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23:
<bibRefCitation id="6333F2F9131ABF0BFED1FB17FDD0469A" author="Gippner, S. &amp; Travers S. L. &amp; Scherz M. D. &amp; Colston T. J. &amp; Lyra M. L. &amp; Mohan A. V. &amp; Multzsch M. &amp; Nielsen S. V. &amp; Rancilhac L. &amp; Glaw F. &amp; Bauer A. M. &amp; Vences M." box="[366,579,1169,1193]" pageId="48" pageNumber="49" pagination="107311" refId="ref37826" refString="Gippner, S., Travers S. L., Scherz M. D., Colston T. J., Lyra M. L., Mohan A. V., Multzsch M., Nielsen S. V., Rancilhac L., Glaw F., Bauer A. M. &amp; Vences M. (2021) A comprehensive phylogeny of dwarf geckos of the genus Lygodactylus, with insights into their systematics and morphological variation. Molecular Phylogenetics and Evolution, 165, 107311. https: // doi. org / 10.1016 / j. ympev. 2021.107311" type="journal article" year="2021">
Gippner
<emphasis id="35D6531A131ABF0BFE78FB14FE67469A" box="[455,500,1169,1193]" italics="true" pageId="48" pageNumber="49">et al</emphasis>
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<paragraph id="071D8F08131ABF0BFF78FAB8FD16444B" blockId="48.[151,1437,1234,2017]" pageId="48" pageNumber="49">
<emphasis id="35D6531A131ABF0BFF78FAB8FED1476B" bold="true" box="[199,322,1342,1368]" pageId="48" pageNumber="49">Diagnosis.</emphasis>
<taxonomicName id="C0A2F48B131ABF0BFEF7FAB8FDDB476B" authority="Vences &amp; Multzsch &amp; Gippner &amp; Miralles &amp; Crottini &amp; Gehring &amp; Rakotoarison &amp; Ratsoavina &amp; Glaw &amp; Scherz, 2022" authorityName="Vences &amp; Multzsch &amp; Gippner &amp; Miralles &amp; Crottini &amp; Gehring &amp; Rakotoarison &amp; Ratsoavina &amp; Glaw &amp; Scherz" authorityYear="2022" box="[328,584,1342,1368]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="hodikazo" status="sp. nov.">
<emphasis id="35D6531A131ABF0BFEF7FAB8FDDB476B" box="[328,584,1342,1368]" italics="true" pageId="48" pageNumber="49">Lygodactylus hodikazo</emphasis>
</taxonomicName>
<emphasis id="35D6531A131ABF0BFDF0FAB8FD3B476B" bold="true" box="[591,680,1342,1368]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="2EE5EE61131ABF0BFDF0FAB8FD3B476B" box="[591,680,1342,1368]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the only
<taxonomicName id="C0A2F48B131ABF0BFC96FAB8FC2F476B" authorityName="Pasteur" authorityYear="1964" box="[809,956,1342,1368]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
<emphasis id="35D6531A131ABF0BFC96FAB8FC2F476B" box="[809,956,1342,1368]" italics="true" pageId="48" pageNumber="49">Domerguella</emphasis>
</taxonomicName>
lineage known from the West of
<collectingCountry id="7FB5CF98131ABF0BFA91FAB8FF28474F" name="Madagascar" pageId="48" pageNumber="49">Madagascar</collectingCountry>
, and only
<specimenCount id="11A44481131ABF0BFE92FAE4FE55474F" box="[301,454,1378,1404]" pageId="48" pageNumber="49" type="generic">one specimen</specimenCount>
is so far known. In our multigene analysis, it forms with high support the sister taxon of the clade of
<taxonomicName id="C0A2F48B131ABF0BFE9BFA01FD9F4793" authorityName=": Pasteur" authorityYear="1965" box="[292,524,1414,1440]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="madagascariensis">
<emphasis id="35D6531A131ABF0BFE9BFA01FD9F4793" box="[292,524,1414,1440]" italics="true" pageId="48" pageNumber="49">L. madagascariensis</emphasis>
</taxonomicName>
and
<taxonomicName id="C0A2F48B131ABF0BFDFAFA01FD3F4793" authorityName="Pasteur &amp; Blanc" authorityYear="1967" box="[581,684,1415,1440]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="petteri">
<emphasis id="35D6531A131ABF0BFDFAFA01FD3F4793" box="[581,684,1415,1440]" italics="true" pageId="48" pageNumber="49">L. petteri</emphasis>
</taxonomicName>
, thus belonging to subclade A4 of
<taxonomicName id="C0A2F48B131ABF0BFB86FA00FB5F4793" authorityName="Pasteur" authorityYear="1964" box="[1081,1228,1414,1440]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
<emphasis id="35D6531A131ABF0BFB86FA00FB5F4793" box="[1081,1228,1414,1440]" italics="true" pageId="48" pageNumber="49">Domerguella</emphasis>
</taxonomicName>
as defined herein. Since these two species are to be considered as distinct species given their sympatric occurrence without allele sharing in the analysed nuclear markers, it is justified to allocate species status also to this western lineage, despite its similarity in morphological characters (scale counts) to
<taxonomicName id="C0A2F48B131ABF0BFC89FA75FB8C443F" authorityName=": Pasteur" authorityYear="1965" box="[822,1055,1522,1548]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="madagascariensis">
<emphasis id="35D6531A131ABF0BFC89FA75FB8C443F" box="[822,1055,1522,1548]" italics="true" pageId="48" pageNumber="49">L. madagascariensis</emphasis>
</taxonomicName>
. The justification for this species thus mainly relies on its phylogenetic position, on the high mitochondrial divergence (&gt;8% 16S distance), and large geographical and ecological distance (only
<taxonomicName id="C0A2F48B131ABF0BFD36F9BCFC8F4467" authorityName="Pasteur" authorityYear="1964" box="[649,796,1594,1620]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
<emphasis id="35D6531A131ABF0BFD36F9BCFC8F4467" box="[649,796,1594,1620]" italics="true" pageId="48" pageNumber="49">Domerguella</emphasis>
</taxonomicName>
in western
<collectingCountry id="7FB5CF98131ABF0BFC1BF9BCFBBE4467" box="[932,1069,1594,1620]" name="Madagascar" pageId="48" pageNumber="49">Madagascar</collectingCountry>
) to
<emphasis id="35D6531A131ABF0BFBE2F9BDFADD4467" box="[1117,1358,1594,1620]" italics="true" pageId="48" pageNumber="49">
<taxonomicName id="C0A2F48B131ABF0BFBE2F9BDFAD94467" authorityName=": Pasteur" authorityYear="1965" box="[1117,1354,1594,1620]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="madagascariensis">L. madagascariensis</taxonomicName>
,
</emphasis>
which morphologically is the most similar species.
</paragraph>
<paragraph id="071D8F08131ABF0AFF78F904FA1B4356" blockId="48.[151,1437,1234,2017]" lastBlockId="49.[151,1437,151,1293]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
<taxonomicName id="C0A2F48B131ABF0BFF78F904FE5444AF" authority="Vences &amp; Multzsch &amp; Gippner &amp; Miralles &amp; Crottini &amp; Gehring &amp; Rakotoarison &amp; Ratsoavina &amp; Glaw &amp; Scherz, 2022" authorityName="Vences &amp; Multzsch &amp; Gippner &amp; Miralles &amp; Crottini &amp; Gehring &amp; Rakotoarison &amp; Ratsoavina &amp; Glaw &amp; Scherz" authorityYear="2022" box="[199,455,1666,1692]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="hodikazo" status="sp. nov.">
<emphasis id="35D6531A131ABF0BFF78F904FE5444AF" box="[199,455,1666,1692]" italics="true" pageId="48" pageNumber="49">Lygodactylus hodikazo</emphasis>
</taxonomicName>
<emphasis id="35D6531A131ABF0BFE72F904FDB544AF" bold="true" box="[461,550,1666,1692]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="2EE5EE61131ABF0BFE72F904FDB544AF" box="[461,550,1666,1692]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be assigned to the subgenus
<taxonomicName id="C0A2F48B131ABF0BFC2DF904FBB644AF" authorityName="Pasteur" authorityYear="1964" box="[914,1061,1666,1692]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
<emphasis id="35D6531A131ABF0BFC2DF904FBB644AF" box="[914,1061,1666,1692]" italics="true" pageId="48" pageNumber="49">Domerguella</emphasis>
</taxonomicName>
morphologically by an undivided mental scale with two postmentals, and absence of a claw on the first finger. Within
<taxonomicName id="C0A2F48B131ABF0BFBF4F920FB4D44F3" authorityName="Pasteur" authorityYear="1964" box="[1099,1246,1702,1728]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
<emphasis id="35D6531A131ABF0BFBF4F920FB4D44F3" box="[1099,1246,1702,1728]" italics="true" pageId="48" pageNumber="49">Domerguella</emphasis>
</taxonomicName>
, the new species is the only lineage known from western
<collectingCountry id="7FB5CF98131ABF0BFDDAF94CFD7D44D7" box="[613,750,1738,1764]" name="Madagascar" pageId="48" pageNumber="49">Madagascar</collectingCountry>
, and belongs to subclade A4. From the species of subclades A1A4 occurring in northern
<collectingCountry id="7FB5CF98131ABF0BFE56F968FDE3453B" box="[489,624,1774,1800]" name="Madagascar" pageId="48" pageNumber="49">Madagascar</collectingCountry>
except
<taxonomicName id="C0A2F48B131ABF0BFD75F969FC21453B" authorityName=": Pasteur" authorityYear="1965" box="[714,946,1774,1800]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="madagascariensis">
<emphasis id="35D6531A131ABF0BFD75F969FC21453B" box="[714,946,1774,1800]" italics="true" pageId="48" pageNumber="49">L. madagascariensis</emphasis>
</taxonomicName>
it differs as follows: from
<taxonomicName id="C0A2F48B131ABF0BFB5BF969FAEB453B" authorityName=": Rosler" authorityYear="2000" box="[1252,1400,1775,1800]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="expectatus">
<emphasis id="35D6531A131ABF0BFB5BF969FAEB453B" box="[1252,1400,1775,1800]" italics="true" pageId="48" pageNumber="49">L. expectatus</emphasis>
</taxonomicName>
by non-enlarged dorsolateral scales (longitudinal count of dorsal scales 231 vs. &lt;170); from
<taxonomicName id="C0A2F48B131ABF0BFBC8F895FB42451F" authorityName=": Rosler" authorityYear="2000" box="[1143,1233,1811,1836]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="rarus">
<emphasis id="35D6531A131ABF0BFBC8F895FB42451F" box="[1143,1233,1811,1836]" italics="true" pageId="48" pageNumber="49">L. rarus</emphasis>
</taxonomicName>
by lack of regular crossbands on tail (vs. presence) and different body shape without elongated limbs (relative hindlimb length 0.44 vs.&gt;0.55); from
<emphasis id="35D6531A131ABF0BFE94F8DDFDFE4547" box="[299,621,1882,1908]" italics="true" pageId="48" pageNumber="49">
<taxonomicName id="C0A2F48B131ABF0BFE94F8DDFE074547" authorityName="Pasteur &amp; Blanc" authorityYear="1967" box="[299,404,1883,1908]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="petteri">L. petteri</taxonomicName>
,
<taxonomicName id="C0A2F48B131ABF0BFE21F8DDFE614547" authorityName="Vences &amp; Multzsch &amp; Gippner &amp; Miralles &amp; Crottini &amp; Gehring &amp; Rakotoarison &amp; Ratsoavina &amp; Glaw &amp; Scherz" authorityYear="2022" box="[414,498,1882,1908]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="salvi">L. salvi</taxonomicName>
,
<taxonomicName id="C0A2F48B131ABF0BFE43F8DDFDFA4547" authorityName="Vences &amp; Multzsch &amp; Gippner &amp; Miralles &amp; Crottini &amp; Gehring &amp; Rakotoarison &amp; Ratsoavina &amp; Glaw &amp; Scherz" authorityYear="2022" box="[508,617,1882,1908]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="roellae">L. roellae</taxonomicName>
,
</emphasis>
and
<taxonomicName id="C0A2F48B131ABF0BFD1DF8DDFD6F4547" authorityName="Vences &amp; Multzsch &amp; Gippner &amp; Miralles &amp; Crottini &amp; Gehring &amp; Rakotoarison &amp; Ratsoavina &amp; Glaw &amp; Scherz" authorityYear="2022" box="[674,764,1882,1908]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="hapei">
<emphasis id="35D6531A131ABF0BFD1DF8DDFD6F4547" box="[674,764,1882,1908]" italics="true" pageId="48" pageNumber="49">L. hapei</emphasis>
</taxonomicName>
by a higher longitudinal count of dorsal scales (231 vs. 159 222), and from
<taxonomicName id="C0A2F48B131ABF0BFEF7F8F9FE2645AB" authorityName="Vences &amp; Multzsch &amp; Gippner &amp; Miralles &amp; Crottini &amp; Gehring &amp; Rakotoarison &amp; Ratsoavina &amp; Glaw &amp; Scherz" authorityYear="2022" box="[328,437,1918,1944]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="roellae">
<emphasis id="35D6531A131ABF0BFEF7F8F9FE2645AB" box="[328,437,1918,1944]" italics="true" pageId="48" pageNumber="49">L. roellae</emphasis>
</taxonomicName>
and
<taxonomicName id="C0A2F48B131ABF0BFE52F8F9FDD945AB" authorityName="Vences &amp; Multzsch &amp; Gippner &amp; Miralles &amp; Crottini &amp; Gehring &amp; Rakotoarison &amp; Ratsoavina &amp; Glaw &amp; Scherz" authorityYear="2022" box="[493,586,1918,1944]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="hapei">
<emphasis id="35D6531A131ABF0BFE52F8F9FDD945AB" box="[493,586,1918,1944]" italics="true" pageId="48" pageNumber="49">L. hapei</emphasis>
</taxonomicName>
also by a higher longitudinal count of ventral scales (109 vs. 8392); from
<taxonomicName id="C0A2F48B131ABF0BFF28F825FE85458F" authorityName="Vences &amp; Multzsch &amp; Gippner &amp; Miralles &amp; Crottini &amp; Gehring &amp; Rakotoarison &amp; Ratsoavina &amp; Glaw &amp; Scherz" authorityYear="2022" box="[151,278,1954,1980]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="tantsaha">
<emphasis id="35D6531A131ABF0BFF28F825FE85458F" box="[151,278,1954,1980]" italics="true" pageId="48" pageNumber="49">L. tantsaha</emphasis>
</taxonomicName>
by a higher number of supralabials (10 vs. 67) and infralabials (8 vs. 56). Genetically, the new species is highly distinct from all species in subclade A5, and we hypothesize that it will differ from many of these by less expressed tubercles at the tail base in males (these tubercles are weakly expressed in the female
<typeStatus id="D81931AA131BBF0AFB47FF11FACC4282" box="[1272,1375,151,177]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
). The new species is morphologically most similar to
<taxonomicName id="C0A2F48B131BBF0AFD0BFF3DFC0E42E6" authorityName=": Pasteur" authorityYear="1965" box="[692,925,187,213]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="madagascariensis">
<emphasis id="35D6531A131BBF0AFD0BFF3DFC0E42E6" box="[692,925,187,213]" italics="true" pageId="49" pageNumber="50">L. madagascariensis</emphasis>
</taxonomicName>
from which it differs by 8.410.5% pairwise distance in the 16S gene, and has different haplotypes in RAG1 and CMOS (although intermixed with haplotypes of
<taxonomicName id="C0A2F48B131BBF0AFF28FE85FEEC432E" authorityName=": Pasteur" authorityYear="1965" box="[151,383,259,285]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="madagascariensis">
<emphasis id="35D6531A131BBF0AFF28FE85FEEC432E" box="[151,383,259,285]" italics="true" pageId="49" pageNumber="50">L. madagascariensis</emphasis>
</taxonomicName>
in the network). The new species also differs as far as known from
<taxonomicName id="C0A2F48B131BBF0AFBC2FE85FAF6432E" authorityName=": Pasteur" authorityYear="1965" box="[1149,1381,259,285]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="madagascariensis">
<emphasis id="35D6531A131BBF0AFBC2FE85FAF6432E" box="[1149,1381,259,285]" italics="true" pageId="49" pageNumber="50">L. madagascariensis</emphasis>
</taxonomicName>
, and indeed from all other
<taxonomicName id="C0A2F48B131BBF0AFE38FEA1FD894372" authorityName="Pasteur" authorityYear="1964" box="[391,538,295,321]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
<emphasis id="35D6531A131BBF0AFE38FEA1FD894372" box="[391,538,295,321]" italics="true" pageId="49" pageNumber="50">Domerguella</emphasis>
</taxonomicName>
, by a high number of supralabials (10 vs.
<quantity id="C05A22ED131BBF0AFC52FEA1FBA04372" box="[1005,1075,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" metricValueMax="2.286" metricValueMin="1.778" pageId="49" pageNumber="50" unit="in" value="8.0" valueMax="9.0" valueMin="7.0">79 in</quantity>
<taxonomicName id="C0A2F48B131BBF0AFB85FEA1FAB24372" authorityName=": Pasteur" authorityYear="1965" box="[1082,1313,295,321]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="madagascariensis">
<emphasis id="35D6531A131BBF0AFB85FEA1FAB24372" box="[1082,1313,295,321]" italics="true" pageId="49" pageNumber="50">L. madagascariensis</emphasis>
</taxonomicName>
and
<quantity id="C05A22ED131BBF0AFAE9FEA0FA0F4372" box="[1366,1436,294,321]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" metricValueMax="2.286" metricValueMin="1.27" pageId="49" pageNumber="50" unit="in" value="7.0" valueMax="9.0" valueMin="5.0">59 in</quantity>
all other species;
<tableCitation id="4A20BAB3131BBF0AFEE5FECDFE3F4356" box="[346,428,331,357]" captionStart="TABLE 1" captionStartId="31.[151,239,459,485]" captionTargetPageId="31" captionText="TABLE 1. Selected morphometric and meristic data for voucher specimens of the Lygodactylus madagascariensis group examined for this study. All morphometric measurements (SVL, TAL, HIL, HW, ED, SED) in mm. All specimens in the table were genotyped by 16S sequences, except for historical types from BMNH, MNHN, and SMF, and two further individuals of L. expectatus. Abbreviations are those described in Materials and Methods section; additional abbreviations and symbols as follows: NA, not assessed or not applicable; * counts and measurements partly or fully taken by M. Puente (Puente et al. 2009); § specimen not genotyped; # tail broken or incomplete; ## tail regenerated. HT (sep) and HT (spi) mark the holotypes of Lygodactylus septemtuberculatus Angel, 1942 and Microscalabotes spinulifer Boettger, 1913, here considered to be junior synonyms of L. miops. First LCDS value for L. expectatus refers to the enlarged dorsolateral scales, the second value to the non-enlarged vertebral scales. See species account of L. fritzi sp. nov for the identity of the historical locality “Senbendrana”." httpUri="http://table.plazi.org/id/53DDDF801335BF24FF28FE4DFD0E408E" pageId="49" pageNumber="50" tableUuid="53DDDF801335BF24FF28FE4DFD0E408E">Table 1</tableCitation>
); this character however requires confirmation by examination of additional specimens.
</paragraph>
</subSubSection>
<subSubSection id="4FB8DC83131BBF0AFF78FEE9FC7A43E2" pageId="49" pageNumber="50" type="etymology">
<paragraph id="071D8F08131BBF0AFF78FEE9FC7A43E2" blockId="49.[151,1437,151,1293]" pageId="49" pageNumber="50">
<emphasis id="35D6531A131BBF0AFF78FEE9FEDD43BA" bold="true" box="[199,334,367,393]" pageId="49" pageNumber="50">Etymology.</emphasis>
The species epithet is derived from the Malagasy words hodi-kazo (= tree bark), in reference to the habitat of most
<taxonomicName id="C0A2F48B131BBF0AFEF8FE15FE49439E" authorityName="Pasteur" authorityYear="1964" box="[327,474,403,429]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
<emphasis id="35D6531A131BBF0AFEF8FE15FE49439E" box="[327,474,403,429]" italics="true" pageId="49" pageNumber="50">Domerguella</emphasis>
</taxonomicName>
(and supposedly also this species) who are typically found on trees in forest and with a color pattern mimicking the bark. The name is used as noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="4FB8DC83131BBF0AFF78FE5CFCFE466A" pageId="49" pageNumber="50" type="description">
<paragraph id="071D8F08131BBF0AFF78FE5CFC2841B2" blockId="49.[151,1437,151,1293]" pageId="49" pageNumber="50">
<emphasis id="35D6531A131BBF0AFF78FE5CFD8143C6" bold="true" box="[199,530,474,501]" pageId="49" pageNumber="50">
Description of the
<typeStatus id="D81931AA131BBF0AFE1BFE5DFD9E43C6" box="[420,525,475,501]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult female, in a good state of preservation (
<figureCitation id="9F99938D131BBF0AFBA3FE5DFBE343C6" box="[1052,1136,474,501]" captionStart="FIGURE 15" captionStartId="25.[151,250,1943,1967]" captionTargetBox="[244,1351,181,1906]" captionTargetId="figure-18@25.[236,1352,181,1919]" captionTargetPageId="25" captionText="FIGURE 15. Dorsal and ventral views of preserved holotypes of new species of the Lygodactylus madagascariensis group named in the current study." figureDoi="http://doi.org/10.5281/zenodo.7040784" httpUri="https://zenodo.org/record/7040784/files/figure.png" pageId="49" pageNumber="50">Fig. 15</figureCitation>
). SVL
<quantity id="C05A22ED131BBF0AFB00FE5DFAB543C6" box="[1215,1318,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="2.94" pageId="49" pageNumber="50" unit="mm" value="29.4">29.4 mm</quantity>
, TAL
<quantity id="C05A22ED131BBF0AFAD4FE5DFF55402A" metricMagnitude="-2" metricUnit="m" metricValue="3.72" pageId="49" pageNumber="50" unit="mm" value="37.2">37.2 mm</quantity>
; for other measurements see
<tableCitation id="4A20BAB3131BBF0AFDAEFE79FDF6402A" box="[529,613,511,537]" captionStart="TABLE 1" captionStartId="31.[151,239,459,485]" captionTargetPageId="31" captionText="TABLE 1. Selected morphometric and meristic data for voucher specimens of the Lygodactylus madagascariensis group examined for this study. All morphometric measurements (SVL, TAL, HIL, HW, ED, SED) in mm. All specimens in the table were genotyped by 16S sequences, except for historical types from BMNH, MNHN, and SMF, and two further individuals of L. expectatus. Abbreviations are those described in Materials and Methods section; additional abbreviations and symbols as follows: NA, not assessed or not applicable; * counts and measurements partly or fully taken by M. Puente (Puente et al. 2009); § specimen not genotyped; # tail broken or incomplete; ## tail regenerated. HT (sep) and HT (spi) mark the holotypes of Lygodactylus septemtuberculatus Angel, 1942 and Microscalabotes spinulifer Boettger, 1913, here considered to be junior synonyms of L. miops. First LCDS value for L. expectatus refers to the enlarged dorsolateral scales, the second value to the non-enlarged vertebral scales. See species account of L. fritzi sp. nov for the identity of the historical locality “Senbendrana”." httpUri="http://table.plazi.org/id/53DDDF801335BF24FF28FE4DFD0E408E" pageId="49" pageNumber="50" tableUuid="53DDDF801335BF24FF28FE4DFD0E408E">Table 1</tableCitation>
. Head and neck long, body broader than head. The distance from the tip of the snout to the anterior border of the eye (
<quantity id="C05A22ED131BBF0AFD27FDA5FD62400E" box="[664,753,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="49" pageNumber="50" unit="mm" value="3.9">3.9 mm</quantity>
) is greater than the interorbital distance anteriorly (
<quantity id="C05A22ED131BBF0AFA8AFDA5FA03400E" box="[1333,1424,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="49" pageNumber="50" unit="mm" value="3.8">3.8 mm</quantity>
), and greater than the distance between the eye and ear opening. Snout covered with granular scales larger than those on the rest of the dorsum. Nostril surrounded by five scales: rostral, first supralabial, and three supranasals. Mental scale undivided; small contact between posterior projection of mental scale and first infralabial; two asymmetrical postmental scales with four postpostmental scales; eight infralabial scales; ten supralabial scales; one internasal scale; granular dorsal scales; dorsum with small, homogeneous, granular and unkeeled scales of similar size to those on trunk, smaller than on head and tail, the scales on limbs can be slightly larger; 231 dorsal scales longitudinally along the body; 109 ventral scales between mental and cloaca; venter with large homogeneous smooth scales; first finger present but very small, not bearing a claw; three pairs of subdigital lamellae on the fourth toe; no dorsolateral tubercles; tail without whorls; small lateral spines at the base of the tail.
</paragraph>
<paragraph id="071D8F08131BBF0AFF78FC0DFF7E4606" blockId="49.[151,1437,151,1293]" pageId="49" pageNumber="50">
Coloration of the
<typeStatus id="D81931AA131BBF0AFE34FC0DFE7E4196" box="[395,493,907,933]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
is only described from the preserved specimen after 15 years in ethanol. The dorsum is fawn to brownish with two symmetrical pairs of brown stripes on the neck. A brownish stripe runs from the snout to the eye and splits into three thinner stripes behind the eye followed by an irregular pattern of brownish stripes on the flanks. The tail is beige with weak patterning. Venter and the snout are uniformly whitish without a distinct pattern.
</paragraph>
<paragraph id="071D8F08131BBF0AFF78FBB8FCFE466A" blockId="49.[151,1437,151,1293]" box="[199,877,1086,1113]" pageId="49" pageNumber="50">
<emphasis id="35D6531A131BBF0AFF78FBB8FEAC466B" bold="true" box="[199,319,1086,1112]" pageId="49" pageNumber="50">Variation.</emphasis>
Only a single individual (the
<typeStatus id="D81931AA131BBF0AFD32FBB9FD62466A" box="[653,753,1087,1113]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
) is known.
</paragraph>
</subSubSection>
<subSubSection id="4FB8DC83131BBF0AFF78FBE4FE014692" pageId="49" pageNumber="50" type="distribution">
<paragraph id="071D8F08131BBF0AFF78FBE4FE014692" blockId="49.[151,1437,151,1293]" pageId="49" pageNumber="50">
<emphasis id="35D6531A131BBF0AFF78FBE4FEF2464F" bold="true" box="[199,353,1122,1148]" pageId="49" pageNumber="50">Distribution.</emphasis>
<taxonomicName id="C0A2F48B131BBF0AFED7FBE5FE78464E" authorityName="Vences &amp; Multzsch &amp; Gippner &amp; Miralles &amp; Crottini &amp; Gehring &amp; Rakotoarison &amp; Ratsoavina &amp; Glaw &amp; Scherz" authorityYear="2022" box="[360,491,1123,1149]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="50" phylum="Chordata" rank="species" species="hodikazo">
<emphasis id="35D6531A131BBF0AFED7FBE5FE78464E" box="[360,491,1123,1149]" italics="true" pageId="49" pageNumber="50">L. hodikazo</emphasis>
</taxonomicName>
is only known from its
<typeStatus id="D81931AA131BBF0AFD47FBE5FCBB464E" box="[760,808,1123,1149]" pageId="49" pageNumber="50">type</typeStatus>
locality, Tsingy de Bemaraha National Park in the West region of
<collectingCountry id="7FB5CF98131BBF0AFEBBFB01FE1E4692" box="[260,397,1159,1185]" name="Madagascar" pageId="49" pageNumber="50">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="4FB8DC83131BBF0AFF78FB2DFCDC473E" pageId="49" pageNumber="50" type="biology_ecology">
<paragraph id="071D8F08131BBF0AFF78FB2DFCDC473E" blockId="49.[151,1437,151,1293]" pageId="49" pageNumber="50">
<emphasis id="35D6531A131BBF0AFF78FB2DFE1546F7" bold="true" box="[199,390,1194,1221]" pageId="49" pageNumber="50">Natural history.</emphasis>
Practically nothing is known of the natural history of this enigmatic species. Its presence in the dry forests of the Tsingy de Bemaraha National Park indicates a tolerance for more arid environments that may set it apart from other
<taxonomicName id="C0A2F48B131BBF0AFE2EFB75FDB7473E" authorityName="Pasteur" authorityYear="1964" box="[401,548,1267,1293]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="49" pageNumber="55" phylum="Chordata" rank="subGenus" subGenus="Domerguella">
<emphasis id="35D6531A131BBF0AFE2EFB75FDB7473E" box="[401,548,1267,1293]" italics="true" pageId="49" pageNumber="50">Domerguella</emphasis>
</taxonomicName>
species (
<bibRefCitation id="6333F2F9131BBF0AFD35FB75FCD7473E" author="Bora, P. &amp; Randrianantoandro, J. C. &amp; Randrianavelona, R. &amp; Hantalalaina, E. F. &amp; Andriantsimanarilafy, R. R. &amp; Rakotondravony, D. &amp; Ramilijaona, O. R. &amp; Vences, M. &amp; Jenkins, R. K. B. &amp; Glaw, F. &amp; Kohler, J." box="[650,836,1267,1293]" pageId="49" pageNumber="50" pagination="111 - 125" refId="ref36289" refString="Bora, P., Randrianantoandro, J. C., Randrianavelona, R., Hantalalaina, E. F., Andriantsimanarilafy, R. R., Rakotondravony, D., Ramilijaona, O. R., Vences, M., Jenkins, R. K. B., Glaw, F. &amp; Kohler, J. (2010) Amphibians and reptiles of the Tsingy de Bemaraha Plateau, Western Madagascar: Checklist, biogeography and conservation. Herpetological Conservation and Biology, 5, 111 - 125." type="journal article" year="2010">Bora et al. 2010</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>