treatments-xml/data/31/6F/87/316F87F3FFBBFFDAFF5FF896C911FEC0.xml

<document id="7A144EA44551D2BD29CA1D781BACC7F7" ID-DOI="10.5281/zenodo.175971" ID-GBIF-Dataset="5d326885-625b-4bb7-a91b-64a4e6679e1a" ID-ISSN="1175-5326" ID-Zenodo-Dep="175971" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459826017394" checkinUser="plazi" docAuthor="Shelley, Rowland M." docDate="2007" docId="316F87F3FFBBFFDAFF5FF896C911FEC0" docLanguage="en" docName="zt01440p068.pdf" docOrigin="Zootaxa 1440" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Arvechamboides ocala Shelley, 2007, new species" docType="treatment" docVersion="9" lastPageNumber="66" masterDocId="CD56FF8BFFB9FFDCFFC8FF8BCC5DFFBF" masterDocTitle="Arvechamboides ocala, n. gen., n. sp., a new aniulinine milliped from peninsular Florida, USA (Julida: Parajulidae)" masterLastPageNumber="68" masterPageNumber="61" pageNumber="63" updateTime="1698222254144" updateUser="plazi">
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<mods:title id="C19A2C4377B7381CA936A8036980A821">Arvechamboides ocala, n. gen., n. sp., a new aniulinine milliped from peninsular Florida, USA (Julida: Parajulidae)</mods:title>
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<mods:namePart id="0872A8C9E040D81E7BA6F3EBC7EFF2A5">Shelley, Rowland M.</mods:namePart>
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<mods:date id="4E00C181C0E6CEE336BCE74150B75A2A">2007</mods:date>
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<treatment id="316F87F3FFBBFFDAFF5FF896C911FEC0" ID-DOI="http://doi.org/10.5281/zenodo.6241125" ID-GBIF-Taxon="119356698" ID-Zenodo-Dep="6241125" LSID="urn:lsid:plazi:treatment:316F87F3FFBBFFDAFF5FF896C911FEC0" httpUri="http://treatment.plazi.org/id/316F87F3FFBBFFDAFF5FF896C911FEC0" lastPageId="6" lastPageNumber="66" pageId="2" pageNumber="63">
<subSubSection id="F1DC656EFFBBFFDEFF5FF896CD44F8E5" pageId="2" pageNumber="63" type="nomenclature">
<paragraph id="B97936E5FFBBFFDEFF5FF896CE1BF888" blockId="2.[151,582,1821,1882]" box="[151,582,1821,1847]" pageId="2" pageNumber="63">
<heading id="E2318189FFBBFFDEFF5FF896CE1BF888" bold="true" box="[151,582,1821,1847]" fontSize="11" level="1" pageId="2" pageNumber="63" reason="1">
<emphasis id="8BB2EAF7FFBBFFDEFF5FF896CE1BF888" bold="true" box="[151,582,1821,1847]" pageId="2" pageNumber="63">
<taxonomicName id="7EC64D66FFBBFFDEFF5FF896CDFAF888" ID-CoL="H3RX" box="[151,423,1821,1847]" class="Diplopoda" family="Parajulidae" genus="Arvechamboides" kingdom="Animalia" order="Julida" pageId="2" pageNumber="63" phylum="Arthropoda" rank="species" species="ocala" status="sp. nov.">
<emphasis id="8BB2EAF7FFBBFFDEFF5FF896CDFAF888" bold="true" box="[151,423,1821,1847]" italics="true" pageId="2" pageNumber="63">Arvechamboides ocala</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="9081578CFFBBFFDEFE7EF896CE1BF888" box="[438,582,1821,1847]" pageId="2" pageNumber="63" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B97936E5FFBBFFDEFF5FF8CBCD44F8E5" blockId="2.[151,582,1821,1882]" box="[151,281,1856,1882]" pageId="2" pageNumber="63">Figs. 2–11.</paragraph>
</subSubSection>
<subSubSection id="F1DC656EFFBBFFDFFF5FF804CFFFFF0D" lastPageId="3" lastPageNumber="64" pageId="2" pageNumber="63" type="materials_examined">
<paragraph id="B97936E5FFBBFFDFFF5FF804CFFFFF0D" blockId="2.[151,1436,1935,2002]" lastBlockId="3.[151,1437,152,2007]" lastPageId="3" lastPageNumber="64" pageId="2" pageNumber="63">
<emphasis id="8BB2EAF7FFBBFFDEFF5FF804CD0BF816" bold="true" box="[151,342,1935,1961]" pageId="2" pageNumber="63">
<typeStatus id="667D8847FFBBFFDEFF5FF804CC89F816" box="[151,212,1935,1961]" pageId="2" pageNumber="63">Type</typeStatus>
specimen.
</emphasis>
Male 
<typeStatus id="667D8847FFBBFFDEFE69F81BCE55F815" box="[417,520,1936,1962]" pageId="2" pageNumber="63" type="holotype">holotype</typeStatus>
(
<collectionCode id="DFD7AE20FFBBFFDEFDDEF81BCE3EF815" box="[534,611,1936,1962]" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="2" pageNumber="63">FSCA</collectionCode>
) collected by C. R. Smith, 
<date id="CD781025FFBBFFDEFC69F81BC839F815" box="[929,1124,1936,1962]" pageId="2" pageNumber="63" value="1977-01-11">11 January 1977</date>
, ca. 21.3 mi (34.1 km) NE 
<taxonomicName id="7EC64D66FFBBFFDEFF5FF833CC83F86D" box="[151,222,1976,2002]" class="Diplopoda" family="Parajulidae" genus="Arvechamboides" kingdom="Animalia" order="Julida" pageId="2" pageNumber="66" phylum="Arthropoda" rank="species" species="ocala">Ocala</taxonomicName>
, 
<quantity id="7E3E9B00FFBBFFDEFF25F833CD1EF86D" box="[237,323,1976,2002]" metricMagnitude="3" metricUnit="m" metricValue="4.02336" metricValueMax="4.828032" metricValueMin="3.218688" pageId="2" pageNumber="63" unit="mi" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mi</quantity>
(3.2–4.8 km) N Florida highway 316 &amp; 1.5–2.5 mi (2.4–4.0 km) E of the Oklawaha River (R24E T12S Sec. 35), 
<taxonomicName id="7EC64D66FFBAFFDFFE68FF13CDB9FF0D" box="[416,484,152,178]" class="Diplopoda" family="Parajulidae" genus="Arvechamboides" kingdom="Animalia" order="Julida" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="ocala">Ocala</taxonomicName>
National Forest, Marion Co., Florida.
</paragraph>
</subSubSection>
<subSubSection id="F1DC656EFFBAFFDFFF0EFF34CF8EFE4B" pageId="3" pageNumber="64" type="diagnosis">
<paragraph id="B97936E5FFBAFFDFFF0EFF34CE93FF65" blockId="3.[151,1437,152,2007]" box="[198,718,191,218]" pageId="3" pageNumber="64">
<emphasis id="8BB2EAF7FFBAFFDFFF0EFF34CD62FF66" bold="true" box="[198,319,191,217]" pageId="3" pageNumber="64">Diagnosis</emphasis>
. With the characters of the genus.
</paragraph>
<paragraph id="B97936E5FFBAFFDFFF0EFF6CCF8EFE4B" blockId="3.[151,1437,152,2007]" pageId="3" pageNumber="64">
<emphasis id="8BB2EAF7FFBAFFDFFF0EFF6CCD10FEBE" bold="true" box="[198,333,231,257]" pageId="3" pageNumber="64">Coloration</emphasis>
. Color in life unknown. Pleuroterga a bland light beige, substantially faded after 29 years in preservative, generally banded and darker on caudal margins with bands completely encircling sclerites, darker on collum and rings 1–8 with faint evidence of middorsal mottling; epiproct and paraprocts slightly darker; ozopores indistinct; legs uniformly light beige. Epicranium darker anteriad, darkest in interantennal region; frons, genae, clypeus, and labrum uniformly lighter; ocelli dark brown, caudalmost row depigmented. Left antenna broken at midlength; right antenna with 7 articles, becoming progressively darker through 6th antennomere, generally darker on distal halves, ultimate article lighter.
</paragraph>
</subSubSection>
<subSubSection id="F1DC656EFFBAFFD8FF0EFD89CD6FFE75" lastPageId="4" lastPageNumber="65" pageId="3" pageNumber="64" type="materials_examined">
<paragraph id="B97936E5FFBAFFDFFF0EFD89CEB1FDFB" blockId="3.[151,1437,152,2007]" pageId="3" pageNumber="64">
<emphasis id="8BB2EAF7FFBAFFDFFF0EFD89CD65FDA3" bold="true" box="[198,312,514,540]" pageId="3" pageNumber="64">
<typeStatus id="667D8847FFBAFFDFFF0EFD89CD65FDA3" box="[198,312,514,540]" pageId="3" pageNumber="64" type="holotype">Holotype</typeStatus>
</emphasis>
. Body relatively long and slender, parallel-sided; 50 rings including epiproct, last 3 rings legless; length ca. 32.3 mm, maximum width 2.1 mm.
</paragraph>
<paragraph id="B97936E5FFBAFFDFFF0EFDD9C86DFC84" blockId="3.[151,1437,152,2007]" pageId="3" pageNumber="64">Head smooth, polished; epicranium glossy. Ocellaria forming inverted triangle, with ca. 48 ocelli arranged in 8 curvilinear rows: 8, 9, 8, 7, 6, 5, 3, 2. Epicranial setae 1–1, clypeal 2–2; no other facial setae detected. Right antenna reaching back to caudal margin of 3rd pleurotergite, becoming progressively more setose distad, 1st antennomere subglobose, 2–5 strongly clavate, 6 subparallel-sided, 7 short and truncate with 4 conical sensory cones, no other sensory structures apparent; relative lengths of antennomeres 2=3&gt;4&gt;5&gt;6&gt;1&gt;7. Ventrodistal corner of mandibular stipes prolonged into broad, apically rounded lobe.</paragraph>
<paragraph id="B97936E5FFBAFFDFFF0EFCC2CEEBFBC2" blockId="3.[151,1437,152,2007]" pageId="3" pageNumber="64">Dorsum smooth, glabrous, &amp; glossy, wholly without setae including epiproct. Collum enlarged, slightly overlapping epicranium, lateral margins sublinear. Pleurotergites virtually identical excepting shorter ones at caudal extremity and 7th &amp; 8th, which are moderately swollen. Ventrolateral striae faint, noticeable on pleurotergites 3–47, extending only slightly dorsad along lateral surfaces, largely obscured by legs. Defensive glands arising on pleurotergite 6, continuing through ring 46. Epiproct (Fig. 2) long &amp; spiniform, overhanging &amp; extending slightly beyond caudal extremities of paraprocts; latter with margins slightly thickened and 2 moderately long submarginal setae arising from rim at 1/3 &amp; 2/3 lengths; hypoproct sublunar, slightly prolonged apically, with one moderately long apical seta.</paragraph>
<paragraph id="B97936E5FFBAFFDFFF0EFB05C941F92E" blockId="3.[151,1437,152,2007]" pageId="3" pageNumber="64">1st leg (Fig. 3) with 6 lightly hirsute podomeres, moderately enlarged and crassate, curving generally anteriomediad, with rounded to subconical &quot;sphaerotrichomes,&quot; some giving rise to hairs, scattered randomly on inner (medial) surfaces of tibia and tarsus, relative lengths of podomeres 5&gt;6&gt;4&gt;2&gt;3&gt;1; coxa narrow, broader mediad, prefemur overlapping medial surface of coxa and touching sternum, femur and postfemur short &amp; clavate, expanding primarily on lateral margins, tibia substantially broader than other podomeres, expanding primarily on lateral margin, tarsus basally broad, narrowing slightly apically, claw short, apically blunt, directed submediad. 2nd legs short &amp; narrow, incorporated into penial apparatus, with 6 articles, margins of ultimate article irregularly indented, forming &quot;pseudoarticles.&quot; Subsequent legs (Fig. 4) subsimilar, with 7 articles, becoming progressively more setose distad, claws shorter than &amp; obscured by overlying hairs of ultimate podomeres, apically acuminate. Penial apparatus (Figs. 5–6) bilobed, curving strongly anteriad at about a right angle, divided for entire length but lobes overlapping and closely appressed to each other, caudal surface with short, apically divided, basal projection angling dorsad and extending caudad to level of 4th sternum (between 3rd legs); 2nd legs extending directly ventrad, becoming progressively more hirsute distad.</paragraph>
<paragraph id="B97936E5FFBAFFD8FF0EF929CF5DFEC5" blockId="3.[151,1437,152,2007]" lastBlockId="4.[151,1437,152,498]" lastPageId="4" lastPageNumber="65" pageId="3" pageNumber="64">Condition of 7th pleurotergite not observed. Sternum of 8th pleurotergite (Figs. 7–8) narrow, extending dorsad well into body cavity, with narrow ventromedial ridge projecting anteriad as triangular lobe, sides elevated and enlarged into anteriorly-directed, dorsally semilunar lobes, extending over 7th pleurotergite and slightly over margin of aperture. Anterior gonopods (Fig. 9) dominated by broad sternum, extending ventrad between telopodites into elongated, triangular, unequally bilobed projection, only slighter shorter than lengths of telopodites. Posterior gonopod (Figs. 10–11) with prefemoral process arising basally on medial surface of prefemur, extending caudad basally with lobed inner margin, curving anterioventrad at 1/3 length caudal to telopodite and extending to around midlength of latter, apically acuminate with rounded, subterminal lobe on anterior (inner) margin; accessory prefemoral process arising distally on lateral surface of prefemur, broadly laminate but narrowing progressively distad to subacuminate tip, directed ventrad and extending to level of distal curvature of telopodite, positioned within curvature arc of latter; telopodite relatively narrow, curving broadly anteriad and narrowing progressively distad, overhanging and extending beyond level of accessory prefemoral process, curving sigmoidally apically to acuminate tip on outer corner; prostatic groove running along medial surface of telopodite to apical opening.</paragraph>
<paragraph id="B97936E5FFBDFFD8FF0EFE0CCD6FFE75" blockId="4.[151,1437,152,498]" pageId="4" pageNumber="65">
<emphasis id="8BB2EAF7FFBDFFD8FF0EFE0CCD75FE1E" bold="true" box="[198,296,391,417]" pageId="4" pageNumber="65">Ecology</emphasis>
. The milliped was recovered from a pitfall trap placed in 4-year-old sand pine (
<taxonomicName id="7EC64D66FFBDFFD8FB30FE03C9CCFE1E" box="[1272,1425,392,417]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="65" phylum="Tracheophyta" rank="species" species="clausa">
<emphasis id="8BB2EAF7FFBDFFD8FB30FE03C9CCFE1E" box="[1272,1425,392,417]" italics="true" pageId="4" pageNumber="65">Pinus clausa</emphasis>
</taxonomicName>
) scrub woods.
</paragraph>
</subSubSection>
<subSubSection id="F1DC656EFFBDFFD8FF0EFE5CCD91F876" pageId="4" pageNumber="65" type="distribution">
<paragraph id="B97936E5FFBDFFD8FF0EFE5CCF57FE4D" blockId="4.[151,1437,152,498]" box="[198,778,471,498]" pageId="4" pageNumber="65">
<emphasis id="8BB2EAF7FFBDFFD8FF0EFE5CCD02FE4E" bold="true" box="[198,351,471,497]" pageId="4" pageNumber="65">Distribution</emphasis>
. Known only from the 
<typeStatus id="667D8847FFBDFFD8FDBAFE53CEF9FE4D" box="[626,676,472,498]" pageId="4" pageNumber="65">type</typeStatus>
locality.
</paragraph>
<paragraph id="B97936E5FFBDFFD8FF5FF8B0CD91F876" blockId="4.[151,1437,1851,1993]" pageId="4" pageNumber="65">
<emphasis id="8BB2EAF7FFBDFFD8FF5FF8B0CD19F8EC" bold="true" box="[151,324,1851,1875]" pageId="4" pageNumber="65">FIGURES 2–6.</emphasis>
<taxonomicName id="7EC64D66FFBDFFD8FE83F8B0CE1DF8ED" box="[331,576,1851,1874]" class="Diplopoda" family="Parajulidae" genus="Arvechamboides" kingdom="Animalia" order="Julida" pageId="4" pageNumber="65" phylum="Arthropoda" rank="species" species="ocala">
<emphasis id="8BB2EAF7FFBDFFD8FE83F8B0CE1DF8ED" box="[331,576,1851,1874]" italics="true" pageId="4" pageNumber="65">Arvechamboides ocala</emphasis>
</taxonomicName>
, somatic features. 2, epiproct, left paraproct, and hypoproct, lateral view of left side. 3, right 1st leg, caudal view. 4, midbody leg on left side, caudal view, coxa omitted. 5, penial apparatus, lateral view of right side. 6, the same, ventral view, position of 2nd legs indicated. Scale line = 0.80 mm for figs. 2 &amp; 4, 0.67 mm for fig. 3, 0.50 mm for figs. 5–6.
</paragraph>
</subSubSection>
<subSubSection id="F1DC656EFFBCFFD9FF5FFA85CF0AFA31" pageId="5" pageNumber="66" type="description">
<paragraph id="B97936E5FFBCFFD9FF5FFA85CF0AFA31" blockId="5.[151,1436,1292,1422]" pageId="5" pageNumber="66">
<emphasis id="8BB2EAF7FFBCFFD9FF5FFA85CD12FA99" bold="true" box="[151,335,1294,1318]" pageId="5" pageNumber="66">FIGURES 7–11.</emphasis>
<taxonomicName id="7EC64D66FFBCFFD9FE9FFA84CE16FA99" box="[343,587,1295,1318]" class="Diplopoda" family="Parajulidae" genus="Arvechamboides" kingdom="Animalia" order="Julida" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="ocala">
<emphasis id="8BB2EAF7FFBCFFD9FE9FFA84CE16FA99" box="[343,587,1295,1318]" italics="true" pageId="5" pageNumber="66">Arvechamboides ocala</emphasis>
</taxonomicName>
, gonopodal features. 7, 8th sternum, ventral view. 8, the same, anterior view. 9, anterior gonopods, anterior view. 10, right posterior gonopod, medial view. 11, the same, lateral view. AT, anterior gonopod telopodite; APFP, accessory prefemoral process; PFP, prefemoral process; PT, posterior gonopod telopodite; S, sternum. Scale line = 1.00 mm for figs. 7–9, 0.83 mm for figs. 10–11.
</paragraph>
</subSubSection>
<subSubSection id="F1DC656EFFBCFFD9FF0EFA4AC867F9BB" pageId="5" pageNumber="66" type="etymology">
<paragraph id="B97936E5FFBCFFD9FF0EFA4AC867F9BB" blockId="5.[151,1437,1473,2028]" pageId="5" pageNumber="66">
<emphasis id="8BB2EAF7FFBCFFD9FF0EFA4ACD16FA64" bold="true" box="[198,331,1473,1499]" pageId="5" pageNumber="66">Etymology</emphasis>
. The specific name is a noun in apposition that references the 
<taxonomicName id="7EC64D66FFBCFFD9FBF6FA49C8DFFA63" box="[1086,1154,1474,1500]" class="Diplopoda" family="Parajulidae" genus="Arvechamboides" kingdom="Animalia" order="Julida" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="ocala">Ocala</taxonomicName>
National Forest, which harbors the only known locality of the species and lies east of the city of 
<taxonomicName id="7EC64D66FFBCFFD9FC26FA61C86BF9BB" box="[1006,1078,1514,1540]" class="Diplopoda" family="Parajulidae" genus="Arvechamboides" kingdom="Animalia" order="Julida" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="ocala">Ocala</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="F1DC656EFFBCFFDAFF0EF99AC911FEC0" lastPageId="6" lastPageNumber="67" pageId="5" pageNumber="66" type="discussion">
<paragraph id="B97936E5FFBCFFD9FF0EF99ACC85F89B" blockId="5.[151,1437,1473,2028]" pageId="5" pageNumber="66">
<emphasis id="8BB2EAF7FFBCFFD9FF0EF99ACD65F994" bold="true" box="[198,312,1553,1579]" pageId="5" pageNumber="66">Remarks</emphasis>
. Not anticipating a new species, much less a new genus, when I found this sample in the FSCA, I did not observe the 7th pleurotergite before dissecting the gonopods and destroying the integrity of this part of the exoskeleton. As 
<taxonomicName id="7EC64D66FFBCFFD9FE65F9EECF25F9C1" authority="Chamberlin, 1938" authorityName="Chamberlin" authorityYear="1938" box="[429,888,1636,1662]" class="Diplopoda" family="Parajulidae" genus="Gyniulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="bufonius">
<emphasis id="8BB2EAF7FFBCFFD9FE65F9EECEDBF9C1" box="[429,646,1637,1662]" italics="true" pageId="5" pageNumber="66">Gyniulus bufonius</emphasis>
(Chamberlin, 1938)
</taxonomicName>
is common in adjacent Alachua Co. (
<bibRefCitation id="DD574B14FFBCFFD9FA8AF9EFCCBDF919" author="Shelley" pageId="5" pageNumber="67" refString="Shelley, R. M. (2000 a) Parajulid studies III. The genus Gyniulus Loomis (Parajulinae: Aniulini). Myriapodologica, 7 (3), 19 - 28." type="journal article" year="2000" yearSuffix="a">
Shelley 2000 
<emphasis id="8BB2EAF7FFBCFFD9FF19F906CCBDF919" box="[209,224,1677,1702]" italics="true" pageId="5" pageNumber="66">a</emphasis>
</bibRefCitation>
, 2001), I expected the Marion Co. parajulid to be this species, and only after extracting the gonopods and seeing an unexpected new form did I notice the condition of the 8th sternum that would have at least suggested a new species of 
<taxonomicName id="7EC64D66FFBCFFD9FE73F968CE07F943" box="[443,602,1763,1788]" class="Diplopoda" family="Parajulidae" genus="Arvechambus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="8BB2EAF7FFBCFFD9FE73F968CE07F943" box="[443,602,1763,1788]" italics="true" pageId="5" pageNumber="66">Arvechambus</emphasis>
</taxonomicName>
. The 7th pleurotergite will thus need to be characterized from another male.
</paragraph>
<paragraph id="B97936E5FFBCFFDAFF0EF8B9C911FEC0" blockId="5.[151,1437,1473,2028]" lastBlockId="6.[151,1436,152,383]" lastPageId="6" lastPageNumber="67" pageId="5" pageNumber="66">
In studying this specimen, I observed the &quot;sphaerotrichomes&quot; (term apparently first employed by 
<bibRefCitation id="DD574B14FFBCFFD9FA93F8B9CD42F8CB" author="Hoffman" pageId="5" pageNumber="67" refString="Hoffman, R. L. (1974) Short studies on dalodesmid millipeds from South Africa and Madagascar (Diplopoda: Polydesmida). The Wasmann Journal of Biology, 32 (2), 221 - 246." type="journal article" year="1974">Hoffman (1974)</bibRefCitation>
for South African 
<taxonomicName id="7EC64D66FFBCFFD9FDC9F8D1CEF4F8CB" box="[513,681,1882,1908]" class="Diplopoda" family="Dalodesmidae" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="66" phylum="Arthropoda" rank="family">Dalodesmidae</taxonomicName>
(
<taxonomicName id="7EC64D66FFBCFFD9FD71F8D1CF09F8CB" box="[697,852,1882,1908]" class="Diplopoda" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="66" phylum="Arthropoda" rank="order">Polydesmida</taxonomicName>
)) on the medial surfaces of the tibiae and tarsi of the first legs. They are difficult to detect in direct medial view on legs that are pallid after years in preservative and are best seen in profile and with incident lighting. As I had not noticed these pustules before, I briefly checked 
<taxonomicName id="7EC64D66FFBCFFD9FF37F858CDAFF853" box="[255,498,2003,2028]" class="Diplopoda" family="Parajulidae" genus="Nesoressa" kingdom="Animalia" order="Julida" pageId="5" pageNumber="66" phylum="Arthropoda" rank="species" species="crawfordi">
<emphasis id="8BB2EAF7FFBCFFD9FF37F858CDAFF853" box="[255,498,2003,2028]" italics="true" pageId="5" pageNumber="66">Nesoressa crawfordi</emphasis>
</taxonomicName>
and representatives of the other aniulinine genera that I have studied (
<taxonomicName id="7EC64D66FFBCFFD9FAF3F858C9C8F853" box="[1339,1429,2003,2028]" class="Diplopoda" family="Parajulidae" genus="Aniulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="8BB2EAF7FFBCFFD9FAF3F858C9C8F853" box="[1339,1429,2003,2028]" italics="true" pageId="5" pageNumber="66">Aniulus</emphasis>
</taxonomicName>
, 
<taxonomicName id="7EC64D66FFBFFFDAFF5FFF13CD28FF0E" authority="(Hakiulus)" baseAuthorityName="Hakiulus" box="[151,373,152,177]" class="Diplopoda" family="Parajulidae" genus="Aniulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="8BB2EAF7FFBFFFDAFF5FFF13CD28FF0E" box="[151,373,152,177]" italics="true" pageId="6" pageNumber="67">Aniulus (Hakiulus)</emphasis>
</taxonomicName>
, 
<taxonomicName id="7EC64D66FFBFFFDAFE4CFF13CDB3FF0E" box="[388,494,152,177]" class="Diplopoda" family="Parajulidae" genus="Gyniulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="8BB2EAF7FFBFFFDAFE4CFF13CDB3FF0E" box="[388,494,152,177]" italics="true" pageId="6" pageNumber="67">Gyniulus</emphasis>
</taxonomicName>
, 
<taxonomicName id="7EC64D66FFBFFFDAFE34FF13CE08FF0E" box="[508,597,152,177]" class="Diplopoda" family="Parajulidae" genus="Oriulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="8BB2EAF7FFBFFFDAFE34FF13CE08FF0E" box="[508,597,152,177]" italics="true" pageId="6" pageNumber="67">Oriulus</emphasis>
</taxonomicName>
, 
<taxonomicName id="7EC64D66FFBFFFDAFDACFF13CEAEFF0E" box="[612,755,152,177]" class="Diplopoda" family="Parajulidae" genus="Pseudojulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="8BB2EAF7FFBFFFDAFDACFF13CEAEFF0E" box="[612,755,152,177]" italics="true" pageId="6" pageNumber="67">Pseudojulus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="7EC64D66FFBFFFDAFCFBFF13CF8FFF0E" box="[819,978,152,177]" class="Diplopoda" family="Parajulidae" genus="Arvechambus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="67" phylum="Arthropoda" rank="genus">
<emphasis id="8BB2EAF7FFBFFFDAFCFBFF13CF8FFF0E" box="[819,978,152,177]" italics="true" pageId="6" pageNumber="67">Arvechambus</emphasis>
</taxonomicName>
(
<bibRefCitation id="DD574B14FFBFFFDAFC2AFF13C8D1FF0E" author="Shelley" box="[994,1164,152,178]" pageId="6" pageNumber="67" refString="Shelley, R. M. (2000 a) Parajulid studies III. The genus Gyniulus Loomis (Parajulinae: Aniulini). Myriapodologica, 7 (3), 19 - 28." type="journal article" year="2000" yearSuffix="a">
Shelley 2000 
<emphasis id="8BB2EAF7FFBFFFDAFBB5FF13C8D1FF0E" box="[1149,1164,152,177]" italics="true" pageId="6" pageNumber="67">a</emphasis>
</bibRefCitation>
, 
<bibRefCitation id="DD574B14FFBFFFDAFB53FF13C8F7FF0E" author="Shelley" box="[1179,1194,152,177]" pageId="6" pageNumber="67" refString="Shelley, R. M. (2000 b) Parajulid studies II. The subgenus Hakiulus Chamberlin (Julida: Parajulidae: Parajulinae: Aniulini). Myriapodologica, 6 (14), 121 - 145." type="journal article" year="2000" yearSuffix="b">
<emphasis id="8BB2EAF7FFBFFFDAFB53FF13C8F7FF0E" box="[1179,1194,152,177]" italics="true" pageId="6" pageNumber="67">b</emphasis>
</bibRefCitation>
, 
<bibRefCitation id="DD574B14FFBFFFDAFB70FF13C95FFF0E" author="Shelley" box="[1208,1282,152,178]" pageId="6" pageNumber="67" refString="Shelley, R. M. (2001 b) A synopsis of the milliped genus Aniulus Chamberlin (Julida: Parajulidae: Parajulinae: Aniulini). Texas Memorial Museum, Speleological Monographs, 5, 73 - 94." type="journal article" year="2001" yearSuffix="b">
2001 
<emphasis id="8BB2EAF7FFBFFFDAFB3BFF13C95FFF0E" box="[1267,1282,152,177]" italics="true" pageId="6" pageNumber="67">b</emphasis>
</bibRefCitation>
, 
<bibRefCitation id="DD574B14FFBFFFDAFAD8FF13C910FF0D" author="Shelley" box="[1296,1357,152,178]" pageId="6" pageNumber="67" refString="Shelley, R. M. (2002) The milliped genus Oriulus Chamberlin (Julida: Parajulidae). Canadian Journal of Zoology, 80, 100 - 109." type="journal article" year="2002">2002</bibRefCitation>
, 
<bibRefCitation id="DD574B14FFBFFFDAFA91FF13C9CAFF0D" author="Shelley" box="[1369,1431,152,178]" lastPageId="6" lastPageNumber="68" pageId="6" pageNumber="67" refString="Shelley, R. M. (2004 [2002]) Parajulid milliped studies V. The genera Pseudojulus Bollman and Arvechambus Causey (Parajulinae: Aniulini). Insecta Mundi, 16 (4), 191 - 204. 1" type="journal article" year="2004">2004</bibRefCitation>
; 
<bibRefCitation id="DD574B14FFBFFFDAFF5FFF4BCD9CFF65" author="Shelley" box="[151,449,192,218]" pageId="6" pageNumber="68" refString="Shelley, R. M., &amp; Medrano, M. F. (2006) Nesoressa crawfordi, n. gen., n. sp., a montane island milliped in New Mexico, USA; proposal of the new tribe Nesoressini and a preliminary cladogram of the lineage &quot; Aniulina &quot; (Julida: Parajulidae). Zootaxa, 1285, 31 - 50." type="journal article" year="2006">Shelley &amp; Medrano 2006</bibRefCitation>
)) and found that they are present in all of them although their sizes and the densities of the &quot;fields&quot; vary. To the best of my knowledge, this constitutes only the third report of &quot;sphaerotrichomes&quot; in the 
<taxonomicName id="7EC64D66FFBFFFDAFF2AFE9BCD34FE95" box="[226,361,272,298]" class="Diplopoda" family="Parajulidae" kingdom="Animalia" order="Julida" pageId="6" pageNumber="67" phylum="Arthropoda" rank="family">Parajulidae</taxonomicName>
, the others being by 
<bibRefCitation id="DD574B14FFBFFFDAFDA8FE9BCF33FE95" author="Verhoeff" box="[608,878,272,298]" pageId="6" pageNumber="68" refString="Verhoeff, K. W. (1926) Chilognathen-Beitrage. Zoologischer Anzeiger, 68 (1 / 2), 57 - 71." type="journal article" year="1926">Verhoeff (1926, fig. 4)</bibRefCitation>
, who called them &quot;Rundhöckerchen&quot; and illustrated them on the 1st leg of 
<taxonomicName id="7EC64D66FFBFFFDAFE3AFEB0CFC1FEEB" authority="Verhoeff, 1926" authorityName="Verhoeff" authorityYear="1926" box="[498,924,314,340]" class="Diplopoda" family="Parajulidae" genus="Mexicoiulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="dampfi">
<emphasis id="8BB2EAF7FFBFFFDAFE3AFEB0CE80FEEB" box="[498,733,315,340]" italics="true" pageId="6" pageNumber="67">Mexicoiulus dampfi</emphasis>
<bibRefCitation id="DD574B14FFBFFFDAFD2FFEB1CFC1FEEB" author="Verhoeff" box="[743,924,314,340]" pageId="6" pageNumber="68" refString="Verhoeff, K. W. (1926) Chilognathen-Beitrage. Zoologischer Anzeiger, 68 (1 / 2), 57 - 71." type="journal article" year="1926">Verhoeff, 1926</bibRefCitation>
</taxonomicName>
, and 
<bibRefCitation id="DD574B14FFBFFFDAFC28FEB1C8B1FEEB" author="Mauries" box="[992,1260,314,340]" pageId="6" pageNumber="67" refString="Mauries, J. - P. (1972) Nouvelle description de Paraiulus olmecus Humbert et Saussure, 1869, espece-type du genre Paraiulus (Diplopoda) conservee au Museum d'Histoire naturelle de Geneve. Revus Suisse de Zoologie, 79 (1), 159 - 165." type="journal article" year="1972">Mauriès (1972, fig. 3)</bibRefCitation>
, who depicted them on the 1st legs of 
<taxonomicName id="7EC64D66FFBFFFDAFE57FEEDCF9EFEC0" authority="Humbert &amp; Saussure, 1869" authorityName="Humbert &amp; Saussure" authorityYear="1869" box="[415,963,357,383]" class="Diplopoda" family="Parajulidae" genus="Parajulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="67" phylum="Arthropoda" rank="species" species="olmecus">
<emphasis id="8BB2EAF7FFBFFFDAFE57FEEDCE27FEC0" box="[415,634,358,383]" italics="true" pageId="6" pageNumber="67">Parajulus olmecus</emphasis>
Humbert &amp; Saussure, 1869
</taxonomicName>
, and termed them &quot;granulations.&quot;
</paragraph>
</subSubSection>
</treatment>
</document>