male B female. Scale bar: 2 mm." figureDoi="10.3897/zookeys.1132.86022.figure19" httpUri="https://binary.pensoft.net/fig/775361" pageId="0" pageNumber="127">, 19</figureCitation>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Wing of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. Scale bar: 2 mm." figureDoi="10.3897/zookeys.1132.86022.figure20" httpUri="https://binary.pensoft.net/fig/775362" pageId="0" pageNumber="127">, 20</figureCitation>
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure21" httpUri="https://binary.pensoft.net/fig/775363" pageId="0" pageNumber="127">, 21</figureCitation>
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Female terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A lateral view B genital frame, ventral view (left = anterior). Scale bars: 0.1 mm" figureDoi="10.3897/zookeys.1132.86022.figure22" httpUri="https://binary.pensoft.net/fig/775364" pageId="0" pageNumber="127">, 22</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="type material">
<paragraph pageId="0" pageNumber="127">Type material examined.</paragraph>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">
<emphasis italics="true" pageId="0" pageNumber="127">Holotype</emphasis>
.
</emphasis>
♂, pinned. Original label: &quot;JAPAN, Fukuoka, Fukuoka-shi, Sawara-ku, Itaya, Mt. Sefuri-san; alt. 970 m; 10 Jun. 2015, D. Kato leg.&quot; &quot;HOLOTYPE 
<taxonomicName authorityName="Kato &amp; Kolcsar" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Parormosia) phalara" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="phalara" subGenus="Parormosia">Ormosia (Parormosia) phalara</taxonomicName>
Kato &amp; 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
, sp. nov. [red label]&quot;; BLKU.
</paragraph>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">
<emphasis italics="true" pageId="0" pageNumber="127">Paratype</emphasis>
s.
</emphasis>
Japan • 1♂; Honshu, Aomori, Hirosaki-shi, Ichinowatari-Yamashita; 
<geoCoordinate degrees="40.53064" direction="north" orientation="latitude" precision="1" value="40.53064">40.53064°N</geoCoordinate>
, 
<geoCoordinate degrees="140.44664" direction="east" orientation="longitude" precision="1" value="140.44664">140.44664°E</geoCoordinate>
; alt. 173 m; 25 Jul. 2014; light trap; D. Kato leg.; BLKU. • 1 ♀; Honshu, Aomori, Nishimeya-mura, Kawaratai, 
<normalizedToken originalValue="Ôkawa-rindô">Okawa-rindo</normalizedToken>
Path; 
<geoCoordinate degrees="40.50062" direction="north" orientation="latitude" precision="1" value="40.50062">40.50062°N</geoCoordinate>
, 
<geoCoordinate degrees="140.20405" direction="east" orientation="longitude" precision="1" value="140.20406">140.20405°E</geoCoordinate>
; alt. 300 m; 25 Jul. - 6 Aug. 2013; Malaise trap; D. Kato and T. Nakamura leg.; BLKU. • 1 ♂; Honshu, Nagano, Iida, Kamimurahodono; 
<geoCoordinate degrees="35.45805" direction="north" orientation="latitude" precision="1" value="35.45805">35.45805°N</geoCoordinate>
, 
<geoCoordinate degrees="138.01166" direction="east" orientation="longitude" precision="1" value="138.01166">138.01166°E</geoCoordinate>
; alt. 1415 m; 3 Aug. 2019; K. Kuroda et al. leg.; EUMJ. • 2 ♂, 1 ♀; Honshu, Nagano; Matsumoto, Azusa lake; 
<geoCoordinate degrees="36.12889" direction="north" orientation="latitude" precision="1" value="36.12889">36.12889°N</geoCoordinate>
, 
<geoCoordinate degrees="137.72512" direction="east" orientation="longitude" precision="1" value="137.72511">137.72512°E</geoCoordinate>
; alt. 1000 m; 21 Jul. 2020; L.-P. 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
leg.; CKLP. • 1 ♀ (BOLD ID: JPCOI002-22); Honshu, Nagano, Kiso, Ohara Shinkai; 
<geoCoordinate degrees="35.83739" direction="north" orientation="latitude" precision="1" value="35.83739">35.83739°N</geoCoordinate>
, 
<geoCoordinate degrees="137.77346" direction="east" orientation="longitude" precision="1" value="137.77345">137.77346°E</geoCoordinate>
; alt. 1220 m, 19 Jul. 2020; L.-P. 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
leg.; CKLP. • 1 ♂; Honshu, Kanagawa, Hakone; approximate coordinates: 
