202 lines
36 KiB
XML
202 lines
36 KiB
XML
<document id="308443216876681599B11E341AB92386" ID-DOI="10.11646/zootaxa.4038.1.1" ID-GBIF-Dataset="066e84f1-206b-40cb-a615-2126696b813d" ID-ISSN="1175-5326" ID-Zenodo-Dep="289876" ID-ZooBank="1B62B78C-AA59-4417-A4FC-1CC9CED745E0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461316934850" checkinUser="plazi" docAuthor="Janák, Jiří & Bordoni, Arnaldo" docDate="2015" docId="039D87F5553EFFC7FF4AF928FCDFB8BD" docLanguage="en" docName="zt04038p094.pdf" docOrigin="Zootaxa 4038 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Afrus Janák & Bordoni, 2015, gen. nov." docType="treatment" docVersion="7" lastPageNumber="7" masterDocId="FFA4FF8D553BFFC1FFDDFFA6FFFFBE25" masterDocTitle="Revision of the genus Thyreocephalus and description of Afrus gen. nov. of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="94" masterPageNumber="1" pageNumber="6" updateTime="1698618126966" updateUser="plazi">
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<mods:titleInfo id="1B6EFA2CD56A7BB3DF42D9A1450A32E9">
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<mods:title id="11A09B670006DD53F3B0495759955E6B">Revision of the genus Thyreocephalus and description of Afrus gen. nov. of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:namePart id="661D58A5310813EA763155C4D78722E9">Janák, Jiří</mods:namePart>
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<mods:roleTerm id="EB04E7228889A30AB2564AD83C34D796">Author</mods:roleTerm>
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<mods:namePart id="23C4A2E03C5CB70DBE24469BD7D8D87D">Bordoni, Arnaldo</mods:namePart>
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<mods:typeOfResource id="601CB59A49736AA263F5AEC3B28E6280">text</mods:typeOfResource>
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<mods:title id="3B2F323DF825E1DA3CC6B14D4F9B6CC6">Zootaxa</mods:title>
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<mods:part id="7DD3E88E06999995E1519D62BD3D188B">
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<mods:date id="BF56F2C29B519D6024A7E49FDC9D506E">2015</mods:date>
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<mods:detail id="B3980618469DF32B2C34C1D259A4F0F5" type="volume">
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<mods:number id="874DCEBE40502AE83F6BFD13D9A42210">4038</mods:number>
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<mods:detail id="7E76F1453160A37206E7BBC1D89434C5" type="issue">
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<mods:number id="9E618660B307EC06BD2D39DA6653CF55">1</mods:number>
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<mods:classification id="1F9C34D7BC992094F54D3FB7BC70F494">journal article</mods:classification>
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<mods:identifier id="401C8E9A3178C142A810D03C0C12A820" type="DOI">10.11646/zootaxa.4038.1.1</mods:identifier>
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<mods:identifier id="FCC621DB5A29CAD8C7FFBE8F739603C0" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="30CA15C33D934C788947DCEAEF58DC49" type="Zenodo-Dep">289876</mods:identifier>
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<treatment id="039D87F5553EFFC7FF4AF928FCDFB8BD" ID-DOI="http://doi.org/10.5281/zenodo.6101599" ID-GBIF-Taxon="127693936" ID-Zenodo-Dep="6101599" LSID="urn:lsid:plazi:treatment:039D87F5553EFFC7FF4AF928FCDFB8BD" httpUri="http://treatment.plazi.org/id/039D87F5553EFFC7FF4AF928FCDFB8BD" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
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<subSubSection id="C32E6568553EFFC4FF4AF928FEAFB88D" box="[151,336,1678,1704]" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph id="8B8B36E3553EFFC4FF4AF928FEAFB88D" blockId="5.[151,336,1678,1704]" box="[151,336,1678,1704]" pageId="5" pageNumber="6">
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<heading id="D0C3818F553EFFC4FF4AF928FEAFB88D" bold="true" box="[151,336,1678,1704]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
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<emphasis id="B940EAF1553EFFC4FF4AF928FEAFB88D" bold="true" box="[151,336,1678,1704]" pageId="5" pageNumber="6">
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<taxonomicName id="4C344D60553EFFC4FF4AF928FF24B88D" ID-CoL="8XJDB" box="[151,219,1678,1704]" class="Insecta" family="Staphylinidae" genus="Afrus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B940EAF1553EFFC4FF4AF928FF24B88D" bold="true" box="[151,219,1678,1704]" italics="true" pageId="5" pageNumber="6">Afrus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A273578A553EFFC4FF3EF928FEAFB88D" box="[227,336,1678,1704]" pageId="5" pageNumber="6" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E6568553EFFC4FF4AF973FD0EB8CE" box="[151,753,1749,1771]" pageId="5" pageNumber="6" type="materials_examined">
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<paragraph id="8B8B36E3553EFFC4FF4AF973FD0EB8CE" blockId="5.[151,753,1749,1771]" box="[151,753,1749,1771]" pageId="5" pageNumber="6">
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<emphasis id="B940EAF1553EFFC4FF4AF973FED9B8CE" bold="true" box="[151,294,1749,1771]" pageId="5" pageNumber="6">
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<typeStatus id="548F8841553EFFC4FF4AF973FF33B8CE" box="[151,204,1749,1771]" pageId="5" pageNumber="6">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C344D60553EFFC4FEF6F973FD0EB8CE" authority="Bernhauer, 1915" authorityName="Bernhauer" authorityYear="1915" box="[299,753,1749,1771]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="spegazzinii">
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<emphasis id="B940EAF1553EFFC4FEF6F973FDBCB8CE" box="[299,579,1749,1771]" italics="true" pageId="5" pageNumber="6">Thyreocephalus spegazzinii</emphasis>
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Bernhauer, 1915
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E6568553EFFC7FF4AF8B1FCDFB8BD" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="description">
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<paragraph id="8B8B36E3553EFFC4FF4AF8B1FC8EB915" blockId="5.[151,1436,1815,2021]" box="[151,881,1815,1840]" pageId="5" pageNumber="6">
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<emphasis id="B940EAF1553EFFC4FF4AF8B1FED6B915" bold="true" box="[151,297,1815,1840]" pageId="5" pageNumber="6">Description.</emphasis>
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Form elongate, stout, of medium size (
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<quantity id="4CCC9B06553EFFC4FD02F8B1FC9AB915" box="[735,869,1815,1840]" metricMagnitude="-3" metricUnit="m" metricValue="8.25" metricValueMax="10.0" metricValueMin="6.5" pageId="5" pageNumber="6" unit="mm" value="8.25" valueMax="10.0" valueMin="6.5">6.5–10 mm</quantity>
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).
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</paragraph>
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<paragraph id="8B8B36E3553EFFC7FF1AF89AFE4FBFF5" blockId="5.[151,1436,1815,2021]" lastBlockId="6.[151,1437,151,1688]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
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Head variably punctate, punctures not forming semi-impressed rows of coalescent grooves, frontal grooves short or moderately long, ocular grooves inconspicuous or absent (
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<figureCitation id="130F2A66553EFFC4FC48F8F9FBFBB95D" box="[917,1028,1887,1912]" captionStart="FIGURES 3 – 7. 3, 4" captionStartId="8.[151,264,1877,1899]" captionTargetBox="[169,1423,213,1814]" captionTargetId="figure@8.[150,1436,193,1828]" captionTargetPageId="8" captionText="FIGURES 3 – 7. 3, 4, Afrus collarti (Cameron); 5 – 7, Afrus spegazzinii (Bernhauer). 3, 5, head and anterior part of pronotum (iop—internal ocular puncture, pop—posterior ocular punctures); 4, 7, lateral part of head (SG—subelliptical groove); 6, lateral part of ventral side of pronotum (SL—superior line, IL—inferior line). Scale 0.5 mm." httpUri="https://zenodo.org/record/289878/files/figure.png" pageId="5" pageNumber="6">Figs. 3, 5</figureCitation>
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). Anterior margin of frons between antennal insertions extended into short and very wide, apically sligthly rounded process, limited at each side by rounded emargination, and slightly impressed dorsally. Eyes small, head with large impression behind the eyes, temples longer than length of eyes seen from above, evenly rounded, laterally with evident subelliptical groove (
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<figureCitation id="130F2A66553DFFC7FF42FF31FEF6BE95" box="[159,265,151,176]" captionStart="FIGURES 3 – 7. 3, 4" captionStartId="8.[151,264,1877,1899]" captionTargetBox="[169,1423,213,1814]" captionTargetId="figure@8.[150,1436,193,1828]" captionTargetPageId="8" captionText="FIGURES 3 – 7. 3, 4, Afrus collarti (Cameron); 5 – 7, Afrus spegazzinii (Bernhauer). 3, 5, head and anterior part of pronotum (iop—internal ocular puncture, pop—posterior ocular punctures); 4, 7, lateral part of head (SG—subelliptical groove); 6, lateral part of ventral side of pronotum (SL—superior line, IL—inferior line). Scale 0.5 mm." httpUri="https://zenodo.org/record/289878/files/figure.png" pageId="6" pageNumber="7">Figs. 4, 7</figureCitation>
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). Antennae geniculate, moderately short, antennal insertions separated from each other by distance about equal to distance separating each insertion from anteromedian margin of eye (
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<figureCitation id="130F2A66553DFFC7FBEEFF1AFB59BEF0" box="[1075,1190,188,213]" captionStart="FIGURES 3 – 7. 3, 4" captionStartId="8.[151,264,1877,1899]" captionTargetBox="[169,1423,213,1814]" captionTargetId="figure@8.[150,1436,193,1828]" captionTargetPageId="8" captionText="FIGURES 3 – 7. 3, 4, Afrus collarti (Cameron); 5 – 7, Afrus spegazzinii (Bernhauer). 3, 5, head and anterior part of pronotum (iop—internal ocular puncture, pop—posterior ocular punctures); 4, 7, lateral part of head (SG—subelliptical groove); 6, lateral part of ventral side of pronotum (SL—superior line, IL—inferior line). Scale 0.5 mm." httpUri="https://zenodo.org/record/289878/files/figure.png" pageId="6" pageNumber="7">Figs. 3, 5</figureCitation>
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), first segment long, thickened towards apex, equal in length to at least the four following segments combined, second segment shorter than third, distal segments more or less transverse (
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<figureCitation id="130F2A66553DFFC7FD36FEA2FCA2BF39" box="[747,861,260,285]" captionStart="FIGURES 1 – 2. 1" captionStartId="7.[151,264,1873,1895]" captionTargetBox="[197,1414,210,1817]" captionTargetId="figure@7.[164,1439,195,1851]" captionTargetPageId="7" captionText="FIGURES 1 – 2. 1, Afrus collarti (Cameron); 2, Afrus spegazzinii (Bernhauer). Scale 1 mm." httpUri="https://zenodo.org/record/289877/files/figure.png" pageId="6" pageNumber="7">Figs. 1, 2</figureCitation>
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). Labrum completely chitinised, short, transverse, lobate apically, with long and strong apical setae (
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<figureCitation id="130F2A66553DFFC7FD0AFE81FCC9BF65" box="[727,822,295,320]" captionStart="FIGURES 9 – 15" captionStartId="10.[151,264,1914,1936]" captionTargetBox="[160,1413,355,1887]" captionTargetId="figure@10.[151,1436,337,1893]" captionTargetPageId="10" captionText="FIGURES 9 – 15. Afrus spegazzinii (Bernhauer); 9, head and pronotum; 10, labrum; 11, male tergite 10; 12, male sternite 9; 13, aedeagus; 14, female genital segment (fp—fused pleurites, s—sternite, ss—supplementary sclerites); 15, metatibia (ac—apical ctenidium)." httpUri="https://zenodo.org/record/289880/files/figure.png" pageId="6" pageNumber="7">Figs. 10</figureCitation>
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,
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<figureCitation id="130F2A66553DFFC7FC99FE8EFC9CBF65" box="[836,867,296,320]" captionStart="FIGURES 16 – 21" captionStartId="12.[151,264,1911,1933]" captionTargetBox="[166,1426,349,1883]" captionTargetId="figure@12.[151,1436,337,1890]" captionTargetPageId="12" captionText="FIGURES 16 – 21. Afrus collarti (Cameron); 16, head and pronotum; 17, labrum; 18, male tergite 10; 19, male sternite 9; 20, aedeagus; 21 female genital segment." httpUri="https://zenodo.org/record/289881/files/figure.png" pageId="6" pageNumber="7">17</figureCitation>
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). Mandible stout, lateral furrow reduced to vague impression at base (
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<figureCitation id="130F2A66553DFFC7FEA4FEEAFDFBBF41" box="[377,516,332,357]" captionStart="FIGURES 3 – 7. 3, 4" captionStartId="8.[151,264,1877,1899]" captionTargetBox="[169,1423,213,1814]" captionTargetId="figure@8.[150,1436,193,1828]" captionTargetPageId="8" captionText="FIGURES 3 – 7. 3, 4, Afrus collarti (Cameron); 5 – 7, Afrus spegazzinii (Bernhauer). 3, 5, head and anterior part of pronotum (iop—internal ocular puncture, pop—posterior ocular punctures); 4, 7, lateral part of head (SG—subelliptical groove); 6, lateral part of ventral side of pronotum (SL—superior line, IL—inferior line). Scale 0.5 mm." httpUri="https://zenodo.org/record/289878/files/figure.png" pageId="6" pageNumber="7">Figs. 3–5, 7</figureCitation>
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). Maxillary palpi moderately long, segment 3 shorter than segment 2, segment 4 longer and somewhat narrower than segment 3, subacute apically (
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<figureCitation id="130F2A66553DFFC7FC70FEC9FC09BFAD" box="[941,1014,367,392]" captionStart="FIGURES 3 – 7. 3, 4" captionStartId="8.[151,264,1877,1899]" captionTargetBox="[169,1423,213,1814]" captionTargetId="figure@8.[150,1436,193,1828]" captionTargetPageId="8" captionText="FIGURES 3 – 7. 3, 4, Afrus collarti (Cameron); 5 – 7, Afrus spegazzinii (Bernhauer). 3, 5, head and anterior part of pronotum (iop—internal ocular puncture, pop—posterior ocular punctures); 4, 7, lateral part of head (SG—subelliptical groove); 6, lateral part of ventral side of pronotum (SL—superior line, IL—inferior line). Scale 0.5 mm." httpUri="https://zenodo.org/record/289878/files/figure.png" pageId="6" pageNumber="7">Fig. 5</figureCitation>
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). Labial palpi moderately long, last segment distinctly longer than segment 2. Mentum short, transverse, pentagonal. Ligula divided. Gula very short, gular sutures contiguous.
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</paragraph>
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<paragraph id="8B8B36E3553DFFC7FF1AFE7BFA8CBC8D" blockId="6.[151,1437,151,1688]" pageId="6" pageNumber="7">
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Pronotum without dorsal rows of punctures, with one big puncture (without seta) close to anterior angles (
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<figureCitation id="130F2A66553DFFC7FAB9FE7AFF20BC3D" captionStart-0="FIGURES 1 – 2. 1" captionStart-1="FIGURES 3 – 7. 3, 4" captionStartId-0="7.[151,264,1873,1895]" captionStartId-1="8.[151,264,1877,1899]" captionTargetBox-0="[197,1414,210,1817]" captionTargetBox-1="[169,1423,213,1814]" captionTargetId-0="figure@7.[164,1439,195,1851]" captionTargetId-1="figure@8.[150,1436,193,1828]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 1 – 2. 1, Afrus collarti (Cameron); 2, Afrus spegazzinii (Bernhauer). Scale 1 mm." captionText-1="FIGURES 3 – 7. 3, 4, Afrus collarti (Cameron); 5 – 7, Afrus spegazzinii (Bernhauer). 3, 5, head and anterior part of pronotum (iop—internal ocular puncture, pop—posterior ocular punctures); 4, 7, lateral part of head (SG—subelliptical groove); 6, lateral part of ventral side of pronotum (SL—superior line, IL—inferior line). Scale 0.5 mm." httpUri-0="https://zenodo.org/record/289877/files/figure.png" httpUri-1="https://zenodo.org/record/289878/files/figure.png" pageId="6" pageNumber="7">Figs. 1–3, 5</figureCitation>
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); antersternal plate with a suture (similar as in
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<figureCitation id="130F2A66553DFFC7FD31FE59FCC0BC3D" box="[748,831,511,536]" captionStart="FIGURES 24 – 28. 24" captionStartId="15.[151,264,1802,1824]" captionTargetBox="[152,1422,209,1768]" captionTargetId="figure@15.[150,1436,193,1781]" captionTargetPageId="15" captionText="FIGURES 24 – 28. 24, Thyreocephalus brunneiventris (Tottenham); 25, 27, T. alluaudi (Fauvel); 26, T. interocularis (Eppelsheim); 28, T. ferox (Harold); 24, 25, head and anterior part of pronotum; 26, ventral side of pronotum (ap—antesternal plate, pr—prosternum, ip—intercoxal process, SL—superior line, IL—inferior line); 27, 28, metatibia (ac—apical ctenidium, sac—subapical ctenidium). Scale 0.5 mm." httpUri="https://zenodo.org/record/289883/files/figure.png" pageId="6" pageNumber="7">Fig. 26</figureCitation>
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); both superior and inferior line of pronotal hypomera strongly developed, superior line turning downwards well before middle, joining or almost joining inferior line next to front margin of procoxae and continuing onto front margin of pronotum (
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<figureCitation id="130F2A66553DFFC7FB8CFDE1FB66BC45" box="[1105,1177,583,608]" captionStart="FIGURES 3 – 7. 3, 4" captionStartId="8.[151,264,1877,1899]" captionTargetBox="[169,1423,213,1814]" captionTargetId="figure@8.[150,1436,193,1828]" captionTargetPageId="8" captionText="FIGURES 3 – 7. 3, 4, Afrus collarti (Cameron); 5 – 7, Afrus spegazzinii (Bernhauer). 3, 5, head and anterior part of pronotum (iop—internal ocular puncture, pop—posterior ocular punctures); 4, 7, lateral part of head (SG—subelliptical groove); 6, lateral part of ventral side of pronotum (SL—superior line, IL—inferior line). Scale 0.5 mm." httpUri="https://zenodo.org/record/289878/files/figure.png" pageId="6" pageNumber="7">Fig. 6</figureCitation>
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). Prosternum elevated medioposteriorly (similar as in
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<figureCitation id="130F2A66553DFFC7FE22FDCAFDACBCA0" box="[511,595,620,645]" captionStart="FIGURES 24 – 28. 24" captionStartId="15.[151,264,1802,1824]" captionTargetBox="[152,1422,209,1768]" captionTargetId="figure@15.[150,1436,193,1781]" captionTargetPageId="15" captionText="FIGURES 24 – 28. 24, Thyreocephalus brunneiventris (Tottenham); 25, 27, T. alluaudi (Fauvel); 26, T. interocularis (Eppelsheim); 28, T. ferox (Harold); 24, 25, head and anterior part of pronotum; 26, ventral side of pronotum (ap—antesternal plate, pr—prosternum, ip—intercoxal process, SL—superior line, IL—inferior line); 27, 28, metatibia (ac—apical ctenidium, sac—subapical ctenidium). Scale 0.5 mm." httpUri="https://zenodo.org/record/289883/files/figure.png" pageId="6" pageNumber="7">Fig. 26</figureCitation>
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), without posteromedian carina, intercoxal process protruding, triangular; epimera present. Mesosternum very short, transverse, widely separating middle coxae. Metasternum very long.
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</paragraph>
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<paragraph id="8B8B36E3553DFFC7FF1AFD12FB0ABD78" blockId="6.[151,1437,151,1688]" pageId="6" pageNumber="7">
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Elytra overlapping at suture (
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<figureCitation id="130F2A66553DFFC7FDCDFD12FD82BCE9" box="[528,637,692,717]" captionStart="FIGURES 1 – 2. 1" captionStartId="7.[151,264,1873,1895]" captionTargetBox="[197,1414,210,1817]" captionTargetId="figure@7.[164,1439,195,1851]" captionTargetPageId="7" captionText="FIGURES 1 – 2. 1, Afrus collarti (Cameron); 2, Afrus spegazzinii (Bernhauer). Scale 1 mm." httpUri="https://zenodo.org/record/289877/files/figure.png" pageId="6" pageNumber="7">Figs. 1, 2</figureCitation>
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). Legs moderately long; protarsi simple (not dilated) in either sex, first four segments gradually becoming shorter, last segment about as long as three preceding segments combined; protibiae with numerous strong spines on outer margin; mesotibiae strongly spinose, with only apical ctenidium; first segment of mesotarsus and metatarsus about equally long as second, last segment about as long as preceding three segments combined; metatibiae spinose on outer margin, with only apical ctenidium (
|
||
<figureCitation id="130F2A66553DFFC7FB49FCE2FB18BD78" box="[1172,1255,836,861]" captionStart="FIGURES 9 – 15" captionStartId="10.[151,264,1914,1936]" captionTargetBox="[160,1413,355,1887]" captionTargetId="figure@10.[151,1436,337,1893]" captionTargetPageId="10" captionText="FIGURES 9 – 15. Afrus spegazzinii (Bernhauer); 9, head and pronotum; 10, labrum; 11, male tergite 10; 12, male sternite 9; 13, aedeagus; 14, female genital segment (fp—fused pleurites, s—sternite, ss—supplementary sclerites); 15, metatibia (ac—apical ctenidium)." httpUri="https://zenodo.org/record/289880/files/figure.png" pageId="6" pageNumber="7">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8B36E3553DFFC7FF1AFCC1FCA5BD81" blockId="6.[151,1437,151,1688]" pageId="6" pageNumber="7">
|
||
Abdomen (
|
||
<figureCitation id="130F2A66553DFFC7FE9AFCC1FE52BDA5" box="[327,429,871,896]" captionStart="FIGURES 1 – 2. 1" captionStartId="7.[151,264,1873,1895]" captionTargetBox="[197,1414,210,1817]" captionTargetId="figure@7.[164,1439,195,1851]" captionTargetPageId="7" captionText="FIGURES 1 – 2. 1, Afrus collarti (Cameron); 2, Afrus spegazzinii (Bernhauer). Scale 1 mm." httpUri="https://zenodo.org/record/289877/files/figure.png" pageId="6" pageNumber="7">Fig. 1, 2</figureCitation>
|
||
) ‘Staphylininae-shaped’ with fourth or fifth segment broadest. Tergite 7 with complete membranous palisade fringe at its posterior margin (
|
||
<figureCitation id="130F2A66553DFFC7FD3DFC2AFCB2BD81" box="[736,845,908,933]" captionStart="FIGURES 1 – 2. 1" captionStartId="7.[151,264,1873,1895]" captionTargetBox="[197,1414,210,1817]" captionTargetId="figure@7.[164,1439,195,1851]" captionTargetPageId="7" captionText="FIGURES 1 – 2. 1, Afrus collarti (Cameron); 2, Afrus spegazzinii (Bernhauer). Scale 1 mm." httpUri="https://zenodo.org/record/289877/files/figure.png" pageId="6" pageNumber="7">Figs. 1, 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8B36E3553DFFC7FF1AFC09FD02BA7D" blockId="6.[151,1437,151,1688]" pageId="6" pageNumber="7">
|
||
<collectingCountry id="F3237673553DFFC7FF1AFC09FEFBBDED" box="[199,260,943,968]" name="Malta" pageId="6" pageNumber="7">Male</collectingCountry>
|
||
. Tergite and sternite 8 of male simple, not modified. Tergite 10 of male genital segment rather narrowly exposed between sclerites of tergite 9, strongly narrowed proximally (
|
||
<figureCitation id="130F2A66553DFFC7FC79FC72FBFFBDC9" box="[932,1024,980,1005]" captionStart="FIGURES 9 – 15" captionStartId="10.[151,264,1914,1936]" captionTargetBox="[160,1413,355,1887]" captionTargetId="figure@10.[151,1436,337,1893]" captionTargetPageId="10" captionText="FIGURES 9 – 15. Afrus spegazzinii (Bernhauer); 9, head and pronotum; 10, labrum; 11, male tergite 10; 12, male sternite 9; 13, aedeagus; 14, female genital segment (fp—fused pleurites, s—sternite, ss—supplementary sclerites); 15, metatibia (ac—apical ctenidium)." httpUri="https://zenodo.org/record/289880/files/figure.png" pageId="6" pageNumber="7">Figs. 11</figureCitation>
|
||
,
|
||
<figureCitation id="130F2A66553DFFC7FBD6FC72FBD6BDC9" box="[1035,1065,980,1004]" captionStart="FIGURES 16 – 21" captionStartId="12.[151,264,1911,1933]" captionTargetBox="[166,1426,349,1883]" captionTargetId="figure@12.[151,1436,337,1890]" captionTargetPageId="12" captionText="FIGURES 16 – 21. Afrus collarti (Cameron); 16, head and pronotum; 17, labrum; 18, male tergite 10; 19, male sternite 9; 20, aedeagus; 21 female genital segment." httpUri="https://zenodo.org/record/289881/files/figure.png" pageId="6" pageNumber="7">18</figureCitation>
|
||
), sclerites of tergite 9 contiguous mediobasally. Sternite 9 of male genital segment asymmetrical (
|
||
<figureCitation id="130F2A66553DFFC7FCA2FC51FC20BA35" box="[895,991,1015,1040]" captionStart="FIGURES 9 – 15" captionStartId="10.[151,264,1914,1936]" captionTargetBox="[160,1413,355,1887]" captionTargetId="figure@10.[151,1436,337,1893]" captionTargetPageId="10" captionText="FIGURES 9 – 15. Afrus spegazzinii (Bernhauer); 9, head and pronotum; 10, labrum; 11, male tergite 10; 12, male sternite 9; 13, aedeagus; 14, female genital segment (fp—fused pleurites, s—sternite, ss—supplementary sclerites); 15, metatibia (ac—apical ctenidium)." httpUri="https://zenodo.org/record/289880/files/figure.png" pageId="6" pageNumber="7">Figs. 12</figureCitation>
|
||
,
|
||
<figureCitation id="130F2A66553DFFC7FC32FC5EFBF2BA35" box="[1007,1037,1016,1040]" captionStart="FIGURES 16 – 21" captionStartId="12.[151,264,1911,1933]" captionTargetBox="[166,1426,349,1883]" captionTargetId="figure@12.[151,1436,337,1890]" captionTargetPageId="12" captionText="FIGURES 16 – 21. Afrus collarti (Cameron); 16, head and pronotum; 17, labrum; 18, male tergite 10; 19, male sternite 9; 20, aedeagus; 21 female genital segment." httpUri="https://zenodo.org/record/289881/files/figure.png" pageId="6" pageNumber="7">19</figureCitation>
|
||
), located centrally. Aedeagus with basal bulbus subovoid, short median lobe, symmetrical parameres and inner sac with shape of variably long and wide ribbon with scales of different size (
|
||
<figureCitation id="130F2A66553DFFC7FDBAFB99FD3BBA7D" box="[615,708,1087,1112]" captionStart="FIGURES 9 – 15" captionStartId="10.[151,264,1914,1936]" captionTargetBox="[160,1413,355,1887]" captionTargetId="figure@10.[151,1436,337,1893]" captionTargetPageId="10" captionText="FIGURES 9 – 15. Afrus spegazzinii (Bernhauer); 9, head and pronotum; 10, labrum; 11, male tergite 10; 12, male sternite 9; 13, aedeagus; 14, female genital segment (fp—fused pleurites, s—sternite, ss—supplementary sclerites); 15, metatibia (ac—apical ctenidium)." httpUri="https://zenodo.org/record/289880/files/figure.png" pageId="6" pageNumber="7">Figs. 13</figureCitation>
|
||
,
|
||
<figureCitation id="130F2A66553DFFC7FD0DFBE6FD0FBA7D" box="[720,752,1088,1112]" captionStart="FIGURES 16 – 21" captionStartId="12.[151,264,1911,1933]" captionTargetBox="[166,1426,349,1883]" captionTargetId="figure@12.[151,1436,337,1890]" captionTargetPageId="12" captionText="FIGURES 16 – 21. Afrus collarti (Cameron); 16, head and pronotum; 17, labrum; 18, male tergite 10; 19, male sternite 9; 20, aedeagus; 21 female genital segment." httpUri="https://zenodo.org/record/289881/files/figure.png" pageId="6" pageNumber="7">20</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8B36E3553DFFC7FF1AFBC2FE2CBA85" blockId="6.[151,1437,151,1688]" pageId="6" pageNumber="7">
|
||
Female. Genital segment with sternite proportionally short and two supplementary sclerites that cover its proximal part (
|
||
<figureCitation id="130F2A66553DFFC7FEE3FB21FE64BA85" box="[318,411,1159,1184]" captionStart="FIGURES 9 – 15" captionStartId="10.[151,264,1914,1936]" captionTargetBox="[160,1413,355,1887]" captionTargetId="figure@10.[151,1436,337,1893]" captionTargetPageId="10" captionText="FIGURES 9 – 15. Afrus spegazzinii (Bernhauer); 9, head and pronotum; 10, labrum; 11, male tergite 10; 12, male sternite 9; 13, aedeagus; 14, female genital segment (fp—fused pleurites, s—sternite, ss—supplementary sclerites); 15, metatibia (ac—apical ctenidium)." httpUri="https://zenodo.org/record/289880/files/figure.png" pageId="6" pageNumber="7">Figs. 14</figureCitation>
|
||
,
|
||
<figureCitation id="130F2A66553DFFC7FE7AFB2EFE39BA85" box="[423,454,1160,1184]" captionStart="FIGURES 16 – 21" captionStartId="12.[151,264,1911,1933]" captionTargetBox="[166,1426,349,1883]" captionTargetId="figure@12.[151,1436,337,1890]" captionTargetPageId="12" captionText="FIGURES 16 – 21. Afrus collarti (Cameron); 16, head and pronotum; 17, labrum; 18, male tergite 10; 19, male sternite 9; 20, aedeagus; 21 female genital segment." httpUri="https://zenodo.org/record/289881/files/figure.png" pageId="6" pageNumber="7">21</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8B36E3553DFFC7FF1AFB0AFCC3BACD" blockId="6.[151,1437,151,1688]" pageId="6" pageNumber="7">
|
||
<emphasis id="B940EAF1553DFFC7FF1AFB0AFE32BAE0" bold="true" box="[199,461,1196,1221]" pageId="6" pageNumber="7">Differential diagnosis.</emphasis>
|
||
This genus differs from other genera of
|
||
<taxonomicName id="4C344D60553DFFC7FC52FB0AFBE3BAE0" box="[911,1052,1196,1221]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="tribe" tribe="Xantholinini">Xantholinini</taxonomicName>
|
||
that lack dorsal rows of punctures on the pronotum in combination of the following characters:
|
||
</paragraph>
|
||
<paragraph id="8B8B36E3553DFFC7FF1AFB52FC26BB15" blockId="6.[151,1437,151,1688]" pageId="6" pageNumber="7">
|
||
1) superior line turning downwards well before middle, joining or almost joining inferior line next to front margin of procoxae and continuing onto front margin of pronotum (
|
||
<figureCitation id="130F2A66553DFFC7FC50FAB1FC2EBB15" box="[909,977,1303,1328]" captionStart="FIGURES 3 – 7. 3, 4" captionStartId="8.[151,264,1877,1899]" captionTargetBox="[169,1423,213,1814]" captionTargetId="figure@8.[150,1436,193,1828]" captionTargetPageId="8" captionText="FIGURES 3 – 7. 3, 4, Afrus collarti (Cameron); 5 – 7, Afrus spegazzinii (Bernhauer). 3, 5, head and anterior part of pronotum (iop—internal ocular puncture, pop—posterior ocular punctures); 4, 7, lateral part of head (SG—subelliptical groove); 6, lateral part of ventral side of pronotum (SL—superior line, IL—inferior line). Scale 0.5 mm." httpUri="https://zenodo.org/record/289878/files/figure.png" pageId="6" pageNumber="7">Fig. 6</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B8B36E3553DFFC7FF1AFA9AFF35BB5D" blockId="6.[151,1437,151,1688]" pageId="6" pageNumber="7">
|
||
2) head with large impression behind the eyes, lateral part of temple with evident subelliptical groove (
|
||
<figureCitation id="130F2A66553DFFC7FA9CFA9AFF3EBB5D" captionStart="FIGURES 3 – 7. 3, 4" captionStartId="8.[151,264,1877,1899]" captionTargetBox="[169,1423,213,1814]" captionTargetId="figure@8.[150,1436,193,1828]" captionTargetPageId="8" captionText="FIGURES 3 – 7. 3, 4, Afrus collarti (Cameron); 5 – 7, Afrus spegazzinii (Bernhauer). 3, 5, head and anterior part of pronotum (iop—internal ocular puncture, pop—posterior ocular punctures); 4, 7, lateral part of head (SG—subelliptical groove); 6, lateral part of ventral side of pronotum (SL—superior line, IL—inferior line). Scale 0.5 mm." httpUri="https://zenodo.org/record/289878/files/figure.png" pageId="6" pageNumber="7">Figs. 3– 5, 7</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B8B36E3553DFFC7FF1AFA22FA69BBB8" blockId="6.[151,1437,151,1688]" box="[199,1430,1412,1437]" pageId="6" pageNumber="7">
|
||
3) metatibia with only apical ctenidium which extends onto inner margin backwards to about middle (
|
||
<figureCitation id="130F2A66553DFFC7FAE8FA22FA77BBB8" box="[1333,1416,1412,1437]" captionStart="FIGURES 9 – 15" captionStartId="10.[151,264,1914,1936]" captionTargetBox="[160,1413,355,1887]" captionTargetId="figure@10.[151,1436,337,1893]" captionTargetPageId="10" captionText="FIGURES 9 – 15. Afrus spegazzinii (Bernhauer); 9, head and pronotum; 10, labrum; 11, male tergite 10; 12, male sternite 9; 13, aedeagus; 14, female genital segment (fp—fused pleurites, s—sternite, ss—supplementary sclerites); 15, metatibia (ac—apical ctenidium)." httpUri="https://zenodo.org/record/289880/files/figure.png" pageId="6" pageNumber="7">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8B36E3553DFFC7FF1AFA01FCDFB8BD" blockId="6.[151,1437,151,1688]" pageId="6" pageNumber="7">
|
||
The sister group of
|
||
<taxonomicName id="4C344D60553DFFC7FE64FA0EFE0ABBE5" box="[441,501,1448,1472]" class="Insecta" family="Staphylinidae" genus="Afrus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B940EAF1553DFFC7FE64FA0EFE0ABBE5" box="[441,501,1448,1472]" italics="true" pageId="6" pageNumber="7">Afrus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A273578A553DFFC7FDDFFA01FD8EBBE5" box="[514,625,1447,1472]" pageId="6" pageNumber="7" rank="genus">
|
||
<emphasis id="B940EAF1553DFFC7FDDFFA01FD8EBBE5" bold="true" box="[514,625,1447,1472]" pageId="6" pageNumber="7">gen. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
could be
|
||
<taxonomicName id="4C344D60553DFFC7FD2DFA01FC5DBBE5" box="[752,930,1447,1472]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B940EAF1553DFFC7FD2DFA01FC5DBBE5" box="[752,930,1447,1472]" italics="true" pageId="6" pageNumber="7">Thyreocephalus</emphasis>
|
||
</taxonomicName>
|
||
, which is supported by the following two characters: superior line turning downwards well before middle, joining or almost joining inferior line next to front margin of procoxae and continuing onto front margin of pronotum (
|
||
<figureCitation id="130F2A66553DFFC7FC12FA49FBE6B82D" box="[975,1049,1519,1544]" captionStart="FIGURES 3 – 7. 3, 4" captionStartId="8.[151,264,1877,1899]" captionTargetBox="[169,1423,213,1814]" captionTargetId="figure@8.[150,1436,193,1828]" captionTargetPageId="8" captionText="FIGURES 3 – 7. 3, 4, Afrus collarti (Cameron); 5 – 7, Afrus spegazzinii (Bernhauer). 3, 5, head and anterior part of pronotum (iop—internal ocular puncture, pop—posterior ocular punctures); 4, 7, lateral part of head (SG—subelliptical groove); 6, lateral part of ventral side of pronotum (SL—superior line, IL—inferior line). Scale 0.5 mm." httpUri="https://zenodo.org/record/289878/files/figure.png" pageId="6" pageNumber="7">Fig. 6</figureCitation>
|
||
) and metatibia with only apical ctenidium which extends onto inner margin backwards to about middle (
|
||
<figureCitation id="130F2A66553DFFC7FC1BF9B2FBE6B808" box="[966,1049,1556,1581]" captionStart="FIGURES 9 – 15" captionStartId="10.[151,264,1914,1936]" captionTargetBox="[160,1413,355,1887]" captionTargetId="figure@10.[151,1436,337,1893]" captionTargetPageId="10" captionText="FIGURES 9 – 15. Afrus spegazzinii (Bernhauer); 9, head and pronotum; 10, labrum; 11, male tergite 10; 12, male sternite 9; 13, aedeagus; 14, female genital segment (fp—fused pleurites, s—sternite, ss—supplementary sclerites); 15, metatibia (ac—apical ctenidium)." httpUri="https://zenodo.org/record/289880/files/figure.png" pageId="6" pageNumber="7">Fig. 15</figureCitation>
|
||
), but this genus differs from
|
||
<taxonomicName id="4C344D60553DFFC7FA82F9B2FA64B809" box="[1375,1435,1556,1580]" class="Insecta" family="Staphylinidae" genus="Afrus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="18" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B940EAF1553DFFC7FA82F9B2FA64B809" box="[1375,1435,1556,1580]" italics="true" pageId="6" pageNumber="7">Afrus</emphasis>
|
||
</taxonomicName>
|
||
by the absence of a subelliptical groove on the lateral parts of the temple and the presence of a subapical ctenidium (
|
||
<figureCitation id="130F2A66553DFFC7FF42F9FAFED7B851" box="[159,296,1628,1653]" captionStart="FIGURES 24 – 28. 24" captionStartId="15.[151,264,1802,1824]" captionTargetBox="[152,1422,209,1768]" captionTargetId="figure@15.[150,1436,193,1781]" captionTargetPageId="15" captionText="FIGURES 24 – 28. 24, Thyreocephalus brunneiventris (Tottenham); 25, 27, T. alluaudi (Fauvel); 26, T. interocularis (Eppelsheim); 28, T. ferox (Harold); 24, 25, head and anterior part of pronotum; 26, ventral side of pronotum (ap—antesternal plate, pr—prosternum, ip—intercoxal process, SL—superior line, IL—inferior line); 27, 28, metatibia (ac—apical ctenidium, sac—subapical ctenidium). Scale 0.5 mm." httpUri="https://zenodo.org/record/289883/files/figure.png" pageId="6" pageNumber="7">Figs. 27, 28</figureCitation>
|
||
). The two hitherto known species of
|
||
<taxonomicName id="4C344D60553DFFC7FD1AF9FAFCFBB851" box="[711,772,1628,1652]" class="Insecta" family="Staphylinidae" genus="Afrus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="18" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B940EAF1553DFFC7FD1AF9FAFCFBB851" box="[711,772,1628,1652]" italics="true" pageId="6" pageNumber="7">Afrus</emphasis>
|
||
</taxonomicName>
|
||
are also smaller (
|
||
<quantity id="4CCC9B06553DFFC7FC17F9FAFBB3B851" box="[970,1100,1628,1653]" metricMagnitude="-3" metricUnit="m" metricValue="8.25" metricValueMax="10.0" metricValueMin="6.5" pageId="6" pageNumber="7" unit="mm" value="8.25" valueMax="10.0" valueMin="6.5">6.5–10 mm</quantity>
|
||
) than
|
||
<taxonomicName id="4C344D60553DFFC7FB4FF9FAFABBB850" box="[1170,1348,1628,1653]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B940EAF1553DFFC7FB4FF9FAFABBB850" box="[1170,1348,1628,1653]" italics="true" pageId="6" pageNumber="7">Thyreocephalus</emphasis>
|
||
</taxonomicName>
|
||
species from Africa south of the Sahara (body length
|
||
<quantity id="4CCC9B06553DFFC7FD4BF9D9FCEBB8BD" box="[662,788,1663,1688]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="3.5" metricValueMin="1.0" pageId="6" pageNumber="7" unit="mm" value="22.5" valueMax="35.0" valueMin="10.0">10–35 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |