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<document id="616319E7FA3D7BD36839C8063AD156C6" ID-DOI="10.11646/zootaxa.4038.1.1" ID-GBIF-Dataset="066e84f1-206b-40cb-a615-2126696b813d" ID-ISSN="1175-5326" ID-Zenodo-Dep="289876" ID-ZooBank="1B62B78C-AA59-4417-A4FC-1CC9CED745E0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461316934850" checkinUser="plazi" docAuthor="Janák, Jiří &amp; Bordoni, Arnaldo" docDate="2015" docId="039D87F55526FFDEFF4AF8F1FCE6BB71" docLanguage="en" docName="zt04038p094.pdf" docOrigin="Zootaxa 4038 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Thyreocephalus alluaudi Fauvel 1907, comb. nov." docType="treatment" docVersion="7" lastPageNumber="32" masterDocId="FFA4FF8D553BFFC1FFDDFFA6FFFFBE25" masterDocTitle="Revision of the genus Thyreocephalus and description of Afrus gen. nov. of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="94" masterPageNumber="1" pageNumber="30" updateTime="1698618126966" updateUser="plazi">
<mods:mods id="18793B31274B111215D2FB9876DFD0A4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A7969EA1F265415B99018A30EC2402DF">
<mods:title id="782981D66BA5EF9E8B0AE44CEE88FE4A">Revision of the genus Thyreocephalus and description of Afrus gen. nov. of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
</mods:titleInfo>
<mods:name id="1167DAC0324D36E7926F7E7022267070" type="personal">
<mods:role id="217F3E5B6C7E90F045311437AEE31FCF">
<mods:roleTerm id="C2978021A9DDAD2A32DAD3A5B24EF1A9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="676938C318316CAB16D9A7718C6A8317">Janák, Jiří</mods:namePart>
</mods:name>
<mods:name id="AEB2D5279631D3B1FB032D4FDD393F20" type="personal">
<mods:role id="57E248AEBDFB65B28137C3792A2B83EC">
<mods:roleTerm id="B6AC0043F5E54D2BA27A7EFD2BBAD967">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4E6B3DB9F3B5E496F108CA8FB4E673DB">Bordoni, Arnaldo</mods:namePart>
</mods:name>
<mods:typeOfResource id="B17B956EE36BB244D236E3E1FF493D67">text</mods:typeOfResource>
<mods:relatedItem id="5BFCE2499019AD0B341F6BC2E804694D" type="host">
<mods:titleInfo id="049DA67C8286ABF2FA69C38952BE247E">
<mods:title id="C96479AC069D375CBCB6009179614EEA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="B195D7586DCA8E52787DD4BA68694963">
<mods:date id="39255AADB1E496EC8762E94348E4CC3B">2015</mods:date>
<mods:detail id="FD623724D1369D06824CCF7262746ACA" type="volume">
<mods:number id="2DBC8783D9C336B1B8C9D8D1F0F5E097">4038</mods:number>
</mods:detail>
<mods:detail id="16DBDF399EE562168B8ADF08029F84D1" type="issue">
<mods:number id="F961642181D6F4B94802418194BE494A">1</mods:number>
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<mods:extent id="611199158ACCC0551A6625790E527A2B" unit="page">
<mods:start id="5C3AEBF94DAF65B44AB2531E09460C09">1</mods:start>
<mods:end id="46970C1CBA95AA4C4059E0D213FCA61A">94</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5604080E0792497BB7D33BD4B65FBEA9">journal article</mods:classification>
<mods:identifier id="BAC434505681A6A72C2C03BF8ECA101B" type="DOI">10.11646/zootaxa.4038.1.1</mods:identifier>
<mods:identifier id="F02CE67F96CD79A3B63A9CB7CC62650F" type="GBIF-Dataset">066e84f1-206b-40cb-a615-2126696b813d</mods:identifier>
<mods:identifier id="CDED22DF686F8B238A36523EA5B53A6B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="05617465729DDB0FCBC762E7ABC8FE75" type="Zenodo-Dep">289876</mods:identifier>
<mods:identifier id="F986499DB5E5170BA7C7537D02DDC43D" type="ZooBank">1B62B78C-AA59-4417-A4FC-1CC9CED745E0</mods:identifier>
</mods:mods>
<treatment id="039D87F55526FFDEFF4AF8F1FCE6BB71" ID-DOI="http://doi.org/10.5281/zenodo.6101635" ID-GBIF-Taxon="127693951" ID-Zenodo-Dep="6101635" LSID="urn:lsid:plazi:treatment:039D87F55526FFDEFF4AF8F1FCE6BB71" httpUri="http://treatment.plazi.org/id/039D87F55526FFDEFF4AF8F1FCE6BB71" lastPageId="31" lastPageNumber="32" pageId="29" pageNumber="30">
<subSubSection id="C32E65685526FFDCFF4AF8F1FED1B9B7" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="8B8B36E35526FFDCFF4AF8F1FCF4B954" blockId="29.[151,779,1879,1938]" box="[151,779,1879,1905]" pageId="29" pageNumber="30">
<heading id="D0C3818F5526FFDCFF4AF8F1FCF4B954" bold="true" box="[151,779,1879,1905]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="B940EAF15526FFDCFF4AF8F1FCF4B954" bold="true" box="[151,779,1879,1905]" pageId="29" pageNumber="30">
<taxonomicName id="4C344D605526FFDCFF4AF8F1FD82B954" ID-CoL="56RTZ" authority="Fauvel, 1907" authorityName="Fauvel" authorityYear="1907" box="[151,637,1879,1905]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="alluaudi" status="comb. nov.">
<emphasis id="B940EAF15526FFDCFF4AF8F1FE3AB954" bold="true" box="[151,453,1879,1905]" italics="true" pageId="29" pageNumber="30">Thyreocephalus alluaudi</emphasis>
(
<bibRefCitation id="EFA54B125526FFDCFE09F8F1FD8AB954" author="Fauvel" box="[468,629,1879,1905]" pageId="29" pageNumber="30" refString="Fauvel, A. (1907) Voyage de M. Ch. Alluaud dans l'Afrique Orientale. Staphylinidae. Revue d'Entomologie, 26, 10 - 70." type="journal article" year="1907">Fauvel, 1907</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A273578A5526FFDCFD58F8F1FCF4B954" box="[645,779,1879,1905]" pageId="29" pageNumber="30">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8B36E35526FFDCFF4AF8DFFED1B9B7" blockId="29.[151,779,1879,1938]" box="[151,302,1913,1938]" pageId="29" pageNumber="30">
(
<figureCitation id="130F2A665526FFDCFF42F8DFFED9B9B7" box="[159,294,1913,1938]" captionStart-0="FIGURE 64" captionStart-1="FIGURES 65 – 69" captionStartId-0="29.[151,250,1745,1767]" captionStartId-1="30.[151,264,1970,1992]" captionTargetBox-0="[152,1432,345,1724]" captionTargetBox-1="[190,1392,206,1939]" captionTargetId-0="figure@29.[151,1436,342,1724]" captionTargetId-1="figure@30.[182,1405,193,1949]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionText-0="FIGURE 64. Distribution of species in Thyreocephalus ater group (● = T. ater (Laporte), ▲ = T. alluaudi (Fauvel), Δ = T. meridionalis sp. nov.)." captionText-1="FIGURES 65 – 69. Thyreocephalus alluaudi (Fauvel); 65, head and pronotum; 66, labrum; 67, male tergite 10; 68, male sternite 9; 69, aedeagus." httpUri-0="https://zenodo.org/record/289891/files/figure.png" httpUri-1="https://zenodo.org/record/289892/files/figure.png" pageId="29" pageNumber="30">Figs. 64–69</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32E65685526FFDCFF4AF867FCD1B9D3" pageId="29" pageNumber="30" type="reference_group">
<paragraph id="8B8B36E35526FFDCFF4AF867FCD1B9D3" blockId="29.[151,1371,1985,2038]" pageId="29" pageNumber="30">
<treatmentCitationGroup id="AB2411CD5526FFDCFF4AF867FCD1B9D3" pageId="29" pageNumber="30">
<taxonomicName id="4C344D605526FFDCFF4AF867FDDFB9F2" ID-CoL="5C5XF" authority="Fauvel, 1907: 34" authorityName="Fauvel" authorityPageNumber="34" authorityYear="1907" box="[151,544,1985,2007]" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B940EAF15526FFDCFF4AF867FE93B9F2" box="[151,364,1985,2007]" italics="true" pageId="29" pageNumber="30">Xantholinus alluaudi</emphasis>
<treatmentCitation id="0A9510F25526FFDCFEAFF867FDDFB9F2" author="Fauvel" box="[370,544,1985,2007]" page="34" pageId="29" pageNumber="30" year="1907">
<bibRefCitation id="EFA54B125526FFDCFEAFF867FE05B9F2" author="Fauvel" box="[370,506,1985,2007]" pageId="29" pageNumber="30" refString="Fauvel, A. (1907) Voyage de M. Ch. Alluaud dans l'Afrique Orientale. Staphylinidae. Revue d'Entomologie, 26, 10 - 70." type="journal article" year="1907">Fauvel, 1907</bibRefCitation>
: 34
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="0A9510F25526FFDCFDF7F867FC7BB9F2" author="Bernhauer" box="[554,900,1985,2007]" page="300" pageId="29" pageNumber="30" year="1914">
<bibRefCitation id="EFA54B125526FFDCFDF7F867FCAEB9F2" author="Bernhauer" box="[554,849,1985,2007]" pageId="29" pageNumber="30" refString="Bernhauer, M. &amp; Schubert, K. (1914) Staphylinidae IV. In: Junk, W. &amp; Schenkling, S. (Eds), Coleopterorum Catalogus, 5 (57). Junk, Berlin, pp. 289 - 408." type="journal article" year="1914">Bernhauer &amp; Schubert, 1914</bibRefCitation>
: 300
</treatmentCitation>
;
<treatmentCitation id="0A9510F25526FFDCFC53F867FB95B9F2" author="Bernhauer" box="[910,1130,1985,2007]" page="64" pageId="29" pageNumber="30" year="1939">
<bibRefCitation id="EFA54B125526FFDCFC53F867FBBAB9F2" author="Bernhauer" box="[910,1093,1985,2007]" pageId="29" pageNumber="30" refString="Bernhauer, M. (1939 a) Beschreibung der neuen Arten. In: Bernhauer, M. &amp; Chapman, W. (Eds.), Coleoptera. XIV. Staphylinidae, Staphylininae. Mission Scientifique de l'Omo 5 (44). Memoires du Museum National d'Histoire Naturelle (n. ser.), 9, 75 - 90." type="journal article" year="1939" yearSuffix="a">Bernhauer, 1939a</bibRefCitation>
: 64
</treatmentCitation>
;
<treatmentCitation id="0A9510F25526FFDCFBA8F867FAA9B9F2" author="Eichelbaum" box="[1141,1366,1985,2007]" page="86" pageId="29" pageNumber="30" year="1910">
<bibRefCitation id="EFA54B125526FFDCFBA8F867FACFB9F2" author="Eichelbaum" box="[1141,1328,1985,2007]" pageId="29" pageNumber="30" refString="Eichelbaum, S. (1910) 7. Coleoptera. 8. Staphylinidae. In: Sjostedt Y., Wissenschaftliche Ergebnisse der schwedischen Zoologischen Expedition nach dem Kilimandjaro, dem Meru und den Umgebenden Massaisteppen Deutsch-Ostafrikas 1905 - 1906, 1 (7), 79 - 94." type="journal article" year="1910">Eichelbaum, 1910</bibRefCitation>
: 86
</treatmentCitation>
.
<taxonomicName id="4C344D605526FFDCFF4AF846FEB9B9D3" box="[151,326,2016,2038]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B940EAF15526FFDCFF4AF846FEB9B9D3" box="[151,326,2016,2038]" italics="true" pageId="29" pageNumber="30">Eulissus alluaudi</emphasis>
</taxonomicName>
;
<treatmentCitation id="0A9510F25526FFDCFE8EF846FDB3B9D3" author="Scheerpeltz" box="[339,588,2016,2038]" page="1316" pageId="29" pageNumber="30" year="1933">
<bibRefCitation id="EFA54B125526FFDCFE8EF846FDF3B9D3" author="Scheerpeltz" box="[339,524,2016,2038]" pageId="29" pageNumber="30" refString="Scheerpeltz, O. (1933) Staphylinidae VII. In: Schenkling, S. (Ed.), Coleopterorum Catalogus, 6 (129). Junk, Berlin, pp. 989 - 1500." type="journal article" year="1933">Scheerpeltz, 1933</bibRefCitation>
: 1316
</treatmentCitation>
;
<treatmentCitation id="0A9510F25526FFDCFD8BF847FCD5B9D3" author="Herman" box="[598,810,2016,2038]" page="3625" pageId="29" pageNumber="30" year="2001">
<bibRefCitation id="EFA54B125526FFDCFD8BF847FD15B9D3" author="Herman" box="[598,746,2016,2038]" pageId="29" pageNumber="30" refString="Herman, L. H. (2001) Catalog of the Staphylinidae (Insecta: Coleoptera). 1758 to the end of the second millennium. VI. Staphylinine group (part 3). Bulletin of the American Museum of Natural History, 265, 3021 - 3840." type="journal article" year="2001">Herman, 2001</bibRefCitation>
: 3625
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF4B666B5525FFDFFF4AF814FEC4B9C2" httpUri="https://zenodo.org/record/289892/files/figure.png" pageId="30" pageNumber="31" targetBox="[190,1392,206,1939]" targetPageId="30">
<paragraph id="8B8B36E35525FFDFFF4AF814FEC4B9C2" blockId="30.[151,1436,1970,2023]" pageId="30" pageNumber="31">
<emphasis id="B940EAF15525FFDFFF4AF814FEB3B9ED" bold="true" box="[151,332,1970,1992]" pageId="30" pageNumber="31">FIGURES 65–69</emphasis>
.
<taxonomicName id="4C344D605525FFDFFE84F814FD4DB9ED" authority="Fauvel" authorityName="Fauvel" box="[345,690,1970,1992]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B940EAF15525FFDFFE84F814FE05B9ED" box="[345,506,1970,1992]" italics="true" pageId="30" pageNumber="31">Thyreocephalus</emphasis>
<emphasis id="B940EAF15525FFDFFE22F814FDABB9ED" box="[511,596,1970,1992]" italics="true" pageId="30" pageNumber="31">alluaudi</emphasis>
(Fauvel)
</taxonomicName>
; 65, head and pronotum; 66, labrum; 67, male tergite 10; 68, male sternite 9; 69, aedeagus.
</paragraph>
</caption>
<subSubSection id="C32E65685524FFDEFF4AFF31FCDFBCE9" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8B8B36E35524FFDEFF4AFF31FD3ABE95" blockId="31.[151,1437,151,1365]" box="[151,709,151,176]" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFF4AFF31FECABE95" bold="true" box="[151,309,151,176]" pageId="31" pageNumber="32">
<typeStatus id="548F88415524FFDEFF4AFF31FF2DBE95" box="[151,210,151,176]" pageId="31" pageNumber="32">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32376735524FFDEFEE1FF31FE5BBE95" box="[316,420,151,176]" name="Tanzania" pageId="31" pageNumber="32">Tanzania</collectingCountry>
, Kilimandjaro, Kiboscho.
</paragraph>
<paragraph id="8B8B36E35524FFDEFF1AFF1AFC94BF89" blockId="31.[151,1437,151,1365]" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFF1AFF1AFE17BEF0" bold="true" box="[199,488,188,213]" pageId="31" pageNumber="32">
<typeStatus id="548F88415524FFDEFF1AFF1AFEFDBEF0" box="[199,258,188,213]" pageId="31" pageNumber="32">Type</typeStatus>
material examined
</emphasis>
.
<typeStatus id="548F88415524FFDEFE2AFF1AFD92BEF0" box="[503,621,188,213]" pageId="31" pageNumber="32" type="lectotype">
<emphasis id="B940EAF15524FFDEFE2AFF1AFD92BEF0" bold="true" box="[503,621,188,213]" pageId="31" pageNumber="32">Lectotype</emphasis>
</typeStatus>
(by present designation) ♂:
<collectingCountry id="F32376735524FFDEFC6DFF1AFBBCBEF0" box="[944,1091,188,213]" name="Tanzania" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFC6DFF1AFBBCBEF0" bold="true" box="[944,1091,188,213]" pageId="31" pageNumber="32">TANZANIA</emphasis>
</collectingCountry>
: “Kiboscho 3, (Kilimanjaro)”, “Ex-Typis”, “Coll. et det. A. Fauvel,
<taxonomicName id="4C344D605524FFDEFDE6FF79FCA3BEDD" authority="Fvl." authorityName="Fvl." box="[571,860,223,248]" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B940EAF15524FFDEFDE6FF79FCD8BEDD" box="[571,807,223,248]" italics="true" pageId="31" pageNumber="32">Xantholinus alluaudi</emphasis>
Fvl.
</taxonomicName>
” (
<collectionCode id="ED25AE265524FFDEFCAAFF46FC2FBEDD" box="[887,976,224,248]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="31" pageNumber="32">IRSNB</collectionCode>
). The specimen is chosen as
<typeStatus id="548F88415524FFDEFACBFF79FA82BEDD" box="[1302,1405,223,248]" pageId="31" pageNumber="32" type="lectotype">lectotype</typeStatus>
of the species. This specimen have additional labels “
<typeStatus id="548F88415524FFDEFD0AFEA2FCA6BF39" box="[727,857,260,284]" pageId="31" pageNumber="32" type="lectotype">Lectotypus</typeStatus>
,
<taxonomicName id="4C344D605524FFDEFCB8FEA2FB15BF38" authority="Fvl. J. Janak" authorityName="Fvl. J. Janak" box="[869,1258,260,285]" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B940EAF15524FFDEFCB8FEA2FBADBF38" box="[869,1106,260,285]" italics="true" pageId="31" pageNumber="32">Xantholinus alluaudi</emphasis>
Fvl. J. Janák
</taxonomicName>
des. 1991” and “
<taxonomicName id="4C344D605524FFDEFF7EFE81FDE8BF65" authority="Fvl." authorityName="Fvl." box="[163,535,295,320]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B940EAF15524FFDEFF7EFE81FE3BBF65" box="[163,452,295,320]" italics="true" pageId="31" pageNumber="32">Thyreocephalus alluaudi</emphasis>
(Fvl.)
</taxonomicName>
, J. Janák det. 1991”.
<typeStatus id="548F88415524FFDEFCE6FE81FC14BF65" box="[827,1003,295,320]" pageId="31" pageNumber="32" type="paralectotype">
<emphasis id="B940EAF15524FFDEFCE6FE81FC14BF65" bold="true" box="[827,1003,295,320]" pageId="31" pageNumber="32">Paralectotypes</emphasis>
</typeStatus>
: 1 ♀:
<collectingCountry id="F32376735524FFDEFB8FFE81FB1ABF65" box="[1106,1253,295,320]" name="Tanzania" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFB8FFE81FB1ABF65" bold="true" box="[1106,1253,295,320]" pageId="31" pageNumber="32">TANZANIA</emphasis>
</collectingCountry>
: “Kibonoto 9, (Kilimanjaro)”;
<specimenCount id="9D32FD6A5524FFDEFE90FEEAFE86BF41" box="[333,377,332,356]" pageId="31" pageNumber="32" type="generic">2 ♂</specimenCount>
: “Nairobi 11, Kilimanjaro”; 1 ♀: “Kibonoto 8, Kilimanjaro” (all
<collectionCode id="ED25AE265524FFDEFBB8FEEAFB43BF41" box="[1125,1212,332,356]" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="31" pageNumber="32">IRSNB</collectionCode>
). All have the label “Coll. et det. A. Fauvel,
<taxonomicName id="4C344D605524FFDEFE63FEC9FD17BFAD" authority="Fvl." authorityName="Fvl." box="[446,744,367,392]" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B940EAF15524FFDEFE63FEC9FD52BFAD" box="[446,685,367,392]" italics="true" pageId="31" pageNumber="32">Xantholinus alluaudi</emphasis>
Fvl.
</taxonomicName>
”, “
<typeStatus id="548F88415524FFDEFCD3FEC9FC47BFAD" box="[782,952,367,392]" pageId="31" pageNumber="32" type="paralectotype">Paralectotypus</typeStatus>
,
<taxonomicName id="4C344D605524FFDEFC1AFEC9FBB1BFAD" box="[967,1102,367,392]" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAF15524FFDEFC1AFEC9FBB1BFAD" box="[967,1102,367,392]" italics="true" pageId="31" pageNumber="32">Xantholinus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C344D605524FFDEFBBCFEC9FA9ABFAD" authority="Fvl., J. Janak" authorityName="Fvl., J. Janak" box="[1121,1381,367,392]" class="Insecta" family="Staphylinidae" genus="Xantholinus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B940EAF15524FFDEFBBCFEC9FB41BFAD" box="[1121,1214,367,392]" italics="true" pageId="31" pageNumber="32">alluaudi</emphasis>
Fvl., J. Janák
</taxonomicName>
des. 1991” and “
<taxonomicName id="4C344D605524FFDEFEC3FE32FD82BF88" authority="Fvl." authorityName="Fvl." box="[286,637,404,429]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B940EAF15524FFDEFEC3FE32FDCBBF88" box="[286,564,404,429]" italics="true" pageId="31" pageNumber="32">Thyreocephalus alluaudi</emphasis>
(Fvl.)
</taxonomicName>
, J. Janák det. 1991”.
</paragraph>
<paragraph id="8B8B36E35524FFDEFF1AFE11FCDFBCE9" blockId="31.[151,1437,151,1365]" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFF1AFE11FD31BFF5" bold="true" box="[199,718,439,464]" pageId="31" pageNumber="32">
Additional material examined.
<collectingCountry id="F32376735524FFDEFDE6FE11FD31BFF5" box="[571,718,439,464]" name="Tanzania" pageId="31" pageNumber="32">TANZANIA</collectingCountry>
</emphasis>
: 1 spec.: Kilimandjaro (
<collectionCode id="ED25AE265524FFDEFC02FE1EFBCABFF5" box="[991,1077,440,464]" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="31" pageNumber="32">MFNB</collectionCode>
); 1 spec.: Kilimancharo, Schira, E. Förster (
<collectionCode id="ED25AE265524FFDEFEC4FE7AFE90BFD1" box="[281,367,476,500]" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="31" pageNumber="32">MFNB</collectionCode>
); 2 spec.: Kilimandjaro, Kibonoto,
<quantity id="4CCC9B065524FFDEFCDAFE7AFC59BFD1" box="[775,934,476,500]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="31" pageNumber="32" unit="m" value="1150.0" valueMax="1300.0" valueMin="1000.0">1000–1300 m</quantity>
, Sjostedt 1905–6 (
<collectionCode id="ED25AE265524FFDEFBA3FE7AFB2CBFD1" box="[1150,1235,476,500]" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="31" pageNumber="32">MFNB</collectionCode>
); 1 spec.: Arusha (
<collectionCode id="ED25AE265524FFDEFF42FDA6FF04BC3D" LSID="urn:lsid:biocol.org:col:34991" box="[159,251,512,536]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="31" pageNumber="32">HNHM</collectionCode>
);
<specimenCount id="9D32FD6A5524FFDEFECBFDA6FEB7BC32" box="[278,328,512,536]" pageId="31" pageNumber="32" type="generic">1 ♂</specimenCount>
: Arusha env.,
<date id="FF8A10235524FFDEFDDEFE59FD82BC3D" box="[515,637,511,536]" pageId="31" pageNumber="32" value="1995-01-01" valueMax="1995-01-07" valueMin="1995-01-01">1–7.i.1995</date>
, Smrž (
<collectionCode id="ED25AE265524FFDEFD3EFDA6FCB9BC3D" box="[739,838,512,536]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="31" pageNumber="32">NHMW</collectionCode>
); 1 spec.: Afr. or., Moschi (
<collectionCode id="ED25AE265524FFDEFB7AFDA6FB02BC3D" box="[1191,1277,512,536]" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="31" pageNumber="32">MFNB</collectionCode>
); 1 spec.: S. Tanganyika, Msamwia,
<quantity id="4CCC9B065524FFDEFE7AFD82FE00BC19" box="[423,511,548,573]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="31" pageNumber="32" unit="m" value="1850.0">1850 m</quantity>
,
<date id="FF8A10235524FFDEFDD1FD82FD9BBC18" box="[524,612,548,573]" pageId="31" pageNumber="32" value="1908-11">xi.1908</date>
, Fromm (
<collectionCode id="ED25AE265524FFDEFD09FD82FCE5BC19" box="[724,794,548,572]" pageId="31" pageNumber="32">ABFI</collectionCode>
); 1 spec.: D.
<collectingCountry id="F32376735524FFDEFC6FFD82FC36BC19" box="[946,969,548,572]" name="Iceland" pageId="31" pageNumber="32">O</collectingCountry>
. Africa,
<collectingCountry id="F32376735524FFDEFBF0FD83FB8DBC18" box="[1069,1138,549,573]" name="Tonga" pageId="31" pageNumber="32">Tonga</collectingCountry>
(
<collectionCode id="ED25AE265524FFDEFB5EFD82FB48BC19" LSID="urn:lsid:biocol.org:col:33211" box="[1155,1207,548,572]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="31" pageNumber="32">DEI</collectionCode>
); 1 ♀:
<collectingCountry id="F32376735524FFDEFAD5FD83FAAEBC18" box="[1288,1361,549,573]" name="Tonga" pageId="31" pageNumber="32">Tonga</collectingCountry>
, D.
<collectingCountry id="F32376735524FFDEFA5CFD82FA67BC19" box="[1409,1432,548,572]" name="Iceland" pageId="31" pageNumber="32">O</collectingCountry>
. Afr., coll. Benningsen (
<collectionCode id="ED25AE265524FFDEFE41FDEEFE2FBC45" LSID="urn:lsid:biocol.org:col:33211" box="[412,464,584,608]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="31" pageNumber="32">DEI</collectionCode>
).
<collectingCountry id="F32376735524FFDEFE39FDE1FC22BC45" box="[484,989,583,608]" name="Democratic Republic of the Congo" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFE39FDE1FC22BC45" bold="true" box="[484,989,583,608]" pageId="31" pageNumber="32">DEMOCRATIC REPUBLIC OF CONGO</emphasis>
</collectingCountry>
: 1 ♀: P. N. U. [Parc National Upemba], Lusinga,
<quantity id="4CCC9B065524FFDEFED8FDCAFE9DBCA1" box="[261,354,620,645]" metricMagnitude="3" metricUnit="m" metricValue="1.76" pageId="31" pageNumber="32" unit="m" value="1760.0">1760 m</quantity>
,
<date id="FF8A10235524FFDEFEAEFDCAFDCABCA0" box="[371,565,620,645]" pageId="31" pageNumber="32" value="1947-11-28" valueMax="1947-12-06" valueMin="1947-11-28">28.xi.–6.xii.1947</date>
, G. F. de Witte (
<collectionCode id="ED25AE265524FFDEFCDBFDCAFCA0BCA1" box="[774,863,620,644]" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="31" pageNumber="32">MRAC</collectionCode>
).
<collectingCountry id="F32376735524FFDEFCABFDCAFC27BCA0" box="[886,984,620,645]" name="Kenya" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFCABFDCAFC27BCA0" bold="true" box="[886,984,620,645]" pageId="31" pageNumber="32">KENYA</emphasis>
</collectingCountry>
:
<specimenCount id="9D32FD6A5524FFDEFC33FDCAFBE1BCA1" box="[1006,1054,620,644]" pageId="31" pageNumber="32" type="generic">1 ♂</specimenCount>
: Loitokitok,
<date id="FF8A10235524FFDEFB61FDCAFAD6BCA0" box="[1212,1321,620,645]" pageId="31" pageNumber="32" value="1989-11-06">6.xi.1989</date>
, Hovorka (
<collectionCode id="ED25AE265524FFDEFF43FD36FEFFBC8D" box="[158,256,656,680]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="31" pageNumber="32">NHMW</collectionCode>
);
<specimenCount id="9D32FD6A5524FFDEFEC8FD36FEC0BC82" box="[277,319,656,680]" pageId="31" pageNumber="32" type="generic">2 ♂</specimenCount>
: Nairobi, Muthaiga,
<date id="FF8A10235524FFDEFDF0FD29FD67BC8D" box="[557,664,655,680]" pageId="31" pageNumber="32" value="2007-04-04">4.iv.2007</date>
, Snížek (
<collectionCode id="ED25AE265524FFDEFCDDFD36FCBEBC8D" box="[768,833,656,680]" pageId="31" pageNumber="32">JJRC</collectionCode>
); 1 ♀: Nairobi, Muthaiga,
<date id="FF8A10235524FFDEFBB3FD36FB2FBC8D" box="[1134,1232,656,680]" pageId="31" pageNumber="32" value="2007-05-09">9.v.2007</date>
, Snížek (
<collectionCode id="ED25AE265524FFDEFAE4FD36FA85BC8D" box="[1337,1402,656,680]" pageId="31" pageNumber="32">JJRC</collectionCode>
); 1 spec.: Nairobi Westlands,
<date id="FF8A10235524FFDEFE66FD12FDD1BCE8" box="[443,558,692,717]" pageId="31" pageNumber="32" value="1976-10-19">19.x.1976</date>
, H. Gonget (
<collectionCode id="ED25AE265524FFDEFD60FD12FCEBBCE9" box="[701,788,692,716]" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="31" pageNumber="32">ZMUC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E65685524FFDEFF1AFD71FDA9BAE1" pageId="31" pageNumber="32" type="description">
<paragraph id="8B8B36E35524FFDEFF1AFD71FC0BBDED" blockId="31.[151,1437,151,1365]" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFF1AFD71FE8BBCD5" bold="true" box="[199,372,727,752]" pageId="31" pageNumber="32">Redescription.</emphasis>
Body length
<quantity id="4CCC9B065524FFDEFDD6FD7EFD78BCCA" box="[523,647,728,752]" metricMagnitude="-2" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="31" pageNumber="32" unit="mm" value="21.5" valueMax="22.0" valueMin="21.0">21–22 mm</quantity>
; length from anterior margin of head to posterior margin of elytra:
<quantity id="4CCC9B065524FFDEFAAFFD7EFF14BD31" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="31" pageNumber="32" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12– 13 mm</quantity>
. Black with genital segment, segment 8 and apical part of segment 7 reddish; elytra sometimes reddish. Body relatively narrow, shiny. Head and pronotum with micro-punctation. Head and pronotum and pertinent punctation as in
<figureCitation id="130F2A665524FFDEFE8FFCE2FE5ABD78" box="[338,421,836,861]" captionStart="FIGURES 65 – 69" captionStartId="30.[151,264,1970,1992]" captionTargetBox="[190,1392,206,1939]" captionTargetId="figure@30.[182,1405,193,1949]" captionTargetPageId="30" captionText="FIGURES 65 – 69. Thyreocephalus alluaudi (Fauvel); 65, head and pronotum; 66, labrum; 67, male tergite 10; 68, male sternite 9; 69, aedeagus." httpUri="https://zenodo.org/record/289892/files/figure.png" pageId="31" pageNumber="32">Fig. 65</figureCitation>
, labrum as in
<figureCitation id="130F2A665524FFDEFD9BFCE2FD65BD78" box="[582,666,836,861]" captionStart="FIGURES 65 – 69" captionStartId="30.[151,264,1970,1992]" captionTargetBox="[190,1392,206,1939]" captionTargetId="figure@30.[182,1405,193,1949]" captionTargetPageId="30" captionText="FIGURES 65 – 69. Thyreocephalus alluaudi (Fauvel); 65, head and pronotum; 66, labrum; 67, male tergite 10; 68, male sternite 9; 69, aedeagus." httpUri="https://zenodo.org/record/289892/files/figure.png" pageId="31" pageNumber="32">Fig. 66</figureCitation>
. Elytra longer and wider than pronotum, posteriad dilated, with less marked humeral angles. Surface with very fine and spaced punctation, arranged in three series, one near suture, one median and one lateral, composed by 7–8 spaced punctures. Abdomen shiny, with feeble traces of transverse micro-striation and fine but deep, sparse punctation, arranged in some series.
</paragraph>
<paragraph id="8B8B36E35524FFDEFF1AFC72FC8DBA10" blockId="31.[151,1437,151,1365]" pageId="31" pageNumber="32">
<collectingCountry id="F32376735524FFDEFF1AFC72FEFBBDC8" box="[199,260,980,1005]" name="Malta" pageId="31" pageNumber="32">Male</collectingCountry>
. Temples with series of fine punctures. Tergite 10 and sternite 9 of male genital segment as in
<figureCitation id="130F2A665524FFDEFAE4FC72FF49BA35" captionStart="FIGURES 65 – 69" captionStartId="30.[151,264,1970,1992]" captionTargetBox="[190,1392,206,1939]" captionTargetId="figure@30.[182,1405,193,1949]" captionTargetPageId="30" captionText="FIGURES 65 – 69. Thyreocephalus alluaudi (Fauvel); 65, head and pronotum; 66, labrum; 67, male tergite 10; 68, male sternite 9; 69, aedeagus." httpUri="https://zenodo.org/record/289892/files/figure.png" pageId="31" pageNumber="32">Figs. 67, 68</figureCitation>
. Aedeagus (
<figureCitation id="130F2A665524FFDEFEE2FC51FE6FBA35" box="[319,400,1015,1040]" captionStart="FIGURES 65 – 69" captionStartId="30.[151,264,1970,1992]" captionTargetBox="[190,1392,206,1939]" captionTargetId="figure@30.[182,1405,193,1949]" captionTargetPageId="30" captionText="FIGURES 65 – 69. Thyreocephalus alluaudi (Fauvel); 65, head and pronotum; 66, labrum; 67, male tergite 10; 68, male sternite 9; 69, aedeagus." httpUri="https://zenodo.org/record/289892/files/figure.png" pageId="31" pageNumber="32">Fig. 69</figureCitation>
) large,
<quantity id="4CCC9B065524FFDEFE38FC5EFDC3BA2A" box="[485,572,1016,1040]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="31" pageNumber="32" unit="mm" value="2.4">2.4 mm</quantity>
long, with long median lobe; parameres asymmetrical; inner sac ribbon-like, proportionally short and narrow, covered with sparse little scales.
</paragraph>
<paragraph id="8B8B36E35524FFDEFF1AFB99FD01BA72" blockId="31.[151,1437,151,1365]" box="[199,766,1087,1112]" pageId="31" pageNumber="32">Female. Temples at most with very fine punctures.</paragraph>
<paragraph id="8B8B36E35524FFDEFF1AFBC2FDA9BAE1" blockId="31.[151,1437,151,1365]" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFF1AFBC2FE30BA58" bold="true" box="[199,463,1124,1149]" pageId="31" pageNumber="32">Differential diagnosis.</emphasis>
<taxonomicName id="4C344D605524FFDEFE0AFBC2FD11BA58" box="[471,750,1124,1149]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B940EAF15524FFDEFE0AFBC2FD11BA58" box="[471,750,1124,1149]" italics="true" pageId="31" pageNumber="32">Thyreocephalus alluaudi</emphasis>
</taxonomicName>
differs from the most similar
<taxonomicName id="4C344D605524FFDEFB9BFBC3FB0FBA58" box="[1094,1264,1124,1149]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="meridionalis" status="sp. nov.">
<emphasis id="B940EAF15524FFDEFB9BFBC3FB0FBA58" box="[1094,1264,1124,1149]" italics="true" pageId="31" pageNumber="32">T. meridionalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273578A5524FFDEFB25FBC2FAACBA58" box="[1272,1363,1124,1149]" pageId="31" pageNumber="32" rank="species">
<emphasis id="B940EAF15524FFDEFB25FBC2FAACBA58" bold="true" box="[1272,1363,1124,1149]" pageId="31" pageNumber="32">sp. nov.</emphasis>
</taxonomicNameLabel>
in the reddish apical part of abdomen, rounded temples, roundly emarginated middle part of labrum and in wider aedeagus with asymmetrical parameres.
</paragraph>
</subSubSection>
<subSubSection id="C32E65685524FFDEFF1AFB69FA97BACD" box="[199,1384,1231,1256]" pageId="31" pageNumber="32" type="distribution">
<paragraph id="8B8B36E35524FFDEFF1AFB69FA97BACD" blockId="31.[151,1437,151,1365]" box="[199,1384,1231,1256]" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFF1AFB69FE9FBACD" bold="true" box="[199,352,1231,1256]" pageId="31" pageNumber="32">Distribution.</emphasis>
The species is distributed in
<collectingCountry id="F32376735524FFDEFD7AFB69FBF7BACD" box="[679,1032,1231,1256]" name="Democratic Republic of the Congo" pageId="31" pageNumber="32">Democratic Republic of Congo</collectingCountry>
,
<collectingCountry id="F32376735524FFDEFBCEFB76FBA3BACD" box="[1043,1116,1232,1256]" name="Kenya" pageId="31" pageNumber="32">Kenya</collectingCountry>
and
<collectingCountry id="F32376735524FFDEFB4EFB69FB07BACD" box="[1171,1272,1231,1256]" name="Tanzania" pageId="31" pageNumber="32">Tanzania</collectingCountry>
(
<figureCitation id="130F2A665524FFDEFADAFB69FAA3BACD" box="[1287,1372,1231,1256]" captionStart="FIGURE 64" captionStartId="29.[151,250,1745,1767]" captionTargetBox="[152,1432,345,1724]" captionTargetId="figure@29.[151,1436,342,1724]" captionTargetPageId="29" captionText="FIGURE 64. Distribution of species in Thyreocephalus ater group (● = T. ater (Laporte), ▲ = T. alluaudi (Fauvel), Δ = T. meridionalis sp. nov.)." httpUri="https://zenodo.org/record/289891/files/figure.png" pageId="31" pageNumber="32">Fig. 64</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E65685524FFDEFF1AFB52FCE6BB71" pageId="31" pageNumber="32" type="discussion">
<paragraph id="8B8B36E35524FFDEFF1AFB52FCE6BB71" blockId="31.[151,1437,151,1365]" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFF1AFB52FECBBB28" bold="true" box="[199,308,1268,1293]" pageId="31" pageNumber="32">Remarks</emphasis>
.
<collectingCountry id="F32376735524FFDEFE99FB52FE9CBB29" box="[324,355,1268,1292]" name="American Samoa" pageId="31" pageNumber="32">As</collectingCountry>
the superior line of pronotal hypomeron is turning downwards well before middle, joining or almost joining inferior line next to front margin of procoxae and continuing onto anterior margin of pronotum, the species was transferred from
<taxonomicName id="4C344D605524FFDEFE02FA9AFDC3BB70" box="[479,572,1340,1365]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAF15524FFDEFE02FA9AFDC3BB70" box="[479,572,1340,1365]" italics="true" pageId="31" pageNumber="32">Eulissus</emphasis>
</taxonomicName>
to
<taxonomicName id="4C344D605524FFDEFDBDFA9AFCEDBB70" box="[608,786,1340,1365]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis id="B940EAF15524FFDEFDBDFA9AFCEDBB70" box="[608,786,1340,1365]" italics="true" pageId="31" pageNumber="32">Thyreocephalus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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