<geoCoordinate degrees="35.23" direction="north" orientation="latitude" precision="555" value="35.23">35.23°N</geoCoordinate>
, 
<geoCoordinate degrees="139.02" direction="east" orientation="longitude" precision="555" value="139.02">139.02°E</geoCoordinate>
; 25 Jul. 1957; light; S. Hisamatsu leg.; EUMJ. • 3 ♂, 1 ♀; Kyushu, Fukuoka, Fukuoka-shi, Sawara-ku, Itaya, Mt. Sefuri-san; alt. 970 m; 10 Jun. 2015; D. Kato leg.; BLKU.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="diagnosis">
<paragraph pageId="0" pageNumber="127">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="127">
General coloration dark brown (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Habitus of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A male B female. Scale bar: 2 mm." figureDoi="10.3897/zookeys.1132.86022.figure19" httpUri="https://binary.pensoft.net/fig/775361" pageId="0" pageNumber="127">19</figureCitation>
). Vertex greyish. Antenna yellowish on pedicel and at least basal segments of flagellum. Mesonotum with ochreous to greyish brown parts. Wing dark brownish tinged, patterned with subhyaline spots on veins, spots free from veins absent. Cell dm closed. Halter yellow. Legs dark brown with narrow yellow areas at tips of femora and bases of tibiae in female, tibiae to tarsi mostly yellow in male. Male terminalia: tergite 9 bearing pair of long membranous lobes at caudal margin. Clasper of gonostylus divided into two arms, ventral arm ~ 3 
<normalizedToken originalValue="×">x</normalizedToken>
as long as curved stout dorsal arm, slender blade-shaped, gradually narrow toward tip. Lobe of gonostylus slender and 3/5 length of clasper, tapered distally, with long setae on ventral margin at distal 2/5. Interbase with mesal-apical lobe bearing two claws. Female terminalia with cercus slender, weakly upcurved distally. Lateral arm of genital fork rounded. Sternite 9 rounded posteriorly.
</paragraph>
<caption doi="10.3897/zookeys.1132.86022.figure19" httpUri="https://binary.pensoft.net/fig/775361" pageId="0" pageNumber="127" start="Figure 19" startId="F19">
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Figure 19.</emphasis>
Habitus of 
<taxonomicName authorityName="Kato &amp; Kolcsar" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Parormosia) phalara" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="phalara" subGenus="Parormosia">Ormosia (Parormosia) phalara</taxonomicName>
Kato &amp; 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
, sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="127">A</emphasis>
male 
<emphasis bold="true" pageId="0" pageNumber="127">B</emphasis>
female. Scale bar: 2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="description">
<paragraph pageId="0" pageNumber="127">Description.</paragraph>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Male.</emphasis>
Body length 3.2-3.6 mm, wing length 4.2-4.7 mm.
</paragraph>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" italics="true" pageId="0" pageNumber="127">Head</emphasis>
: covered with yellow and black setae. Vertex grey to dark grey, dark brownish on posterolateral part. Eyes relatively large and widely separated, ~ 4/5 as wide as narrowest point of vertex, ~ 1/2 length of head including rostrum in dorsal view. Rostrum dark brown, ~ 1/2 length of eye in lateral view. Palpus dark brown, 5-segmented, ca. as long as head, palpomere 1 globular and small, palpomeres 2-5 cylindrical, slenderer in palpomeres 2 and 5. Labellum dark brown. Antenna 3 
<normalizedToken originalValue="×">x</normalizedToken>
as long as head; 16-segmented, scape dark brown, 2 
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide, narrower basally; pedicel pale to dusky yellow, roughly globular, 1/2 of length of scape; flagellomeres pale to dusky yellow, sometimes weakly dark on distal segments, oval on basal one or two segments, long cylindrical on distal segments, each flagellomere with one or two verticils, longest one at most 2 
<normalizedToken originalValue="×">x</normalizedToken>
as long as corresponding segment, gradually shorter toward distal segment, sensilla abundant especially in ventral side, at most 1/2 as long as each flagellomere.
</paragraph>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" italics="true" pageId="0" pageNumber="127">Thorax</emphasis>
: covered with yellow to dark brown setae. Antepronotum dark brown, pale at caudal margin; postpronotum dusky yellow. Mesonotum ochreous to greyish brown, dark brown at anterior and lateral margins, sometimes with three indistinctly dark stripes just anterior to transverse suture. Prescutal pit dark brown, roughly oval with narrower inner end. Tuberculate pit distinct, situated at anterior 1/3 to 1/2 between anterior margin of mesonotum and prescutal pit. Pleuron dark brown, variegated with grey pruinosity (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Habitus of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A male B female. Scale bar: 2 mm." figureDoi="10.3897/zookeys.1132.86022.figure19" httpUri="https://binary.pensoft.net/fig/775361" pageId="0" pageNumber="127">19A</figureCitation>
). Wing (Fig. 
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Wing of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. Scale bar: 2 mm." figureDoi="10.3897/zookeys.1132.86022.figure20" httpUri="https://binary.pensoft.net/fig/775362" pageId="0" pageNumber="127">20</figureCitation>
) tinged with dark brown, subhyaline on prearcular region, patterned with subhyaline spots restricted to vicinity of veins, without ones free from veins; spot each at MA, Rs origin, crossvein sc-r, outer end of cell dm, and tips of all longitudinal veins; spot at each tip of R1, and R4 to CuA smaller; cord seamed with subhyaline; relatively narrow, 3.3 
<normalizedToken originalValue="×">x</normalizedToken>
as long as wide; Sc ending at level of R2; crossvein sc-r distinct, situated at level of middle of Rs; R2+3+4 2/7 length of R3; R2 situated between 1/2-1 
<normalizedToken originalValue="×">x</normalizedToken>
lengths of itself distal to fork of R2+3+4; M4 0.6-0.7 
<normalizedToken originalValue="×">x</normalizedToken>
as long as M3+4; cell dm closed, 0.7-0.8 
<normalizedToken originalValue="×">x</normalizedToken>
as long as cell m1+2; wing margin between tips of CuP and A1 2.5-3 
<normalizedToken originalValue="×">x</normalizedToken>
as long as that between tips of CuP and CuA; A1 curved posteriorly near middle. Halter white to dusky yellow, slightly brownish at base, ~ 1/2 length of thorax (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Habitus of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A male B female. Scale bar: 2 mm." figureDoi="10.3897/zookeys.1132.86022.figure19" httpUri="https://binary.pensoft.net/fig/775361" pageId="0" pageNumber="127">19</figureCitation>
). Legs with coxae dark brown; trochanters dusky yellow on fore pair, brown to dark brown on mid and hind pairs; femora dark brown, bases and tips narrowly yellow; tibiae yellow, tips weakly brownish; tarsi yellow, turning to brown to dark brown toward tip distal to middle of tarsomere 1 (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Habitus of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A male B female. Scale bar: 2 mm." figureDoi="10.3897/zookeys.1132.86022.figure19" httpUri="https://binary.pensoft.net/fig/775361" pageId="0" pageNumber="127">19A</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1132.86022.figure20" httpUri="https://binary.pensoft.net/fig/775362" pageId="0" pageNumber="127" start="Figure 20" startId="F20">
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Figure 20.</emphasis>
Wing of 
<taxonomicName authorityName="Kato &amp; Kolcsar" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Parormosia) phalara" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="phalara" subGenus="Parormosia">Ormosia (Parormosia) phalara</taxonomicName>
Kato &amp; 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
, sp. nov. Scale bar: 2 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" italics="true" pageId="0" pageNumber="127">Abdomen</emphasis>
: dark brown, densely covered with brown setae; yellowish setae present on genital part.
</paragraph>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" italics="true" pageId="0" pageNumber="127">Male terminalia</emphasis>
(Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure21" httpUri="https://binary.pensoft.net/fig/775363" pageId="0" pageNumber="127">21</figureCitation>
): Tergite 9 with pair of largely membranous, tongue-shaped lobes at caudal margin, ~ 1/3 length of remainder of tergite 9; tergite 9 slightly longer than wide including caudal lobe (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure21" httpUri="https://binary.pensoft.net/fig/775363" pageId="0" pageNumber="127">21A</figureCitation>
). Sternite 9 slightly and widely concave at middle of posterior margin (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure21" httpUri="https://binary.pensoft.net/fig/775363" pageId="0" pageNumber="127">21B</figureCitation>
). Gonocoxite oval, ca. as long as tergite 9, posteroventral margin not produced beyond base of clasper of gonostylus (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure21" httpUri="https://binary.pensoft.net/fig/775363" pageId="0" pageNumber="127">21A</figureCitation>
). Gonocoxal apodeme long, connected to each other, forming bridge, central part jointed with anteromedial part of interbase (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure21" httpUri="https://binary.pensoft.net/fig/775363" pageId="0" pageNumber="127">21D</figureCitation>
). Clasper of gonostylus dark, ~ 1.4 
<normalizedToken originalValue="×">x</normalizedToken>
longer than gonocoxite, divided into two arms; dorsal arm stout, rounded at tip, curved ventrally in apical view (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure21" httpUri="https://binary.pensoft.net/fig/775363" pageId="0" pageNumber="127">21C</figureCitation>
) and tip directed distally in dorsal view (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure21" httpUri="https://binary.pensoft.net/fig/775363" pageId="0" pageNumber="127">21A</figureCitation>
), distal part densely covered with black microscopic setae; ventral arm ~ 3 
<normalizedToken originalValue="×">x</normalizedToken>
as long as dorsal arm, slender blade-shaped, gradually narrow toward tip, acute at tip (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure21" httpUri="https://binary.pensoft.net/fig/775363" pageId="0" pageNumber="127">21C</figureCitation>
). Lobe of gonostylus slender, tapered distally and curved dorsally, 3/5 length of clasper, ventral margin with several long setae at distal 2/5 (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure21" httpUri="https://binary.pensoft.net/fig/775363" pageId="0" pageNumber="127">21C</figureCitation>
). Interbases fused basally, dorsolateral part roundly produced in lateral view; mesal-apical lobe slender claw-shaped, curved and directed posterodorsally, bearing smaller curved spine arising from ventral surface of mesal-apical lobe (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure21" httpUri="https://binary.pensoft.net/fig/775363" pageId="0" pageNumber="127">21D, E</figureCitation>
). Paramere wide, distinctly shorter than interbase (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure21" httpUri="https://binary.pensoft.net/fig/775363" pageId="0" pageNumber="127">21D, E</figureCitation>
). Aedeagus slender and cylindrical, extreme tip and subapical region slightly widened, tip extending beyond tip of interbase (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure21" httpUri="https://binary.pensoft.net/fig/775363" pageId="0" pageNumber="127">21D, E</figureCitation>
). Aedeagal sheath covering aedeagus except apical part, posterior end ~ 5 
<normalizedToken originalValue="×">x</normalizedToken>
as wide as aedeagus (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure21" httpUri="https://binary.pensoft.net/fig/775363" pageId="0" pageNumber="127">21D</figureCitation>
) at this point and produced dorsally near tip and produced ventrally at distal 3/5 (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure21" httpUri="https://binary.pensoft.net/fig/775363" pageId="0" pageNumber="127">21E</figureCitation>
). Sperm pump bacilliform in dorsal view, anterior end situated at level of 1/3 of paramere (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure21" httpUri="https://binary.pensoft.net/fig/775363" pageId="0" pageNumber="127">21D</figureCitation>
). Ejaculatory apodeme developed, dorsoventrally compressed, fin-like plate, ca. as long as diameter of sperm pump (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure21" httpUri="https://binary.pensoft.net/fig/775363" pageId="0" pageNumber="127">21E</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1132.86022.figure21" httpUri="https://binary.pensoft.net/fig/775363" pageId="0" pageNumber="127" start="Figure 21" startId="F21">
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Figure 21.</emphasis>
Male terminalia of 
<taxonomicName authorityName="Kato &amp; Kolcsar" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Parormosia) phalara" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="phalara" subGenus="Parormosia">Ormosia (Parormosia) phalara</taxonomicName>
Kato &amp; 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
, sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="127">A</emphasis>
dorsal view 
<emphasis bold="true" pageId="0" pageNumber="127">B</emphasis>
sternite 9, ventral view 
<emphasis bold="true" pageId="0" pageNumber="127">C</emphasis>
gonostylus, outer surface 
<emphasis bold="true" pageId="0" pageNumber="127">D</emphasis>
aedeagal complex, dorsal view (left gonocoxal apodeme omitted) 
<emphasis bold="true" pageId="0" pageNumber="127">E</emphasis>
aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Female</emphasis>
. Body length 3.6-4.8 mm, wing length 4.2-5.0 mm. Generally resembling male (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Habitus of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A male B female. Scale bar: 2 mm." figureDoi="10.3897/zookeys.1132.86022.figure19" httpUri="https://binary.pensoft.net/fig/775361" pageId="0" pageNumber="127">19B</figureCitation>
) except, antenna shorter, ~ 2 
<normalizedToken originalValue="×">x</normalizedToken>
length of head; each flagellomere with ca. eight verticils on each of basal segments, fewer on each of distal segments, longest one at most 1.5 
<normalizedToken originalValue="×">x</normalizedToken>
as long as corresponding segment. Tibiae to tarsi dark brown, bases of tibiae narrowly yellow (Fig. 
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Habitus of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A male B female. Scale bar: 2 mm." figureDoi="10.3897/zookeys.1132.86022.figure19" httpUri="https://binary.pensoft.net/fig/775361" pageId="0" pageNumber="127">19B</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" italics="true" pageId="0" pageNumber="127">Female terminalia</emphasis>
(Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Female terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A lateral view B genital frame, ventral view (left = anterior). Scale bars: 0.1 mm" figureDoi="10.3897/zookeys.1132.86022.figure22" httpUri="https://binary.pensoft.net/fig/775364" pageId="0" pageNumber="127">22</figureCitation>
): dark brown, cercus and hypogynial valve amber-colored, weakly dark on basal parts. Tergites 8 and 9 fused. Cercus weakly upcurved distally, 1.6 
<normalizedToken originalValue="×">x</normalizedToken>
longer than tergite 10; hypogynial valve relatively stout, 1.5 
<normalizedToken originalValue="×">x</normalizedToken>
as long as sternite 8, gradually narrowed toward tip, basal part 1.4 
<normalizedToken originalValue="×">x</normalizedToken>
as wide as that of cercus, tip ending at level of basal 3/5 of cercus (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Female terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A lateral view B genital frame, ventral view (left = anterior). Scale bars: 0.1 mm" figureDoi="10.3897/zookeys.1132.86022.figure22" httpUri="https://binary.pensoft.net/fig/775364" pageId="0" pageNumber="127">22A</figureCitation>
). Genital frame with genital fork widened posteriorly, anterior part slender; lateral arm of genital fork weakly produced, rounded, situated at posterior of genital fork; sternite 9 rounded posteriorly; long and arched groove present posterior to genital opening, lateral part curved anteriorly (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Female terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A lateral view B genital frame, ventral view (left = anterior). Scale bars: 0.1 mm" figureDoi="10.3897/zookeys.1132.86022.figure22" httpUri="https://binary.pensoft.net/fig/775364" pageId="0" pageNumber="127">22B</figureCitation>
). Three spermathecal ducts present, basal parts blackened (Fig. 
<figureCitation captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Female terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A lateral view B genital frame, ventral view (left = anterior). Scale bars: 0.1 mm" figureDoi="10.3897/zookeys.1132.86022.figure22" httpUri="https://binary.pensoft.net/fig/775364" pageId="0" pageNumber="127">22B</figureCitation>
). Spermathecae indistinct.
</paragraph>
<caption doi="10.3897/zookeys.1132.86022.figure22" httpUri="https://binary.pensoft.net/fig/775364" pageId="0" pageNumber="127" start="Figure 22" startId="F22">
<paragraph pageId="0" pageNumber="127">
<emphasis bold="true" pageId="0" pageNumber="127">Figure 22.</emphasis>
Female terminalia of 
<taxonomicName authorityName="Kato &amp; Kolcsar" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Parormosia) phalara" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="phalara" subGenus="Parormosia">Ormosia (Parormosia) phalara</taxonomicName>
Kato &amp; 
<normalizedToken originalValue="Kolcsár">Kolcsar</normalizedToken>
, sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="127">A</emphasis>
lateral view 
<emphasis bold="true" pageId="0" pageNumber="127">B</emphasis>
genital frame, ventral view (left = anterior). Scale bars: 0.1 mm
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="etymology">
<paragraph pageId="0" pageNumber="127">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="127">
The name of this species, 
<taxonomicName authorityName="Kato &amp; Kolcsar" authorityYear="2022" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia phalara" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="phalara">
<emphasis italics="true" pageId="0" pageNumber="127">Ormosia phalara</emphasis>
</taxonomicName>
, is from the Greek 
<emphasis italics="true" pageId="0" pageNumber="127">phalaros</emphasis>
, meaning white-spotted. The name is deemed to be a Latinized feminine adjective in nominative singular.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="distribution">
<paragraph pageId="0" pageNumber="127">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="127">
Japan (Honshu and Kyushu) (Fig. 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Distribution map of Japanese species of Ormosia (Oreophila) and Ormosia (Parormosia). A Ormosia (Oreophila) confluenta Alexander, 1922 (red), Ormosia (Oreophila) komazawai Kato &amp; Kolcsar, sp. nov. (blue) B Ormosia (Oreophila) sootryeni Lackschewitz, 1935 C Ormosia (Parormosia) diversipes Alexander, 1919 D Ormosia (Parormosia) nippoalpina Alexander, 1941 (red), Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. (blue)." figureDoi="10.3897/zookeys.1132.86022.figure5" httpUri="https://binary.pensoft.net/fig/775347" pageId="0" pageNumber="127">5D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="127" type="remarks">
<paragraph pageId="0" pageNumber="127">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="127">
This species resembles a Chinese species, 
<taxonomicName authorityName="Alexander" authorityYear="1936" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Parormosia) angustaurata" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="angustaurata" subGenus="Parormosia">Ormosia (Parormosia) angustaurata</taxonomicName>
Alexander, 1936 (
<bibRefCitation author="Alexander, CP" journalOrPublisher="Philippine Journal of Science" pageId="0" pageNumber="127" pagination="323 - 360" refId="B6" refString="Alexander, CP, 1936b. New or little-known Tipulidae from eastern Asia (Diptera). XXXI. Philippine Journal of Science 60: 323 - 360" title="New or little-known Tipulidae from eastern Asia (Diptera). XXXI." volume="60" year="1936 b">Alexander 1936b</bibRefCitation>
), but is distinguished from it by the following characters: scape dark brown (yellow in 
<taxonomicName authorityName="Alexander" authorityYear="1936" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Parormosia) angustaurata" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="angustaurata" subGenus="Parormosia">Ormosia (Parormosia) angustaurata</taxonomicName>
); wing with subhyaline spots restricted to vicinity of veins, without smaller spots free from veins (with at least a few smaller spots free from veins in addition to ones on veins in 
<taxonomicName authorityName="Alexander" authorityYear="1936" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Parormosia) angustaurata" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="angustaurata" subGenus="Parormosia">Ormosia (Parormosia) angustaurata</taxonomicName>
); mesal-apical lobe slender with additional spine near tip (Fig. 
<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Male terminalia of Ormosia (Parormosia) phalara Kato &amp; Kolcsar, sp. nov. A dorsal view B sternite 9, ventral view C gonostylus, outer surface D aedeagal complex, dorsal view (left gonocoxal apodeme omitted) E aedeagal complex, lateral view (left = dorsal). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1132.86022.figure21" httpUri="https://binary.pensoft.net/fig/775363" pageId="0" pageNumber="127">21D, E</figureCitation>
) (stout and without additional spine in 
<taxonomicName authorityName="Alexander" authorityYear="1936" class="Insecta" family="Limoniidae" genus="Ormosia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ormosia (Parormosia) angustaurata" order="Diptera" pageId="0" pageNumber="127" phylum="Arthropoda" rank="species" species="angustaurata" subGenus="Parormosia">Ormosia (Parormosia) angustaurata</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>