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<mods:title id="22FEC7195B76B397110F637DA00E8499">The Mecistocephalidae of the Japanese and Taiwanese islands (Chilopoda: Geophilomorpha)</mods:title>
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<subSubSection id="C32EA3CE547F954E36FE7C8AFDEFFB6C" box="[151,564,1194,1220]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8B8BF045547F954E36FE7C8AFDEFFB6C" blockId="12.[151,564,1194,1255]" box="[151,564,1194,1220]" pageId="12" pageNumber="13">
<heading id="D0C34729547F954E36FE7C8AFDEFFB6C" bold="true" box="[151,564,1194,1220]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<taxonomicName id="4C348BC6547F954E36FE7C8AFDEFFB6C" ID-CoL="5VSJS" authority="(Takakuwa, 1934)" baseAuthorityName="Takakuwa" baseAuthorityYear="1934" box="[151,564,1194,1220]" class="Chilopoda" family="Mecistocephalidae" genus="Arrup" kingdom="Animalia" order="Geophilomorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547F954E36FE7C8AFDEFFB6C" bold="true" box="[151,564,1194,1220]" pageId="12" pageNumber="13">
<emphasis id="B9402C57547F954E36FE7C8AFE96FB6C" bold="true" box="[151,333,1194,1220]" italics="true" pageId="12" pageNumber="13">Arrup dentatus</emphasis>
(Takakuwa, 1934)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8B8BF045547F954E36FE7CEDFEFAFB4F" blockId="12.[151,564,1194,1255]" box="[151,289,1229,1255]" pageId="12" pageNumber="13">
<figureCitation id="130FECC0547F954E36FE7CEDFEFAFB4F" box="[151,289,1229,1255]" captionStart="FIGURES 1013" captionStartId="13.[151,269,1796,1820]" captionTargetBox="[267,1321,480,1745]" captionTargetId="figure-143@13.[258,1329,439,1772]" captionTargetPageId="13" captionText="FIGURES 1013. Arrup dentatus, female from Mt. Soranuma­dake, Hokkaido: 10, head, dorsal; 11, anterior part of head, ventral (maxillary complex removed); 12, maxillary complex, ventral; 13, forcipular segment and left forcipule, dorsal (poison calyx indicated by solid lines, setae not drawn). Areolation not drawn." pageId="12" pageNumber="13">Figs. 1013</figureCitation>
</paragraph>
<paragraph id="8B8BF045547F954E36FE7D3DFBD6FA1C" blockId="12.[151,1297,1309,1844]" pageId="12" pageNumber="13">
[1]
<taxonomicName id="4C348BC6547F954E36AE7D3DFDA8FA9C" authority="Takakuwa 1934 a" authorityName=": Takakuwa" authorityYear="1934" box="[199,627,1309,1332]" class="Chilopoda" family="Mecistocephalidae" genus="Prolamnonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547F954E36AE7D3DFE6CFA9C" box="[199,439,1309,1332]" italics="true" pageId="12" pageNumber="13">Prolamnonyx dentatus</emphasis>
<bibRefCitation id="EFA58DB4547F954E37D47D3DFDA8FA9C" author="Takakuwa, Y." box="[445,627,1309,1332]" pageId="12" pageNumber="13" pagination="355 - 363" refId="ref55473" refString="Takakuwa, Y. (1934 a) Neue Japanische Mecistocephalidae. Annotationes Zoologicae Japonenses, 14, 355 - 363." type="journal article" year="1934">Takakuwa 1934a</bibRefCitation>
</taxonomicName>
— Annot. Zool.
<collectingCountry id="F323B0D5547F954E35427D3DFCB6FA9C" box="[811,877,1309,1332]" name="Japan" pageId="12" pageNumber="13">Japon</collectingCountry>
. 14: 359; fig. 78 (original description)
<taxonomicName id="4C348BC6547F954E36AE7D1DFDA0FAFC" authority=": Takakuwa 1934 b" authorityName=": Takakuwa" authorityYear="1934" box="[199,635,1341,1364]" class="Chilopoda" family="Mecistocephalidae" genus="Prolamnonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547F954E36AE7D1DFE6CFAFC" box="[199,439,1341,1364]" italics="true" pageId="12" pageNumber="13">Prolamnonyx dentatus</emphasis>
:
<bibRefCitation id="EFA58DB4547F954E37AD7D1DFDA0FAFC" author="Takakuwa, Y." box="[452,635,1341,1364]" pageId="12" pageNumber="13" pagination="706 - 712" refId="ref55495" refString="Takakuwa, Y. (1934 b) The family Mecistocephalidae of Japan, I. Botany and Zoology [= Shokubutsu - oyobi - Dobutsu], 2, 706 - 712." type="journal article" year="1934">Takakuwa 1934b</bibRefCitation>
</taxonomicName>
— Bot. &amp; Zool.
<collectingRegion id="49F03EA7547F954E355B7D1DFCAEFAFC" box="[818,885,1341,1364]" country="Japan" name="Tokyo" pageId="12" pageNumber="13">Tokyo</collectingRegion>
2: 707 (key)
<taxonomicName id="4C348BC6547F954E36AE7D7DFDA2FADC" authority=": Takakuwa 1934 c" authorityName=": Takakuwa" authorityYear="1934" box="[199,633,1373,1396]" class="Chilopoda" family="Mecistocephalidae" genus="Prolamnonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547F954E36AE7D7DFE6DFADC" box="[199,438,1373,1396]" italics="true" pageId="12" pageNumber="13">Prolamnonyx dentatus</emphasis>
:
<bibRefCitation id="EFA58DB4547F954E37AD7D7DFDA2FADC" author="Takakuwa, Y." box="[452,633,1373,1396]" pageId="12" pageNumber="13" pagination="878 - 884" refId="ref55529" refString="Takakuwa, Y. (1934 c) Japanese Mecistocephalidae II. Botany and Zoology [= Shokubutsu - oyobi - Dobutsu], 2, 878 - 884." type="journal article" year="1934">Takakuwa 1934c</bibRefCitation>
</taxonomicName>
— Bot. &amp; Zool.
<collectingRegion id="49F03EA7547F954E35587D7DFCAEFADC" box="[817,885,1373,1396]" country="Japan" name="Tokyo" pageId="12" pageNumber="13">Tokyo</collectingRegion>
2: 883; fig. 2733
<taxonomicName id="4C348BC6547F954E36AE7D5DFDA1FA3C" authority=": Takakuwa 1938 a" authorityName=": Takakuwa" authorityYear="1938" box="[199,634,1405,1428]" class="Chilopoda" family="Mecistocephalidae" genus="Prolamnonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547F954E36AE7D5DFE6DFA3C" box="[199,438,1405,1428]" italics="true" pageId="12" pageNumber="13">Prolamnonyx dentatus</emphasis>
:
<bibRefCitation id="EFA58DB4547F954E37AD7D5DFDA1FA3C" author="Takakuwa, Y." box="[452,634,1405,1428]" pageId="12" pageNumber="13" pagination="353 - 358" refId="ref55804" refString="Takakuwa, Y. (1938 a) Verzeichnis der Japanischen und Mandschureischen Chilopodenarten. Annotationes Zoologicae Japonenses, 17, 353 - 358." type="journal article" year="1938">Takakuwa 1938a</bibRefCitation>
</taxonomicName>
— Annot. Zool.
<collectingCountry id="F323B0D5547F954E35587D5DFCA9FA3C" box="[817,882,1405,1428]" name="Japan" pageId="12" pageNumber="13">Japon</collectingCountry>
. 17: 355
<taxonomicName id="4C348BC6547F954E36AE7DBDFDA0FA1C" authority=": Takakuwa 1938 b" authorityName=": Takakuwa" authorityYear="1938" box="[199,635,1437,1460]" class="Chilopoda" family="Mecistocephalidae" genus="Prolamnonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547F954E36AE7DBDFE6CFA1C" box="[199,439,1437,1460]" italics="true" pageId="12" pageNumber="13">Prolamnonyx dentatus</emphasis>
:
<bibRefCitation id="EFA58DB4547F954E37AC7DBDFDA0FA1C" author="Takakuwa, Y." box="[453,635,1437,1460]" pageId="12" pageNumber="13" pagination="2023 - 2032" refId="ref55829" refString="Takakuwa, Y. (1938 b) Key to the Japanese species of Scolopendromorpha and Geophilomorpha. Botany and Zoology [= Shokubutsu - oyobi - Dobutsu], 6, 2023 - 2032" type="journal article" year="1938">Takakuwa 1938b</bibRefCitation>
</taxonomicName>
— Bot. &amp; Zool.
<collectingRegion id="49F03EA7547F954E355A7DBDFCADFA1C" box="[819,886,1437,1460]" country="Japan" name="Tokyo" pageId="12" pageNumber="13">Tokyo</collectingRegion>
6: 2030 (key)
</paragraph>
<paragraph id="8B8BF045547F954E36FE7D9DFC72F9DC" blockId="12.[151,1297,1309,1844]" pageId="12" pageNumber="13">
[2]
<taxonomicName id="4C348BC6547F954E36AE7D9DFDB5FA7C" authority=": Takakuwa 1940" authorityName="Takakuwa" authorityYear="1940" box="[199,622,1469,1492]" class="Chilopoda" family="Mecistocephalidae" genus="Prolamnonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547F954E36AE7D9DFE6CFA7C" box="[199,439,1469,1492]" italics="true" pageId="12" pageNumber="13">Prolamnonyx dentatus</emphasis>
:
<bibRefCitation id="EFA58DB4547F954E37AD7D9DFDB5FA7C" author="Takakuwa, Y." box="[452,622,1469,1492]" pageId="12" pageNumber="13" refId="ref55954" refString="Takakuwa, Y. (1940) Fauna Nipponica 9 (8, 1). Geophilomorpha. Sanseido, Tokyo, 156 pp." type="book" year="1940">Takakuwa 1940</bibRefCitation>
</taxonomicName>
— Fauna
<collectingCountry id="F323B0D5547F954E34B77D9DFCEAFA7C" box="[734,817,1469,1492]" name="Japan" pageId="12" pageNumber="13">Nippon</collectingCountry>
. 9: 94 (key); fig. 98100
<taxonomicName id="4C348BC6547F954E36AE7DFDFDB5FA5C" authority=": Takakuwa 1943" authorityName="Takakuwa" authorityYear="1943" box="[199,622,1501,1524]" class="Chilopoda" family="Mecistocephalidae" genus="Prolamnonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547F954E36AE7DFDFE6DFA5C" box="[199,438,1501,1524]" italics="true" pageId="12" pageNumber="13">Prolamnonyx dentatus</emphasis>
:
<bibRefCitation id="EFA58DB4547F954E37AD7DFDFDB5FA5C" author="Takakuwa, Y." box="[452,622,1501,1524]" pageId="12" pageNumber="13" pagination="147 - 213" refId="ref56084" refString="Takakuwa, Y. (1943) Distribution of chilopods and diplopods in Japan. Bulletin of the Biogeographical Society of Japan, 13, 147 - 213." type="journal article" year="1943">Takakuwa 1943</bibRefCitation>
</taxonomicName>
— Bull. Biogeogr. Soc.
<collectingCountry id="F323B0D5547F954E351C7DFDFC69FA5C" box="[885,946,1501,1524]" name="Japan" pageId="12" pageNumber="13">Japan</collectingCountry>
13: 165
<taxonomicName id="4C348BC6547F954E36AE7DDDFD96F9BC" authority=": Attems 1947" authorityName="Attems" authorityYear="1947" box="[199,589,1533,1556]" class="Chilopoda" family="Mecistocephalidae" genus="Prolamnonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547F954E36AE7DDDFE6DF9BC" box="[199,438,1533,1556]" italics="true" pageId="12" pageNumber="13">Prolamnonyx dentatus</emphasis>
:
<bibRefCitation id="EFA58DB4547F954E37AD7DDDFD96F9BC" author="Attems, C. G." box="[452,589,1533,1556]" pageId="12" pageNumber="13" pagination="50 - 149" refId="ref52509" refString="Attems, C. G. (1947) Neue Geophilomorpha des Wiener Museums. Annalen des Naturhistorischen Museums, Wien, 55 (1944 - 47), 50 - 149." type="journal article" year="1947">Attems 1947</bibRefCitation>
</taxonomicName>
— Ann. Naturhistor. Mus. Wien 55: 105 (key)
<taxonomicName id="4C348BC6547F954E36AE7E3DFDA8F99C" authority=": Takashima 1949" authorityName="Takashima" authorityYear="1949" box="[199,627,1565,1588]" class="Chilopoda" family="Mecistocephalidae" genus="Prolamnonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547F954E36AE7E3DFE6DF99C" box="[199,438,1565,1588]" italics="true" pageId="12" pageNumber="13">Prolamnonyx dentatus</emphasis>
:
<bibRefCitation id="EFA58DB4547F954E37AD7E3DFDA8F99C" author="Takashima, H." box="[452,627,1565,1588]" pageId="12" pageNumber="13" pagination="8 - 15" refId="ref56166" refString="Takashima, H. (1949) The general view of Japanese Myriapods. Acta Arachnologica, 11, 8 - 15." type="journal article" year="1949">Takashima 1949</bibRefCitation>
</taxonomicName>
— Acta Arachn. 11: 13
<taxonomicName id="4C348BC6547F954E36AE7E1DFCD9F9FC" authority=": Takashima &amp; Shinohara 1952" authorityName="Takashima &amp; Shinohara" authorityYear="1952" box="[199,770,1597,1620]" class="Chilopoda" family="Mecistocephalidae" genus="Prolamnonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547F954E36AE7E1DFE6DF9FC" box="[199,438,1597,1620]" italics="true" pageId="12" pageNumber="13">Prolamnonyx dentatus</emphasis>
:
<bibRefCitation id="EFA58DB4547F954E37AD7E1DFCD9F9FC" author="Takashima, H. &amp; Shinohara, K." box="[452,770,1597,1620]" pageId="12" pageNumber="13" pagination="3 - 17" refId="ref56189" refString="Takashima, H. &amp; Shinohara, K. (1952) The centipede-fauna of the Tokyo District. Acta Arachnologica, 13, 3 - 17." type="journal article" year="1952">Takashima &amp; Shinohara 1952</bibRefCitation>
</taxonomicName>
— Acta Arachn. 13: 9 (key)
<taxonomicName id="4C348BC6547F954E36AE7E7DFE20F9DC" authority=": Crabill 1964" authorityName="Crabill" authorityYear="1964" box="[199,507,1629,1652]" class="Chilopoda" family="Mecistocephalidae" genus="Arrup" kingdom="Animalia" order="Geophilomorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547F954E36AE7E7DFEBCF9DC" box="[199,359,1629,1652]" italics="true" pageId="12" pageNumber="13">Arrup dentatus</emphasis>
:
<bibRefCitation id="EFA58DB4547F954E371C7E7DFE20F9DC" author="Crabill, R. E." box="[373,507,1629,1652]" pageId="12" pageNumber="13" pagination="161 - 170" refId="ref53240" refString="Crabill, R. E. (1964) A revised interpretation of the primitive centipede genus Arrup, with redescription of its type-species and list of known species. Proceedings of the Biological Society of Washington, 77, 161 - 170." type="journal article" year="1964">Crabill 1964</bibRefCitation>
</taxonomicName>
— Proc. Biol. Soc. Washington 77: 166
</paragraph>
<paragraph id="8B8BF045547F954E36FE7E5DFC28F93C" blockId="12.[151,1297,1309,1844]" box="[151,1011,1661,1684]" pageId="12" pageNumber="13">
[
<materialsCitation id="3B5CFA18547F954E36F67E5DFC6CF93C" box="[159,951,1661,1684]" collectorName="Mem. Nat. Sc. Mus." location="Tokyo" pageId="12" pageNumber="13" specimenCount="1" stateProvince="Tokyo">
3]
<taxonomicName id="4C348BC6547F954E36AE7E5DFDB7F93C" authority=": Shinohara 1972" authorityName="Shinohara" authorityYear="1972" box="[199,620,1661,1684]" class="Chilopoda" family="Mecistocephalidae" genus="Prolamnonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547F954E36AE7E5DFE6CF93C" box="[199,439,1661,1684]" italics="true" pageId="12" pageNumber="13">Prolamnonyx dentatus</emphasis>
:
<bibRefCitation id="EFA58DB4547F954E37AD7E5DFDB7F93C" author="Shinohara, K." box="[452,620,1661,1684]" pageId="12" pageNumber="13" pagination="65 - 73" refId="ref55213" refString="Shinohara, K. (1972) Some chilopods of the Hidaka Mountain Range in Hokkaido, Northern Japan. Memoirs of the National Science Museum, Tokyo, 5, 65 - 73." type="journal article" year="1972">Shinohara 1972</bibRefCitation>
</taxonomicName>
<collectorName id="26C19593547F954E34FC7E5DFCB6F93C" box="[661,877,1661,1684]" pageId="12" pageNumber="13">Mem. Nat. Sc. Mus.</collectorName>
<collectingRegion id="49F03EA7547F954E351A7E5DFC6CF93C" box="[883,951,1661,1684]" country="Japan" name="Tokyo" pageId="12" pageNumber="13">Tokyo</collectingRegion>
</materialsCitation>
5: 66
</paragraph>
<paragraph id="8B8BF045547F954E36FE7EBDFC99F89C" blockId="12.[151,1297,1309,1844]" pageId="12" pageNumber="13">
[4]
<taxonomicName id="4C348BC6547F954E36AE7EBDFD9EF91C" authority=": Titova 1975" authorityName="Titova" authorityYear="1975" box="[199,581,1693,1716]" class="Chilopoda" family="Mecistocephalidae" genus="Prolamnonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547F954E36AE7EBDFE6DF91C" box="[199,438,1693,1716]" italics="true" pageId="12" pageNumber="13">Prolamnonyx dentatus</emphasis>
:
<bibRefCitation id="EFA58DB4547F954E37AD7EBDFD9EF91C" author="Titova, L. P." box="[452,581,1693,1716]" pageId="12" pageNumber="13" pagination="39 - 48" refId="ref56253" refString="Titova, L. P. (1975) Geophilids of the family Mecistocephalidae (Chilopoda) in the Fauna of the USSR. Zoologichesckii Zhurnal, 54, 39 - 48." type="journal article" year="1975">Titova 1975</bibRefCitation>
</taxonomicName>
— Zool. Zh. 54: 47 (redescription, key); fig. 4b
<taxonomicName id="4C348BC6547F954E36AE7E9DFDB4F97C" authority=": Murakami 1993" authorityName="Murakami" authorityYear="1993" box="[199,623,1725,1748]" class="Chilopoda" family="Mecistocephalidae" genus="Prolamnonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547F954E36AE7E9DFE6DF97C" box="[199,438,1725,1748]" italics="true" pageId="12" pageNumber="13">Prolamnonyx dentatus</emphasis>
:
<bibRefCitation id="EFA58DB4547F954E37AD7E9DFDB4F97C" author="Murakami, Y." box="[452,623,1725,1748]" pageId="12" pageNumber="13" pagination="95 - 107" refId="ref54511" refString="Murakami, Y. (1993) Arthropoda: Diplopoda, Pauropoda, Chilopoda, Symphyla. In: Environmental Agency Japan (Ed.), List of species of Animals in Japan: Invertebrates I. The Shizen-Kankyo-Kenkyu Center, Tokyo, 5, 95 - 107." type="journal article" year="1993">Murakami 1993</bibRefCitation>
</taxonomicName>
— List Species Anim.
<collectingCountry id="F323B0D5547F954E35037E9DFC7CF97C" box="[874,935,1725,1748]" name="Japan" pageId="12" pageNumber="13">Japan</collectingCountry>
5: 105
<taxonomicName id="4C348BC6547F954E36AE7EFDFDB7F95C" authority=": Shinohara 1999" authorityName="Shinohara" authorityYear="1999" box="[199,620,1757,1780]" class="Chilopoda" family="Mecistocephalidae" genus="Prolamnonyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Geophilomorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547F954E36AE7EFDFE6DF95C" box="[199,438,1757,1780]" italics="true" pageId="12" pageNumber="13">Prolamnonyx dentatus</emphasis>
:
<bibRefCitation id="EFA58DB4547F954E37AD7EFDFDB7F95C" author="Shinohara, K." box="[452,620,1757,1780]" pageId="12" pageNumber="13" pagination="693 - 709" refId="ref55374" refString="Shinohara, K. (1999) Chilopoda. In: Aoki, J. (Ed.), Pictorial keys to soil animals of Japan. Tokay University Press, Tokyo, 693 - 709." type="book chapter" year="1999">Shinohara 1999</bibRefCitation>
</taxonomicName>
— Pictorial Keys Soil Anim.
<collectingCountry id="F323B0D5547F954E35C77EFDFC34F95C" box="[942,1007,1757,1780]" name="Japan" pageId="12" pageNumber="13">Japan</collectingCountry>
: 709
<taxonomicName id="4C348BC6547F954E36AE7EDDFDE1F8BC" authority=": Bonato et al. 2003" authorityName="Bonato" authorityYear="2003" box="[199,570,1789,1812]" class="Chilopoda" family="Mecistocephalidae" genus="Arrup" kingdom="Animalia" order="Geophilomorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547F954E36AE7EDDFEBCF8BC" box="[199,359,1789,1812]" italics="true" pageId="12" pageNumber="13">Arrup dentatus</emphasis>
:
<bibRefCitation id="EFA58DB4547F954E371C7EDDFDE1F8BC" author="Bonato, L. &amp; Foddai, D. &amp; Minelli, A." box="[373,570,1789,1812]" pageId="12" pageNumber="13" pagination="539 - 579" refId="ref52693" refString="Bonato, L., Foddai, D. &amp; Minelli, A. (2003) Evolutionary trends and patterns in centipede segment number based on a cladistic analysis of Mecistocephalidae (Chilopoda: Geophilomorpha). Systematic Entomology, 28, 539 - 579." type="journal article" year="2003">
Bonato
<emphasis id="B9402C57547F954E37A07EDDFE24F8BC" box="[457,511,1789,1812]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2003
</bibRefCitation>
</taxonomicName>
— Syst. Entom. 28: 550, 551
<taxonomicName id="4C348BC6547F954E36AE7F3DFDECF89C" authority=": Foddai et al. 2003" authorityName="Foddai" authorityYear="2003" box="[199,567,1821,1844]" class="Chilopoda" family="Mecistocephalidae" genus="Arrup" kingdom="Animalia" order="Geophilomorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547F954E36AE7F3DFEBCF89C" box="[199,359,1821,1844]" italics="true" pageId="12" pageNumber="13">Arrup dentatus</emphasis>
:
<bibRefCitation id="EFA58DB4547F954E371C7F3DFDECF89C" author="Foddai, D. &amp; Bonato, L. &amp; Pereira, L. A. &amp; Minelli, A." box="[373,567,1821,1844]" pageId="12" pageNumber="13" refId="ref53487" refString="Foddai, D., Bonato, L., Pereira, L. A. &amp; Minelli, A., 2003 Phylogeny and systematics of the Arrupinae (Chilopoda Geophilomorpha Mecistocephalidae) with description of a new dwarfed species. Journal of Natural History, 37,1247 - 1267." type="book" year="2003">
Foddai
<emphasis id="B9402C57547F954E37AF7F3DFE27F89C" box="[454,508,1821,1844]" italics="true" pageId="12" pageNumber="13">et al.</emphasis>
2003
</bibRefCitation>
</taxonomicName>
— J. Nat. Hist. 37: 1261
</paragraph>
<paragraph id="8B8BF045547F954E36FE7F46FA46F867" blockId="12.[151,1437,1893,1999]" pageId="12" pageNumber="13">
<emphasis id="B9402C57547F954E36FE7F46FED5F8D7" box="[151,270,1894,1919]" italics="true" pageId="12" pageNumber="13">Diagnosis</emphasis>
. An
<taxonomicName id="4C348BC6547F954E37207F46FE36F8D7" box="[329,493,1893,1919]" class="Chilopoda" family="Mecistocephalidae" genus="Arrup" kingdom="Animalia" order="Geophilomorpha" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9402C57547F954E37207F46FE54F8D7" box="[329,399,1894,1919]" italics="true" pageId="12" pageNumber="13">Arrup</emphasis>
species
</taxonomicName>
with body length reaching about
<quantity id="4CCC5DA0547F954E35147F45FC60F8D7" box="[893,955,1893,1919]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="13" unit="cm" value="2.0">2 cm</quantity>
. Frontal line curved. Distal article of the telopodite of the second maxillae with a claw. Forcipular article III with a well developed tooth. Poison calyx only reaching the distal part of forcipular article I both in males and in females. Telopodites of the last leg­
</paragraph>
<paragraph id="8B8BF045547E954F36FE78B8FF09FF72" blockId="13.[151,1436,152,418]" pageId="13" pageNumber="14">bearing segments swollen in males, slender in females. Sternum of the last leg­bearing segment wider than long.</paragraph>
<paragraph id="8B8BF045547E954F36AF78C8FBF1FEAA" blockId="13.[151,1436,152,418]" box="[198,1066,232,258]" pageId="13" pageNumber="14">
<emphasis id="B9402C57547E954F36AF78C8FEBCFEA9" box="[198,359,232,257]" italics="true" pageId="13" pageNumber="14">Type material</emphasis>
.
<typeStatus id="548F4EE7547E954F371F78C8FE3CFEAA" box="[374,487,232,258]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
: male,
<quantity id="4CCC5DA0547E954F345378C8FD57FEAA" box="[570,652,232,258]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="13" pageNumber="14" unit="mm" value="20.0">20 mm</quantity>
long, subadult (
<bibRefCitation id="EFA58DB4547E954F352078C8FBC5FEAA" author="Takakuwa, Y." box="[841,1054,232,258]" pageId="13" pageNumber="14" pagination="355 - 363" refId="ref55473" refString="Takakuwa, Y. (1934 a) Neue Japanische Mecistocephalidae. Annotationes Zoologicae Japonenses, 14, 355 - 363." type="journal article" year="1934">Takakuwa, 1934a</bibRefCitation>
).
</paragraph>
<paragraph id="8B8BF045547E954F36AF7930FBC8FE82" blockId="13.[151,1436,152,418]" box="[198,1043,272,298]" pageId="13" pageNumber="14">
<materialsCitation id="3B5CFA18547E954F36AF7930FBD4FE82" box="[198,1039,272,298]" location="Zyozankei" pageId="13" pageNumber="14" specimenCount="1" stateProvince="Hokkaido" typeStatus="holotype">
<emphasis id="B9402C57547E954F36AF7930FE80FE81" box="[198,347,272,297]" italics="true" pageId="13" pageNumber="14">
<typeStatus id="548F4EE7547E954F36AF7930FF26FE81" box="[198,253,272,297]" pageId="13" pageNumber="14">Type</typeStatus>
locality
</emphasis>
.
<location id="8EEBA69E547E954F37047930FE35FE82" LSID="urn:lsid:plazi:treatment:039D4153547F954C36FE7C8AFCB1FB50:8EEBA69E547E954F37047930FE35FE82" box="[365,494,272,298]" name="Zyozankei" pageId="13" pageNumber="14" stateProvince="Hokkaido">Zyôzankei</location>
(bei Sapporo),
<collectingRegion id="49F03EA7547E954F34C67930FCF9FE82" box="[687,802,272,298]" country="Japan" name="Hokkaido" pageId="13" pageNumber="14">Hokkaido</collectingRegion>
(
<bibRefCitation id="EFA58DB4547E954F35587930FBDCFE82" author="Takakuwa, Y." box="[817,1031,272,298]" pageId="13" pageNumber="14" pagination="355 - 363" refId="ref55473" refString="Takakuwa, Y. (1934 a) Neue Japanische Mecistocephalidae. Annotationes Zoologicae Japonenses, 14, 355 - 363." type="journal article" year="1934">Takakuwa, 1934a</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BF045547E954F36AF7918FB9BFEFA" blockId="13.[151,1436,152,418]" box="[198,1088,312,338]" pageId="13" pageNumber="14">
<emphasis id="B9402C57547E954F36AF7918FDD2FEF9" box="[198,521,312,337]" italics="true" pageId="13" pageNumber="14">Depository of type material</emphasis>
. The
<typeStatus id="548F4EE7547E954F34247918FD6FFEFA" box="[589,692,312,338]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
is probably lost (see Discussion).
</paragraph>
<paragraph id="8B8BF045547E954F36AF7940FB68FE0A" blockId="13.[151,1436,152,418]" pageId="13" pageNumber="14">
<emphasis id="B9402C57547E954F36AF7940FE7DFED1" box="[198,422,352,377]" italics="true" pageId="13" pageNumber="14">Material examined</emphasis>
.
<specimenCount id="9D323BCC547E954F37DC7940FD97FED2" box="[437,588,352,378]" pageId="13" pageNumber="14" type="generic">3 specimens</specimenCount>
:
<specimenCount id="9D323BCC547E954F34317940FD70FED2" box="[600,683,352,378]" pageId="13" pageNumber="14" type="male">1 male</specimenCount>
, adult (
<quantity id="4CCC5DA0547E954F356D7940FC86FED2" box="[772,861,352,378]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="13" pageNumber="14" unit="mm" value="17.0">17 mm</quantity>
), and
<materialsCitation id="3B5CFA18547E954F35CC7940FB74FE0A" collectingDate="1985-05-21" collectionCode="AM, TT" collectorName="M. Ohara" location="Sapporo-shi" municipality="Mt. Soranuma-dake" pageId="13" pageNumber="14" specimenCount="2" specimenCount-female="2" stateProvince="Hokkaido">
<specimenCount id="9D323BCC547E954F35CC7940FBC1FED2" box="[933,1050,352,378]" pageId="13" pageNumber="14" type="female">2 females</specimenCount>
, subadult (19 and
<quantity id="4CCC5DA0547E954F329F7940FA95FED2" box="[1270,1358,352,378]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="13" pageNumber="14" unit="mm" value="21.0">21 mm</quantity>
), from
<collectingMunicipality id="6BEF6A3F547E954F36FE79A8FE58FE0A" box="[151,387,392,418]" pageId="13" pageNumber="14">Mt. Soranuma­dake</collectingMunicipality>
,
<location id="8EEBA69E547E954F37E679A8FDFBFE0A" LSID="urn:lsid:plazi:treatment:039D4153547F954C36FE7C8AFCB1FB50:8EEBA69E547E954F37E679A8FDFBFE0A" box="[399,544,392,418]" municipality="Mt. Soranuma-dake" name="Sapporo-shi" pageId="13" pageNumber="14" stateProvince="Hokkaido">Sapporo­shi</location>
,
<collectingRegion id="49F03EA7547E954F344279A8FD79FE0A" box="[555,674,392,418]" country="Japan" name="Hokkaido" pageId="13" pageNumber="14">Hokkaido</collectingRegion>
,
<date id="FF8AD685547E954F34C779A8FCF3FE0A" box="[686,808,392,418]" pageId="13" pageNumber="14" value="1985-05-21">
<collectingDate id="EFCE2F6D547E954F34C779A8FCF3FE0A" box="[686,808,392,418]" pageId="13" pageNumber="14" value="1985-05-21">21.V.1985</collectingDate>
</date>
,
<collectorName id="26C19593547E954F355C79A8FC7DFE0A" box="[821,934,392,418]" pageId="13" pageNumber="14">M. Ohara</collectorName>
leg., coll.
<collectionCode id="ED256880547E954F324879A8FB8BFE0A" box="[1057,1104,392,418]" country="Australia" name="Australian Museum" pageId="13" pageNumber="14" type="Museum">AM</collectionCode>
and
<collectionCode id="ED256880547E954F32E379A8FB74FE0A" box="[1162,1199,392,418]" pageId="13" pageNumber="14">TT</collectionCode>
</materialsCitation>
.
</paragraph>
<caption id="DF4BA0CD547E954F36FE7F24FBF9F8C9" pageId="13" pageNumber="14" startId="13.[151,269,1796,1820]" targetBox="[267,1321,480,1745]" targetPageId="13">
<paragraph id="8B8BF045547E954F36FE7F24FBF9F8C9" blockId="13.[151,1436,1796,1889]" pageId="13" pageNumber="14">
<emphasis id="B9402C57547E954F36FE7F24FEBBF8B4" bold="true" box="[151,352,1796,1820]" pageId="13" pageNumber="14">FIGURES 1013.</emphasis>
<taxonomicName id="4C348BC6547E954F37037F25FDD6F8B4" box="[362,525,1797,1820]" class="Chilopoda" family="Mecistocephalidae" genus="Arrup" kingdom="Animalia" order="Geophilomorpha" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547E954F37037F25FDD6F8B4" box="[362,525,1797,1820]" italics="true" pageId="13" pageNumber="14">Arrup dentatus</emphasis>
</taxonomicName>
, female from Mt. Soranuma­dake, Hokkaido: 10, head, dorsal; 11, anterior part of head, ventral (maxillary complex removed); 12, maxillary complex, ventral; 13, forcipular segment and left forcipule, dorsal (poison calyx indicated by solid lines, setae not drawn). Areolation not drawn.
</paragraph>
</caption>
<paragraph id="8B8BF045547D954C36AF78B8FDE1FCF2" blockId="14.[151,1438,152,1272]" pageId="14" pageNumber="15">
<emphasis id="B9402C57547D954C36AF78B8FE8BFF19" box="[198,336,152,177]" italics="true" pageId="14" pageNumber="15">Description</emphasis>
. Body length of adults reaching at least about
<quantity id="4CCC5DA0547D954C35CF78B8FC32FF1A" box="[934,1001,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="15" unit="cm" value="2.0">2 cm</quantity>
. Body colour yellow, without dark patches. Head 1.21.3 times as long as wide; frontal line curved. Antennae 2.12.5 times longer than the head width; apical sensilla about
<quantity id="4CCC5DA0547D954C378978C8FDF5FEAA" box="[480,558,232,258]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="15" unit="m" value="10.0">10 m</quantity>
long. Clypeal setae: a median pair anterior to the plagulae and a group on each side of the clypeus, reaching the anterior corners. Clypeal ratio between about 2.0 and 2.5. Labrum: anterior ala triangular, medial margin reduced to a vertex; internal margins of side­pieces parallel, not touching each other; posterior margin of each side­piece sinuous, slightly convex close to the internal and external ends. First maxillae: anterior corners of coxosternum not projecting; each medial projection only slightly longer than wide, the internal margin with very short spines, the distal lobe about as long as wide; each telopodite about 2.5 times as long as wide, the distal lobe attenuated; hyaline scales on the dorsal sides of medial projections. Second maxillae: article I of the telopodite about 2.0 times as long as wide; article III about 2.5 times as long as wide, with a few setae only; apical claw present, quite stout. Forcipular segment: width to length ratio of exposed part of coxosternum about 1.11.3; cerrus absent; each pleuron with a distinct dorsal ridge, scapular point not reaching the anterior margin of coxosternum. Forcipules: article I about 1.5 times as long as wide, distal tooth well developed; article II almost without teeth; article III with a well developed tooth; tarsungulum with one basal, obtuse tooth; poison calyx only reaching the distal part of article I, in both sexes. Last leg­bearing segment: sternum sub­triangular, about 1.51.7 times as wide as long; telopodites more swollen in males than in females, with dense short setae on the ventral side, as dense in male as in female; one short apical spine.
</paragraph>
<paragraph id="8B8BF045547D954C36AF7B48FDB3FC2A" blockId="14.[151,1438,152,1272]" box="[198,616,872,898]" pageId="14" pageNumber="15">
<emphasis id="B9402C57547D954C36AF7B48FDB9FC29" box="[198,610,872,897]" italics="true" pageId="14" pageNumber="15">Distribution in the considered area</emphasis>
.
</paragraph>
<paragraph id="8B8BF045547D954C36AF7BB2FD70FBAB" blockId="14.[151,1438,152,1272]" pageId="14" pageNumber="15">
<materialsCitation id="3B5CFA18547D954C36AF7BB2FD7CFBAB" county="Wakkanai" location="Konbu" municipality="Engaru-cho" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Hokkaido">
<collectingRegion id="49F03EA7547D954C36AF7BB2FEE5FC04" box="[198,318,914,940]" country="Japan" name="Hokkaido" pageId="14" pageNumber="15">Hokkaido</collectingRegion>
: Zyôzankei
<superScript id="7C415D0D547D954C37B57BB0FE34FC36" attach="left" box="[476,495,912,926]" fontSize="6" pageId="14" pageNumber="15">[1]</superScript>
[currently Jyozankei] (
<typeStatus id="548F4EE7547D954C35607BB2FCE6FC04" box="[777,829,914,940]" pageId="14" pageNumber="15">type</typeStatus>
locality);
<collectingCounty id="62EA88C9547D954C35D57BB2FBE8FC04" box="[956,1075,914,940]" pageId="14" pageNumber="15">Wakkanai</collectingCounty>
,
<collectingMunicipality id="6BEF6A3F547D954C322A7BB2FB17FC04" box="[1091,1228,914,940]" pageId="14" pageNumber="15">Engaru­cho</collectingMunicipality>
(Monbetsu­gun),
<location id="8EEBA69E547D954C36FE7B9DFF3CFC7F" LSID="urn:lsid:plazi:treatment:039D4153547F954C36FE7C8AFCB1FB50:8EEBA69E547D954C36FE7B9DFF3CFC7F" box="[151,231,957,983]" county="Wakkanai" municipality="Engaru-cho" name="Konbu" pageId="14" pageNumber="15" stateProvince="Hokkaido">Konbu</location>
(Niseko, Niseko­cho, Abuta­gun)
<superScript id="7C415D0D547D954C34E97B9BFD48FC61" attach="left" box="[640,659,955,969]" fontSize="6" pageId="14" pageNumber="15">[2]</superScript>
(transcribed from Japanese);
<location id="8EEBA69E547D954C32687B9DFB66FC7F" LSID="urn:lsid:plazi:treatment:039D4153547F954C36FE7C8AFCB1FB50:8EEBA69E547D954C32687B9DFB66FC7F" box="[1025,1213,957,983]" county="Wakkanai" municipality="Engaru-cho" name="Poroshiri-sanso" pageId="14" pageNumber="15" stateProvince="Hokkaido">Poroshiri­sansô</location>
at the foot of
<location id="8EEBA69E547D954C331B7B9DFE96FBAB" LSID="urn:lsid:plazi:treatment:039D4153547F954C36FE7C8AFCB1FB50:8EEBA69E547D954C331B7B9DFE96FBAB" county="Wakkanai" municipality="Engaru-cho" name="Mt. Poroshiri-dake'" pageId="14" pageNumber="15" stateProvince="Hokkaido">Mt. Poroshiri­dake</location>
<superScript id="7C415D0D547D954C37247BC5FEBBFC5B" attach="left" box="[333,352,997,1011]" fontSize="6" pageId="14" pageNumber="15">[3]</superScript>
;
<location id="8EEBA69E547D954C37067BC9FD83FBAB" LSID="urn:lsid:plazi:treatment:039D4153547F954C36FE7C8AFCB1FB50:8EEBA69E547D954C37067BC9FD83FBAB" box="[367,600,1001,1027]" county="Wakkanai" municipality="Engaru-cho" name="Mt. Soranuma-dake" pageId="14" pageNumber="15" stateProvince="Hokkaido">Mt. Soranuma­dake</location>
(new)
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BF045547D954C36AF7C34FED8FBF0" blockId="14.[151,1438,152,1272]" pageId="14" pageNumber="15">
<emphasis id="B9402C57547D954C36AF7C34FE6CFB85" box="[198,439,1044,1069]" italics="true" pageId="14" pageNumber="15">General distribution</emphasis>
.
<materialsCitation id="3B5CFA18547D954C37A17C33FF31FBF0" location="Kuril Islands" municipality="Sikhote-Alin mountains" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Hokkaido">
The species is known from
<collectingRegion id="49F03EA7547D954C357B7C33FC53FB85" box="[786,904,1043,1069]" country="Japan" name="Hokkaido" pageId="14" pageNumber="15">Hokkaido</collectingRegion>
, the
<collectingMunicipality id="6BEF6A3F547D954C35AB7C33FB07FB85" box="[962,1244,1043,1069]" pageId="14" pageNumber="15">Sikhote­Alin mountains</collectingMunicipality>
<superScript id="7C415D0D547D954C328E7C30FB21FBB6" attach="right" box="[1255,1274,1040,1054]" fontSize="6" pageId="14" pageNumber="15">[4]</superScript>
and the
<location id="8EEBA69E547D954C33097C33FF31FBF0" LSID="urn:lsid:plazi:treatment:039D4153547F954C36FE7C8AFCB1FB50:8EEBA69E547D954C33097C33FF31FBF0" municipality="Sikhote-Alin mountains" name="Kuril Islands" pageId="14" pageNumber="15" stateProvince="Hokkaido">Kuril Islands</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8B8BF045547D954C36837C1BFF27FBE1" blockId="14.[151,1438,152,1272]" box="[234,252,1083,1097]" pageId="14" pageNumber="15">
<superScript id="7C415D0D547D954C36837C1BFF27FBE1" attach="left" box="[234,252,1083,1097]" fontSize="6" pageId="14" pageNumber="15">[4]</superScript>
</paragraph>
<paragraph id="8B8BF045547D954C36AF7C47FCB1FB50" blockId="14.[151,1438,152,1272]" pageId="14" pageNumber="15">
<emphasis id="B9402C57547D954C36AF7C47FEF6FB28" box="[198,301,1127,1152]" italics="true" pageId="14" pageNumber="15">Remarks</emphasis>
.
<taxonomicName id="4C348BC6547D954C37547C47FE2BFB28" box="[317,496,1127,1152]" class="Chilopoda" family="Mecistocephalidae" genus="Arrup" kingdom="Animalia" order="Geophilomorpha" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="dentatus">
<emphasis id="B9402C57547D954C37547C47FE2BFB28" box="[317,496,1127,1152]" italics="true" pageId="14" pageNumber="15">Arrup dentatus</emphasis>
</taxonomicName>
was described as new in two different papers published by the same author in the same year (
<bibRefCitation id="EFA58DB4547D954C372E7CAEFDC2FB00" author="Takakuwa, Y." box="[327,537,1166,1192]" pageId="14" pageNumber="15" pagination="355 - 363" refId="ref55473" refString="Takakuwa, Y. (1934 a) Neue Japanische Mecistocephalidae. Annotationes Zoologicae Japonenses, 14, 355 - 363." type="journal article" year="1934">Takakuwa, 1934a</bibRefCitation>
,
<bibRefCitation id="EFA58DB4547D954C344F7CAEFDA9FB00" author="Takakuwa, Y." box="[550,626,1166,1192]" pageId="14" pageNumber="15" pagination="878 - 884" refId="ref55529" refString="Takakuwa, Y. (1934 c) Japanese Mecistocephalidae II. Botany and Zoology [= Shokubutsu - oyobi - Dobutsu], 2, 878 - 884." type="journal article" year="1934">1934c</bibRefCitation>
). Because we were not able to assess the actual date of publication of these papers, we provisionally consider the description by
<bibRefCitation id="EFA58DB4547D954C353F7C96FBE9FB78" author="Takakuwa, Y." box="[854,1074,1206,1232]" pageId="14" pageNumber="15" pagination="355 - 363" refId="ref55473" refString="Takakuwa, Y. (1934 a) Neue Japanische Mecistocephalidae. Annotationes Zoologicae Japonenses, 14, 355 - 363." type="journal article" year="1934">Takakuwa (1934a)</bibRefCitation>
as the original one, following the opinion of other authors (
<bibRefCitation id="EFA58DB4547D954C37857CFEFD57FB50" author="Crabill, R. E." box="[492,652,1246,1272]" pageId="14" pageNumber="15" pagination="161 - 170" refId="ref53240" refString="Crabill, R. E. (1964) A revised interpretation of the primitive centipede genus Arrup, with redescription of its type-species and list of known species. Proceedings of the Biological Society of Washington, 77, 161 - 170." type="journal article" year="1964">Crabill, 1964</bibRefCitation>
;
<bibRefCitation id="EFA58DB4547D954C34F17CFEFC86FB50" author="Shinohara, K." box="[664,861,1246,1272]" pageId="14" pageNumber="15" pagination="65 - 73" refId="ref55213" refString="Shinohara, K. (1972) Some chilopods of the Hidaka Mountain Range in Hokkaido, Northern Japan. Memoirs of the National Science Museum, Tokyo, 5, 65 - 73." type="journal article" year="1972">Shinohara, 1972</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>