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<document id="0F77BB7E6C2522B500B3763F873F9EA4" ID-CLB-Dataset="4235" ID-DOI="10.11646/zootaxa.3868.1.1" ID-GBIF-Dataset="c0b3c91a-f100-4743-afcc-d4e77ff11d9e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4930657" ID-ZooBank="ACC2AFCA-549B-4E2E-9E33-BF229FA3348A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623445173730" checkinUser="felipe" docAuthor="Almeida, Ana C. S., Souza, Facelucia B. C., Menegola, Carla M. S., Sanner, Joann &amp; Vieira, Leandro M." docDate="2014" docId="F20487E3FFDC2125F5FFFF1CFC113F53" docLanguage="en" docName="zt03868p061.pdf" docOrigin="Zootaxa 3868 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cigclisula porosa" docType="treatment" docVersion="10" lastPageNumber="20" masterDocId="0E3DFF9BFFCE2136F568FF84FF893B66" masterDocTitle="Taxonomic review of the family Colatooeciidae Winston, 2005 (Bryozoa, Cheilostomata), with description of seven new species" masterLastPageNumber="61" masterPageNumber="1" pageNumber="19" updateTime="1699013965909" updateUser="ExternalLinkService">
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<mods:title id="4325B684C12E8062E0231C63F41FC66E">Taxonomic review of the family Colatooeciidae Winston, 2005 (Bryozoa, Cheilostomata), with description of seven new species</mods:title>
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<mods:namePart id="8C6E539B51CD1AB310679B1552A9873A">Almeida, Ana C. S.</mods:namePart>
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<mods:namePart id="D7AB458ADAFB17833612F22AB61472E7">Souza, Facelucia B. C.</mods:namePart>
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<mods:namePart id="524B4713800FD431FF5224C170F46272">Menegola, Carla M. S.</mods:namePart>
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<mods:namePart id="4A0E409735171E664C9793A883DB0D1D">Sanner, Joann</mods:namePart>
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<mods:namePart id="D94A1F994D33AC67177E8B05774C73DF">Vieira, Leandro M.</mods:namePart>
<mods:nameIdentifier id="1CEE106E6BBF5E1C62F9158D387C970E" type="email">carol.salmeida@gmail.com</mods:nameIdentifier>
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<mods:title id="B53DEDE3A26311D3E82F0F9EEA4B7D07">Zootaxa</mods:title>
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<mods:date id="D4765227327330B14F03157DBBD57539">2014</mods:date>
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<mods:number id="521AB1A72F553C46B52864AA07A87AD4">3868</mods:number>
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<treatment id="F20487E3FFDC2125F5FFFF1CFC113F53" ID-DOI="http://doi.org/10.5281/zenodo.5523129" ID-GBIF-Taxon="188465080" ID-Zenodo-Dep="5523129" LSID="urn:lsid:plazi:treatment:F20487E3FFDC2125F5FFFF1CFC113F53" httpUri="http://treatment.plazi.org/id/F20487E3FFDC2125F5FFFF1CFC113F53" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<subSubSection id="32B7657EFFDC2124F5FFFF1CFD183BD4" box="[151,657,152,178]" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="7A1236F5FFDC2124F5FFFF1CFD183BD4" blockId="18.[151,657,152,211]" box="[151,657,152,178]" pageId="18" pageNumber="19">
<heading id="215A8199FFDC2124F5FFFF1CFD183BD4" bold="true" box="[151,657,152,178]" fontSize="11" level="2" pageId="18" pageNumber="19" reason="4">
<taxonomicName id="BDAD4D76FFDC2124F5FFFF1CFD183BD4" ID-CoL="VBMS" authority="(Canu &amp; Bassler, 1930)" baseAuthorityName="Canu &amp; Bassler" baseAuthorityYear="1930" box="[151,657,152,178]" class="Gymnolaemata" family="Colatooeciidae" genus="Cigclisula" kingdom="Animalia" order="Cheilostomatida" pageId="18" pageNumber="19" phylum="Bryozoa" rank="species" species="porosa">
<emphasis id="48D9EAE7FFDC2124F5FFFF1CFD183BD4" bold="true" box="[151,657,152,178]" pageId="18" pageNumber="19">
<emphasis id="48D9EAE7FFDC2124F5FFFF1CFEE03BD4" bold="true" box="[151,361,152,178]" italics="true" pageId="18" pageNumber="19">Cigclisula porosa</emphasis>
(Canu &amp; Bassler, 1930)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="32B7657EFFDC2124F5FFFF3EFED33BB5" box="[151,346,186,211]" pageId="18" pageNumber="19" type="description">
<paragraph id="7A1236F5FFDC2124F5FFFF3EFED33BB5" blockId="18.[151,657,152,211]" box="[151,346,186,211]" pageId="18" pageNumber="19">
(
<figureCitation id="E2962A70FFDC2124F5F7FF3EFF7B3BB5" box="[159,242,186,211]" captionStart="FIGURE 10" captionStartId="18.[151,250,1932,1954]" captionTargetBox="[161,1432,462,1915]" captionTargetId="figure-141@18.[153,1437,459,1924]" captionTargetPageId="18" captionText="FIGURE 10. Cigclisula porosa (Canu &amp; Bassler, 1930). A–F, USNM 8514, holotype, Galápagos Islands. A, part of encrusting colony. B, group of zooids. C, primary orifice. D, unusually narrow primary orifice. E, F, ovicelled zooids and elliptical frontal avicularia. Scale bars: A = 500 µm; B = 200 µm; C, D = 100 µm; E, F = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.4930679" httpUri="https://zenodo.org/record/4930679/files/figure.png" pageId="18" pageNumber="19">Fig. 10</figureCitation>
,
<tableCitation id="372F034EFFDC2124F595FF3EFED83BB5" box="[253,337,186,211]" captionStart="TABLE 2" captionStartId="13.[151,239,1427,1450]" captionTargetPageId="13" captionText="TABLE 2. Morphometric data for Cigclisula species studied (in mm)." httpUri="http://table.plazi.org/id/2ED2667DFFC3213BF5FFFA17FC0A3ECC" pageId="18" pageNumber="19" tableUuid="2ED2667DFFC3213BF5FFFA17FC0A3ECC">Table 2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="32B7657EFFDC2124F5FFFE86FB283A51" pageId="18" pageNumber="19" type="reference_group">
<paragraph id="7A1236F5FFDC2124F5FFFE86FC6D3A7E" blockId="18.[151,1185,258,311]" box="[151,996,258,280]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="5ABD11DBFFDC2124F5FFFE86FC6D3A7E" box="[151,996,258,280]" pageId="18" pageNumber="19">
<treatmentCitation id="FB0C10E4FFDC2124F5FFFE86FDF83A7E" author="Canu, F. &amp; Bassler, R. S." box="[151,625,258,280]" page="39" pageId="18" pageNumber="19" year="1930">
<taxonomicName id="BDAD4D76FFDC2124F5FFFE86FDF83A7E" authority="Canu &amp; Bassler 1930 a: 39" authorityName="F. Canu &amp; R. Bassler" authorityPageNumber="39" authorityYear="1930" box="[151,625,258,280]" class="Gymnolaemata" family="Lepraliellidae" genus="Holoporella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="18" pageNumber="19" phylum="Bryozoa" rank="species" species="porosa">
<emphasis id="48D9EAE7FFDC2124F5FFFE86FED43A7E" box="[151,349,258,280]" italics="true" pageId="18" pageNumber="19">Holoporella porosa</emphasis>
<bibRefCitation id="1E3C4B04FFDC2124F40BFE86FDF83A7E" author="Canu, F. &amp; Bassler, R. S." box="[355,625,258,280]" pageId="18" pageNumber="19" pagination="1 - 92" refId="ref30220" refString="Canu, F. &amp; Bassler, R. S. (1930 a) The Bryozoan Fauna of the Galapagos Islands. Proceedings of United States National Museum, Smithsonian Institution, Washington, 76 (2810), 1 - 92." type="journal article" year="1930">Canu &amp; Bassler 1930a: 39</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 6, figs. 7–8. [Galápagos Islands]
</treatmentCitationGroup>
</paragraph>
<paragraph id="7A1236F5FFDC2124F5FFFEA6FB283A51" blockId="18.[151,1185,258,311]" box="[151,1185,289,311]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="5ABD11DBFFDC2124F5FFFEA6FB283A51" box="[151,1185,289,311]" pageId="18" pageNumber="19">
<treatmentCitation id="FB0C10E4FFDC2124F5FFFEA6FDB03A51" author="Osburn, R. C." box="[151,569,289,311]" page="503" pageId="18" pageNumber="19" year="1952">
<taxonomicName id="BDAD4D76FFDC2124F5FFFEA6FDB03A51" authority=": Osburn, 1952: 503" authorityName="Osburn" authorityPageNumber="503" authorityYear="1952" box="[151,569,289,311]" class="Gymnolaemata" family="Colatooeciidae" genus="Trematooecia" kingdom="Animalia" order="Cheilostomatida" pageId="18" pageNumber="19" phylum="Bryozoa" rank="species" species="porosa">
<emphasis id="48D9EAE7FFDC2124F5FFFEA6FEE53A51" box="[151,364,290,311]" italics="true" pageId="18" pageNumber="19">Trematooecia porosa</emphasis>
:
<materialsCitation id="CAC53CA8FFDC2124F412FEA5FDB03A51" ID-GBIF-Occurrence="3357128336" box="[378,569,289,311]" location="Galapagos Islands" pageId="18" pageNumber="19" specimenCount="1" stateProvince="California">
<bibRefCitation id="1E3C4B04FFDC2124F412FEA5FDB03A51" author="Osburn, R. C." box="[378,569,289,311]" pageId="18" pageNumber="19" pagination="271 - 611" refId="ref31697" refString="Osburn, R. C. (1952) Bryozoa of the Pacific coast of America, part 2, Cheilostomata - Ascophora. Report of the Allan Hancock Pacific Expeditions, 14, 271 - 611." type="journal article" year="1952">Osburn, 1952: 503</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
, pl. 60, figs. 8–9. [Gulf of
<collectingRegion id="B869F817FFDC2124F620FEA5FC263A51" box="[840,943,289,311]" country="United States of America" name="California" pageId="18" pageNumber="19">California</collectingRegion>
and
<location id="7F72602EFFDC2124F688FEA5FB153A51" LSID="urn:lsid:plazi:treatment:F20487E3FFDC2125F5FFFF1CFC113F53:7F72602EFFDC2124F688FEA5FB153A51" box="[992,1180,289,311]" name="Galapagos Islands" pageId="18" pageNumber="19" stateProvince="California">Galápagos Islands</location>
]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="32B7657EFFDC2124F5FFFEE0FDEC3AF9" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="7A1236F5FFDC2124F5FFFEE0FDEC3AF9" blockId="18.[151,1436,356,416]" pageId="18" pageNumber="19">
<emphasis id="48D9EAE7FFDC2124F5FFFEE0FE0A3A1B" bold="true" box="[151,387,356,381]" pageId="18" pageNumber="19">Material examined.</emphasis>
<materialsCitation id="CAC53CA8FFDC2124F4E5FEE0FDEC3AF9" ID-GBIF-Occurrence="3357128375" collectionCode="USNM" collectorName="Albatross Station, D." location="Galapagos Islands" municipality="Albatross Station" pageId="18" pageNumber="19" specimenCode="USNM 8514" specimenCount="61" typeStatus="holotype">
<emphasis id="48D9EAE7FFDC2124F4E5FEE0FE743A1B" box="[397,509,356,381]" italics="true" pageId="18" pageNumber="19">
<typeStatus id="A5168857FFDC2124F4E5FEE0FE7E3A1B" box="[397,503,356,381]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
:
</emphasis>
<specimenCode id="2A0B9E8EFFDC2124F76FFEE0FD143A1B" box="[519,669,356,381]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="18" pageNumber="19" type="Museum">USNM 8514</specimenCode>
,
<taxonomicName id="BDAD4D76FFDC2124F7C3FEE0FB053A1B" authority="F. Canu &amp; R. Bassler" authorityName="F. Canu &amp; R. Bassler" authorityYear="1930" box="[683,1164,356,381]" class="Gymnolaemata" family="Lepraliellidae" genus="Holoporella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="18" pageNumber="19" phylum="Bryozoa" rank="species" species="porosa">
<emphasis id="48D9EAE7FFDC2124F7C3FEE0FC003A1A" box="[683,905,356,381]" italics="true" pageId="18" pageNumber="19">Holoporella porosa</emphasis>
F. Canu &amp; R. Bassler
</taxonomicName>
det., 1930a,
<location id="7F72602EFFDC2124F04EFEE0FF603AC6" LSID="urn:lsid:plazi:treatment:F20487E3FFDC2125F5FFFF1CFC113F53:7F72602EFFDC2124F04EFEE0FF603AC6" municipality="Albatross Station" name="Galapagos Islands" pageId="18" pageNumber="19">Galápagos Islands</location>
,
<collectorName id="D7585323FFDC2124F59CFE03FE543AC6" box="[244,477,391,416]" pageId="18" pageNumber="19">
<collectingMunicipality id="9A76AC8FFFDC2124F59CFE03FE313AC6" box="[244,440,391,416]" pageId="18" pageNumber="19">Albatross Station</collectingMunicipality>
, D.
</collectorName>
2815,
<specimenCount id="6CABFD7CFFDC2124F74DFE03FDEC3AF9" box="[549,613,391,416]" pageId="18" pageNumber="19" type="generic">61 m.</specimenCount>
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="2ED2667DFFDC2124F5FFF808FC473CB9" ID-DOI="http://doi.org/10.5281/zenodo.4930679" ID-Zenodo-Dep="4930679" httpUri="https://zenodo.org/record/4930679/files/figure.png" pageId="18" pageNumber="19" startId="18.[151,250,1932,1954]" targetBox="[161,1432,462,1915]" targetPageId="18">
<paragraph id="7A1236F5FFDC2124F5FFF808FC473CB9" blockId="18.[151,1436,1932,2015]" pageId="18" pageNumber="19">
<emphasis id="48D9EAE7FFDC2124F5FFF808FE963CC4" bold="true" box="[151,287,1932,1954]" pageId="18" pageNumber="19">FIGURE 10.</emphasis>
<taxonomicName id="BDAD4D76FFDC2124F44DF808FD593CC4" authority="(Canu &amp; Bassler, 1930)" baseAuthorityName="Canu &amp; Bassler" baseAuthorityYear="1930" box="[293,720,1932,1954]" class="Gymnolaemata" family="Colatooeciidae" genus="Cigclisula" kingdom="Animalia" order="Cheilostomatida" pageId="18" pageNumber="19" phylum="Bryozoa" rank="species" species="porosa">
<emphasis id="48D9EAE7FFDC2124F44DF808FE513CC4" box="[293,472,1932,1954]" italics="true" pageId="18" pageNumber="19">Cigclisula porosa</emphasis>
(Canu &amp; Bassler, 1930)
</taxonomicName>
.
<emphasis id="48D9EAE7FFDC2124F7B2F808FC843CC4" bold="true" box="[730,781,1932,1954]" pageId="18" pageNumber="19">A–F,</emphasis>
USNM 8514, holotype, Galápagos Islands.
<emphasis id="48D9EAE7FFDC2124F1A3F808FB6D3CC4" bold="true" box="[1227,1252,1932,1954]" pageId="18" pageNumber="19">A,</emphasis>
part of encrusting colony.
<emphasis id="48D9EAE7FFDC2124F58EF82FFF743CA7" bold="true" box="[230,253,1963,1985]" pageId="18" pageNumber="19">B,</emphasis>
group of zooids.
<emphasis id="48D9EAE7FFDC2124F4C7F82FFE413CA7" bold="true" box="[431,456,1963,1985]" pageId="18" pageNumber="19">C,</emphasis>
primary orifice.
<emphasis id="48D9EAE7FFDC2124F71BF82FFD023CA7" bold="true" box="[627,651,1963,1985]" pageId="18" pageNumber="19">D,</emphasis>
unusually narrow primary orifice.
<emphasis id="48D9EAE7FFDC2124F684F82FFB973CA7" bold="true" box="[1004,1054,1963,1985]" pageId="18" pageNumber="19">E, F,</emphasis>
ovicelled zooids and elliptical frontal avicularia. Scale bars: A = 500 µm; B = 200 µm; C, D = 100 µm; E, F = 200 µm.
</paragraph>
</caption>
<subSubSection id="32B7657EFFDD2125F5AFFF13FAC63AEE" pageId="19" pageNumber="20" type="description">
<paragraph id="7A1236F5FFDD2125F5AFFF13FAC63AEE" blockId="19.[151,1438,151,1077]" pageId="19" pageNumber="20">
<emphasis id="48D9EAE7FFDD2125F5AFFF13FED03BD6" bold="true" box="[199,345,151,176]" pageId="19" pageNumber="20">Description.</emphasis>
Colony encrusting, uni- to multilaminar. Zooids hexagonal, becoming polygonal with growth, longer than wide, delimited by distinct grooves. Frontal shield heavily calcified, uniformly punctured by 20–50 pseudopores, minutely granular. Primary orifice large relative to frontal shield, centered, somewhat hoof-shaped, mostly as long as wide, sunken, with arch-shaped anter and concave poster, separated by 2 proximolateral downcurved condyles. Secondary orifice non-tubercular, slightly raised, surrounded by pseudopores. Frontal avicularia small, elliptical, often single (rarely 2 per zooid), placed at zooidal margins. Ooecium subglobose, minutely granular, inclined toward zooid surface; ectooecium with longitudinal band of 33–48 irregular pseudopores.
</paragraph>
</subSubSection>
<subSubSection id="32B7657EFFDD2125F5AFFE10FCC43F76" pageId="19" pageNumber="20" type="discussion">
<paragraph id="7A1236F5FFDD2125F5AFFE10FCF5385E" blockId="19.[151,1438,151,1077]" pageId="19" pageNumber="20">
<emphasis id="48D9EAE7FFDD2125F5AFFE10FEB23ACB" bold="true" box="[199,315,404,429]" pageId="19" pageNumber="20">Remarks.</emphasis>
<taxonomicName id="BDAD4D76FFDD2125F424FE10FD953ACA" baseAuthorityName="Canu &amp; Bassler" baseAuthorityYear="1930" box="[332,540,404,429]" class="Gymnolaemata" family="Colatooeciidae" genus="Cigclisula" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="20" phylum="Bryozoa" rank="species" species="porosa">
<emphasis id="48D9EAE7FFDD2125F424FE10FD953ACA" box="[332,540,404,429]" italics="true" pageId="19" pageNumber="20">Cigclisula porosa</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="1E3C4B04FFDD2125F67BFE10FBC93ACB" author="Canu, F. &amp; Bassler, R. S." box="[787,1088,404,429]" pageId="19" pageNumber="20" pagination="1 - 92" refId="ref30220" refString="Canu, F. &amp; Bassler, R. S. (1930 a) The Bryozoan Fauna of the Galapagos Islands. Proceedings of United States National Museum, Smithsonian Institution, Washington, 76 (2810), 1 - 92." type="journal article" year="1930">Canu &amp; Bassler (1930a)</bibRefCitation>
in the same publication as
<taxonomicName id="BDAD4D76FFDD2125F5FFFE3CFD7E3AB6" authority="(Canu &amp; Bassler, 1930)" baseAuthorityName="Canu &amp; Bassler" baseAuthorityYear="1930" box="[151,759,439,464]" class="Gymnolaemata" family="Colatooeciidae" genus="Trematooecia" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="20" phylum="Bryozoa" rank="species" species="hexagonalis">
<emphasis id="48D9EAE7FFDD2125F5FFFE3CFE4F3AB6" box="[151,454,439,464]" italics="true" pageId="19" pageNumber="20">Trematooecia hexagonalis</emphasis>
(Canu &amp; Bassler, 1930)
</taxonomicName>
. While the ectooecium of
<taxonomicName id="BDAD4D76FFDD2125F122FE3CFB363AB6" baseAuthorityName="Canu &amp; Bassler" baseAuthorityYear="1930" box="[1098,1215,440,464]" class="Gymnolaemata" family="Colatooeciidae" genus="Cigclisula" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="20" phylum="Bryozoa" rank="species" species="porosa">
<emphasis id="48D9EAE7FFDD2125F122FE3CFB363AB6" box="[1098,1215,440,464]" italics="true" pageId="19" pageNumber="20">C. porosa</emphasis>
</taxonomicName>
was described as pseudoporous,
<bibRefCitation id="1E3C4B04FFDD2125F42DFE58FDEB3A93" author="Canu, F. &amp; Bassler, R. S." box="[325,610,476,501]" pageId="19" pageNumber="20" pagination="1 - 92" refId="ref30220" refString="Canu, F. &amp; Bassler, R. S. (1930 a) The Bryozoan Fauna of the Galapagos Islands. Proceedings of United States National Museum, Smithsonian Institution, Washington, 76 (2810), 1 - 92." type="journal article" year="1930">Canu &amp; Bassler (1930a)</bibRefCitation>
did not recognize ooecia in the
<typeStatus id="A5168857FFDD2125F68FFE58FBC03A93" box="[999,1097,476,501]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
of
<taxonomicName id="BDAD4D76FFDD2125F110FE59FA963A93" baseAuthorityName="Canu &amp; Bassler" baseAuthorityYear="1930" box="[1144,1311,476,501]" class="Gymnolaemata" family="Colatooeciidae" genus="Trematooecia" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="20" phylum="Bryozoa" rank="species" species="hexagonalis">
<emphasis id="48D9EAE7FFDD2125F110FE59FA963A93" box="[1144,1311,476,501]" italics="true" pageId="19" pageNumber="20">T. hexagonalis</emphasis>
</taxonomicName>
, but some authors (
<bibRefCitation id="1E3C4B04FFDD2125F59FFE7BFE04397E" author="Osburn, R. C." box="[247,397,511,536]" pageId="19" pageNumber="20" pagination="271 - 611" refId="ref31697" refString="Osburn, R. C. (1952) Bryozoa of the Pacific coast of America, part 2, Cheilostomata - Ascophora. Report of the Allan Hancock Pacific Expeditions, 14, 271 - 611." type="journal article" year="1952">Osburn 1952</bibRefCitation>
;
<bibRefCitation id="1E3C4B04FFDD2125F4F0FE7BFD90397E" author="Soule, J. D." box="[408,537,511,536]" pageId="19" pageNumber="20" pagination="1 - 66" refId="ref32035" refString="Soule, J. D. (1961) Results of the Puritan-American Museum of Natural History expedition to western Mexico. 13. Ascophoran Cheilostomata (Bryozoa) of the Gulf of California. American Museum Novitates, 2053, 1 - 66." type="journal article" year="1961">Soule 1961</bibRefCitation>
;
<bibRefCitation id="1E3C4B04FFDD2125F74DFE7BFC8E397E" author="Soule, D. F. &amp; Soule, J. D." box="[549,775,511,536]" pageId="19" pageNumber="20" pagination="1 - 56" refId="ref32080" refString="Soule, D. F. &amp; Soule, J. D. (1964) The Ectoprocta (Bryozoa) of Scammon's Lagoon, Baja California, Mexico. American Museum Novitates, 2199, 1 - 56." type="journal article" year="1964">Soule &amp; Soule 1964</bibRefCitation>
) described the ectooecium of this species as pseudoporous, designating these specimens as
<taxonomicName id="BDAD4D76FFDD2125F496FDA1FD2B395B" baseAuthorityName="Canu &amp; Bassler" baseAuthorityYear="1930" box="[510,674,548,573]" class="Gymnolaemata" family="Colatooeciidae" genus="Trematooecia" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="20" phylum="Bryozoa" rank="species" species="hexagonalis">
<emphasis id="48D9EAE7FFDD2125F496FDA1FD2B395B" box="[510,674,548,573]" italics="true" pageId="19" pageNumber="20">T. hexagonalis</emphasis>
</taxonomicName>
. Examination of the
<typeStatus id="A5168857FFDD2125F6FAFDA0FC7D395B" box="[914,1012,548,573]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
of
<taxonomicName id="BDAD4D76FFDD2125F174FDA1FB36395B" baseAuthorityName="Canu &amp; Bassler" baseAuthorityYear="1930" box="[1052,1215,548,573]" class="Gymnolaemata" family="Colatooeciidae" genus="Trematooecia" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="20" phylum="Bryozoa" rank="species" species="hexagonalis">
<emphasis id="48D9EAE7FFDD2125F174FDA1FB36395B" box="[1052,1215,548,573]" italics="true" pageId="19" pageNumber="20">T. hexagonalis</emphasis>
</taxonomicName>
, however, revealed that the ectooecium has single median membranous area (see remarks under
<taxonomicName id="BDAD4D76FFDD2125F162FDCDFB273906" baseAuthorityName="Canu &amp; Bassler" baseAuthorityYear="1930" box="[1034,1198,583,608]" class="Gymnolaemata" family="Colatooeciidae" genus="Trematooecia" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="20" phylum="Bryozoa" rank="species" species="hexagonalis">
<emphasis id="48D9EAE7FFDD2125F162FDCDFB273906" box="[1034,1198,583,608]" italics="true" pageId="19" pageNumber="20">T. hexagonalis</emphasis>
</taxonomicName>
) contrasting with an irregularly pseudoporous ectooecium, which is distinct in
<emphasis id="48D9EAE7FFDD2125F643FDE8FC2839E2" box="[811,929,620,644]" italics="true" pageId="19" pageNumber="20">
<taxonomicName id="BDAD4D76FFDD2125F643FDE8FC1439E2" baseAuthorityName="Canu &amp; Bassler" baseAuthorityYear="1930" box="[811,925,620,644]" class="Gymnolaemata" family="Colatooeciidae" genus="Cigclisula" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="20" phylum="Bryozoa" rank="species" species="porosa">C. porosa</taxonomicName>
.
</emphasis>
Besides the ectooecium and tubercles, other differences between these two species include the diameter of frontal pseudopores, larger in
<taxonomicName id="BDAD4D76FFDD2125F04CFD14FA1539CE" baseAuthorityName="Canu &amp; Bassler" baseAuthorityYear="1930" box="[1316,1436,656,680]" class="Gymnolaemata" family="Colatooeciidae" genus="Cigclisula" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="20" phylum="Bryozoa" rank="species" species="porosa">
<emphasis id="48D9EAE7FFDD2125F04CFD14FA1539CE" box="[1316,1436,656,680]" italics="true" pageId="19" pageNumber="20">C. porosa</emphasis>
</taxonomicName>
(
<quantity id="BD559B10FFDD2125F5F6FD30FEE839AB" box="[158,353,692,717]" metricMagnitude="-5" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="3.2" metricValueMin="1.5" pageId="19" pageNumber="20" unit="mm" value="0.0235" valueMax="0.032" valueMin="0.015">
<geoCoordinate id="1F995032FFDD2125F5F6FD30FF5439AB" box="[158,221,692,717]" degrees="0.015" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="55" value="0.015">0.015</geoCoordinate>
<geoCoordinate id="1F995032FFDD2125F5B5FD30FEA539AB" box="[221,300,692,717]" degrees="0.032" direction="west" orientation="longitude" pageId="19" pageNumber="20" precision="55" value="-0.032">–0.032</geoCoordinate>
mm
</quantity>
) than in
<taxonomicName id="BDAD4D76FFDD2125F4ACFD31FDEE39AB" baseAuthorityName="Canu &amp; Bassler" baseAuthorityYear="1930" box="[452,615,692,717]" class="Gymnolaemata" family="Colatooeciidae" genus="Trematooecia" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="20" phylum="Bryozoa" rank="species" species="hexagonalis">
<emphasis id="48D9EAE7FFDD2125F4ACFD31FDEE39AB" box="[452,615,692,717]" italics="true" pageId="19" pageNumber="20">T. hexagonalis</emphasis>
</taxonomicName>
(
<quantity id="BD559B10FFDD2125F71FFD30FCB539AB" box="[631,828,692,717]" metricMagnitude="-5" metricUnit="m" metricValue="1.4" metricValueMax="1.7999999999999998" metricValueMin="1.0" pageId="19" pageNumber="20" unit="mm" value="0.013999999999999999" valueMax="0.018" valueMin="0.01">
<geoCoordinate id="1F995032FFDD2125F71FFD30FD3E39AB" box="[631,695,692,717]" degrees="0.010" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="55" value="0.01">0.010</geoCoordinate>
<geoCoordinate id="1F995032FFDD2125F7DFFD30FC8D39AB" box="[695,772,692,717]" degrees="0.018" direction="west" orientation="longitude" pageId="19" pageNumber="20" precision="55" value="-0.018">–0.018</geoCoordinate>
mm
</quantity>
), and the size of frontal avicularia, larger in
<taxonomicName id="BDAD4D76FFDD2125F046FD30FA1539AA" baseAuthorityName="Canu &amp; Bassler" baseAuthorityYear="1930" box="[1326,1436,692,716]" class="Gymnolaemata" family="Colatooeciidae" genus="Cigclisula" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="20" phylum="Bryozoa" rank="species" species="porosa">
<emphasis id="48D9EAE7FFDD2125F046FD30FA1539AA" box="[1326,1436,692,716]" italics="true" pageId="19" pageNumber="20">C. porosa</emphasis>
</taxonomicName>
(
<quantity id="BD559B10FFDD2125F5F6FD53FEE83989" box="[158,353,727,752]" metricMagnitude="-5" metricUnit="m" metricValue="7.500000000000001" metricValueMax="8.200000000000001" metricValueMin="6.800000000000001" pageId="19" pageNumber="20" unit="mm" value="0.075" valueMax="0.082" valueMin="0.068">
<geoCoordinate id="1F995032FFDD2125F5F6FD53FF543996" box="[158,221,727,752]" degrees="0.068" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="55" value="0.068">0.068</geoCoordinate>
<geoCoordinate id="1F995032FFDD2125F5B5FD53FEA53996" box="[221,300,727,752]" degrees="0.082" direction="west" orientation="longitude" pageId="19" pageNumber="20" precision="55" value="-0.082">–0.082</geoCoordinate>
mm
</quantity>
long,
<quantity id="BD559B10FFDD2125F4C4FD53FDE43989" box="[428,621,727,752]" metricMagnitude="-5" metricUnit="m" metricValue="7.25" metricValueMax="7.5" metricValueMin="7" pageId="19" pageNumber="20" unit="mm" value="0.0725" valueMax="0.075" valueMin="0.07">
<geoCoordinate id="1F995032FFDD2125F4C4FD53FE623996" box="[428,491,727,752]" degrees="0.070" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="55" value="0.07">0.070</geoCoordinate>
<geoCoordinate id="1F995032FFDD2125F484FD53FDB13996" box="[492,568,727,752]" degrees="0.075" direction="west" orientation="longitude" pageId="19" pageNumber="20" precision="55" value="-0.075">–0.075</geoCoordinate>
mm
</quantity>
wide) than in
<taxonomicName id="BDAD4D76FFDD2125F67FFD5DFC353996" baseAuthorityName="Canu &amp; Bassler" baseAuthorityYear="1930" box="[791,956,727,752]" class="Gymnolaemata" family="Colatooeciidae" genus="Trematooecia" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="20" phylum="Bryozoa" rank="species" species="hexagonalis">
<emphasis id="48D9EAE7FFDD2125F67FFD5DFC353996" box="[791,956,727,752]" italics="true" pageId="19" pageNumber="20">T. hexagonalis</emphasis>
</taxonomicName>
(
<quantity id="BD559B10FFDD2125F6A5FD53FB193989" box="[973,1168,727,752]" metricMagnitude="-5" metricUnit="m" metricValue="3.65" metricValueMax="4.5" metricValueMin="2.8" pageId="19" pageNumber="20" unit="mm" value="0.0365" valueMax="0.045" valueMin="0.028">
<geoCoordinate id="1F995032FFDD2125F6A5FD53FB843996" box="[973,1037,727,752]" degrees="0.028" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="55" value="0.028">0.028</geoCoordinate>
<geoCoordinate id="1F995032FFDD2125F165FD53FBD53996" box="[1037,1116,727,752]" degrees="0.045" direction="west" orientation="longitude" pageId="19" pageNumber="20" precision="55" value="-0.045">–0.045</geoCoordinate>
mm
</quantity>
long,
<quantity id="BD559B10FFDD2125F1B3FD53FA153989" box="[1243,1436,727,752]" metricMagnitude="-5" metricUnit="m" metricValue="3.35" metricValueMax="3.9" metricValueMin="2.8" pageId="19" pageNumber="20" unit="mm" value="0.0335" valueMax="0.039" valueMin="0.028">
<geoCoordinate id="1F995032FFDD2125F1B3FD53FA933996" box="[1243,1306,727,752]" degrees="0.028" direction="north" orientation="latitude" pageId="19" pageNumber="20" precision="55" value="0.028">0.028</geoCoordinate>
<geoCoordinate id="1F995032FFDD2125F072FD53FAEE3996" box="[1306,1383,727,752]" degrees="0.039" direction="west" orientation="longitude" pageId="19" pageNumber="20" precision="55" value="-0.039">–0.039</geoCoordinate>
mm
</quantity>
wide). Specimens previously reported as
<taxonomicName id="BDAD4D76FFDD2125F70AFD79FC8D3873" baseAuthorityName="Canu &amp; Bassler" baseAuthorityYear="1930" box="[610,772,764,789]" class="Gymnolaemata" family="Colatooeciidae" genus="Trematooecia" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="20" phylum="Bryozoa" rank="species" species="hexagonalis">
<emphasis id="48D9EAE7FFDD2125F70AFD79FC8D3873" box="[610,772,764,789]" italics="true" pageId="19" pageNumber="20">T. hexagonalis</emphasis>
</taxonomicName>
from the Pacific (
<bibRefCitation id="1E3C4B04FFDD2125F6A6FD78FBEB3873" author="Osburn, R. C." box="[974,1122,764,789]" pageId="19" pageNumber="20" pagination="271 - 611" refId="ref31697" refString="Osburn, R. C. (1952) Bryozoa of the Pacific coast of America, part 2, Cheilostomata - Ascophora. Report of the Allan Hancock Pacific Expeditions, 14, 271 - 611." type="journal article" year="1952">Osburn 1952</bibRefCitation>
;
<bibRefCitation id="1E3C4B04FFDD2125F105FD78FB673873" author="Soule, J. D." box="[1133,1262,764,789]" pageId="19" pageNumber="20" pagination="1 - 66" refId="ref32035" refString="Soule, J. D. (1961) Results of the Puritan-American Museum of Natural History expedition to western Mexico. 13. Ascophoran Cheilostomata (Bryozoa) of the Gulf of California. American Museum Novitates, 2053, 1 - 66." type="journal article" year="1961">Soule 1961</bibRefCitation>
;
<bibRefCitation id="1E3C4B04FFDD2125F191FD78FF59385E" author="Soule, D. F. &amp; Soule, J. D." pageId="19" pageNumber="20" pagination="1 - 56" refId="ref32080" refString="Soule, D. F. &amp; Soule, J. D. (1964) The Ectoprocta (Bryozoa) of Scammon's Lagoon, Baja California, Mexico. American Museum Novitates, 2199, 1 - 56." type="journal article" year="1964">Soule &amp; Soule 1964</bibRefCitation>
) are here reassigned to
<taxonomicName id="BDAD4D76FFDD2125F4B1FC9BFD20385E" authority="Almeida &amp; Souza &amp; Menegola &amp; Sanner &amp; Vieira, 2014" authorityName="Almeida &amp; Souza &amp; Menegola &amp; Sanner &amp; Vieira" authorityYear="2014" box="[473,681,799,824]" class="Gymnolaemata" family="Colatooeciidae" genus="Cigclisula" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="20" phylum="Bryozoa" rank="species" species="osburni" status="sp. nov.">
<emphasis id="48D9EAE7FFDD2125F4B1FC9BFD20385E" box="[473,681,799,824]" italics="true" pageId="19" pageNumber="20">Cigclisula osburni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="53EA579CFFDD2125F7D8FCA4FD653851" box="[688,748,800,823]" pageId="19" pageNumber="20" rank="species">n. sp.</taxonomicNameLabel>
(see below).
</paragraph>
<paragraph id="7A1236F5FFDD2125F5AFFCC0FCC43F76" blockId="19.[151,1438,151,1077]" pageId="19" pageNumber="20">
<taxonomicName id="BDAD4D76FFDD2125F5AFFCC0FE06383A" baseAuthorityName="Canu &amp; Bassler" baseAuthorityYear="1930" box="[199,399,836,861]" class="Gymnolaemata" family="Colatooeciidae" genus="Cigclisula" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="20" phylum="Bryozoa" rank="species" species="porosa">
<emphasis id="48D9EAE7FFDD2125F5AFFCC0FE06383A" box="[199,399,836,861]" italics="true" pageId="19" pageNumber="20">Cigclisula porosa</emphasis>
</taxonomicName>
is characterized by encrusting colonies with the zooidal frontal shield uniformly punctured by 20–50 pseudopores, primary orifice almost rounded with distolateral condyles, secondary orifice nontubercular, elliptical frontal avicularia and ectooecium with 33–48 irregular pseudopores. This species resembles
<taxonomicName id="BDAD4D76FFDD2125F5FFFC34FE9338AE" authorityName=": Livingstone" authorityYear="1926" box="[151,282,943,968]" class="Gymnolaemata" family="Bryocryptellidae" genus="Porella" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="20" phylum="Bryozoa" rank="species" species="areolata">
<emphasis id="48D9EAE7FFDD2125F5FFFC34FE9338AE" box="[151,282,943,968]" italics="true" pageId="19" pageNumber="20">C. areolata</emphasis>
</taxonomicName>
in having encrusting colonies, a uniformly punctured frontal shield and non-tubercular secondary orifice, but differs in the the hoof-shaped primary orifice (transversely D-shaped in
<taxonomicName id="BDAD4D76FFDD2125F157FC50FB37388B" authorityName=": Livingstone" authorityYear="1926" box="[1087,1214,980,1005]" class="Gymnolaemata" family="Bryocryptellidae" genus="Porella" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="20" phylum="Bryozoa" rank="species" species="areolata">
<emphasis id="48D9EAE7FFDD2125F157FC50FB37388B" box="[1087,1214,980,1005]" italics="true" pageId="19" pageNumber="20">C. areolata</emphasis>
</taxonomicName>
) and in the absence of suboral and interzooidal avicularia (present in
<taxonomicName id="BDAD4D76FFDD2125F7D6FC7CFCB43F76" authorityName=": Livingstone" authorityYear="1926" box="[702,829,1015,1040]" class="Gymnolaemata" family="Bryocryptellidae" genus="Porella" kingdom="Animalia" order="Cheilostomatida" pageId="19" pageNumber="20" phylum="Bryozoa" rank="species" species="areolata">
<emphasis id="48D9EAE7FFDD2125F7D6FC7CFCB43F76" box="[702,829,1015,1040]" italics="true" pageId="19" pageNumber="20">C. areolata</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="32B7657EFFDD2125F5AFFB98FC113F53" box="[199,920,1052,1077]" pageId="19" pageNumber="20" type="distribution">
<paragraph id="7A1236F5FFDD2125F5AFFB98FC113F53" blockId="19.[151,1438,151,1077]" box="[199,920,1052,1077]" pageId="19" pageNumber="20">
<emphasis id="48D9EAE7FFDD2125F5AFFB98FEE93F53" bold="true" box="[199,352,1052,1077]" pageId="19" pageNumber="20">Distribution.</emphasis>
Pacific: Gulf of
<collectingRegion id="B869F817FFDD2125F774FB98FD073F53" box="[540,654,1052,1077]" country="United States of America" name="California" pageId="19" pageNumber="20">California</collectingRegion>
and Galápagos Islands.
</paragraph>
</subSubSection>
</treatment>
</document>
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