470 lines
52 KiB
XML
470 lines
52 KiB
XML
<document id="59DE28BE06BD8340D0B33C230453AADF" ID-DOI="10.1111/j.1096-3642.2005.00189.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5435260" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630434122956" checkinUser="diego" docAuthor="Daugeron, Christophe & Grootaert, Patrick" docDate="2005" docId="039D7838FFF1FFE53C43DFFDFCADF92C" docLanguage="en" docName="j.1096-3642.2005.00189.x.pdf" docOrigin="Zoological Journal of the Linnean Society 145 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2005.00189.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Empis (Coptophlebia) nahaeoensis Daugeron & Grootaert (Daugeron & Grootaert 2003" docType="treatment" docVersion="4" lastPageNumber="381" masterDocId="FFA40040FFD8FFCF3F28DF3EFFC4FFD9" masterDocTitle="Phylogenetic systematics of the Empis (Coptophlebia) hyalea-group (Insecta: Diptera: Empididae)" masterLastPageNumber="391" masterPageNumber="339" pageNumber="380" updateTime="1699269823662" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="832028611214F092B4472093A0EE84F3">Phylogenetic systematics of the Empis (Coptophlebia) hyalea-group (Insecta: Diptera: Empididae)</mods:title>
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<mods:namePart id="3F60A1CE9AB2EB34B91BD514DAF4C0CD">Daugeron, Christophe</mods:namePart>
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<mods:namePart id="4BBEB88BC8809EC270A343821DE6276E">Grootaert, Patrick</mods:namePart>
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<mods:title id="DEFAABBD10E2461F1D99AED9AC738580">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="2AF53CEDF9A179D3DE61BD211F44E3E2">
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<mods:date id="351785CEA70A6829196585357FBB4923">2005</mods:date>
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<mods:number id="9B9C0BD65D861C00F9FF175E8C7D839A">2005-11-30</mods:number>
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<treatment id="039D7838FFF1FFE53C43DFFDFCADF92C" ID-DOI="http://doi.org/10.5281/zenodo.5490726" ID-Zenodo-Dep="5490726" LSID="urn:lsid:plazi:treatment:039D7838FFF1FFE53C43DFFDFCADF92C" httpUri="http://treatment.plazi.org/id/039D7838FFF1FFE53C43DFFDFCADF92C" lastPageId="42" lastPageNumber="381" pageId="41" pageNumber="380">
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<subSubSection id="C32E9AA5FFF1FFE63C43DFFDFA65FEBC" pageId="41" pageNumber="380" type="nomenclature">
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<paragraph id="8B8BC92EFFF1FFE63C43DFFDFAB5FF00" blockId="41.[875,1393,194,218]" box="[875,1393,194,218]" pageId="41" pageNumber="380">
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<heading id="D0C37E42FFF1FFE63C43DFFDFAB5FF00" box="[875,1393,194,218]" centered="true" fontSize="9" level="2" pageId="41" pageNumber="380" reason="2">
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<taxonomicName id="4C34B2ADFFF1FFE63C43DFFDFAE5FF00" ID-CoL="8BZ6Q" authorityName="Daugeron & Grootaert (Daugeron & Grootaert" authorityYear="2003" box="[875,1313,194,218]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="385" phylum="Arthropoda" rank="species" species="nahaeoensis" subGenus="Coptophlebia">
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<emphasis id="B940153CFFF1FFE63C43DFFDFC77FF00" box="[875,947,195,218]" italics="true" pageId="41" pageNumber="380">EMPIS</emphasis>
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(
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<emphasis id="B940153CFFF1FFE63CECDFFCFBB5FF00" box="[964,1137,194,218]" italics="true" pageId="41" pageNumber="380">COPTOPHLEBIA</emphasis>
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)
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<emphasis id="B940153CFFF1FFE63BAADFF8FAE5FF00" box="[1154,1313,198,217]" italics="true" pageId="41" pageNumber="380">NAHAEOENSIS</emphasis>
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</taxonomicName>
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CLADE
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</heading>
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</paragraph>
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<paragraph id="8B8BC92EFFF1FFE63C12DFCDFA65FEBC" blockId="41.[826,1442,243,357]" pageId="41" pageNumber="380">
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<emphasis id="B940153CFFF1FFE63C12DFCDFC7AFED1" box="[826,958,243,264]" italics="true" pageId="41" pageNumber="380">Monophyly:</emphasis>
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The monophyly of this clade is supported by two characters: epandrial lamellae narrowly separated dorsally [character 26(1)], and epandrium with a regular dorsal row of very strong, long bristles [33(1)].
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E9AA5FFF1FFE53C12DEB2FA4BFEEC" lastPageId="42" lastPageNumber="381" pageId="41" pageNumber="380" type="distribution">
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<paragraph id="8B8BC92EFFF1FFE63C12DEB2FC75FE06" blockId="41.[826,1442,396,479]" pageId="41" pageNumber="380">
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<emphasis id="B940153CFFF1FFE63C12DEB2FBF1FE78" box="[826,1077,396,417]" italics="true" pageId="41" pageNumber="380">
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Distribution (
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<figureCitation id="130FD5ABFFF1FFE63CFFDEB2FBE3FE78" box="[983,1063,396,417]" captionStart="Figure 40" captionStartId="46.[142,220,1290,1309]" captionTargetBox="[375,1188,203,1249]" captionTargetId="figure-290@46.[362,1194,193,1256]" captionTargetPageId="46" captionText="Figure 40. Distribution of the Empis (Coptophlebia) nahaeoensis clade: E. (C.) incurva sp. nov. = 2; E. (C.) kuaensis sp. nov. = 4; E. (C.) nahaeoensis Daugeron & Grootaert = 3; E. (C.) persimilis Frey = 1; E. (C.) pseudonahaeoensis sp. nov. = 3." figureDoi="http://doi.org/10.5281/zenodo.5435346" httpUri="https://zenodo.org/record/5435346/files/figure.png" pageId="41" pageNumber="380">Fig. 40</figureCitation>
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):
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</emphasis>
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This group is recorded from South-east Asia [
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<collectingCountry id="F32389BEFFF1FFE63B3DDE95FB98FE18" box="[1045,1116,427,449]" name="China" pageId="41" pageNumber="380">China</collectingCountry>
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(
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<collectingRegion id="49F007CCFFF1FFE63B5FDE95FB10FE18" box="[1143,1236,427,449]" country="China" name="Yunnan" pageId="41" pageNumber="380">Yunnan</collectingRegion>
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),
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<collectingCountry id="F32389BEFFF1FFE63BDBDE95FAA7FE19" box="[1267,1379,427,448]" name="Myanmar" pageId="41" pageNumber="380">Myanmar</collectingCountry>
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and
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<collectingCountry id="F32389BEFFF1FFE63C12DEF7FC63FE06" box="[826,935,457,479]" name="Thailand" pageId="41" pageNumber="380">Thailand</collectingCountry>
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].
