142 lines
16 KiB
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142 lines
16 KiB
XML
<document id="ECF2314211CB94C2C21BA9DE2812A58B" ID-DOI="10.5281/zenodo.4651080" ID-ISSN="1638-9395" ID-Zenodo-Dep="4651080" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617195131926" checkinUser="felipe" docAuthor="Bianucci, Giovanni, Lambert, Olivier & Post, Klaas" docDate="2007" docId="550B87C3C158FF8DFC8A77FAFCCC04A1" docLanguage="en" docName="Geodiversitas.29.4.561-618.pdf" docOrigin="Geodiversitas 29 (4)" docStyle="DocumentStyle:94524207C6714CF92F879FE43E306202.3:Geodiversitas.2000-2008.journal_article.type2" docStyleId="94524207C6714CF92F879FE43E306202" docStyleName="Geodiversitas.2000-2008.journal_article.type2" docStyleVersion="3" docTitle="Odontoceti Flower 1867" docType="treatment" docVersion="3" lastPageNumber="601" masterDocId="A932FFBBC17EFFA5FFC47277FFBE056E" masterDocTitle="A high diversity in fossil beaked whales (Mammalia, Odontoceti, Ziphiidae) recovered by trawling from the sea floor off South Africa" masterLastPageNumber="618" masterPageNumber="561" pageNumber="599" updateTime="1698897450673" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:mods id="D4AB93BE28355874C36BCEA9A8B206BE" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A11525BA0BE02157F82C37AF2636A1F9">
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<mods:title id="2161AD237E7AA597FD3FD615DFF351B0">A high diversity in fossil beaked whales (Mammalia, Odontoceti, Ziphiidae) recovered by trawling from the sea floor off South Africa</mods:title>
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</mods:titleInfo>
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<mods:name id="DBBD80BD8C58BA34679BDF7146E44B91" type="personal">
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<mods:role id="2729B06C45649C5EA36443213452184E">
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<mods:roleTerm id="90C6014F613DD2C0A59888E79886DEC3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CFAD4508B184B4BD9A0792125C379FC9">Bianucci, Giovanni</mods:namePart>
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<mods:affiliation id="21A1799BCED096038276C0E70CBBB408">Dipartimento di Scienze della Terra, Università di Pisa, via S. Maria, 53, I- 56126 Pisa (Italy) bianucci @ dst. unipi. it</mods:affiliation>
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</mods:name>
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<mods:name id="2A128DB7DDCC4CF9AF211C30302CD995" type="personal">
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<mods:role id="544F9B67A35B4ADE13FB1EF9CBFC6338">
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<mods:roleTerm id="48E8E9685FECC4ABC1620700BDB4A7A0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3D8B09AD13477C0B32249E5F02D9A2E1">Lambert, Olivier</mods:namePart>
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<mods:affiliation id="C5DDF9C9DB748FFCB40EAF5EF22DD908">Institut royal des Sciences naturelles de Belgique, Département de Paléontologie, rue Vautier, 29, B- 1000 Brussels (Belgium) olivier. lambert @ naturalsciences. be</mods:affiliation>
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</mods:name>
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<mods:name id="533240541A975CE1EB5D8B89B5C9EC9E" type="personal">
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<mods:role id="C00EA072F0D6DC2C54CE517A0ACA085D">
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<mods:roleTerm id="1BD5231560AA23D7B52AE6E0688B2F7F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6A1FE3B525319E599877EEBE36A2C460">Post, Klaas</mods:namePart>
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<mods:affiliation id="3087447ADB5853D15E485DFBA6221963">Natuurhistorisch Museum Rotterdam, P. O. Box 23452, NL- 3001 Rotterdam (The Netherlands) klaas @ fiskano. nl</mods:affiliation>
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<mods:nameIdentifier id="7CFCEC1C9FFC258DD3C0B29517D2AC47" type="email">klaas@fiskano.nl</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="85D83608AE5E2D2E07413C3CB65FC067">text</mods:typeOfResource>
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<mods:title id="39B9FEA9A50A8BED03874A848A1FA934">Geodiversitas</mods:title>
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</mods:titleInfo>
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<mods:part id="1E6DB38DDCD855A8DC861FF64660C7F4">
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<mods:date id="E39BDAFA0A46CEEFEAC496E79FD792A6">2007</mods:date>
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<mods:detail id="6E13C2D810C833251E6621F8A9CD06BA" type="volume">
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<mods:number id="ECC158290FF96994D43DCC3ADFF141FD">29</mods:number>
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</mods:detail>
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<mods:detail id="6B8119420335DF40799DB920A153C207" type="issue">
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<mods:number id="699F68124FE5438957546EBC0A5A51E2">4</mods:number>
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</mods:detail>
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<mods:extent id="55C1DB7DC25B7364782E376091349CBD" unit="page">
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<mods:start id="804F4CFCDA64C74002ACAA0BFD7688EE">561</mods:start>
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<mods:end id="6C97061C91DAA7E5E8DF333EF0EBF2BA">618</mods:end>
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</mods:extent>
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<mods:classification id="68439F8AF8D7C6E4848B4A243CCA1A84">journal article</mods:classification>
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<mods:identifier id="F7E6D9B61C6E976790FBD826ABC61E5F" type="DOI">10.5281/zenodo.4651080</mods:identifier>
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<mods:identifier id="F6A6F553ADEAAC7AF7383B4FBEC0233C" type="ISSN">1638-9395</mods:identifier>
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<mods:identifier id="3E55D82CCA05720E0C843627170A6612" type="Zenodo-Dep">4651080</mods:identifier>
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<mods:identifier id="4C79991B10AC4E5C364511E5EE7A49CB" type="ZooBank">FA38C827-6C5F-4B70-B306-F30C90801A2F</mods:identifier>
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</mods:mods>
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<treatment id="550B87C3C158FF8DFC8A77FAFCCC04A1" LSID="urn:lsid:plazi:treatment:550B87C3C158FF8DFC8A77FAFCCC04A1" httpUri="http://treatment.plazi.org/id/550B87C3C158FF8DFC8A77FAFCCC04A1" lastPageId="40" lastPageNumber="601" pageId="38" pageNumber="599">
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<subSubSection id="95B8655EC158FF83FC8A77FAFBE50344" pageId="38" pageNumber="599" type="nomenclature">
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<paragraph id="DD1D36D5C158FF83FC8A77FAFBAF00C8" blockId="38.[846,1041,1420,1447]" box="[846,1041,1420,1447]" pageId="38" pageNumber="599">
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<heading id="865581B9C158FF83FC8A77FAFBAF00C8" box="[846,1041,1420,1447]" centered="true" fontSize="11" level="2" pageId="38" pageNumber="599" reason="2">
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<taxonomicName id="1AA24D56C158FF83FC8A77FAFC7500C9" ID-CoL="62397" authorityName="Flower" authorityYear="1867" box="[846,971,1421,1447]" class="Mammalia" kingdom="Animalia" order="Cetacea" pageId="38" pageNumber="565" phylum="Chordata" rank="subOrder" subOrder="Odontoceti">Odontoceti</taxonomicName>
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indet.
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</heading>
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</paragraph>
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<paragraph id="DD1D36D5C158FF83FD6177B3FBE50344" blockId="38.[677,1210,1475,1578]" pageId="38" pageNumber="599">
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REFERRED SPECIMENS. — SAM PQ 2717, isolated rostrum, trawled off Saldanha Bay, west coast of
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<collectingCountry id="A5B57645C158FF83FBF077A9FB0B0098" box="[1076,1205,1502,1526]" name="South Africa" pageId="38" pageNumber="599">South Africa</collectingCountry>
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, Atlantic Ocean, depth of
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<quantity id="1A5A9B30C158FF83FC55778FFC6E037E" box="[913,976,1528,1552]" metricMagnitude="2" metricUnit="m" metricValue="3.66" pageId="38" pageNumber="599" unit="m" value="366.0">366 m</quantity>
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; SAM PQ 2719, isolated rostrum, trawled off the South African coast.
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</paragraph>
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</subSubSection>
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<subSubSection id="95B8655EC158FF8DFD61743DFCCC04A1" lastPageId="40" lastPageNumber="601" pageId="38" pageNumber="599" type="description">
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<paragraph id="DD1D36D5C158FF83FD61743DFB3B030B" blockId="38.[677,1210,1610,1700]" box="[677,1157,1610,1637]" pageId="38" pageNumber="599">
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DESCRIPTION AND DISCUSSION (
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<figureCitation id="45992A50C158FF83FC39743CFBEA030B" box="[1021,1108,1611,1637]" captionStart="FIG" captionStartId="39.[156,167,1680,1697]" captionTargetBox="[115,1210,250,1560]" captionTargetId="figure-18@39.[136,1180,223,1569]" captionTargetPageId="39" captionText="FIG. 31. — Rostrum of Odontoceti indet. (SAM PQ 2717): A, dorsal view; B, lateral view; C, ventral view. Scale bar: 10 cm." figureDoi="http://doi.org/10.5281/zenodo.4651149" httpUri="https://zenodo.org/record/4651149/files/figure.png" pageId="38" pageNumber="599">FIGS 31</figureCitation>
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;
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<figureCitation id="45992A50C158FF83FBA4743CFBC3030B" box="[1120,1149,1611,1637]" captionStart="FIG" captionStartId="41.[113,124,1635,1652]" captionTargetBox="[132,1194,227,1592]" captionTargetId="figure-18@41.[125,1214,224,1621]" captionTargetPageId="41" captionText="FIG. 32. — A-C, rostrum of Odontoceti indet. (SAM PQ 2717); A, detail of the alveolar groove; B, anterior view; C, anterolateral view; D-G, rostrum of Odontoceti indet. (SAM PQ 2719); D, dorsal view; E, lateral view; F, anterior view; G, anterolateral view. Scale bars: 10 cm." figureDoi="http://doi.org/10.5281/zenodo.4651151" httpUri="https://zenodo.org/record/4651151/files/figure.png" pageId="38" pageNumber="599">32</figureCitation>
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)
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</paragraph>
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<paragraph id="DD1D36D5C158FF83FD61741EFB0703CA" blockId="38.[677,1210,1610,1700]" pageId="38" pageNumber="599">The isolated rostra SAM PQ 2717 and SAM PQ 2719, preserved for a length of respectively 519</paragraph>
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<paragraph id="DD1D36D5C159FF82FF6672ACFD780591" blockId="39.[162,710,219,255]" box="[162,710,219,255]" pageId="39" pageNumber="600">A premaxilla maxilla thickened vomer</paragraph>
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<caption id="89DD665DC159FF82FF5874E7FB2E03CF" ID-DOI="http://doi.org/10.5281/zenodo.4651149" ID-Zenodo-Dep="4651149" box="[156,1168,1680,1697]" httpUri="https://zenodo.org/record/4651149/files/figure.png" pageId="39" pageNumber="600" startId="39.[156,167,1680,1697]" targetBox="[115,1210,250,1560]" targetPageId="39">
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<paragraph id="DD1D36D5C159FF82FF5874E7FB2E03CF" blockId="39.[156,1168,1541,1697]" box="[156,1168,1680,1697]" pageId="39" pageNumber="600">
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FIG. 31. — Rostrum of
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<taxonomicName id="1AA24D56C159FF82FE9C74E7FE0B03CF" authorityName="Flower" authorityYear="1867" box="[344,437,1680,1697]" class="Mammalia" kingdom="Animalia" order="Cetacea" pageId="39" pageNumber="565" phylum="Chordata" rank="subOrder" subOrder="Odontoceti">Odontoceti</taxonomicName>
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indet. (SAM PQ 2717):
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<emphasis id="EFD6EAC7C159FF82FDBD74E7FD3803CF" bold="true" box="[633,646,1680,1697]" pageId="39" pageNumber="600">A</emphasis>
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, dorsal view;
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<emphasis id="EFD6EAC7C159FF82FD3D74E7FCB803CF" bold="true" box="[761,774,1680,1697]" pageId="39" pageNumber="600">B</emphasis>
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, lateral view;
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<emphasis id="EFD6EAC7C159FF82FCBC74E7FC3803CF" bold="true" box="[888,902,1680,1697]" pageId="39" pageNumber="600">C</emphasis>
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, ventral view. Scale bar: 10 cm.
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</paragraph>
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</caption>
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<paragraph id="DD1D36D5C156FF8DFFB572AFFF67047C" blockId="40.[113,647,216,1700]" pageId="40" pageNumber="601">
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and
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<quantity id="1A5A9B30C156FF8DFF6572AFFEBC059C" box="[161,258,216,243]" metricMagnitude="-1" metricUnit="m" metricValue="5.47" pageId="40" pageNumber="601" unit="mm" value="547.0">547 mm</quantity>
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, are characterized by a very unusual structure.
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</paragraph>
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<paragraph id="DD1D36D5C156FF8DFF4C7360FDA50783" blockId="40.[113,647,216,1700]" pageId="40" pageNumber="601">
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The premaxilla is weakly modified compared to the general odontocete plan. It occupies the margins of the mesorostral groove in dorsal view and the upturned apex of the rostrum, ventrally appearing on the first third of the rostrum. At the apex of the maxilla, the rostrum has a width of respectively 60 and
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<quantity id="1A5A9B30C156FF8DFF0173A2FEAB049E" box="[197,277,469,496]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="40" pageNumber="601" unit="mm" value="45.0">45 mm</quantity>
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for SAM PQ 2717 and SAM PQ 2719 and a height of respectively 56 and
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<quantity id="1A5A9B30C156FF8DFDF47382FD3D0761" box="[560,643,501,527]" metricMagnitude="-2" metricUnit="m" metricValue="4.4" pageId="40" pageNumber="601" unit="mm" value="44.0">44 mm</quantity>
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. The mesorostral groove is partially filled by the thickened lateral walls of the vomer, leaving an open median V-shaped groove. The development of the vomer in the mesorostral groove combined with high-density bone tissue is mostly observed in ziphiids (see above). Ventrally, the exposure of the vomer is respectively 290 and
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<quantity id="1A5A9B30C156FF8DFE7270A4FDA90783" box="[438,535,723,749]" metricMagnitude="-1" metricUnit="m" metricValue="2.4" pageId="40" pageNumber="601" unit="mm" value="240.0">240 mm</quantity>
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.
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</paragraph>
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<paragraph id="DD1D36D5C156FF8DFF4C7085FEA000C8" blockId="40.[113,647,216,1700]" pageId="40" pageNumber="601">
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The maxilla is highly modified; the lateral margin is considerably elevated laterodorsally along the posterior two thirds of the rostrum. This lateral maxillary crest is robust at its base, progressively thinning dorsally. The maximum height of the crest is mid-way along the rostrum.The two crests anteriorly delimit a wide and deep basin, ending at some distance before the apex of the maxilla. The inner wall of the crest is striated with vascularization sulci, best seen in SAM PQ 2717. On this specimen, the outer wall of the crest, continuous with the wide ventral surface of the rostrum, bears a shallow alveolar groove punctuated by a series of rounded protuberances interpreted as unerupted teeth. No alveolar groove is observed in SAM PQ 2719. The basin is considerably larger in SAM PQ 2717 (maximum width and height of the rostrum respectively 230 and
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<quantity id="1A5A9B30C156FF8DFF59777AFF470049" box="[157,249,1293,1319]" metricMagnitude="-1" metricUnit="m" metricValue="2.01" pageId="40" pageNumber="601" unit="mm" value="201.0">201 mm</quantity>
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), a difference that might correspond to sexual dimorphism, as seen in
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<taxonomicName id="1AA24D56C156FF8DFE67775AFD380029" authorityName="Linnaeus" authorityYear="1758" box="[419,646,1325,1351]" class="Mammalia" family="Physeteridae" genus="Physeter" kingdom="Animalia" order="Cetacea" pageId="40" pageNumber="601" phylum="Chordata" rank="species" species="macrocephalus">
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<emphasis id="EFD6EAC7C156FF8DFE67775AFD380029" box="[419,646,1325,1351]" italics="true" pageId="40" pageNumber="601">Physeter macrocephalus</emphasis>
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</taxonomicName>
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for the supracranial basin, in
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<taxonomicName id="1AA24D56C156FF8DFE62773BFDDF0008" authorityName="Cuvier" authorityYear="1823" box="[422,609,1356,1382]" class="Mammalia" family="Hyperoodontidae" genus="Ziphius" kingdom="Animalia" order="Cetacea" pageId="40" pageNumber="601" phylum="Chordata" rank="species" species="cavirostris">
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<emphasis id="EFD6EAC7C156FF8DFE62773BFDDF0008" box="[422,609,1356,1382]" italics="true" pageId="40" pageNumber="601">Ziphius cavirostris</emphasis>
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</taxonomicName>
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for the prenarial basin, and in
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<taxonomicName id="1AA24D56C156FF8DFE48771BFDB600E8" authorityName="Lacepede" authorityYear="1804" box="[396,520,1388,1414]" class="Mammalia" family="Hyperoodontidae" genus="Hyperoodon" kingdom="Animalia" order="Cetacea" pageId="40" pageNumber="601" phylum="Chordata" rank="genus">
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<emphasis id="EFD6EAC7C156FF8DFE48771BFDB600E8" box="[396,520,1388,1414]" italics="true" pageId="40" pageNumber="601">Hyperoodon</emphasis>
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</taxonomicName>
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spp. for the maxillary crests.
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</paragraph>
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<paragraph id="DD1D36D5C156FF8DFF4C77DCFCCC04A1" blockId="40.[113,647,216,1700]" lastBlockId="40.[677,1210,216,464]" pageId="40" pageNumber="601">
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Similarities with
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<taxonomicName id="1AA24D56C156FF8DFE8E77DCFE7400AB" authorityName="Lacepede" authorityYear="1804" box="[330,458,1451,1477]" class="Mammalia" family="Hyperoodontidae" genus="Hyperoodon" kingdom="Animalia" order="Cetacea" pageId="40" pageNumber="601" phylum="Chordata" rank="genus">
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<emphasis id="EFD6EAC7C156FF8DFE8E77DCFE7400AB" box="[330,458,1451,1477]" italics="true" pageId="40" pageNumber="601">Hyperoodon</emphasis>
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</taxonomicName>
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spp. at the level of the maxillary crests are functionally rejected: the high maxillary crests of the latter do not form a basin. If the prenarial basin of
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<taxonomicName id="1AA24D56C156FF8DFDEB747CFF0E032A" authorityName="Cuvier" authorityYear="1823" class="Mammalia" family="Hyperoodontidae" genus="Ziphius" kingdom="Animalia" order="Cetacea" pageId="40" pageNumber="601" phylum="Chordata" rank="species" species="cavirostris">
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<emphasis id="EFD6EAC7C156FF8DFDEB747CFF0E032A" italics="true" pageId="40" pageNumber="601">Z. cavirostris</emphasis>
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</taxonomicName>
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is differently built, the supracranial basin of the physeterids bearing the spermaceti organ is relatively similar, especially in
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<taxonomicName id="1AA24D56C156FF8DFE14741DFDC103ED" authorityName="Linnaeus" authorityYear="1758" box="[464,639,1641,1667]" class="Mammalia" family="Physeteridae" genus="Physeter" kingdom="Animalia" order="Cetacea" pageId="40" pageNumber="601" phylum="Chordata" rank="species" species="macrocephalus">
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<emphasis id="EFD6EAC7C156FF8DFE14741DFDC103ED" box="[464,639,1641,1667]" italics="true" pageId="40" pageNumber="601">P. macrocephalus</emphasis>
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</taxonomicName>
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. Significant thickening of the vomer has not been described in physeterids until now and we therefore may not refer with certainty these relatively small specimens to the
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<taxonomicName id="1AA24D56C156FF8DFC7A7360FBE8045F" authorityName="J.E.Gray" authorityYear="1821" box="[958,1110,279,305]" class="Mammalia" family="Physeteridae" kingdom="Animalia" order="Cetacea" pageId="40" pageNumber="601" phylum="Chordata" rank="family">Physeteridae</taxonomicName>
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. On the other hand, their maxilla differs significantly from the known ziphiids. Future information about the bony nares and additional cranial characters would be crucial to solve the familial affinities of this strange whale.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |