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<document id="6A7481040B5372D435AED5ECA0C18357" ID-DOI="10.11646/zootaxa.4038.1.1" ID-GBIF-Dataset="066e84f1-206b-40cb-a615-2126696b813d" ID-ISSN="1175-5326" ID-Zenodo-Dep="289876" ID-ZooBank="1B62B78C-AA59-4417-A4FC-1CC9CED745E0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461316934850" checkinUser="plazi" docAuthor="Janák, Jiří &amp; Bordoni, Arnaldo" docDate="2015" docId="039D87F55524FFE0FF4AFA3BFB85BA59" docLanguage="en" docName="zt04038p094.pdf" docOrigin="Zootaxa 4038 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Thyreocephalus meridionalis Janák &amp; Bordoni, 2015, sp. nov." docType="treatment" docVersion="7" lastPageNumber="34" masterDocId="FFA4FF8D553BFFC1FFDDFFA6FFFFBE25" masterDocTitle="Revision of the genus Thyreocephalus and description of Afrus gen. nov. of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="94" masterPageNumber="1" pageNumber="32" updateTime="1698618126966" updateUser="plazi">
<mods:mods id="617E2F3B9E8B7E586631F6AB0C6A13BF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D2547A580D019EA6F1DCD4580A21BE4F">
<mods:title id="66A68E46E059B64394D68AB3FB716BCD">Revision of the genus Thyreocephalus and description of Afrus gen. nov. of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
</mods:titleInfo>
<mods:name id="75C200F4AD65D5435F43F2FFDC05D6FB" type="personal">
<mods:role id="577D1A8303D0E6465D2961432A00ED1A">
<mods:roleTerm id="8D9C3A2AA9AA24C2A390ABEC611E4C07">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FEF944AD0AF4594646A016C4FA78B3BA">Janák, Jiří</mods:namePart>
</mods:name>
<mods:name id="FABD0393D2C2FC9FABC318938406694A" type="personal">
<mods:role id="5D5E5E6D1895CF27421FD8E0EC3979B1">
<mods:roleTerm id="FC5861DA69F72590EAEAB1918F4382FE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="70B21A2F3D8D29004269F67CECF02AA9">Bordoni, Arnaldo</mods:namePart>
</mods:name>
<mods:typeOfResource id="73CE61D80A158C54899AC3162F1CB03A">text</mods:typeOfResource>
<mods:relatedItem id="4528E5DA5113C4CD27DA94D0D2321B75" type="host">
<mods:titleInfo id="FAE9CD216A22A8909B1606E8D23D5DB8">
<mods:title id="327DA167C11F71AD34401A9DB8715E4F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="519852F186B388A62D29151AC63C51D2">
<mods:date id="6B4A726687E7259A7BF422944444ECE2">2015</mods:date>
<mods:detail id="E81B067EFECCCB9647BB9C58ACE2FAD2" type="volume">
<mods:number id="4348B55AB367DDAF092905D502FDA114">4038</mods:number>
</mods:detail>
<mods:detail id="8A9E53AAB15A11E2311292DD2754207A" type="issue">
<mods:number id="D1894E082C0A51EBA6614F9C679E3594">1</mods:number>
</mods:detail>
<mods:extent id="7351CB20E83CAE778BCAD113D40C51D5" unit="page">
<mods:start id="D3A8D79AA75A657C8528AB092117C72C">1</mods:start>
<mods:end id="47EE66CC9DE6A0932382C6C8FCA3A84D">94</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F6E675B4F3CD7C8D35F81C10C6E08FD8">journal article</mods:classification>
<mods:identifier id="C7B127C2AA941FBC7D3D3C189075B904" type="DOI">10.11646/zootaxa.4038.1.1</mods:identifier>
<mods:identifier id="EC4547660AA12096A64E949118E0D4A0" type="GBIF-Dataset">066e84f1-206b-40cb-a615-2126696b813d</mods:identifier>
<mods:identifier id="2ECB2E7DE60F42AF80784EF04913CD6F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EC8BBCFF94D09C331A01368A4FB23F3B" type="Zenodo-Dep">289876</mods:identifier>
<mods:identifier id="9446AE8F8B0E8A58CDED1E36158FBC35" type="ZooBank">1B62B78C-AA59-4417-A4FC-1CC9CED745E0</mods:identifier>
</mods:mods>
<treatment id="039D87F55524FFE0FF4AFA3BFB85BA59" ID-DOI="http://doi.org/10.5281/zenodo.6101637" ID-GBIF-Taxon="127693950" ID-Zenodo-Dep="6101637" LSID="urn:lsid:plazi:treatment:039D87F55524FFE0FF4AFA3BFB85BA59" httpUri="http://treatment.plazi.org/id/039D87F55524FFE0FF4AFA3BFB85BA59" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32">
<subSubSection id="C32E65685524FFDEFF4AFA3BFE6FBBFD" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8B8B36E35524FFDEFF4AFA3BFDA3BB92" blockId="31.[151,604,1437,1496]" box="[151,604,1437,1463]" pageId="31" pageNumber="32">
<heading id="D0C3818F5524FFDEFF4AFA3BFDA3BB92" bold="true" box="[151,604,1437,1463]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
<emphasis id="B940EAF15524FFDEFF4AFA3BFDA3BB92" bold="true" box="[151,604,1437,1463]" pageId="31" pageNumber="32">
<taxonomicName id="4C344D605524FFDEFF4AFA3BFE07BB92" ID-CoL="56RX9" box="[151,504,1437,1463]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="meridionalis" status="sp. nov.">
<emphasis id="B940EAF15524FFDEFF4AFA3BFE07BB92" bold="true" box="[151,504,1437,1463]" italics="true" pageId="31" pageNumber="32">Thyreocephalus meridionalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273578A5524FFDEFE23FA3BFDA3BB92" box="[510,604,1437,1463]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8B36E35524FFDEFF4AFA19FE6FBBFD" blockId="31.[151,604,1437,1496]" box="[151,400,1471,1496]" pageId="31" pageNumber="32">
(
<figureCitation id="130F2A665524FFDEFF42FA19FF03BBFD" box="[159,252,1471,1496]" captionStart="FIGURE 64" captionStartId="29.[151,250,1745,1767]" captionTargetBox="[152,1432,345,1724]" captionTargetId="figure@29.[151,1436,342,1724]" captionTargetPageId="29" captionText="FIGURE 64. Distribution of species in Thyreocephalus ater group (● = T. ater (Laporte), ▲ = T. alluaudi (Fauvel), Δ = T. meridionalis sp. nov.)." httpUri="https://zenodo.org/record/289891/files/figure.png" pageId="31" pageNumber="32">Figs. 64</figureCitation>
,
<figureCitation id="130F2A665524FFDEFED5FA66FEAEBBFD" box="[264,337,1472,1496]" captionStart="FIGURES 70 – 74" captionStartId="32.[151,264,1919,1941]" captionTargetBox="[167,1421,211,1883]" captionTargetId="figure@32.[151,1436,195,1897]" captionTargetPageId="32" captionText="FIGURES 70 – 74. Thyreocephalus meridionalis sp. nov.; 70, head and pronotum; 71, labrum; 72, male tergite 10; 73, male sternite 9; 74, aedeagus." httpUri="https://zenodo.org/record/289893/files/figure.png" pageId="31" pageNumber="32">70–74</figureCitation>
,
<figureCitation id="130F2A665524FFDEFE80FA66FE77BBFD" box="[349,392,1472,1496]" captionStart="FIGURES 238 – 240. 238, 239" captionStartId="90.[151,264,1811,1833]" captionTargetBox="[170,1419,213,1777]" captionTargetId="figure@90.[150,1436,193,1790]" captionTargetPageId="90" captionText="FIGURES 238 – 240. 238, 239, Tsingidi, Republic of Congo (type locality of Thyreocephalus tsingidianus sp. nov. and T. pseudoafricanus sp. nov.); 238, forest; 239, pitfall traps; 240, Mariepskop, South Africa (type locality of T. meridionalis sp. nov.) — arrow indicates a place where the specimen was collected." httpUri="https://zenodo.org/record/289931/files/figure.png" pageId="31" pageNumber="32">240</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32E65685524FFE0FF4AF9A1FBFFBF65" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32" type="materials_examined">
<paragraph id="8B8B36E35524FFDEFF4AF9A1FC9CB83A" blockId="31.[151,1437,1543,2037]" box="[151,867,1543,1568]" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFF4AF9A1FECAB805" bold="true" box="[151,309,1543,1568]" pageId="31" pageNumber="32">
<typeStatus id="548F88415524FFDEFF4AF9A1FF2DB805" box="[151,210,1543,1568]" pageId="31" pageNumber="32">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F32376735524FFDEFEE1F9A1FE60B805" box="[316,415,1543,1568]" name="Namibia" pageId="31" pageNumber="32">Namibia</collectingCountry>
, Popa Falls,
<geoCoordinate id="EE0050245524FFDEFDF3F9AEFD3DB805" box="[558,706,1543,1568]" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="15" value="-18.12111">18° 07' 16'' S</geoCoordinate>
,
<geoCoordinate id="EE0050245524FFDEFD12F9AEFC9CB83A" box="[719,867,1543,1568]" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="15" value="21.580833">21° 34' 51'' E</geoCoordinate>
</paragraph>
<paragraph id="8B8B36E35524FFE0FF1AF98AFBFFBF65" blockId="31.[151,1437,1543,2037]" lastBlockId="33.[151,1437,151,1149]" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFF1AF98AFE00B860" bold="true" box="[199,511,1580,1605]" pageId="31" pageNumber="32">
<typeStatus id="548F88415524FFDEFF1AF98AFEFDB860" box="[199,258,1580,1605]" pageId="31" pageNumber="32">Type</typeStatus>
material.
<typeStatus id="548F88415524FFDEFE4FF98AFE00B860" box="[402,511,1580,1605]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
</emphasis>
♂:
<collectingCountry id="F32376735524FFDEFD9DF98AFD40B860" box="[576,703,1580,1605]" name="Namibia" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFD9DF98AFD40B860" bold="true" box="[576,703,1580,1605]" pageId="31" pageNumber="32">NAMIBIA</emphasis>
</collectingCountry>
: “
<collectingCountry id="F32376735524FFDEFD39F98AFCB5B860" box="[740,842,1580,1605]" name="Namibia" pageId="31" pageNumber="32">Namibia</collectingCountry>
, Popa Falls,
<geoCoordinate id="EE0050245524FFDEFC27F98AFB85B860" box="[1018,1146,1580,1605]" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="15" value="-18.12111">18°07'16''S</geoCoordinate>
,
<geoCoordinate id="EE0050245524FFDEFB4CF98AFAECB860" box="[1169,1299,1580,1605]" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="15" value="21.580833">21°34'51''E</geoCoordinate>
, M. Uhlig
<date id="FF8A10235524FFDEFF4AF9F6FEDCB84D" box="[151,291,1616,1640]" pageId="31" pageNumber="32" value="1993-12-13">13.XII.1993</date>
”, “
<typeStatus id="548F88415524FFDEFE8DF9E9FE38B84D" box="[336,455,1615,1640]" pageId="31" pageNumber="32" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C344D605524FFDEFE04F9E9FCD5B84D" box="[473,810,1615,1640]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="meridionalis" status="sp. nov.">
<emphasis id="B940EAF15524FFDEFE04F9E9FCD5B84D" box="[473,810,1615,1640]" italics="true" pageId="31" pageNumber="32">Thyreocephalus meridionalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273578A5524FFDEFCE0F9F6FC5EB842" box="[829,929,1616,1639]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
, Janák &amp; Bordoni det. 2015” (
<collectionCode id="ED25AE265524FFDEFAE6F9F6FA6EB84D" box="[1339,1425,1616,1640]" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="31" pageNumber="32">MFNB</collectionCode>
).
<typeStatus id="548F88415524FFDEFF4AF9D2FEFBB8A8" box="[151,260,1652,1677]" pageId="31" pageNumber="32" type="paratype">
<emphasis id="B940EAF15524FFDEFF4AF9D2FEFBB8A8" bold="true" box="[151,260,1652,1677]" pageId="31" pageNumber="32">Paratype</emphasis>
</typeStatus>
s:
<collectingCountry id="F32376735524FFDEFEFCF9D2FE3EB8A8" box="[289,449,1652,1677]" name="Botswana" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFEFCF9D2FE3EB8A8" bold="true" box="[289,449,1652,1677]" pageId="31" pageNumber="32">BOTSWANA</emphasis>
</collectingCountry>
: 1 ♀: “
<collectingCountry id="F32376735524FFDEFDFDF9D2FD6BB8A9" box="[544,660,1652,1676]" name="Botswana" pageId="31" pageNumber="32">Botswana</collectingCountry>
.
<quantity id="4CCC9B065524FFDEFD7EF9D2FD11B8A8" box="[675,750,1652,1677]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="31" pageNumber="32" unit="km" value="10.0">10 km</quantity>
N of Maun, J. Moravec” (
<collectionCode id="ED25AE265524FFDEFBF2F9D2FB8FB8A9" box="[1071,1136,1652,1676]" pageId="31" pageNumber="32">JJRC</collectionCode>
);
<collectingCountry id="F32376735524FFDEFB5AF9D2FAACB8A8" box="[1159,1363,1652,1677]" name="Mozambique" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFB5AF9D2FAACB8A8" bold="true" box="[1159,1363,1652,1677]" pageId="31" pageNumber="32">MOZAMBIQUE</emphasis>
</collectingCountry>
: 1 ♀: “
<collectingCountry id="F32376735524FFDEFF7CF931FEC3B895" box="[161,316,1687,1712]" name="Mozambique" pageId="31" pageNumber="32">Mozambique</collectingCountry>
: Zambesia,
<quantity id="4CCC9B065524FFDEFE12F931FDE2B895" box="[463,541,1687,1712]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="31" pageNumber="32" unit="km" value="50.0">50 km</quantity>
S Gurué,
<geoCoordinate id="EE0050245524FFDEFD79F931FD01B895" box="[676,766,1687,1712]" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="555" value="-15.47">15.47 S</geoCoordinate>
–
<geoCoordinate id="EE0050245524FFDEFCD3F93EFC8FB895" box="[782,880,1688,1712]" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="555" value="37.97">37.97 E</geoCoordinate>
”, “
<date id="FF8A10235524FFDEFC44F931FBF0B895" box="[921,1039,1687,1712]" pageId="31" pageNumber="32" value="2006-08-12">8.12.2006</date>
; E-Y: 3746, on mushrooms, leg. Gussman, Müller” (
<collectionCode id="ED25AE265524FFDEFEABF91AFE48B8F1" box="[374,439,1724,1748]" pageId="31" pageNumber="32">JJRC</collectionCode>
).
<collectingCountry id="F32376735524FFDEFE17F91AFDB5B8F0" box="[458,586,1724,1749]" name="Namibia" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFE17F91AFDB5B8F0" bold="true" box="[458,586,1724,1749]" pageId="31" pageNumber="32">NAMIBIA</emphasis>
</collectingCountry>
: 1 ♀: “
<collectingCountry id="F32376735524FFDEFD43F91AFCFBB8F0" box="[670,772,1724,1749]" name="Namibia" pageId="31" pageNumber="32">Namibia</collectingCountry>
, East Caprivi, Katima Mulilo, Zambezi riv., 19–
<date id="FF8A10235524FFDEFAFEF91AFA67B8F0" box="[1315,1432,1724,1749]" pageId="31" pageNumber="32" value="1999-01-25">25.1.1999</date>
,
<quantity id="4CCC9B065524FFDEFF4AF979FF1BB8D2" box="[151,228,1759,1784]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="31" pageNumber="32" unit="m" value="950.0">950 m</quantity>
, Rudolf Kmeco leg.” (
<collectionCode id="ED25AE265524FFDEFE2DF946FDCEB8DD" box="[496,561,1760,1784]" pageId="31" pageNumber="32">JJRC</collectionCode>
); 1 ♀: “
<collectingCountry id="F32376735524FFDEFD4EF979FD09B8DD" box="[659,758,1759,1784]" name="Namibia" pageId="31" pageNumber="32">Namibia</collectingCountry>
bor. or., Katima Mulilo, Caprivi zipfel, 15–
<date id="FF8A10235524FFDEFB25F979FA92B8DD" box="[1272,1389,1759,1784]" pageId="31" pageNumber="32" value="1995-01-24">24.1.1995</date>
, M. Snížek lgt.” (
<collectionCode id="ED25AE265524FFDEFEF3F8A2FE90B939" box="[302,367,1796,1820]" pageId="31" pageNumber="32">JJRC</collectionCode>
);
<collectingCountry id="F32376735524FFDEFE59F8A2FE08B938" box="[388,503,1796,1821]" name="Zambia" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFE59F8A2FE08B938" bold="true" box="[388,503,1796,1821]" pageId="31" pageNumber="32">ZAMBIA</emphasis>
</collectingCountry>
: 1 ♀: “NO
<collectingCountry id="F32376735524FFDEFD5EF8A2FD23B938" box="[643,732,1796,1821]" name="Zambia" pageId="31" pageNumber="32">Zambia</collectingCountry>
, E Mkushi env., Snizek &amp; Tichy 16.18.
<date id="FF8A10235524FFDEFB7DF8A2FAF6B938" box="[1184,1289,1796,1821]" pageId="31" pageNumber="32" value="2004-12">XII.2004</date>
” (
<collectionCode id="ED25AE265524FFDEFAF8F8A2FA86B939" box="[1317,1401,1796,1820]" pageId="31" pageNumber="32">GROB</collectionCode>
); 1 spec.: “NW
<collectingCountry id="F32376735524FFDEFEFCF881FE85B965" box="[289,378,1831,1856]" name="Zambia" pageId="31" pageNumber="32">Zambia</collectingCountry>
, Mutumbwe, B. &amp; P. Malec
<date id="FF8A10235524FFDEFD66F881FCADB965" box="[699,850,1831,1856]" pageId="31" pageNumber="32" value="2005-01-24" valueMax="2005-01-30" valueMin="2005-01-24">24–30.I.2005</date>
” (
<collectionCode id="ED25AE265524FFDEFCB1F88EFC3EB965" box="[876,961,1832,1856]" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="31" pageNumber="32">MFNB</collectionCode>
);
<specimenCount id="9D32FD6A5524FFDEFC08F88EFC00B91A" box="[981,1023,1832,1856]" pageId="31" pageNumber="32" type="generic">1 ♂</specimenCount>
: “
<collectingCountry id="F32376735524FFDEFBC5F881FB8CB965" box="[1048,1139,1831,1856]" name="Zambia" pageId="31" pageNumber="32">Zambia</collectingCountry>
, Livingstone Victoria Fals [sic!], 26–
<date id="FF8A10235524FFDEFED6F8EAFE73B940" box="[267,396,1868,1893]" pageId="31" pageNumber="32" value="1994-12-31">31.12.1994</date>
, Lgt. Snizek” (
<collectionCode id="ED25AE265524FFDEFDE4F8EAFD85B941" box="[569,634,1868,1892]" pageId="31" pageNumber="32">JJRC</collectionCode>
); 1 spec.: “
<collectingCountry id="F32376735524FFDEFD24F8EAFCABB940" box="[761,852,1868,1893]" name="Zambia" pageId="31" pageNumber="32">Zambia</collectingCountry>
, Kapiri, Mposhi,
<quantity id="4CCC9B065524FFDEFBC0F8EAFB8CB940" box="[1053,1139,1868,1893]" metricMagnitude="5" metricUnit="m" metricValue="1.6" pageId="31" pageNumber="32" unit="km" value="160.0">160 km</quantity>
N Lusaka, B. &amp; P. Malec
<date id="FF8A10235524FFDEFF4AF8C9FF00B9AD" box="[151,255,1903,1928]" pageId="31" pageNumber="32" value="2005-02-02">2.II.2005</date>
” (
<collectionCode id="ED25AE265524FFDEFEFFF8D6FE87B9AD" box="[290,376,1904,1928]" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="31" pageNumber="32">MFNB</collectionCode>
);
<collectingCountry id="F32376735524FFDEFE4EF8C9FDCCB9AD" box="[403,563,1903,1928]" name="Zimbabwe" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFE4EF8C9FDCCB9AD" bold="true" box="[403,563,1903,1928]" pageId="31" pageNumber="32">ZIMBABWE</emphasis>
</collectingCountry>
: 1 ♀: “
<collectingCountry id="F32376735524FFDEFD7DF8C9FCF1B9AD" box="[672,782,1903,1928]" name="Zimbabwe" pageId="31" pageNumber="32">Rhodesia</collectingCountry>
, Atlantica, Ecol. Res. Station, nr. Salisbury [Harare], 25:III:1970, at light, Coll. J. R. Clover &amp; A. C. Kistner Field No 1883” (
<collectionCode id="ED25AE265524FFDEFC11F832FBDCB989" LSID="urn:lsid:biocol.org:col:34795" box="[972,1059,1940,1964]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="31" pageNumber="32">FMNH</collectionCode>
);
<specimenCount id="9D32FD6A5524FFDEFBEAF832FB9EB989" box="[1079,1121,1940,1964]" pageId="31" pageNumber="32" type="generic">1 ♂</specimenCount>
: “
<collectingCountry id="F32376735524FFDEFBA6F832FB0AB988" box="[1147,1269,1940,1965]" name="Zimbabwe" pageId="31" pageNumber="32">Zimbabwe</collectingCountry>
W, Bulawayo, Shangani, Naletale ruins,
<date id="FF8A10235524FFDEFE60F81EFDC1B9F5" box="[445,574,1976,2000]" pageId="31" pageNumber="32" value="2012-12-20">20.12.2012</date>
, Snížek (
<collectionCode id="ED25AE265524FFDEFD75F81EFD16B9F5" box="[680,745,1976,2000]" pageId="31" pageNumber="32">JJRC</collectionCode>
);
<specimenCount id="9D32FD6A5524FFDEFD22F81EFCD5B9F5" box="[767,810,1976,2000]" pageId="31" pageNumber="32" type="generic">1 ♂</specimenCount>
: “
<collectingCountry id="F32376735524FFDEFC98F81EFC3DB9F4" box="[837,962,1976,2001]" name="Zimbabwe" pageId="31" pageNumber="32">Zimbabwe</collectingCountry>
, Kyle Recr. Park at Lake Mutiri Kwi, M. Uhlig
<date id="FF8A10235524FFDEFF00F87AFE7AB9D0" box="[221,389,2012,2037]" pageId="31" pageNumber="32" value="1995-12-02" valueMax="1995-12-04" valueMin="1995-12-02">2–4. XII. 1995</date>
,
<geoCoordinate id="EE0050245524FFDEFE4AF87AFE17B9D1" box="[407,488,2012,2036]" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="555" value="-20.13">20.13S</geoCoordinate>
,
<geoCoordinate id="EE0050245524FFDEFE26F87AFDB2B9D1" box="[507,589,2012,2036]" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="555" value="31.0">31.00E</geoCoordinate>
” (
<collectionCode id="ED25AE265524FFDEFDBAF87AFD52B9D1" box="[615,685,2012,2036]" pageId="31" pageNumber="32">ABFI</collectionCode>
);
<collectingCountry id="F32376735524FFDEFD1DF87AFC69B9D0" box="[704,918,2012,2037]" name="South Africa" pageId="31" pageNumber="32">
<emphasis id="B940EAF15524FFDEFD1DF87AFC69B9D0" bold="true" box="[704,918,2012,2037]" pageId="31" pageNumber="32">SOUTH AFRICA</emphasis>
</collectingCountry>
: Mpumalanga: 1 ♀: “
<collectingCountry id="F32376735524FFDEFB57F87AFB3CB9D1" box="[1162,1219,2012,2036]" name="South Africa" pageId="31" pageNumber="32">RSA</collectingCountry>
(N), Mpumalanga,
<geoCoordinate id="EE005024551AFFE0FF4AFF31FEFEBE95" box="[151,257,151,176]" direction="south" orientation="latitude" pageId="33" pageNumber="34" precision="5" value="-24.5851">24.5851S</geoCoordinate>
/
<geoCoordinate id="EE005024551AFFE0FECCFF31FE80BE95" box="[273,383,151,176]" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="5" value="30.8633">30.8633E</geoCoordinate>
, Mariepskop (forestry office),
<quantity id="4CCC9B06551AFFE0FD0AFF3EFCD1BE8A" box="[727,814,152,176]" metricMagnitude="3" metricUnit="m" metricValue="1.33" pageId="33" pageNumber="34" unit="m" value="1330.0">1330 m</quantity>
, tree trunk,
<date id="FF8A1023551AFFE0FC69FF31FBCBBE95" box="[948,1076,151,176]" pageId="33" pageNumber="34" value="2012-11-16">16.11.2012</date>
, leg. P. Jałoszyński” (
<collectionCode id="ED25AE26551AFFE0FAFAFF3EFA84BE95" box="[1319,1403,152,176]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="33" pageNumber="34">TMSA</collectionCode>
); 1 ♀: “S. Afr.; Kruger Nat. Pk, Skukuza – Sabi River,
<geoCoordinate id="EE005024551AFFE0FD0FFF1AFCDABEF1" box="[722,805,188,213]" direction="south" orientation="latitude" pageId="33" pageNumber="34" precision="555" value="-24.57">24.57 S</geoCoordinate>
–
<geoCoordinate id="EE005024551AFFE0FCE9FF1AFC70BEF1" box="[820,911,188,212]" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="555" value="31.42">31.42 E</geoCoordinate>
”, “
<date id="FF8A1023551AFFE0FC6DFF1AFBD8BEF0" box="[944,1063,188,213]" pageId="33" pageNumber="34" value="1995-02-22">22.2.1995</date>
: E-Y: 3111, riverinefor. litter, leg. Endrödy-Younga” (
<collectionCode id="ED25AE26551AFFE0FEABFF46FE35BEDD" box="[374,458,224,248]" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="33" pageNumber="34">TMSA</collectionCode>
); 1 spec.: “
<collectingCountry id="F3237673551AFFE0FD8FFF79FD13BEDD" box="[594,748,223,248]" name="South Africa" pageId="33" pageNumber="34">South Africa</collectingCountry>
, Transvaal, Duiwelskloof,
<geoCoordinate id="EE005024551AFFE0FBFAFF46FB87BEDD" box="[1063,1144,224,248]" direction="south" orientation="latitude" pageId="33" pageNumber="34" precision="555" value="-23.42">23.42S</geoCoordinate>
,
<geoCoordinate id="EE005024551AFFE0FB55FF79FB24BEDD" box="[1160,1243,223,248]" direction="east" orientation="longitude" pageId="33" pageNumber="34" precision="555" value="30.06">30.06E</geoCoordinate>
, V. M. Uys
<date id="FF8A1023551AFFE0FAAFFF46FEFCBF39" pageId="33" pageNumber="34" value="1987-01-12" valueMax="1987-01-14" valueMin="1987-01-12">12– 14.I.1987</date>
” (
<collectionCode id="ED25AE26551AFFE0FEC2FEA2FE8FBF39" LSID="urn:lsid:biocol.org:col:34248" box="[287,368,260,284]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34248" name="Agricultural Research Council-Plant Protection Research Institute" pageId="33" pageNumber="34">SANC</collectionCode>
); 1 spec.: “Mp' home (?) Transv., M. Knothe” (
<collectionCode id="ED25AE26551AFFE0FC56FEA2FC1EBF39" box="[907,993,260,284]" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="33" pageNumber="34">MFNB</collectionCode>
). The label “
<typeStatus id="548F8841551AFFE0FBACFEA2FB1DBF39" box="[1137,1250,260,284]" pageId="33" pageNumber="34" type="paratype">Paratypus</typeStatus>
<taxonomicName id="4C344D60551AFFE0FB37FEA2FEDBBF65" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="meridionalis" status="sp. nov.">
<emphasis id="B940EAF1551AFFE0FB37FEA2FEDBBF65" italics="true" pageId="33" pageNumber="34">Thyreocephalus meridionalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273578A551AFFE0FEF6FE8EFE7BBF1A" box="[299,388,296,319]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
, Janák &amp; Bordoni det. 2015” was added to all
<typeStatus id="548F8841551AFFE0FC50FE8EFC03BF65" box="[909,1020,296,320]" pageId="33" pageNumber="34" type="paratype">paratypes</typeStatus>
.
</paragraph>
</subSubSection>
<caption id="DF4B666B551BFFE1FF4AF8D9FE74B996" httpUri="https://zenodo.org/record/289893/files/figure.png" pageId="32" pageNumber="33" targetBox="[167,1421,211,1883]" targetPageId="32">
<paragraph id="8B8B36E3551BFFE1FF4AF8D9FE74B996" blockId="32.[151,1435,1919,1971]" pageId="32" pageNumber="33">
<emphasis id="B940EAF1551BFFE1FF4AF8D9FEB1B9B0" bold="true" box="[151,334,1919,1941]" pageId="32" pageNumber="33">FIGURES 70–74</emphasis>
.
<taxonomicName id="4C344D60551BFFE1FE80F8D9FD7AB9B0" box="[349,645,1919,1941]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="meridionalis" status="sp. nov.">
<emphasis id="B940EAF1551BFFE1FE80F8D9FD7AB9B0" box="[349,645,1919,1941]" italics="true" pageId="32" pageNumber="33">Thyreocephalus meridionalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273578A551BFFE1FD50F8D9FD20B9B0" box="[653,735,1919,1941]" pageId="32" pageNumber="33" rank="species">
<emphasis id="B940EAF1551BFFE1FD50F8D9FD20B9B0" bold="true" box="[653,735,1919,1941]" pageId="32" pageNumber="33">sp. nov.</emphasis>
</taxonomicNameLabel>
; 70, head and pronotum; 71, labrum; 72, male tergite 10; 73, male sternite 9; 74, aedeagus.
</paragraph>
</caption>
<subSubSection id="C32E6568551AFFE0FF1AFEEAFD1ABD79" pageId="33" pageNumber="34" type="description">
<paragraph id="8B8B36E3551AFFE0FF1AFEEAFE0BBC45" blockId="33.[151,1437,151,1149]" pageId="33" pageNumber="34">
<emphasis id="B940EAF1551AFFE0FF1AFEEAFEA6BF40" bold="true" box="[199,345,332,357]" pageId="33" pageNumber="34">Description.</emphasis>
Body about
<quantity id="4CCC9B06551AFFE0FE31FEEAFD98BF41" box="[492,615,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="33" pageNumber="34" unit="mm" value="23.5" valueMax="25.0" valueMin="22.0">22–25 mm</quantity>
long; length from anterior margin of head to posterior margin of elytra:
<quantity id="4CCC9B06551AFFE0FF4AFED6FEE7BFA2" box="[151,280,368,392]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="33" pageNumber="34" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">12–14 mm</quantity>
. Black with reddish genital segment, exceptionally tip of abdomen beginning from apical part of segment 7 reddish. Elytra with feebly bluish reflections. Head and pronotum and related punctation as in
<figureCitation id="130F2A66551AFFE0FA99FE32FA68BF89" box="[1348,1431,404,429]" captionStart="FIGURES 70 – 74" captionStartId="32.[151,264,1919,1941]" captionTargetBox="[167,1421,211,1883]" captionTargetId="figure@32.[151,1436,195,1897]" captionTargetPageId="32" captionText="FIGURES 70 – 74. Thyreocephalus meridionalis sp. nov.; 70, head and pronotum; 71, labrum; 72, male tergite 10; 73, male sternite 9; 74, aedeagus." httpUri="https://zenodo.org/record/289893/files/figure.png" pageId="33" pageNumber="34">Fig. 70</figureCitation>
; labrum as in
<figureCitation id="130F2A66551AFFE0FEF4FE11FE83BFF5" box="[297,380,439,464]" captionStart="FIGURES 70 – 74" captionStartId="32.[151,264,1919,1941]" captionTargetBox="[167,1421,211,1883]" captionTargetId="figure@32.[151,1436,195,1897]" captionTargetPageId="32" captionText="FIGURES 70 – 74. Thyreocephalus meridionalis sp. nov.; 70, head and pronotum; 71, labrum; 72, male tergite 10; 73, male sternite 9; 74, aedeagus." httpUri="https://zenodo.org/record/289893/files/figure.png" pageId="33" pageNumber="34">Fig. 71</figureCitation>
. Head and pronotum with very minute, sparse micro-punctation. Pronotum with sinuate sides. Elytra sub-quadrangular, slightly dilated posteriad, with marked humeral angles. Surface with fine punctation bearing long, brown setae, arranged in three series of very sparse punctures, one near suture of 3–4, one median of 4–5 more coarse and one lateral of 5–6 punctures. Abdomen with extremely fine and dense, transverse microstriation and sparse punctation.
</paragraph>
<paragraph id="8B8B36E3551AFFE0FF1AFDCAFD3BBCE9" blockId="33.[151,1437,151,1149]" pageId="33" pageNumber="34">
<collectingCountry id="F3237673551AFFE0FF1AFDCAFEFBBCA0" box="[199,260,620,645]" name="Malta" pageId="33" pageNumber="34">Male</collectingCountry>
. Temples with fine punctures. Tergite 10 and sternite 9 of the male genital segment as in
<figureCitation id="130F2A66551AFFE0FAD0FDCAFA67BCA1" box="[1293,1432,620,645]" captionStart="FIGURES 70 – 74" captionStartId="32.[151,264,1919,1941]" captionTargetBox="[167,1421,211,1883]" captionTargetId="figure@32.[151,1436,195,1897]" captionTargetPageId="32" captionText="FIGURES 70 – 74. Thyreocephalus meridionalis sp. nov.; 70, head and pronotum; 71, labrum; 72, male tergite 10; 73, male sternite 9; 74, aedeagus." httpUri="https://zenodo.org/record/289893/files/figure.png" pageId="33" pageNumber="34">Figs. 72, 73</figureCitation>
. Aedeagus (
<figureCitation id="130F2A66551AFFE0FEC3FD29FE85BC8D" box="[286,378,655,680]" captionStart="FIGURES 70 – 74" captionStartId="32.[151,264,1919,1941]" captionTargetBox="[167,1421,211,1883]" captionTargetId="figure@32.[151,1436,195,1897]" captionTargetPageId="32" captionText="FIGURES 70 – 74. Thyreocephalus meridionalis sp. nov.; 70, head and pronotum; 71, labrum; 72, male tergite 10; 73, male sternite 9; 74, aedeagus." httpUri="https://zenodo.org/record/289893/files/figure.png" pageId="33" pageNumber="34">Fig. 74</figureCitation>
)
<quantity id="4CCC9B06551AFFE0FE4EFD36FDFEBC82" box="[403,513,656,680]" metricMagnitude="-3" metricUnit="m" metricValue="1.88" pageId="33" pageNumber="34" unit="mm" value="1.88">1.88 mm</quantity>
long, sub-ovoid, with long median lobe of characteristic apex; parameres symmetrical; inner sac shorter than in
<taxonomicName id="4C344D60551AFFE0FD99FD13FD43BCE8" box="[580,700,692,717]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B940EAF1551AFFE0FD99FD13FD43BCE8" box="[580,700,692,717]" italics="true" pageId="33" pageNumber="34">T. alluaudi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8B36E3551AFFE0FF1AFD71FCB8BCCA" blockId="33.[151,1437,151,1149]" box="[199,839,727,752]" pageId="33" pageNumber="34">Female. Temples impunctate or with very fine punctures.</paragraph>
<paragraph id="8B8B36E3551AFFE0FF1AFD5AFD1ABD79" blockId="33.[151,1437,151,1149]" pageId="33" pageNumber="34">
<emphasis id="B940EAF1551AFFE0FF1AFD5AFE32BD30" bold="true" box="[199,461,764,789]" pageId="33" pageNumber="34">Differential diagnosis.</emphasis>
<taxonomicName id="4C344D60551AFFE0FE09FD5AFCE5BD30" box="[468,794,764,789]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="meridionalis" status="sp. nov.">
<emphasis id="B940EAF1551AFFE0FE09FD5AFCE5BD30" box="[468,794,764,789]" italics="true" pageId="33" pageNumber="34">Thyreocephalus meridionalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A273578A551AFFE0FCFDFD5AFC85BD30" box="[800,890,764,789]" pageId="33" pageNumber="34" rank="species">
<emphasis id="B940EAF1551AFFE0FCFDFD5AFC85BD30" bold="true" box="[800,890,764,789]" pageId="33" pageNumber="34">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from the most similar
<taxonomicName id="4C344D60551AFFE0FB1AFD5BFAC0BD30" box="[1223,1343,764,789]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B940EAF1551AFFE0FB1AFD5BFAC0BD30" box="[1223,1343,764,789]" italics="true" pageId="33" pageNumber="34">T. alluaudi</emphasis>
</taxonomicName>
in more robust body, in angular temples, longer pronotum, different shape of labrum with sharply emarginated middle part and narrower aedeagus with symmetrical parameres.
</paragraph>
</subSubSection>
<subSubSection id="C32E6568551AFFE0FF1AFCC1FE7EBDC9" pageId="33" pageNumber="34" type="etymology">
<paragraph id="8B8B36E3551AFFE0FF1AFCC1FEE6BD80" blockId="33.[151,1437,151,1149]" pageId="33" pageNumber="34">
<emphasis id="B940EAF1551AFFE0FF1AFCC1FEB1BDA5" bold="true" box="[199,334,871,896]" pageId="33" pageNumber="34">Etymology.</emphasis>
The specific epithet refers to the Latin
<taxonomicName id="4C344D60551AFFE0FCD4FCC1FC69BDA5" box="[777,918,871,896]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="B940EAF1551AFFE0FCD4FCC1FC69BDA5" box="[777,918,871,896]" italics="true" pageId="33" pageNumber="34">meridionalis</emphasis>
</taxonomicName>
- e (southern), in relation to the distribution of the species.
</paragraph>
<paragraph id="8B8B36E3551AFFE0FF1AFC09FE7EBDC9" blockId="33.[151,1437,151,1149]" pageId="33" pageNumber="34">
<emphasis id="B940EAF1551AFFE0FF1AFC09FEC7BDED" bold="true" box="[199,312,943,968]" pageId="33" pageNumber="34">Bionomy.</emphasis>
Some specimens were collected in mushrooms and in forest litter. One specimen was collected on a tree trunk (
<figureCitation id="130F2A66551AFFE0FECFFC72FE8ABDC9" box="[274,373,980,1005]" captionStart="FIGURES 238 – 240. 238, 239" captionStartId="90.[151,264,1811,1833]" captionTargetBox="[170,1419,213,1777]" captionTargetId="figure@90.[150,1436,193,1790]" captionTargetPageId="90" captionText="FIGURES 238 – 240. 238, 239, Tsingidi, Republic of Congo (type locality of Thyreocephalus tsingidianus sp. nov. and T. pseudoafricanus sp. nov.); 238, forest; 239, pitfall traps; 240, Mariepskop, South Africa (type locality of T. meridionalis sp. nov.) — arrow indicates a place where the specimen was collected." httpUri="https://zenodo.org/record/289931/files/figure.png" pageId="33" pageNumber="34">Fig. 240</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E6568551AFFE0FF1AFC51FE80BA10" pageId="33" pageNumber="34" type="distribution">
<paragraph id="8B8B36E3551AFFE0FF1AFC51FE80BA10" blockId="33.[151,1437,151,1149]" pageId="33" pageNumber="34">
<emphasis id="B940EAF1551AFFE0FF1AFC51FE9FBA35" bold="true" box="[199,352,1015,1040]" pageId="33" pageNumber="34">Distribution.</emphasis>
The new species is distributed in
<collectingCountry id="F3237673551AFFE0FD38FC5EFCA9BA2A" box="[741,854,1016,1039]" name="Botswana" pageId="33" pageNumber="34">Botswana</collectingCountry>
,
<collectingCountry id="F3237673551AFFE0FCBEFC51FC05BA35" box="[867,1018,1015,1040]" name="Mozambique" pageId="33" pageNumber="34">Mozambique</collectingCountry>
,
<collectingCountry id="F3237673551AFFE0FBDBFC51FB96BA35" box="[1030,1129,1015,1040]" name="Namibia" pageId="33" pageNumber="34">Namibia</collectingCountry>
,
<collectingCountry id="F3237673551AFFE0FBABFC51FAF6BA35" box="[1142,1289,1015,1040]" name="South Africa" pageId="33" pageNumber="34">South Africa</collectingCountry>
,
<collectingCountry id="F3237673551AFFE0FACBFC51FA94BA35" box="[1302,1387,1015,1040]" name="Zambia" pageId="33" pageNumber="34">Zambia</collectingCountry>
and
<collectingCountry id="F3237673551AFFE0FF4AFBBAFEF0BA10" box="[151,271,1052,1077]" name="Zimbabwe" pageId="33" pageNumber="34">Zimbabwe</collectingCountry>
(
<figureCitation id="130F2A66551AFFE0FEC3FBBAFE8DBA10" box="[286,370,1052,1077]" captionStart="FIGURE 64" captionStartId="29.[151,250,1745,1767]" captionTargetBox="[152,1432,345,1724]" captionTargetId="figure@29.[151,1436,342,1724]" captionTargetPageId="29" captionText="FIGURE 64. Distribution of species in Thyreocephalus ater group (● = T. ater (Laporte), ▲ = T. alluaudi (Fauvel), Δ = T. meridionalis sp. nov.)." httpUri="https://zenodo.org/record/289891/files/figure.png" pageId="33" pageNumber="34">Fig. 64</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32E6568551AFFE0FF1AFB99FB85BA59" pageId="33" pageNumber="34" type="discussion">
<paragraph id="8B8B36E3551AFFE0FF1AFB99FB85BA59" blockId="33.[151,1437,151,1149]" pageId="33" pageNumber="34">
<emphasis id="B940EAF1551AFFE0FF1AFB99FEC4BA7D" bold="true" box="[199,315,1087,1112]" pageId="33" pageNumber="34">Remarks.</emphasis>
A specimen from
<collectingCountry id="F3237673551AFFE0FDD6FB99FD79BA7D" box="[523,646,1087,1112]" name="Zimbabwe" pageId="33" pageNumber="34">Zimbabwe</collectingCountry>
, Shangani has the same colorations as
<taxonomicName id="4C344D60551AFFE0FBE5FBE7FB4EBA7D" box="[1080,1201,1087,1112]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="alluaudi">
<emphasis id="B940EAF1551AFFE0FBE5FBE7FB4EBA7D" box="[1080,1201,1087,1112]" italics="true" pageId="33" pageNumber="34">T. alluaudi</emphasis>
</taxonomicName>
, but other characters such as shape of the head, pronotum, labrum and aedeagus are typical of
<taxonomicName id="4C344D60551AFFE0FC14FBC3FB8DBA58" box="[969,1138,1124,1149]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="meridionalis">
<emphasis id="B940EAF1551AFFE0FC14FBC3FB8DBA58" box="[969,1138,1124,1149]" italics="true" pageId="33" pageNumber="34">T. meridionalis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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