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</paragraph>
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<paragraph id="8B8BC92EFFF1FFE63C12DD39FAC5FD4E" blockId="41.[826,1442,519,663]" pageId="41" pageNumber="380">
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<emphasis id="B940153CFFF1FFE63C12DD39FC62FDC5" box="[826,934,519,540]" italics="true" pageId="41" pageNumber="380">Remarks:</emphasis>
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Even if
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<taxonomicName id="4C34B2ADFFF1FFE63B23DD39FB12FDC5" authority="Frey" authorityName="Frey" authorityYear="1953" box="[1035,1238,519,540]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="380" phylum="Arthropoda" rank="species" species="persimilis">
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<emphasis id="B940153CFFF1FFE63B23DD39FB5EFDC5" box="[1035,1178,519,540]" italics="true" pageId="41" pageNumber="380">E. persimilis</emphasis>
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Frey
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</taxonomicName>
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is not included in the phylogenetic analysis and not reviewed taxonomically, a preliminary morphological study of this species permits its inclusion in the
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<taxonomicName id="4C34B2ADFFF1FFE63BE1DD5DFA66FDA1" authorityName="Daugeron & Grootaert (Daugeron & Grootaert" authorityYear="2003" box="[1225,1442,611,632]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="380" phylum="Arthropoda" rank="species" species="nahaeoensis" subGenus="Coptophlebia">
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<emphasis id="B940153CFFF1FFE63BE1DD5DFB26FDA1" box="[1225,1250,611,632]" italics="true" pageId="41" pageNumber="380">E.</emphasis>
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(
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<emphasis id="B940153CFFF1FFE63BD9DD5DFACDFDA1" box="[1265,1289,611,632]" italics="true" pageId="41" pageNumber="380">C.</emphasis>
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)
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<emphasis id="B940153CFFF1FFE63A30DD5DFA66FDA1" box="[1304,1442,611,632]" italics="true" pageId="41" pageNumber="380">nahaeoensis</emphasis>
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</taxonomicName>
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clade as well as in the key given below.
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</paragraph>
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<table id="F9343B8EFFF100303F8ADDF0FA67FA82" box="[162,1443,718,1371]" pageId="41" pageNumber="380">
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<paragraph id="8B8BC92EFFF1FFE63E16DDDCFA50FCAF" blockId="41.[318,1289,738,762]" lastBlockId="41.[179,1428,779,1356]" pageId="41" pageNumber="380">
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KEY TO SPECIES OF THE
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<taxonomicName id="4C34B2ADFFF1FFE63D72DDDDFBD4FD20" authorityName="Daugeron & Grootaert (Daugeron & Grootaert" authorityYear="2003" box="[602,1040,738,762]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="385" phylum="Arthropoda" rank="species" species="nahaeoensis" subGenus="Coptophlebia">
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<emphasis id="B940153CFFF1FFE63D72DDDDFD67FD20" box="[602,675,739,762]" italics="true" pageId="41" pageNumber="380">EMPIS</emphasis>
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(
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<emphasis id="B940153CFFF1FFE63D9CDDDCFCA4FD20" box="[692,864,738,762]" italics="true" pageId="41" pageNumber="380">COPTOPHLEBIA</emphasis>
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)
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<emphasis id="B940153CFFF1FFE63C59DDD8FBD4FD20" box="[881,1040,742,761]" italics="true" pageId="41" pageNumber="380">NAHAEOENSIS</emphasis>
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</taxonomicName>
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CLADE (MALES ONLY) 1. Fore femur with a strong to spine-like posteroventral bristle apically, cercus with 1 very strong spine-like curved bristle laterally (
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<figureCitation id="130FD5ABFFF1FFE63E74DC16FE68FCE5" box="[348,428,808,828]" captionStart="Figure 37" captionStartId="43.[162,240,1411,1430]" captionTargetBox="[397,1200,201,1376]" captionTargetId="figure-207@43.[384,1216,188,1387]" captionTargetPageId="43" captionText="Figure 37. The Empis (Coptophlebia) nahaeoensis clade, E. (C.) kuaensis sp. nov. male hypopygium in lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435340" httpUri="https://zenodo.org/record/5435340/files/figure.png" pageId="41" pageNumber="380">Figs 37</figureCitation>
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,
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<figureCitation id="130FD5ABFFF1FFE63E90DC17FE17FCE5" box="[440,467,809,828]" captionStart="Figure 39" captionStartId="45.[162,240,919,938]" captionTargetBox="[172,769,203,868]" captionTargetId="figure-608@45.[166,774,195,886]" captionTargetPageId="45" captionText="Figure 39. The Empis (Coptophlebia) nahaeoensis clade, male of E. (C.) pseudonahaeoensis sp. nov. epandrium and cercus in lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435344" httpUri="https://zenodo.org/record/5435344/files/figure.png" pageId="41" pageNumber="380">39</figureCitation>
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;
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<bibRefCitation id="EFA5B4DFFFF1FFE63EF6DC17FCDFFCE5" author="Daugeron C & Grootaert P" box="[478,795,809,828]" pageId="41" pageNumber="380" pagination="167 - 179" refId="ref32630" refString="Daugeron C, Grootaert P. 2003 a. The Empis (Coptophlebia) hyalea - group from Thailand, with a discussion on the world distribution of this species-group (Diptera: Empididae: Empidinae). European Journal of Entomology 100: 167 - 179." type="journal article" year="2003">Daugeron & Grootaert, 2003a</bibRefCitation>
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: fig. 6) ...................................................................................... 2 Fore femur without a strong to spine-like posteroventral bristles apically, cercus without a very strong spine-like curved bristle laterally........................................................................................................................................................ 4
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</paragraph>
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<paragraph id="8B8BC92EFFF1FFE63F9BDCBFFA50FC35" blockId="41.[179,1428,779,1356]" pageId="41" pageNumber="380">
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2. Fore femur with a short anteroventral spine apically, fore tarsomeres 2–5 with short dorsal bristles......................... ..............................................................................................................
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<emphasis id="B940153CFFF1FFE63CA7DCA0FAD9FC68" bold="true" box="[911,1309,925,945]" pageId="41" pageNumber="380">
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<taxonomicName id="4C34B2ADFFF1FFE63CA7DCA0FB79FC69" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[911,1213,925,945]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="380" phylum="Arthropoda" rank="species" species="pseudonahaeoensis" status="sp. nov." subGenus="Coptophlebia">
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<emphasis id="B940153CFFF1FFE63CA7DCA0FC62FC68" bold="true" box="[911,934,926,945]" italics="true" pageId="41" pageNumber="380">E.</emphasis>
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(
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<emphasis id="B940153CFFF1FFE63C9EDCA0FC09FC68" bold="true" box="[950,973,926,945]" italics="true" pageId="41" pageNumber="380">C.</emphasis>
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)
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<emphasis id="B940153CFFF1FFE63CF4DCA3FB79FC69" bold="true" box="[988,1213,925,944]" italics="true" pageId="41" pageNumber="380">pseudonahaeoensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A273A847FFF1FFE63BEEDCA0FAD9FC68" box="[1222,1309,926,945]" pageId="41" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
(
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<collectingCountry id="F32389BEFFF1FFE63A04DCA0FA48FC68" box="[1324,1420,926,945]" name="Thailand" pageId="41" pageNumber="380">Thailand</collectingCountry>
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) Fore femur without a short anteroventral spine apically or with a single bristle, fore tarsomeres 2–5 with long dorsal bristles.................................................................................................................................................................................. 3
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</paragraph>
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<paragraph id="8B8BC92EFFF1FFE63F9BDCC8FA56FB0F" blockId="41.[179,1428,779,1356]" pageId="41" pageNumber="380">
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3. Fore tarsomeres 2–5 more or less rounded, with numerous dorsal bristles, anterodorsal rounded projection of epandrium without short spine-like apically rounded spines, epandrium with a long apical prolongation bearing
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<emphasis id="B940153CFFF1FFE63A4CDB2AFAB0FBFF" box="[1380,1396,1044,1062]" italics="true" pageId="41" pageNumber="380">c.</emphasis>
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15 strong, long bristles, with a second fan of strong, long bristles ventrally (
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<figureCitation id="130FD5ABFFF1FFE63CFCDB0EFBDAFB9D" box="[980,1054,1072,1092]" captionStart="Figure 37" captionStartId="43.[162,240,1411,1430]" captionTargetBox="[397,1200,201,1376]" captionTargetId="figure-207@43.[384,1216,188,1387]" captionTargetPageId="43" captionText="Figure 37. The Empis (Coptophlebia) nahaeoensis clade, E. (C.) kuaensis sp. nov. male hypopygium in lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435340" httpUri="https://zenodo.org/record/5435340/files/figure.png" pageId="41" pageNumber="380">Fig. 37</figureCitation>
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) ......................................................... ..................................................................................................................................
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<emphasis id="B940153CFFF1FFE63B21DB70FAD9FBB8" bold="true" box="[1033,1309,1101,1121]" pageId="41" pageNumber="380">
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||
<taxonomicName id="4C34B2ADFFF1FFE63B21DB70FB79FBB9" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[1033,1213,1101,1121]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="380" phylum="Arthropoda" rank="species" species="kuaensis" status="sp. nov." subGenus="Coptophlebia">
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<emphasis id="B940153CFFF1FFE63B21DB70FBE4FBB8" bold="true" box="[1033,1056,1102,1121]" italics="true" pageId="41" pageNumber="380">E.</emphasis>
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(
|
||
<emphasis id="B940153CFFF1FFE63B18DB70FB83FBB8" bold="true" box="[1072,1095,1102,1121]" italics="true" pageId="41" pageNumber="380">C.</emphasis>
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)
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<emphasis id="B940153CFFF1FFE63B7EDB73FB79FBB9" bold="true" box="[1110,1213,1101,1120]" italics="true" pageId="41" pageNumber="380">kuaensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A273A847FFF1FFE63BEEDB70FAD9FBB8" box="[1222,1309,1102,1121]" pageId="41" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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(
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<collectingCountry id="F32389BEFFF1FFE63A04DB70FA48FBB8" box="[1324,1420,1102,1121]" name="Thailand" pageId="41" pageNumber="380">Thailand</collectingCountry>
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) Fore tarsomeres 2–5 not rounded, with dorsal bristles less numerous, anterodorsal rounded projection of epandrium with 2 short spine-like apically rounded spines, epandrium with a short apical projection bearing
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<emphasis id="B940153CFFF1FFE63BC9DBB7FB35FB42" box="[1249,1265,1161,1179]" italics="true" pageId="41" pageNumber="380">c.</emphasis>
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10 strong, long bristles, without a second fan of bristles ventrally (
|
||
<bibRefCitation id="EFA5B4DFFFF1FFE63DC7DB98FBE9FB60" author="Daugeron C & Grootaert P" box="[751,1069,1190,1209]" pageId="41" pageNumber="380" pagination="167 - 179" refId="ref32630" refString="Daugeron C, Grootaert P. 2003 a. The Empis (Coptophlebia) hyalea - group from Thailand, with a discussion on the world distribution of this species-group (Diptera: Empididae: Empidinae). European Journal of Entomology 100: 167 - 179." type="journal article" year="2003">Daugeron & Grootaert, 2003a</bibRefCitation>
|
||
: fig. 6).............................................
|
||
<emphasis id="B940153CFFF1FFE63FFDDBFAFC9AFB0F" box="[213,862,1220,1238]" italics="true" pageId="41" pageNumber="380">.......................................................................................................</emphasis>
|
||
<taxonomicName id="4C34B2ADFFF1FFE63C48DBFAFA56FB0F" authority="Daugeron & Grootaert" authorityName="Daugeron & Grootaert (Daugeron & Grootaert" authorityYear="2003" box="[864,1426,1219,1239]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="380" phylum="Arthropoda" rank="species" species="nahaeoensis" subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFF1FFE63C48DBFAFCB3FB0F" box="[864,887,1220,1238]" italics="true" pageId="41" pageNumber="380">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFF1FFE63CACDBFDFC5FFB0F" box="[900,923,1219,1238]" italics="true" pageId="41" pageNumber="380">C.</emphasis>
|
||
)
|
||
<emphasis id="B940153CFFF1FFE63C80DBFDFBE3FB0F" box="[936,1063,1219,1238]" italics="true" pageId="41" pageNumber="380">nahaeoensis</emphasis>
|
||
Daugeron & Grootaert (
|
||
<collectingCountry id="F32389BEFFF1FFE63A03DBFDFA4FFB0F" box="[1323,1419,1219,1238]" name="Thailand" pageId="41" pageNumber="380">Thailand</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8B8BC92EFFF1FFE63F9BDBDFFA57FA95" blockId="41.[179,1428,779,1356]" pageId="41" pageNumber="380">
|
||
4. Phallus bent at right angle apically (
|
||
<figureCitation id="130FD5ABFFF1FFE63D72DBDEFD72FB2A" box="[602,694,1248,1267]" captionStart="Figure 36" captionStartId="42.[142,220,1075,1094]" captionTargetBox="[153,747,202,1039]" captionTargetId="figure-624@42.[144,752,185,1054]" captionTargetPageId="42" captionText="Figure 36. The Empis (Coptophlebia) nahaeoensis clade, male of E. (C.) incurva sp. nov. A, epandrium and cercus in lateral view; B, hypandrium and phallus in lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435338" httpUri="https://zenodo.org/record/5435338/files/figure.png" pageId="41" pageNumber="380">Fig. 36B</figureCitation>
|
||
), epandrium with 2 anterodorsal brushes of very strong, rather long, apically rounded bristles (
|
||
<figureCitation id="130FD5ABFFF1FFE63ECEDBC0FD84FAC8" box="[486,576,1278,1297]" captionStart="Figure 36" captionStartId="42.[142,220,1075,1094]" captionTargetBox="[153,747,202,1039]" captionTargetId="figure-624@42.[144,752,185,1054]" captionTargetPageId="42" captionText="Figure 36. The Empis (Coptophlebia) nahaeoensis clade, male of E. (C.) incurva sp. nov. A, epandrium and cercus in lateral view; B, hypandrium and phallus in lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435338" httpUri="https://zenodo.org/record/5435338/files/figure.png" pageId="41" pageNumber="380">Fig. 36A</figureCitation>
|
||
).............................................................................
|
||
<emphasis id="B940153CFFF1FFE63B1CDBC0FAF9FAC8" bold="true" box="[1076,1341,1278,1297]" pageId="41" pageNumber="380">
|
||
<taxonomicName id="4C34B2ADFFF1FFE63B1CDBC0FB19FAC8" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[1076,1245,1278,1297]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="380" phylum="Arthropoda" rank="species" species="incurva" status="sp. nov." subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFF1FFE63B1CDBC0FB8FFAC8" bold="true" box="[1076,1099,1278,1297]" italics="true" pageId="41" pageNumber="380">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFF1FFE63B73DBC0FBB6FAC8" bold="true" box="[1115,1138,1278,1297]" italics="true" pageId="41" pageNumber="380">C.</emphasis>
|
||
)
|
||
<emphasis id="B940153CFFF1FFE63BAADBC0FB19FAC8" bold="true" box="[1154,1245,1278,1297]" italics="true" pageId="41" pageNumber="380">incurva</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A273A847FFF1FFE63BCDDBC0FAF9FAC8" box="[1253,1341,1278,1297]" pageId="41" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<collectingCountry id="F32389BEFFF1FFE63A63DBC0FA48FAC8" box="[1355,1420,1278,1297]" name="China" pageId="41" pageNumber="380">China</collectingCountry>
|
||
) Phallus long, slender, not bent, epandrium with 2 dorsal single protuberances bearing distinct bristles
|
||
<emphasis id="B940153CFFF1FFE63FFDDA07FBE3FA92" box="[213,1063,1337,1355]" italics="true" pageId="41" pageNumber="380">.......................................................................................................................................</emphasis>
|
||
<taxonomicName id="4C34B2ADFFF1FFE63B06DA07FA57FA95" authority="Frey" authorityName="Frey" authorityYear="1953" box="[1070,1427,1336,1356]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="380" phylum="Arthropoda" rank="species" species="persimilis" subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFF1FFE63B06DA07FB81FA92" box="[1070,1093,1337,1355]" italics="true" pageId="41" pageNumber="380">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFF1FFE63B7ADA07FBADFA95" box="[1106,1129,1337,1356]" italics="true" pageId="41" pageNumber="380">C.</emphasis>
|
||
)
|
||
<emphasis id="B940153CFFF1FFE63B5EDA06FB19FA92" box="[1142,1245,1336,1355]" italics="true" pageId="41" pageNumber="380">persimilis</emphasis>
|
||
Frey (
|
||
<collectingCountry id="F32389BEFFF1FFE63A0CDA07FA48FA95" box="[1316,1420,1337,1356]" name="Myanmar" pageId="41" pageNumber="380">Myanmar</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</table>
|
||
<caption id="DF4B99A6FFF1FFE63F8AD9D8FED1F8B7" ID-DOI="http://doi.org/10.5281/zenodo.5435336" ID-Zenodo-Dep="5435336" httpUri="https://zenodo.org/record/5435336/files/figure.png" pageId="41" pageNumber="380" startId="41.[162,240,1766,1785]" targetBox="[333,1270,1448,1733]" targetPageId="41">
|
||
<paragraph id="8B8BC92EFFF1FFE63F8AD9D8FED1F8B7" blockId="41.[162,1442,1766,1902]" pageId="41" pageNumber="380">
|
||
<emphasis id="B940153CFFF1FFE63F8AD9D8FED3F920" bold="true" box="[162,279,1766,1785]" pageId="41" pageNumber="380">Figure 35.</emphasis>
|
||
Distribution of the
|
||
<taxonomicName id="4C34B2ADFFF1FFE63D09D9D8FCAFF920" authorityName="De Meijere" authorityYear="1907" box="[545,875,1766,1785]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="380" phylum="Arthropoda" rank="species" species="jacobsoni" subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFF1FFE63D09D9D8FDA0F921" box="[545,612,1766,1784]" italics="true" pageId="41" pageNumber="380">Empis</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFF1FFE63D5BD9D8FD39F920" box="[627,765,1766,1785]" italics="true" pageId="41" pageNumber="380">Coptophlebia</emphasis>
|
||
)
|
||
<emphasis id="B940153CFFF1FFE63C22D9D8FCAFF920" box="[778,875,1766,1785]" italics="true" pageId="41" pageNumber="380">jacobsoni</emphasis>
|
||
</taxonomicName>
|
||
clade:
|
||
<taxonomicName id="4C34B2ADFFF1FFE63CFFD9D8FAD9F920" authority="De Meijere" authorityName="De Meijere" authorityYear="1907" box="[983,1309,1766,1785]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="380" phylum="Arthropoda" rank="species" species="jacobsoni" subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFF1FFE63CFFD9D8FC2BF921" box="[983,1007,1766,1784]" italics="true" pageId="41" pageNumber="380">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFF1FFE63CD4D9D8FBD6F920" box="[1020,1042,1766,1785]" italics="true" pageId="41" pageNumber="380">C.</emphasis>
|
||
)
|
||
<emphasis id="B940153CFFF1FFE63B37D9D8FB44F920" box="[1055,1152,1766,1785]" italics="true" pageId="41" pageNumber="380">jacobsoni</emphasis>
|
||
De Meijere
|
||
</taxonomicName>
|
||
= 6 (Java);
|
||
<taxonomicName id="4C34B2ADFFF1FFE63F8AD83DFE9CF8CF" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[162,344,1795,1814]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="380" phylum="Arthropoda" rank="species" species="lachaisei" status="sp. nov." subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFF1FFE63F8AD83DFF7DF8CF" bold="true" box="[162,185,1795,1814]" italics="true" pageId="41" pageNumber="380">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFF1FFE63FEFD83DFF1BF8CF" bold="true" box="[199,223,1795,1814]" italics="true" pageId="41" pageNumber="380">C.</emphasis>
|
||
) lachaisei
|
||
</taxonomicName>
|
||
<emphasis id="B940153CFFF1FFE63E76D83DFE77F8CF" bold="true" box="[350,435,1795,1814]" pageId="41" pageNumber="380">
|
||
<taxonomicNameLabel id="A273A847FFF1FFE63E76D83DFE77F8CF" box="[350,435,1795,1814]" pageId="41" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
= 1 (Ivory Coast);
|
||
<taxonomicName id="4C34B2ADFFF1FFE63D59D83DFD2AF8CF" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[625,750,1795,1814]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="380" phylum="Arthropoda" rank="species" species="loici" status="sp. nov." subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFF1FFE63D59D83DFD4CF8CF" bold="true" box="[625,648,1795,1814]" italics="true" pageId="41" pageNumber="380">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFF1FFE63DBDD83DFD68F8CF" bold="true" box="[661,684,1795,1814]" italics="true" pageId="41" pageNumber="380">C.</emphasis>
|
||
) loici
|
||
</taxonomicName>
|
||
<emphasis id="B940153CFFF1FFE63DDBD83DFC8DF8CF" bold="true" box="[755,841,1795,1814]" pageId="41" pageNumber="380">
|
||
<taxonomicNameLabel id="A273A847FFF1FFE63DDBD83DFC8DF8CF" box="[755,841,1795,1814]" pageId="41" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
= 2 (Central African Republic);
|
||
<taxonomicName id="4C34B2ADFFF1FFE63BBAD83DFABFF8CF" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[1170,1403,1795,1814]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="380" phylum="Arthropoda" rank="species" species="menglunensis" status="sp. nov." subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFF1FFE63BBAD83DFB6DF8CF" bold="true" box="[1170,1193,1795,1814]" italics="true" pageId="41" pageNumber="380">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFF1FFE63B9ED83DFB0AF8CF" bold="true" box="[1206,1230,1795,1814]" italics="true" pageId="41" pageNumber="380">C.</emphasis>
|
||
) menglunensis
|
||
</taxonomicName>
|
||
<emphasis id="B940153CFFF1FFE63AA9D83DFF16F8EA" bold="true" pageId="41" pageNumber="380">
|
||
<taxonomicNameLabel id="A273A847FFF1FFE63AA9D83DFF16F8EA" pageId="41" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
= 3 (China);
|
||
<taxonomicName id="4C34B2ADFFF1FFE63E7FD81FFD95F8EA" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[343,593,1824,1844]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="380" phylum="Arthropoda" rank="species" species="mengyangensis" status="sp. nov." subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFF1FFE63E7FD81FFEAAF8ED" bold="true" box="[343,366,1825,1844]" italics="true" pageId="41" pageNumber="380">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFF1FFE63E53D81EFE56F8EA" bold="true" box="[379,402,1824,1843]" italics="true" pageId="41" pageNumber="380">C.</emphasis>
|
||
) mengyangensis
|
||
</taxonomicName>
|
||
<emphasis id="B940153CFFF1FFE63D7ED81EFD68F8EA" bold="true" box="[598,684,1824,1843]" pageId="41" pageNumber="380">
|
||
<taxonomicNameLabel id="A273A847FFF1FFE63D7ED81EFD68F8EA" box="[598,684,1824,1843]" pageId="41" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
= 3 (China);
|
||
<taxonomicName id="4C34B2ADFFF1FFE63C19D81FFBE7F8EA" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[817,1059,1824,1844]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="380" phylum="Arthropoda" rank="species" species="cameronsensis" status="sp. nov." subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFF1FFE63C19D81FFC8CF8ED" bold="true" box="[817,840,1825,1844]" italics="true" pageId="41" pageNumber="380">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFF1FFE63C7DD81EFCA8F8EA" bold="true" box="[853,876,1824,1843]" italics="true" pageId="41" pageNumber="380">C.</emphasis>
|
||
) cameronsensis
|
||
</taxonomicName>
|
||
<emphasis id="B940153CFFF1FFE63B01D81EFBBAF8EA" bold="true" box="[1065,1150,1824,1843]" pageId="41" pageNumber="380">
|
||
<taxonomicNameLabel id="A273A847FFF1FFE63B01D81EFBBAF8EA" box="[1065,1150,1824,1843]" pageId="41" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
= 4 (Malaysia);
|
||
<taxonomicName id="4C34B2ADFFF1FFE63A0CD81FFF3DF889" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="380" phylum="Arthropoda" rank="species" species="nimbaensis" status="sp. nov." subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFF1FFE63A0CD81FFAFFF8ED" bold="true" box="[1316,1339,1825,1844]" italics="true" pageId="41" pageNumber="380">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFF1FFE63A60D81EFA9BF8EA" bold="true" box="[1352,1375,1824,1843]" italics="true" pageId="41" pageNumber="380">C.</emphasis>
|
||
) nimbaensis
|
||
</taxonomicName>
|
||
<emphasis id="B940153CFFF1FFE63E2CD800FEA4F888" bold="true" box="[260,352,1854,1873]" pageId="41" pageNumber="380">
|
||
<taxonomicNameLabel id="A273A847FFF1FFE63E2CD800FEA4F888" box="[260,352,1854,1873]" pageId="41" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
= 1 (Guinea);
|
||
<taxonomicName id="4C34B2ADFFF1FFE63ED5D800FCFDF888" authority="Daugeron" authorityName="Daugeron" authorityYear="2002" box="[509,825,1854,1873]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="380" phylum="Arthropoda" rank="species" species="nondouensis" subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFF1FFE63ED5D800FDD0F889" box="[509,532,1854,1872]" italics="true" pageId="41" pageNumber="380">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFF1FFE63D09D800FDFCF888" box="[545,568,1854,1873]" italics="true" pageId="41" pageNumber="380">C.</emphasis>
|
||
)
|
||
<emphasis id="B940153CFFF1FFE63D6DD800FD03F888" box="[581,711,1854,1873]" italics="true" pageId="41" pageNumber="380">nondouensis</emphasis>
|
||
Daugeron
|
||
</taxonomicName>
|
||
= 7 (New Caledonia);
|
||
<taxonomicName id="4C34B2ADFFF1FFE63B06D800FAD2F889" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[1070,1302,1853,1873]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="41" pageNumber="380" phylum="Arthropoda" rank="species" species="padangensis" status="sp. nov." subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFF1FFE63B06D800FB81F888" bold="true" box="[1070,1093,1854,1873]" italics="true" pageId="41" pageNumber="380">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFF1FFE63B70D800FBABF888" bold="true" box="[1112,1135,1854,1873]" italics="true" pageId="41" pageNumber="380">C.</emphasis>
|
||
) padangensis
|
||
</taxonomicName>
|
||
<emphasis id="B940153CFFF1FFE63A09D800FAB9F888" bold="true" box="[1313,1405,1854,1873]" pageId="41" pageNumber="380">
|
||
<taxonomicNameLabel id="A273A847FFF1FFE63A09D800FAB9F888" box="[1313,1405,1854,1873]" pageId="41" pageNumber="380" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
= 5 (Sumatra).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4B99A6FFF2FFE53FA6DB0DFE9BFB46" ID-DOI="http://doi.org/10.5281/zenodo.5435338" ID-Zenodo-Dep="5435338" httpUri="https://zenodo.org/record/5435338/files/figure.png" pageId="42" pageNumber="381" startId="42.[142,220,1075,1094]" targetBox="[153,747,202,1039]" targetPageId="42">
|
||
<paragraph id="8B8BC92EFFF2FFE53FA6DB0DFE9BFB46" blockId="42.[142,759,1075,1183]" pageId="42" pageNumber="381">
|
||
<emphasis id="B940153CFFF2FFE53FA6DB0DFEC7FB9F" bold="true" box="[142,259,1075,1094]" pageId="42" pageNumber="381">Figure 36.</emphasis>
|
||
The
|
||
<taxonomicName id="4C34B2ADFFF2FFE53E69DB0AFD74FB9F" authorityName="Daugeron & Grootaert (Daugeron & Grootaert" authorityYear="2003" box="[321,688,1075,1095]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="381" phylum="Arthropoda" rank="species" species="nahaeoensis" subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFF2FFE53E69DB0AFE40FB9F" box="[321,388,1076,1094]" italics="true" pageId="42" pageNumber="381">Empis</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFF2FFE53EBEDB0DFDE4FB9F" box="[406,544,1075,1094]" italics="true" pageId="42" pageNumber="381">Coptophlebia</emphasis>
|
||
)
|
||
<emphasis id="B940153CFFF2FFE53D19DB0DFD74FB9F" box="[561,688,1075,1094]" italics="true" pageId="42" pageNumber="381">nahaeoensis</emphasis>
|
||
</taxonomicName>
|
||
clade, male of
|
||
<taxonomicName id="4C34B2ADFFF2FFE53FCADB6FFE4DFBBD" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[226,393,1105,1124]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="381" phylum="Arthropoda" rank="species" species="incurva" status="sp. nov." subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFF2FFE53FCADB6FFF3DFBBD" bold="true" box="[226,249,1105,1124]" italics="true" pageId="42" pageNumber="381">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFF2FFE53E20DB6FFEDBFBBD" bold="true" box="[264,287,1105,1124]" italics="true" pageId="42" pageNumber="381">C.</emphasis>
|
||
) incurva
|
||
</taxonomicName>
|
||
<emphasis id="B940153CFFF2FFE53EB9DB6FFE2CFBBD" bold="true" box="[401,488,1105,1124]" pageId="42" pageNumber="381">
|
||
<taxonomicNameLabel id="A273A847FFF2FFE53EB9DB6FFE2CFBBD" box="[401,488,1105,1124]" pageId="42" pageNumber="381" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A, epandrium and cercus in lateral view; B, hypandrium and phallus in lateral view. Scale bar = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8BC92EFFF2FFE53FEDDBDFFD7BFB20" blockId="42.[197,703,1249,1305]" box="[197,703,1249,1273]" pageId="42" pageNumber="381">
|
||
<taxonomicName id="4C34B2ADFFF2FFE53FEDDBDFFE25FB20" authority="" box="[197,481,1249,1273]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="42" pageNumber="387" phylum="Arthropoda" rank="subGenus" subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFF2FFE53FEDDBDFFED5FB21" bold="true" box="[197,273,1249,1273]" italics="true" pageId="42" pageNumber="381">EMPIS</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFF2FFE53E09DBDFFE1DFB21" bold="true" box="[289,473,1249,1273]" italics="true" pageId="42" pageNumber="381">COPTOPHLEBIA</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicName id="4C34B2ADFFF2FFE53EC2DBDBFD97FB21" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[490,595,1253,1272]" pageId="42" pageNumber="381" rank="species" species="incurva" status="sp. nov.">INCURVA</taxonomicName>
|
||
<emphasis id="B940153CFFF2FFE53D73DBDBFD7BFB20" bold="true" box="[603,703,1249,1273]" pageId="42" pageNumber="381">
|
||
<taxonomicNameLabel id="A273A847FFF2FFE53D73DBDBFD7BFB20" box="[603,703,1249,1273]" pageId="42" pageNumber="381" rank="species">SP. NOV.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B8BC92EFFF2FFE53EB8DA3FFE30FAC0" blockId="42.[197,703,1249,1305]" box="[400,500,1281,1305]" pageId="42" pageNumber="381">
|
||
(
|
||
<figureCitation id="130FD5ABFFF2FFE53EB0DA3FFE28FAC0" box="[408,492,1281,1305]" captionStart="Figure 36" captionStartId="42.[142,220,1075,1094]" captionTargetBox="[153,747,202,1039]" captionTargetId="figure-624@42.[144,752,185,1054]" captionTargetPageId="42" captionText="Figure 36. The Empis (Coptophlebia) nahaeoensis clade, male of E. (C.) incurva sp. nov. A, epandrium and cercus in lateral view; B, hypandrium and phallus in lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435338" httpUri="https://zenodo.org/record/5435338/files/figure.png" pageId="42" pageNumber="381">FIG. 36</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B8BC92EFFF2FFE53FA6DA0DFDCEF9D9" blockId="42.[142,758,1330,1536]" pageId="42" pageNumber="381">
|
||
<emphasis id="B940153CFFF2FFE53FA6DA0DFEF7FA9E" box="[142,307,1330,1352]" italics="true" pageId="42" pageNumber="381">Type material:</emphasis>
|
||
<materialsCitation id="3B5CC373FFF2FFE53E68DA0CFD35FA5C" collectingDate="1999-03-08" collectionCode="KBIN" collectorName="P. Grootaert" country="China" location="Mengyang" municipality="Jinhong" pageId="42" pageNumber="381" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
|
||
<typeStatus id="548F778CFFF2FFE53E68DA0CFE6DFA91" box="[320,425,1330,1352]" pageId="42" pageNumber="381" type="holotype">Holotype</typeStatus>
|
||
: male,
|
||
<collectingCountry id="F32389BEFFF2FFE53ED7DA0CFD82FA91" box="[511,582,1330,1352]" name="China" pageId="42" pageNumber="381">China</collectingCountry>
|
||
,
|
||
<collectingRegion id="49F007CCFFF2FFE53D7DDA0CFF7AFABE" country="China" name="Yunnan" pageId="42" pageNumber="381">Yunnan Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BEF5354FFF2FFE53FFADA6FFEF4FABE" box="[210,304,1361,1383]" pageId="42" pageNumber="381">Jinhong</collectingMunicipality>
|
||
,
|
||
<location id="8EEB9FF5FFF2FFE53E6CDA6FFE79FABF" LSID="urn:lsid:plazi:treatment:039D7838FFF1FFE53C43DFFDFCADF92C:8EEB9FF5FFF2FFE53E6CDA6FFE79FABF" box="[324,445,1361,1382]" country="China" municipality="Jinhong" name="Mengyang" pageId="42" pageNumber="381" stateProvince="Yunnan">Mengyang</location>
|
||
,
|
||
<date id="FF8AEFEEFFF2FFE53EF9DA6FFDF8FABE" box="[465,572,1361,1383]" pageId="42" pageNumber="381" value="1999-03-08">
|
||
<collectingDate id="EFCE1606FFF2FFE53EF9DA6FFDF8FABE" box="[465,572,1361,1383]" pageId="42" pageNumber="381" value="1999-03-08">8.iii.1999</collectingDate>
|
||
</date>
|
||
, primary rainforest,
|
||
<collectorName id="26C1ACF8FFF2FFE53FC9DA51FEA0FA5D" box="[225,356,1391,1412]" pageId="42" pageNumber="381">P.Grootaert</collectorName>
|
||
(sample 99060) (
|
||
<collectionCode id="ED2551EBFFF2FFE53D1FDA51FDB8FA5C" box="[567,636,1391,1413]" pageId="42" pageNumber="381">KBIN</collectionCode>
|
||
–IRScNB)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B5CC373FFF2FFE53FA6DAB0FD35FA7D" box="[142,753,1422,1444]" collectingDate="1999-03-08" collectionCode="KBIN" collectorName="P. Grootaert" country="China" location="Mengyang" municipality="Jinhong" pageId="42" pageNumber="381" specimenCount="9" specimenCount-female="1" specimenCount-male="8" stateProvince="Yunnan" typeStatus="paratype">
|
||
<typeStatus id="548F778CFFF2FFE53FA6DAB0FEC0FA7A" box="[142,260,1422,1443]" pageId="42" pageNumber="381" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D3202A7FFF2FFE53E38DAB0FEA8FA7D" box="[272,364,1422,1444]" pageId="42" pageNumber="381" type="male">8 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D3202A7FFF2FFE53E51DAB0FE19FA7D" box="[377,477,1422,1444]" pageId="42" pageNumber="381" type="female">1 female</specimenCount>
|
||
, same data as holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B5CC373FFF2FFE53FA6DA93FE47FA1A" box="[142,387,1453,1475]" collectingDate="1999-03-08" collectionCode="KBIN" collectorName="P. Grootaert" country="China" location="Mengyang" municipality="Jinhong" pageId="42" pageNumber="381" specimenCount="17" specimenCount-male="17" stateProvince="Yunnan" typeStatus="paratype">
|
||
<specimenCount id="9D3202A7FFF2FFE53FA6DA93FF3EFA1A" box="[142,250,1453,1475]" pageId="42" pageNumber="381" type="male">17 males</specimenCount>
|
||
, same data
|
||
</materialsCitation>
|
||
<materialsCitation id="3B5CC373FFF2FFE53EA6DA93FE8AFA38" collectingDate="1999-03-11" collectionCode="KBIN" country="China" pageId="42" pageNumber="381" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
|
||
but
|
||
<date id="FF8AEFEEFFF2FFE53EE8DA93FDF3FA1A" box="[448,567,1453,1475]" pageId="42" pageNumber="381" value="1999-03-11">
|
||
<collectingDate id="EFCE1606FFF2FFE53EE8DA93FDF3FA1A" box="[448,567,1453,1475]" pageId="42" pageNumber="381" value="1999-03-11">11.iii.1999</collectingDate>
|
||
</date>
|
||
(sample 99090) (
|
||
<collectionCode id="ED2551EBFFF2FFE53FBDDAF5FF1DFA38" box="[149,217,1483,1505]" pageId="42" pageNumber="381">KBIN</collectionCode>
|
||
–IRScNB)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B5CC373FFF2FFE53E77DAF5FDFAFA38" box="[351,574,1483,1505]" collectingDate="1999-03-11" collectionCode="KBIN" country="China" pageId="42" pageNumber="381" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
|
||
<specimenCount id="9D3202A7FFF2FFE53E77DAF5FE76FA38" box="[351,434,1483,1505]" pageId="42" pageNumber="381" type="male">1 male</specimenCount>
|
||
, same data
|
||
</materialsCitation>
|
||
<materialsCitation id="3B5CC373FFF2FFE53D63DAF5FDC2F9D9" collectingDate="1999-03-11" collectionCode="KBIN" country="China" pageId="42" pageNumber="381" specimenCount="1" stateProvince="Yunnan" typeStatus="paratype">
|
||
but
|
||
<date id="FF8AEFEEFFF2FFE53D56DAF5FD31FA38" box="[638,757,1483,1505]" pageId="42" pageNumber="381" value="1999-03-11">
|
||
<collectingDate id="EFCE1606FFF2FFE53D56DAF5FD31FA38" box="[638,757,1483,1505]" pageId="42" pageNumber="381" value="1999-03-11">11.iii.1999</collectingDate>
|
||
</date>
|
||
(sample 99091) (
|
||
<collectionCode id="ED2551EBFFF2FFE53E64DAD4FE55F9D9" box="[332,401,1514,1536]" pageId="42" pageNumber="381">KBIN</collectionCode>
|
||
–IRScNB)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8BC92EFFF2FFE53FA6D919FD4EF92C" blockId="42.[142,758,1575,1781]" pageId="42" pageNumber="381">
|
||
<materialsCitation id="3B5CC373FFF2FFE53FA6D919FD41F92C" collectingDate="1999-03-08" collectingDateMax="1999-03-11" collectingDateMin="1999-03-08" collectionCode="KBIN" collectorName="P. Grootaert" country="China" county="Because" location="Mengyang" municipality="Jinhong" pageId="42" pageNumber="381" specimenCount="6" specimenCount-male="6" stateProvince="Yunnan" typeStatus="paratype">
|
||
<emphasis id="B940153CFFF2FFE53FA6D919FEFBF9E5" box="[142,319,1575,1596]" italics="true" pageId="42" pageNumber="381">Other material:</emphasis>
|
||
<collectingCounty id="62EAB1A2FFF2FFE53E64D916FE6EF9E4" box="[332,426,1576,1597]" pageId="42" pageNumber="381">Because</collectingCounty>
|
||
their abdomen is missing or genitalia are in poor condition, the following male specimens are not considered as
|
||
<typeStatus id="548F778CFFF2FFE53D3AD95BFD43F9A3" box="[530,647,1637,1658]" pageId="42" pageNumber="381" type="paratype">paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D3202A7FFF2FFE53DBDD95BFD35F9A2" box="[661,753,1637,1659]" pageId="42" pageNumber="381" type="male">2 males</specimenCount>
|
||
,
|
||
<collectingCountry id="F32389BEFFF2FFE53FA6D9BDFF11F940" box="[142,213,1667,1689]" name="China" pageId="42" pageNumber="381">China</collectingCountry>
|
||
,
|
||
<collectingRegion id="49F007CCFFF2FFE53FDFD9BAFE10F940" box="[247,468,1667,1689]" country="China" name="Yunnan" pageId="42" pageNumber="381">Yunnan Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6BEF5354FFF2FFE53EDFD9BDFD91F940" box="[503,597,1667,1689]" pageId="42" pageNumber="381">Jinhong</collectingMunicipality>
|
||
,
|
||
<location id="8EEB9FF5FFF2FFE53D50D9BAFD36F940" LSID="urn:lsid:plazi:treatment:039D7838FFF1FFE53C43DFFDFCADF92C:8EEB9FF5FFF2FFE53D50D9BAFD36F940" box="[632,754,1668,1689]" country="China" county="Because" municipality="Jinhong" name="Mengyang" pageId="42" pageNumber="381" stateProvince="Yunnan">Mengyang</location>
|
||
,
|
||
<date id="FF8AEFEEFFF2FFE53FA6D99CFF3DF961" box="[142,249,1698,1720]" pageId="42" pageNumber="381" value="1999-03-08">
|
||
<collectingDate id="EFCE1606FFF2FFE53FA6D99CFF3DF961" box="[142,249,1698,1720]" pageId="42" pageNumber="381" value="1999-03-08">8.iii.1999</collectingDate>
|
||
</date>
|
||
, primary rainforest,
|
||
<collectorName id="26C1ACF8FFF2FFE53D2FD99CFD4EF96E" box="[519,650,1698,1719]" pageId="42" pageNumber="381">P.Grootaert</collectorName>
|
||
(sample 99060) (
|
||
<collectionCode id="ED2551EBFFF2FFE53FDCD9FFFEFCF90E" box="[244,312,1729,1751]" pageId="42" pageNumber="381">KBIN</collectionCode>
|
||
–IRScNB);
|
||
<specimenCount id="9D3202A7FFF2FFE53EEBD9FFFDE3F90E" box="[451,551,1729,1751]" pageId="42" pageNumber="381" type="male">4 males</specimenCount>
|
||
, same data but
|
||
<date id="FF8AEFEEFFF2FFE53FA6D9E1FEC1F92C" box="[142,261,1759,1781]" pageId="42" pageNumber="381" value="1999-03-11">
|
||
<collectingDate id="EFCE1606FFF2FFE53FA6D9E1FEC1F92C" box="[142,261,1759,1781]" pageId="42" pageNumber="381" value="1999-03-11">11.iii.1999</collectingDate>
|
||
</date>
|
||
(sample 99090) (
|
||
<collectionCode id="ED2551EBFFF2FFE53EE4D9E1FDD5F92C" box="[460,529,1759,1781]" pageId="42" pageNumber="381">KBIN</collectionCode>
|
||
–IRScNB)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8BC92EFFF2FFE53FA6D823FE6DF8A9" blockId="42.[142,758,1821,1904]" pageId="42" pageNumber="381">
|
||
<emphasis id="B940153CFFF2FFE53FA6D823FEC8F8EB" box="[142,268,1821,1842]" italics="true" pageId="42" pageNumber="381">Etymology:</emphasis>
|
||
From the Latin word
|
||
<emphasis id="B940153CFFF2FFE53D0BD823FD42F8EB" box="[547,646,1821,1842]" italics="true" pageId="42" pageNumber="381">incurvus</emphasis>
|
||
meaning
|
||
<emphasis id="B940153CFFF2FFE53FA6D805FF1EF889" box="[142,218,1851,1872]" italics="true" pageId="42" pageNumber="381">curved</emphasis>
|
||
in reference to the shape of the phallus, which is strongly bent apically.
|
||
</paragraph>
|
||
<paragraph id="8B8BC92EFFF2FFE53C0EDFFAFA4BFEEC" blockId="42.[806,1423,195,1781]" pageId="42" pageNumber="381">
|
||
<emphasis id="B940153CFFF2FFE53C0EDFFAFC6AFF00" box="[806,942,196,217]" italics="true" pageId="42" pageNumber="381">Description:</emphasis>
|
||
A blackish species of middle size; epandrium with a characteristic anterodorsal double brush of strong bristles, a row of 8 very strong, long dorsal bristles apically, phallus bent at right angle apically.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32E9AA5FFF2FFE53C0EDE00FCADF92C" pageId="42" pageNumber="381" type="description">
|
||
<paragraph id="8B8BC92EFFF2FFE53C0EDE00FC0EFA38" blockId="42.[806,1423,195,1781]" pageId="42" pageNumber="381">
|
||
MALE: HEAD. Occiput blackish with grey dusting. Face, palpus, antenna black. First flagellomere conical, second and third flagellomeres aristiform. Proboscis blackish. Labrum length
|
||
<emphasis id="B940153CFFF2FFE53B9CDEA5FB01FE69" box="[1204,1221,411,432]" italics="true" pageId="42" pageNumber="381">c.</emphasis>
|
||
1.5× head height. Labium rather thick, well sclerotized, with numerous short bristles. THORAX blackish, scutum subshiny, prescutellar depression, scutellum dusty. All bristles black. Antepronotum with some lateral bristles. Proepisternum, prosternum with 1 strong, rather long bristle. Acrostichals apparently uniserial, fine. Dorsocentrals uniserial, ending with 2 strong, long bristles in prescutellar depression. Anterior and posterior spiracles blackish. LEGS brownish. Fore tibia with dorsal row of
|
||
<emphasis id="B940153CFFF2FFE53C5CDDF3FC41FD3B" box="[884,901,717,738]" italics="true" pageId="42" pageNumber="381">c.</emphasis>
|
||
20 short bristles, ventrally covered with very fine, short bristly hairs; fore tarsomeres with distinct bristles at tip. Mid femur with some strong, long ventral bristles in basal third; mid tibia anteroventrally covered with short bristles, 1 strong, long ventral at basal quarter, 2 strong, long dorsals (at middle, apically); mid tarsomeres with apical circlet of distinct bristles. Hind femur with some distinct bristles postero- and anterodorsally; hind tibia with numerous strong, long ventral, antero- and posterodorsal bristles; first three hind tarsomeres thick; first two hind tarsomeres covered with numerous strong, long dorsal bristles; other hind tarsomeres with apical circlet of strong bristles. WING (length = 2.75) faintly tinged with brown. A1 abbreviated. Halter black. ABDOMEN black except for first five sternites brownish to yellowish, with tergite 8 slightly more sclerotized anterolaterally, distinct bristles at base and at posterior margin of tergites; sternite 8 with strong posterior bristles. HYPOPYGIUM (
|
||
<figureCitation id="130FD5ABFFF2FFE53CE3DA2DFBD9FAF0" box="[971,1053,1299,1321]" captionStart="Figure 36" captionStartId="42.[142,220,1075,1094]" captionTargetBox="[153,747,202,1039]" captionTargetId="figure-624@42.[144,752,185,1054]" captionTargetPageId="42" captionText="Figure 36. The Empis (Coptophlebia) nahaeoensis clade, male of E. (C.) incurva sp. nov. A, epandrium and cercus in lateral view; B, hypandrium and phallus in lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435338" httpUri="https://zenodo.org/record/5435338/files/figure.png" pageId="42" pageNumber="381">Fig. 36</figureCitation>
|
||
).
|
||
<taxonomicName id="4C34B2ADFFF2FFE53B1ADA2DFB45FAF1" box="[1074,1153,1299,1320]" class="Insecta" family="Kateretidae" genus="Cercus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="345" phylum="Arthropoda" rank="genus">Cercus</taxonomicName>
|
||
with anterodorsal protuberance bearing brush of numerous fine bristles. Epandrium with pair of dorsal projections bearing brush of strong, long, apically rounded bristles; row of 8 strong, long dorsal bristles at tip. Hypandrium membranous ventrally. Phallus thick at base, bent at right angle apically.
|
||
</paragraph>
|
||
<paragraph id="8B8BC92EFFF2FFE53C17DAD4FCADF92C" blockId="42.[806,1423,195,1781]" pageId="42" pageNumber="381">FEMALE: Similar to male except for following characters: anteroventral ommatidia somewhat enlarged, frons as wide as face, shiny. Legs with all bristles shorter; mid femur with short ventral subpennate bristles at middle; hind femur with rather long dorsal pennation except apically, ventral pennation except basally; hind tibia with short ventral subpennate bristles on basal half. Tergites subshiny except for last three.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |