treatments-xml/data/81/65/9C/81659C1DFFDFFF84D6B6F92DFCA8D478.xml
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<document id="48A685835F387B34F2F532C87F06AF42" ID-ISSN="1175-5326" ID-ZooBank="1C88D39B-92D0-4045-8E96-59A820FD14B5" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629811198716" checkinUser="felipe" docAuthor="Rafael, J. A. &amp; Câmara, J. T." docDate="2012" docId="81659C1DFFDFFF84D6B6F92DFCA8D478" docLanguage="en" docName="zt03488p079.pdf" docOrigin="Zootaxa 3488" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Empis spinifera Bezzi 1909" docType="treatment" docVersion="2" lastPageNumber="79" masterDocId="7D5CE465FFD2FF94D621FF8BFFA7D644" masterDocTitle="Revision of Neotropical species of Empididae (Diptera) described by Mario Bezzi. XI. The species described in Empis Linnaeus" masterLastPageNumber="79" masterPageNumber="63" pageNumber="76" updateTime="1699115580821" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="09EE4A73E19857AE70D137B891BBC967">Revision of Neotropical species of Empididae (Diptera) described by Mario Bezzi. XI. The species described in Empis Linnaeus</mods:title>
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<mods:namePart id="CE15665BC65BEECA30A5CA7D8B523317">Rafael, J. A.</mods:namePart>
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<subSubSection id="41D67E80FFDFFF99D6B6F92DFE07D084" box="[151,416,1702,1728]" pageId="13" pageNumber="76" type="nomenclature">
<paragraph id="09732D0BFFDFFF99D6B6F92DFE07D084" blockId="13.[151,416,1702,1761]" box="[151,416,1702,1728]" pageId="13" pageNumber="76">
<heading id="523B9A67FFDFFF99D6B6F92DFE07D084" bold="true" box="[151,416,1702,1728]" fontSize="11" level="1" pageId="13" pageNumber="76" reason="1">
<taxonomicName id="CECC5688FFDFFF99D6B6F92DFE07D084" ID-CoL="6F6YJ" authority="Bezzi" authorityName="Bezzi" authorityYear="1909" box="[151,416,1702,1728]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="76" phylum="Arthropoda" rank="species" species="spinifera">
<emphasis id="3BB8F119FFDFFF99D6B6F92DFE07D084" bold="true" box="[151,416,1702,1728]" pageId="13" pageNumber="76">
<emphasis id="3BB8F119FFDFFF99D6B6F92DFEF1D084" bold="true" box="[151,342,1702,1728]" italics="true" pageId="13" pageNumber="76">Empis spinifera</emphasis>
Bezzi
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="41D67E80FFDFFF99D6B6F942FEF0D0A5" box="[151,343,1737,1761]" pageId="13" pageNumber="76" type="description">
<paragraph id="09732D0BFFDFFF99D6B6F942FEF0D0A5" blockId="13.[151,416,1702,1761]" box="[151,343,1737,1761]" pageId="13" pageNumber="76">
(
<figureCitation id="91F7318EFFDFFF99D6BEF942FE80D0A5" box="[159,295,1737,1761]" captionStart="FIGURES 7077" captionStartId="14.[151,263,1276,1299]" captionTargetBox="[267,1300,216,1255]" captionTargetId="figure-345@14.[266,1320,193,1255]" captionTargetPageId="14" captionText="FIGURES 7077. Empis spinifera Bezzi, lectotype ♂. 70, head, lateral view; 71; thorax, dorsal view; 72, left foreleg, posterior view; 73, right midleg, anterior view; 74, right hindleg, anterior view; 75, terminalia, lateral view; 76, epandrium, dorsal view; 77, schematic of ejaculatory apodeme, posterior view. Abbreviations: EJAP = ejaculatory apodeme; CE = cercus; EP = epandrium; Ph = phallus; Hyp = hypandrium; St = sternite; Scales: Figures 70, 7274 = 1.0 mm; 71 = 0.5 mm; 7576 = 0.2 mm." pageId="13" pageNumber="76">Figs. 7077</figureCitation>
,
<figureCitation id="91F7318EFFDFFF99D713F942FEE8D0A5" box="[306,335,1737,1761]" captionStart="FIGURES 7885" captionStartId="15.[151,263,962,985]" captionTargetBox="[306,1281,194,941]" captionTargetId="figure-441@15.[306,1281,193,941]" captionTargetPageId="15" captionText="FIGURES 7885. Wings. 78, Empis angustipennis Bezzi, lectotype ♀; 79, E. chiragra Bezzi, lectotype ♂; 80, E. induta Bezzi, female holotype; 81, E. leucostigma Bezzi, holotype ♀. 82, E. liodes Bezzi, holotype ♀. 83, E. lucidilabris Bezzi, ♀ from Peru; 84, E. penniventris Bezzi, holotype ♀. 85, E. spinifera Bezzi, lectotype ♂. Length of each wing informed along the text." pageId="13" pageNumber="76">85</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="41D67E80FFDFFF99D6B6F899FD81D102" pageId="13" pageNumber="76" type="reference_group">
<paragraph id="09732D0BFFDFFF99D6B6F899FD81D102" blockId="13.[151,1436,1809,1862]" pageId="13" pageNumber="76">
<treatmentCitationGroup id="29DC0A25FFDFFF99D6B6F899FD81D102" pageId="13" pageNumber="76">
<treatmentCitation id="886D0B1AFFDFFF99D6B6F899FE57D163" author="Bezzi, M." box="[151,496,1809,1831]" page="354" pageId="13" pageNumber="76" year="1909">
<taxonomicName id="CECC5688FFDFFF99D6B6F899FE57D163" ID-CoL="6F6YJ" authority="Bezzi, 1909: 354" authorityName="Bezzi" authorityPageNumber="354" authorityYear="1909" box="[151,496,1809,1831]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="76" phylum="Arthropoda" rank="species" species="spinifera">
<emphasis id="3BB8F119FFDFFF99D6B6F899FE90D163" box="[151,311,1809,1831]" italics="true" pageId="13" pageNumber="76">Empis spinifera</emphasis>
<bibRefCitation id="6D5D50FAFFDFFF99D71EF89AFE57D163" author="Bezzi, M." box="[319,496,1809,1831]" pageId="13" pageNumber="76" pagination="298 - 407" refId="ref10982" refString="Bezzi, M. (1909) Beitrage zur Kenntniss der sudamerikanischen Dipterenfauna auf Grund der Sammelergebnisse einer Reise in Chile, Peru und Bolivia, ausgefuhrt in den Jahren 1902 - 1904 von W. Schnuse. Fam. Empididae. Nova Acta Academiae Caesareae Leopoldino-Carolinae Germanicae Naturae Curiosorum, 91, 298 - 407." type="journal article" year="1909">Bezzi, 1909: 354</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="886D0B1AFFDFFF99D7DAF89AFD77D163" author="Melander, A. L." box="[507,720,1809,1831]" page="173" pageId="13" pageNumber="76" year="1928">
<bibRefCitation id="6D5D50FAFFDFFF99D7DAF89AFD77D163" author="Melander, A. L." box="[507,720,1809,1831]" pageId="13" pageNumber="76" refId="ref11303" refString="Melander, A. L. (1928) Diptera, Fam. Empididae. In: Wytsman, O. (Ed.), Genera Insectorum Fasc. 185. Louis Desmet-Verteneuil, Bruxelles, 434 pp." type="book" year="1928">Melander, 1928: 173</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="886D0B1AFFDFFF99D570F89AFBA2D163" author="Collin, J. E." box="[849,1029,1809,1831]" page="231" pageId="13" pageNumber="76" year="1933">
<bibRefCitation id="6D5D50FAFFDFFF99D570F89AFBA2D163" author="Collin, J. E." box="[849,1029,1809,1831]" pageId="13" pageNumber="76" refId="ref11039" refString="Collin, J. E. (1933) Empididae. Diptera of Patagonia and South Chile. Volume 4, London, British Museum of Natural History, London. Vol. 4, 334 pp., 74 figs." type="book" year="1933">Collin, 1933: 231</bibRefCitation>
</treatmentCitation>
(citation);
<treatmentCitation id="886D0B1AFFDFFF99D256F89AFABCD163" author="Smith, K. G. V." box="[1143,1307,1809,1831]" page="24" pageId="13" pageNumber="76" year="1967">
<bibRefCitation id="6D5D50FAFFDFFF99D256F89AFABCD163" author="Smith, K. G. V." box="[1143,1307,1809,1831]" pageId="13" pageNumber="76" refId="ref11556" refString="Smith, K. G. V. (1967) Family Empididae. In: Museu de Zoologia da Universidade de Sao Paulo (Ed.), A Catalogue of Diptera of Americas South of the United States. Sao Paulo, Vol. 39, 67 pp." type="book" year="1967">Smith, 1967: 24</bibRefCitation>
</treatmentCitation>
(catalogue);
<treatmentCitation id="886D0B1AFFDFFF99D6E6F8BBFE00D102" author="Yang, D. &amp; Zhang K. &amp; Yao G. &amp; Zhang J." box="[199,423,1840,1862]" page="140" pageId="13" pageNumber="76" year="2007">
<bibRefCitation id="6D5D50FAFFDFFF99D6E6F8BBFE00D102" author="Yang, D. &amp; Zhang K. &amp; Yao G. &amp; Zhang J." box="[199,423,1840,1862]" pageId="13" pageNumber="76" refId="ref11607" refString="Yang, D., Zhang K., Yao G. &amp; Zhang J. (2007). World Catalog of Empididae (Insecta: Diptera). Beijing, China Agricultural University Press, vi + 599 pp." type="book" year="2007">
Yang
<emphasis id="3BB8F119FFDFFF99D720F8BAFE92D102" box="[257,309,1840,1862]" italics="true" pageId="13" pageNumber="76">et al.</emphasis>
, 2007: 140
</bibRefCitation>
</treatmentCitation>
(catalogue).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="41D67E80FFDFFF9BD6B6F8F9FDE1D169" lastPageId="15" lastPageNumber="78" pageId="13" pageNumber="76" type="diagnosis">
<paragraph id="09732D0BFFDFFF99D6B6F8F9FA8DD196" blockId="13.[151,1436,1906,2002]" pageId="13" pageNumber="76">
<emphasis id="3BB8F119FFDFFF99D6B6F8F9FEB6D1CE" box="[151,273,1906,1931]" inLineHeading="true" pageId="13" pageNumber="76" reason="1">
<emphasis id="3BB8F119FFDFFF99D6B6F8F9FEADD1CF" bold="true" box="[151,266,1906,1931]" pageId="13" pageNumber="76">Diagnosis</emphasis>
.
</emphasis>
Brown, shiny; legs with long setae, most notably on hind pair; scape, pedicel, halter and legs yellow, except coxae and trochanters brown and hind pair mostly brown; propleura with proepisternal and proepimeral setae; wing slightly yellow, pterostigma brown; epandrium with a thin acuminate dorsoapical projection.
</paragraph>
<caption id="5DB37D83FFDCFF9AD6B6FB77FF62D3C8" pageId="14" pageNumber="77" startId="14.[151,263,1276,1299]" targetBox="[267,1300,216,1255]" targetPageId="14">
<paragraph id="09732D0BFFDCFF9AD6B6FB77FF62D3C8" blockId="14.[151,1436,1276,1420]" pageId="14" pageNumber="77">
<emphasis id="3BB8F119FFDCFF9AD6B6FB77FEF4D357" bold="true" box="[151,339,1276,1299]" pageId="14" pageNumber="77">FIGURES 7077.</emphasis>
<taxonomicName id="CECC5688FFDCFF9AD778FB76FD9DD356" authority="Bezzi" authorityName="Bezzi" authorityYear="1909" box="[345,570,1276,1298]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="spinifera">
<emphasis id="3BB8F119FFDCFF9AD778FB76FE5FD356" box="[345,504,1276,1298]" italics="true" pageId="14" pageNumber="77">Empis spinifera</emphasis>
Bezzi
</taxonomicName>
, lectotype ♂.
<emphasis id="3BB8F119FFDCFF9AD4E7FB77FD78D357" bold="true" box="[710,735,1276,1299]" pageId="14" pageNumber="77">70</emphasis>
, head, lateral view;
<emphasis id="3BB8F119FFDCFF9AD58AFB77FC63D357" bold="true" box="[939,964,1276,1299]" pageId="14" pageNumber="77">71</emphasis>
; thorax, dorsal view;
<emphasis id="3BB8F119FFDCFF9AD280FB77FB1DD357" bold="true" box="[1185,1210,1276,1299]" pageId="14" pageNumber="77">72</emphasis>
, left foreleg, posterior view;
<emphasis id="3BB8F119FFDCFF9AD6F4FA91FF49D375" bold="true" box="[213,238,1306,1329]" pageId="14" pageNumber="77">73</emphasis>
, right midleg, anterior view;
<emphasis id="3BB8F119FFDCFF9AD438FA91FD95D375" bold="true" box="[537,562,1306,1329]" pageId="14" pageNumber="77">74</emphasis>
, right hindleg, anterior view;
<emphasis id="3BB8F119FFDCFF9AD543FA90FCDCD374" bold="true" box="[866,891,1307,1328]" pageId="14" pageNumber="77">75</emphasis>
, terminalia, lateral view;
<emphasis id="3BB8F119FFDCFF9AD2A1FA91FB3ED375" bold="true" box="[1152,1177,1306,1329]" pageId="14" pageNumber="77">76</emphasis>
, epandrium, dorsal view;
<emphasis id="3BB8F119FFDCFF9AD6B6FAB1FF17D30B" bold="true" box="[151,176,1338,1359]" pageId="14" pageNumber="77">77</emphasis>
, schematic of ejaculatory apodeme, posterior view. Abbreviations:
<emphasis id="3BB8F119FFDCFF9AD55FFAB2FC1AD314" bold="true" box="[894,957,1337,1360]" pageId="14" pageNumber="77">EJAP</emphasis>
= ejaculatory apodeme;
<emphasis id="3BB8F119FFDCFF9AD2EBFAB2FB4AD314" bold="true" box="[1226,1261,1337,1360]" pageId="14" pageNumber="77">CE</emphasis>
= cercus;
<emphasis id="3BB8F119FFDCFF9AD342FAB1FA24D30B" bold="true" box="[1379,1411,1338,1359]" pageId="14" pageNumber="77">EP</emphasis>
= epandrium;
<emphasis id="3BB8F119FFDCFF9AD733FAD3FE88D329" bold="true" box="[274,303,1368,1389]" pageId="14" pageNumber="77">Ph</emphasis>
= phallus;
<emphasis id="3BB8F119FFDCFF9AD784FAD3FE74D329" bold="true" box="[421,467,1368,1389]" pageId="14" pageNumber="77">Hyp</emphasis>
= hypandrium;
<emphasis id="3BB8F119FFDCFF9AD45BFAD3FD36D32B" bold="true" box="[634,657,1368,1391]" pageId="14" pageNumber="77">St</emphasis>
= sternite; Scales: Figures
<emphasis id="3BB8F119FFDCFF9AD588FAD3FBA8D32B" bold="true" box="[937,1039,1368,1391]" pageId="14" pageNumber="77">70, 7274</emphasis>
= 1.0 mm;
<emphasis id="3BB8F119FFDCFF9AD2AAFAD3FB03D32B" bold="true" box="[1163,1188,1368,1391]" pageId="14" pageNumber="77">71</emphasis>
= 0.5 mm;
<emphasis id="3BB8F119FFDCFF9AD33EFAD3FAF9D32B" bold="true" box="[1311,1374,1368,1391]" pageId="14" pageNumber="77">7576</emphasis>
= 0.2 mm.
</paragraph>
</caption>
<paragraph id="09732D0BFFDCFF9AD6E6FA32FE50D0B5" blockId="14.[151,1436,1465,2030]" pageId="14" pageNumber="77">
<emphasis id="3BB8F119FFDCFF9AD6E6FA32FED0D396" bold="true" box="[199,375,1465,1490]" pageId="14" pageNumber="77">Re-description</emphasis>
.
<typeStatus id="D67793A9FFDCFF9AD7ABFA31FE5ED396" box="[394,505,1466,1490]" pageId="14" pageNumber="77" type="lectotype">Lectotype</typeStatus>
<emphasis id="3BB8F119FFDCFF9AD424FA31FD99D396" bold="true" box="[517,574,1466,1490]" pageId="14" pageNumber="77">male</emphasis>
. Body length
<quantity id="CE3480EEFFDCFF9AD4CDFA32FCEDD395" box="[748,842,1465,1489]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="77" unit="mm" value="4.5">4.5 mm</quantity>
; wing length
<quantity id="CE3480EEFFDCFF9AD5D1FA32FBE9D395" box="[1008,1102,1465,1489]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="14" pageNumber="77" unit="mm" value="4.5">4.5 mm</quantity>
. Holoptic; dorsal ommatidia larger. Ocellar tubercle and frontal triangle brown with faint grey pruinescence. Ocellus glassy, yellow; with anterior ocellar setae apparently convergent and 2 small posterior setae. Face bare, half as high as frons, subquadrate, brown, shiny, protuberant at apex (
<figureCitation id="91F7318EFFDCFF9AD49DF9AEFCB6D079" box="[700,785,1573,1597]" captionStart="FIGURES 7077" captionStartId="14.[151,263,1276,1299]" captionTargetBox="[267,1300,216,1255]" captionTargetId="figure-345@14.[266,1320,193,1255]" captionTargetPageId="14" captionText="FIGURES 7077. Empis spinifera Bezzi, lectotype ♂. 70, head, lateral view; 71; thorax, dorsal view; 72, left foreleg, posterior view; 73, right midleg, anterior view; 74, right hindleg, anterior view; 75, terminalia, lateral view; 76, epandrium, dorsal view; 77, schematic of ejaculatory apodeme, posterior view. Abbreviations: EJAP = ejaculatory apodeme; CE = cercus; EP = epandrium; Ph = phallus; Hyp = hypandrium; St = sternite; Scales: Figures 70, 7274 = 1.0 mm; 71 = 0.5 mm; 7576 = 0.2 mm." pageId="14" pageNumber="77">Fig. 70</figureCitation>
), with narrow grey inverted U-shaped pruinescence spot. Postcranium (
<figureCitation id="91F7318EFFDCFF9AD712F9C2FE23D025" box="[307,388,1609,1633]" captionStart="FIGURES 7077" captionStartId="14.[151,263,1276,1299]" captionTargetBox="[267,1300,216,1255]" captionTargetId="figure-345@14.[266,1320,193,1255]" captionTargetPageId="14" captionText="FIGURES 7077. Empis spinifera Bezzi, lectotype ♂. 70, head, lateral view; 71; thorax, dorsal view; 72, left foreleg, posterior view; 73, right midleg, anterior view; 74, right hindleg, anterior view; 75, terminalia, lateral view; 76, epandrium, dorsal view; 77, schematic of ejaculatory apodeme, posterior view. Abbreviations: EJAP = ejaculatory apodeme; CE = cercus; EP = epandrium; Ph = phallus; Hyp = hypandrium; St = sternite; Scales: Figures 70, 7274 = 1.0 mm; 71 = 0.5 mm; 7576 = 0.2 mm." pageId="14" pageNumber="77">Fig. 70</figureCitation>
) brown with fine grey pruinescence, uniserial postocular setae, uniserial long occipital setae, 3 setae between both rows placed below middle. Gena and postgena with various long, slender setae. Antenna with scape and pedicel yellow; postpedicel coniform, velvety matte-black, twice as long as scape and pedicel together, twice as long as stylus. Proboscis roughly 2.5X head height; labrum yellow, shiny; labium brown. Palpus curved dorsally, yellow to light brown.
</paragraph>
<paragraph id="09732D0BFFDCFF9AD6E6F976FC8BD1AA" blockId="14.[151,1436,1465,2030]" pageId="14" pageNumber="77">
Thorax (
<figureCitation id="91F7318EFFDCFF9AD709F976FEDDD151" box="[296,378,1789,1813]" captionStart="FIGURES 7077" captionStartId="14.[151,263,1276,1299]" captionTargetBox="[267,1300,216,1255]" captionTargetId="figure-345@14.[266,1320,193,1255]" captionTargetPageId="14" captionText="FIGURES 7077. Empis spinifera Bezzi, lectotype ♂. 70, head, lateral view; 71; thorax, dorsal view; 72, left foreleg, posterior view; 73, right midleg, anterior view; 74, right hindleg, anterior view; 75, terminalia, lateral view; 76, epandrium, dorsal view; 77, schematic of ejaculatory apodeme, posterior view. Abbreviations: EJAP = ejaculatory apodeme; CE = cercus; EP = epandrium; Ph = phallus; Hyp = hypandrium; St = sternite; Scales: Figures 70, 7274 = 1.0 mm; 71 = 0.5 mm; 7576 = 0.2 mm." pageId="14" pageNumber="77">Fig. 71</figureCitation>
) brown, shiny, finely grey pruinescent. Chaetotaxy: 67 uniserial antepronotals; 1 robust and 810 weak postpronotals; 23 proepisternals; 4 proepimerals, one of them more robust; acrostichal setae difficult to observe due to invagination of median portion of scutum caused by pin, but present, uniserial; dorsocentral also shifted by scutum deformation, long, biserial; 1 presutural intra-alar; 2 postsutural intra-alars, one of these directly posterior transverse suture and other near scutellum; 1 presutural supra-alar; 2 postsutural supra-alars, 1 small anterior and 1 longer posterior; 3 robust posterior and 56 weak anterior notopleurals; 1 postalar; 2 scutellars, apical pair more robust, crossed; roughly 20 laterotergitals.
</paragraph>
<caption id="5DB37D83FFDDFF9BD6B6FC49FAFFD251" pageId="15" pageNumber="78" startId="15.[151,263,962,985]" targetBox="[306,1281,194,941]" targetPageId="15">
<paragraph id="09732D0BFFDDFF9BD6B6FC49FAFFD251" blockId="15.[151,1436,962,1046]" pageId="15" pageNumber="78">
<emphasis id="3BB8F119FFDDFF9BD6B6FC49FEF4D59D" bold="true" box="[151,339,962,985]" pageId="15" pageNumber="78">FIGURES 7885.</emphasis>
Wings.
<emphasis id="3BB8F119FFDDFF9BD787FC49FE18D59D" bold="true" box="[422,447,962,985]" pageId="15" pageNumber="78">78</emphasis>
,
<taxonomicName id="CECC5688FFDDFF9BD7EDFC48FD78D59C" authority="Bezzi" authorityName="Bezzi" authorityYear="1909" box="[460,735,962,984]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="78" phylum="Arthropoda" rank="species" species="angustipennis">
<emphasis id="3BB8F119FFDDFF9BD7EDFC48FD39D59C" box="[460,670,963,984]" italics="true" pageId="15" pageNumber="78">Empis angustipennis</emphasis>
Bezzi
</taxonomicName>
, lectotype ♀;
<emphasis id="3BB8F119FFDDFF9BD54DFC49FC22D59D" bold="true" box="[876,901,962,985]" pageId="15" pageNumber="78">79</emphasis>
,
<taxonomicName id="CECC5688FFDDFF9BD5B0FC4FFBE0D59C" authority="Bezzi" authorityName="Bezzi" authorityYear="1909" box="[913,1095,962,985]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="78" phylum="Arthropoda" rank="species" species="chiragra">
<emphasis id="3BB8F119FFDDFF9BD5B0FC4FFBA1D59D" box="[913,1030,963,985]" italics="true" pageId="15" pageNumber="78">E. chiragra</emphasis>
Bezzi
</taxonomicName>
, lectotype ♂;
<emphasis id="3BB8F119FFDDFF9BD2F5FC49FB4AD59D" bold="true" box="[1236,1261,962,985]" pageId="15" pageNumber="78">80</emphasis>
,
<taxonomicName id="CECC5688FFDDFF9BD2DBFC4FFA3FD59C" authority="Bezzi" authorityName="Bezzi" authorityYear="1909" box="[1274,1432,962,985]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="78" phylum="Arthropoda" rank="species" species="induta">
<emphasis id="3BB8F119FFDDFF9BD2DBFC4FFAF2D59D" box="[1274,1365,963,985]" italics="true" pageId="15" pageNumber="78">E. induta</emphasis>
Bezzi
</taxonomicName>
, female holotype;
<emphasis id="3BB8F119FFDDFF9BD766FC6AFEC7D5BC" bold="true" box="[327,352,993,1016]" pageId="15" pageNumber="78">81</emphasis>
,
<taxonomicName id="CECC5688FFDDFF9BD74CFC69FDE2D5B3" authority="Bezzi" authorityName="Bezzi" authorityYear="1909" box="[365,581,993,1015]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="78" phylum="Arthropoda" rank="species" species="leucostigma">
<emphasis id="3BB8F119FFDDFF9BD74CFC69FDA4D5B3" box="[365,515,993,1015]" italics="true" pageId="15" pageNumber="78">E. leucostigma</emphasis>
Bezzi
</taxonomicName>
, holotype ♀.
<emphasis id="3BB8F119FFDDFF9BD4EDFC6AFD42D5BC" bold="true" box="[716,741,993,1016]" pageId="15" pageNumber="78">82</emphasis>
,
<taxonomicName id="CECC5688FFDDFF9BD4D3FC69FC2AD5B3" authority="Bezzi" authorityName="Bezzi" authorityYear="1909" box="[754,909,993,1015]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="78" phylum="Arthropoda" rank="species" species="liodes">
<emphasis id="3BB8F119FFDDFF9BD4D3FC69FCEDD5B3" box="[754,842,993,1015]" italics="true" pageId="15" pageNumber="78">E. liodes</emphasis>
Bezzi
</taxonomicName>
, holotype ♀.
<emphasis id="3BB8F119FFDDFF9BD235FC6AFB8AD5BC" bold="true" box="[1044,1069,993,1016]" pageId="15" pageNumber="78">83</emphasis>
,
<taxonomicName id="CECC5688FFDDFF9BD21BFC69FAABD5B3" authority="Bezzi" authorityName="Bezzi" authorityYear="1905" box="[1082,1292,993,1015]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="78" phylum="Arthropoda" rank="species" species="lucidilabris">
<emphasis id="3BB8F119FFDDFF9BD21BFC69FB6CD5B3" box="[1082,1227,993,1015]" italics="true" pageId="15" pageNumber="78">E. lucidilabris</emphasis>
Bezzi
</taxonomicName>
, ♀ from Peru;
<emphasis id="3BB8F119FFDDFF9BD6B6FC74FF17D252" bold="true" box="[151,176,1023,1046]" pageId="15" pageNumber="78">84</emphasis>
,
<taxonomicName id="CECC5688FFDDFF9BD69CFB8AFE3FD252" authority="Bezzi" authorityName="Bezzi" authorityYear="1909" box="[189,408,1024,1046]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="78" phylum="Arthropoda" rank="species" species="penniventris">
<emphasis id="3BB8F119FFDDFF9BD69CFB8AFEF1D252" box="[189,342,1025,1046]" italics="true" pageId="15" pageNumber="78">E. penniventris</emphasis>
Bezzi
</taxonomicName>
, holotype ♀.
<emphasis id="3BB8F119FFDDFF9BD400FC74FD9DD252" bold="true" box="[545,570,1023,1046]" pageId="15" pageNumber="78">85</emphasis>
,
<taxonomicName id="CECC5688FFDDFF9BD466FB8AFCA7D252" authority="Bezzi" authorityName="Bezzi" authorityYear="1909" box="[583,768,1024,1046]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="78" phylum="Arthropoda" rank="species" species="spinifera">
<emphasis id="3BB8F119FFDDFF9BD466FB8AFD1AD252" box="[583,701,1024,1046]" italics="true" pageId="15" pageNumber="78">E. spinifera</emphasis>
Bezzi
</taxonomicName>
, lectotype ♂. Length of each wing informed along the text.
</paragraph>
</caption>
<paragraph id="09732D0BFFDDFF9BD6E6FBC8FAF0D386" blockId="15.[151,1436,1090,2017]" pageId="15" pageNumber="78">
Legs (
<figureCitation id="91F7318EFFDDFF9BD72EFBC9FE3FD21E" box="[271,408,1090,1114]" captionStart="FIGURES 7077" captionStartId="14.[151,263,1276,1299]" captionTargetBox="[267,1300,216,1255]" captionTargetId="figure-345@14.[266,1320,193,1255]" captionTargetPageId="14" captionText="FIGURES 7077. Empis spinifera Bezzi, lectotype ♂. 70, head, lateral view; 71; thorax, dorsal view; 72, left foreleg, posterior view; 73, right midleg, anterior view; 74, right hindleg, anterior view; 75, terminalia, lateral view; 76, epandrium, dorsal view; 77, schematic of ejaculatory apodeme, posterior view. Abbreviations: EJAP = ejaculatory apodeme; CE = cercus; EP = epandrium; Ph = phallus; Hyp = hypandrium; St = sternite; Scales: Figures 70, 7274 = 1.0 mm; 71 = 0.5 mm; 7576 = 0.2 mm." pageId="15" pageNumber="78">Figs. 7274</figureCitation>
) with long setae, especially hind pair which bears largest and thickest setae; legs mainly yellow, except all coxae and trochanters brown, hind tibia brown with yellow base. Foreleg (
<figureCitation id="91F7318EFFDDFF9BD2E3FBECFABED23B" box="[1218,1305,1127,1151]" captionStart="FIGURES 7077" captionStartId="14.[151,263,1276,1299]" captionTargetBox="[267,1300,216,1255]" captionTargetId="figure-345@14.[266,1320,193,1255]" captionTargetPageId="14" captionText="FIGURES 7077. Empis spinifera Bezzi, lectotype ♂. 70, head, lateral view; 71; thorax, dorsal view; 72, left foreleg, posterior view; 73, right midleg, anterior view; 74, right hindleg, anterior view; 75, terminalia, lateral view; 76, epandrium, dorsal view; 77, schematic of ejaculatory apodeme, posterior view. Abbreviations: EJAP = ejaculatory apodeme; CE = cercus; EP = epandrium; Ph = phallus; Hyp = hypandrium; St = sternite; Scales: Figures 70, 7274 = 1.0 mm; 71 = 0.5 mm; 7576 = 0.2 mm." pageId="15" pageNumber="78">Fig. 72</figureCitation>
): tibia with series of 5 short equidistant anterodorsal setae; tarsomere 1 with distal row of more robust setae and 1 subbasal anterodorsal; remaining tarsomeres with 1 short antero- and posteroventral apically. Midleg (
<figureCitation id="91F7318EFFDDFF9BD295FB24FAADD283" box="[1204,1290,1199,1223]" captionStart="FIGURES 7077" captionStartId="14.[151,263,1276,1299]" captionTargetBox="[267,1300,216,1255]" captionTargetId="figure-345@14.[266,1320,193,1255]" captionTargetPageId="14" captionText="FIGURES 7077. Empis spinifera Bezzi, lectotype ♂. 70, head, lateral view; 71; thorax, dorsal view; 72, left foreleg, posterior view; 73, right midleg, anterior view; 74, right hindleg, anterior view; 75, terminalia, lateral view; 76, epandrium, dorsal view; 77, schematic of ejaculatory apodeme, posterior view. Abbreviations: EJAP = ejaculatory apodeme; CE = cercus; EP = epandrium; Ph = phallus; Hyp = hypandrium; St = sternite; Scales: Figures 70, 7274 = 1.0 mm; 71 = 0.5 mm; 7576 = 0.2 mm." pageId="15" pageNumber="78">Fig. 73</figureCitation>
): femur with roughly 12 posteroventral setae; tibia with 3 dorsal on distal two-thirds, 1 posterodorsal medially, 4 ventral on distal two-thirds, 1 antero- and posteroventral and 1 posterodorsal distally; tarsomere 1 with complete distal row of more robust setae in addition to 2 anterodorsals on basal half, 2 ventral medially, 1 subbasal posteroventral; remaining tarsomeres same as fore pair. Hindleg (
<figureCitation id="91F7318EFFDDFF9BD4F3FAB4FC8ED313" box="[722,809,1343,1367]" captionStart="FIGURES 7077" captionStartId="14.[151,263,1276,1299]" captionTargetBox="[267,1300,216,1255]" captionTargetId="figure-345@14.[266,1320,193,1255]" captionTargetPageId="14" captionText="FIGURES 7077. Empis spinifera Bezzi, lectotype ♂. 70, head, lateral view; 71; thorax, dorsal view; 72, left foreleg, posterior view; 73, right midleg, anterior view; 74, right hindleg, anterior view; 75, terminalia, lateral view; 76, epandrium, dorsal view; 77, schematic of ejaculatory apodeme, posterior view. Abbreviations: EJAP = ejaculatory apodeme; CE = cercus; EP = epandrium; Ph = phallus; Hyp = hypandrium; St = sternite; Scales: Figures 70, 7274 = 1.0 mm; 71 = 0.5 mm; 7576 = 0.2 mm." pageId="15" pageNumber="78">Fig. 74</figureCitation>
): femur with anterior, antero- and posteroventral rows, 34 anterodorsals on distal third in addition to distal row of more robust setae; tibia with antero- and posterodorsal series of rather long setae, 4 short anteroventral medially, 1 short posteroventral apically; tarsomere 1 with antero- and posterodorsal rows longer than antero- and posteroventral rows; remaining tarsomeres same as fore pair.
</paragraph>
<paragraph id="09732D0BFFDDFF9BD6E6FA44FF4ED075" blockId="15.[151,1436,1090,2017]" pageId="15" pageNumber="78">
Wing (
<figureCitation id="91F7318EFFDDFF9BD735FA44FEC1D3A3" box="[276,358,1487,1511]" captionStart="FIGURES 7885" captionStartId="15.[151,263,962,985]" captionTargetBox="[306,1281,194,941]" captionTargetId="figure-441@15.[306,1281,193,941]" captionTargetPageId="15" captionText="FIGURES 7885. Wings. 78, Empis angustipennis Bezzi, lectotype ♀; 79, E. chiragra Bezzi, lectotype ♂; 80, E. induta Bezzi, female holotype; 81, E. leucostigma Bezzi, holotype ♀. 82, E. liodes Bezzi, holotype ♀. 83, E. lucidilabris Bezzi, ♀ from Peru; 84, E. penniventris Bezzi, holotype ♀. 85, E. spinifera Bezzi, lectotype ♂. Length of each wing informed along the text." pageId="15" pageNumber="78">Fig. 85</figureCitation>
) light yellow with faint yellow pterostigma, veins brown; base of costal vein with 1 robust seta (not represented in figure). Vein R
<subScript id="95482F4EFFDDFF9BD406F989FD88D054" attach="left" box="[551,559,1538,1552]" fontSize="6" pageId="15" pageNumber="78">1</subScript>
bare dorsally, not dilated towards apex; vein A
<subScript id="95482F4EFFDDFF9BD27BF989FBC5D054" attach="left" box="[1114,1122,1538,1552]" fontSize="6" pageId="15" pageNumber="78">1</subScript>
evanescent at apex. Halter yellow.
</paragraph>
<paragraph id="09732D0BFFDDFF9BD6E6F9B6FB8BD03D" blockId="15.[151,1436,1090,2017]" pageId="15" pageNumber="78">Abdomen brown, shiny, with narrow grey band on posterior margin of tergites; lateral and posterior setae on tergites distinctly longer. Basal sternites yellow, distal ones same color as tergites.</paragraph>
<paragraph id="09732D0BFFDDFF9BD6E6F90EFAB7D0A1" blockId="15.[151,1436,1090,2017]" pageId="15" pageNumber="78">
Terminalia concolorous with tergites. Epandrium (
<figureCitation id="91F7318EFFDDFF9BD52BF90EFC30D0D9" box="[778,919,1669,1693]" captionStart="FIGURES 7077" captionStartId="14.[151,263,1276,1299]" captionTargetBox="[267,1300,216,1255]" captionTargetId="figure-345@14.[266,1320,193,1255]" captionTargetPageId="14" captionText="FIGURES 7077. Empis spinifera Bezzi, lectotype ♂. 70, head, lateral view; 71; thorax, dorsal view; 72, left foreleg, posterior view; 73, right midleg, anterior view; 74, right hindleg, anterior view; 75, terminalia, lateral view; 76, epandrium, dorsal view; 77, schematic of ejaculatory apodeme, posterior view. Abbreviations: EJAP = ejaculatory apodeme; CE = cercus; EP = epandrium; Ph = phallus; Hyp = hypandrium; St = sternite; Scales: Figures 70, 7274 = 1.0 mm; 71 = 0.5 mm; 7576 = 0.2 mm." pageId="15" pageNumber="78">Figs. 75, 76</figureCitation>
) with thin acuminate dorsoapical projection; hypandrium (
<figureCitation id="91F7318EFFDDFF9BD715F922FE2AD085" box="[308,397,1705,1729]" captionStart="FIGURES 7077" captionStartId="14.[151,263,1276,1299]" captionTargetBox="[267,1300,216,1255]" captionTargetId="figure-345@14.[266,1320,193,1255]" captionTargetPageId="14" captionText="FIGURES 7077. Empis spinifera Bezzi, lectotype ♂. 70, head, lateral view; 71; thorax, dorsal view; 72, left foreleg, posterior view; 73, right midleg, anterior view; 74, right hindleg, anterior view; 75, terminalia, lateral view; 76, epandrium, dorsal view; 77, schematic of ejaculatory apodeme, posterior view. Abbreviations: EJAP = ejaculatory apodeme; CE = cercus; EP = epandrium; Ph = phallus; Hyp = hypandrium; St = sternite; Scales: Figures 70, 7274 = 1.0 mm; 71 = 0.5 mm; 7576 = 0.2 mm." pageId="15" pageNumber="78">Fig. 75</figureCitation>
) with series of long ventral setae and bifid, downcurved apex; cercus subtrapezoidal; ejaculatory apodeme (
<figureCitation id="91F7318EFFDDFF9BD7AEF946FE47D0A1" box="[399,480,1741,1765]" captionStart="FIGURES 7077" captionStartId="14.[151,263,1276,1299]" captionTargetBox="[267,1300,216,1255]" captionTargetId="figure-345@14.[266,1320,193,1255]" captionTargetPageId="14" captionText="FIGURES 7077. Empis spinifera Bezzi, lectotype ♂. 70, head, lateral view; 71; thorax, dorsal view; 72, left foreleg, posterior view; 73, right midleg, anterior view; 74, right hindleg, anterior view; 75, terminalia, lateral view; 76, epandrium, dorsal view; 77, schematic of ejaculatory apodeme, posterior view. Abbreviations: EJAP = ejaculatory apodeme; CE = cercus; EP = epandrium; Ph = phallus; Hyp = hypandrium; St = sternite; Scales: Figures 70, 7274 = 1.0 mm; 71 = 0.5 mm; 7576 = 0.2 mm." pageId="15" pageNumber="78">Fig. 77</figureCitation>
) flattened with lateral expansions, tetra-lamellar shape in posterior view.
</paragraph>
<paragraph id="09732D0BFFDDFF9BD6E6F979FE05D14D" blockId="15.[151,1436,1090,2017]" box="[199,418,1777,1802]" pageId="15" pageNumber="78">
<emphasis id="3BB8F119FFDDFF9BD6E6F979FE82D14D" box="[199,293,1778,1802]" inLineHeading="true" pageId="15" pageNumber="78" reason="1">
<emphasis id="3BB8F119FFDDFF9BD6E6F979FEB9D14E" bold="true" box="[199,286,1778,1802]" pageId="15" pageNumber="78">Female</emphasis>
.
</emphasis>
Unknown.
</paragraph>
<paragraph id="09732D0BFFDDFF9BD6E6F89EFDE1D169" blockId="15.[151,1436,1090,2017]" box="[199,582,1813,1838]" pageId="15" pageNumber="78">
<emphasis id="3BB8F119FFDDFF9BD6E6F89EFDA3D169" box="[199,516,1813,1838]" inLineHeading="true" pageId="15" pageNumber="78" reason="1">
<emphasis id="3BB8F119FFDDFF9BD6E6F89EFE5AD16A" bold="true" box="[199,509,1813,1838]" pageId="15" pageNumber="78">Geographical distribution</emphasis>
.
</emphasis>
<collectingCountry id="71DB6D9BFFDDFF9BD42AF89DFDE5D169" box="[523,578,1814,1837]" name="Peru" pageId="15" pageNumber="78">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="41D67E80FFDDFF9BD6E6F8B2FC4ED1DD" pageId="15" pageNumber="78" type="materials_examined">
<paragraph id="09732D0BFFDDFF9BD6E6F8B2FD89D131" blockId="15.[151,1436,1090,2017]" pageId="15" pageNumber="78">
<emphasis id="3BB8F119FFDDFF9BD6E6F8B2FE11D115" box="[199,438,1849,1874]" inLineHeading="true" pageId="15" pageNumber="78" reason="1">
<emphasis id="3BB8F119FFDDFF9BD6E6F8B2FE08D116" bold="true" box="[199,431,1849,1874]" pageId="15" pageNumber="78">Material examined</emphasis>
.
</emphasis>
<materialsCitation id="B9A42756FFDDFF9BD7E5F8B2FD89D131" collectingDate="1903-08-10" country="Peru" elevation="2500" location="Paso de Lares" pageId="15" pageNumber="78" specimenCount="1" specimenCount-male="1" stateProvince="Cusco" typeStatus="lectotype">
<emphasis id="3BB8F119FFDDFF9BD7E5F8B2FDD7D116" bold="true" box="[452,624,1849,1874]" pageId="15" pageNumber="78">
<typeStatus id="D67793A9FFDDFF9BD7E5F8B2FDD7D116" box="[452,624,1849,1874]" pageId="15" pageNumber="78" type="lectotype">LECTOTYPE</typeStatus>
</emphasis>
<specimenCount id="1FCAE682FFDDFF9BD45CF8B1FD35D115" box="[637,658,1850,1873]" pageId="15" pageNumber="78" type="male"></specimenCount>
(here designated). “
<collectingCountry id="71DB6D9BFFDDFF9BD5A7F8B2FC77D115" box="[902,976,1849,1873]" name="Peru" pageId="15" pageNumber="78">PERU</collectingCountry>
[
<collectingRegion id="CB08E3E9FFDDFF9BD5C5F8B2FB94D115" box="[996,1075,1849,1873]" country="Peru" name="Cusco" pageId="15" pageNumber="78">Cuzco</collectingRegion>
], Laristhal (=
<location id="0C137BD0FFDDFF9BD2C2F8B1FA37D115" LSID="urn:lsid:plazi:treatment:81659C1DFFDFFF84D6B6F92DFCA8D478:0C137BD0FFDDFF9BD2C2F8B1FA37D115" box="[1251,1424,1849,1873]" country="Peru" name="Paso de Lares" pageId="15" pageNumber="78" stateProvince="Cusco">Paso de Lares</location>
),
<date id="7D720BCBFFDDFF9BD6B6F8D6FE87D131" box="[151,288,1885,1909]" pageId="15" pageNumber="78" value="1903-08-10">
<collectingDate id="6D36F223FFDDFF9BD6B6F8D6FE87D131" box="[151,288,1885,1909]" pageId="15" pageNumber="78" value="1903-08-10">10.viii.1903</collectingDate>
</date>
,
<quantity id="CE3480EEFFDDFF9BD70DF8D6FE6ED131" box="[300,457,1885,1909]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="15" pageNumber="78" unit="m" value="2500.0" valueMax="3000.0" valueMin="2000.0">
<elevation id="82E1CA38FFDDFF9BD70DF8D6FE6ED131" box="[300,457,1885,1909]" metricMagnitude="3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="15" pageNumber="78" unit="m" value="2500.0" valueMax="3000.0" valueMin="2000.0">20003000 m</elevation>
</quantity>
” (SMT).
</materialsCitation>
</paragraph>
<paragraph id="09732D0BFFDDFF9BD6E6F809FC4ED1DD" blockId="15.[151,1436,1090,2017]" box="[199,1001,1921,1946]" pageId="15" pageNumber="78">
<emphasis id="3BB8F119FFDDFF9BD6E6F809FE1CD1DD" box="[199,443,1921,1946]" inLineHeading="true" pageId="15" pageNumber="78" reason="1">
<emphasis id="3BB8F119FFDDFF9BD6E6F809FE13D1DE" bold="true" box="[199,436,1921,1946]" pageId="15" pageNumber="78">
<typeStatus id="D67793A9FFDDFF9BD6E6F809FE9AD1DE" box="[199,317,1922,1946]" pageId="15" pageNumber="78" type="lectotype">Lectotype</typeStatus>
condition
</emphasis>
.
</emphasis>
Left wing on microslides. Terminalia in glycerin.
</paragraph>
</subSubSection>
<subSubSection id="41D67E80FFDDFF84D6E6F82DFCA8D478" lastPageId="16" lastPageNumber="79" pageId="15" pageNumber="78" type="discussion">
<paragraph id="09732D0BFFDDFF9BD6E6F82DFE08D1A5" blockId="15.[151,1436,1090,2017]" pageId="15" pageNumber="78">
<emphasis id="3BB8F119FFDDFF9BD6E6F82DFE9CD1F9" box="[199,315,1958,1982]" inLineHeading="true" pageId="15" pageNumber="78" reason="1">
<emphasis id="3BB8F119FFDDFF9BD6E6F82DFE93D1FA" bold="true" box="[199,308,1958,1982]" pageId="15" pageNumber="78">Remarks</emphasis>
.
</emphasis>
This species was described based on
<specimenCount id="1FCAE682FFDDFF9BD52EF82DFCDAD1F9" box="[783,893,1957,1981]" pageId="15" pageNumber="78" type="male">two male</specimenCount>
specimens from the same locality. Only
<specimenCount id="1FCAE682FFDDFF9BD355F82DFEA6D1A5" pageId="15" pageNumber="78" type="generic">one specimen</specimenCount>
was examined.
</paragraph>
<paragraph id="09732D0BFFC2FF84D6E6FF1CFCACD77B" blockId="16.[151,1436,151,319]" pageId="16" pageNumber="79">
<emphasis id="3BB8F119FFC2FF84D6E6FF1CFEEBD6EB" box="[199,332,151,176]" inLineHeading="true" pageId="16" pageNumber="79" reason="1">
<emphasis id="3BB8F119FFC2FF84D6E6FF1CFEE2D6F4" bold="true" box="[199,325,151,176]" pageId="16" pageNumber="79">Discussion</emphasis>
.
</emphasis>
The male specimen of
<taxonomicName id="CECC5688FFC2FF84D47CFF13FCB7D6F4" authorityName="Bezzi" authorityYear="1909" box="[605,784,152,176]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="79" phylum="Arthropoda" rank="species" species="spinifera">
<emphasis id="3BB8F119FFC2FF84D47CFF13FCB7D6F4" box="[605,784,152,176]" italics="true" pageId="16" pageNumber="79">Empis spinifera</emphasis>
</taxonomicName>
was identified as part of group II in
<bibRefCitation id="6D5D50FAFFC2FF84D29FFF1CFACDD6EB" author="Collin, J. E." box="[1214,1386,151,175]" pageId="16" pageNumber="79" refId="ref11039" refString="Collin, J. E. (1933) Empididae. Diptera of Patagonia and South Chile. Volume 4, London, British Museum of Natural History, London. Vol. 4, 334 pp., 74 figs." type="book" year="1933">Collins (1933)</bibRefCitation>
key through the same couplets of
<taxonomicName id="CECC5688FFC2FF84D7FEFF36FD3ED690" authorityName="Bezzi" authorityYear="1909" box="[479,665,189,212]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="79" phylum="Arthropoda" rank="species" species="angustipennis">
<emphasis id="3BB8F119FFC2FF84D7FEFF36FD3ED690" box="[479,665,189,212]" italics="true" pageId="16" pageNumber="79">E. angustipennis</emphasis>
</taxonomicName>
as presented above. Further it keys out to couplet (19) of
<taxonomicName id="CECC5688FFC2FF84D33CFF36FA3BD690" authorityName="Bigot" authorityYear="1889" box="[1309,1436,189,212]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="79" phylum="Arthropoda" rank="species" species="macrura">
<emphasis id="3BB8F119FFC2FF84D33CFF36FA3BD690" box="[1309,1436,189,212]" italics="true" pageId="16" pageNumber="79">E. macrura</emphasis>
</taxonomicName>
through: (16) “thorax blackish without distinct stripes”; (16b) “face bare, abdomen dark” and (19) “legs entirely tawny-yellow (yellow in examined specimen). They are not conspecific because the male terminalia are entirely different (compare
<bibRefCitation id="6D5D50FAFFC2FF84D74FFEACFE58D77B" author="Collin, J. E." box="[366,511,295,319]" pageId="16" pageNumber="79" refId="ref11039" refString="Collin, J. E. (1933) Empididae. Diptera of Patagonia and South Chile. Volume 4, London, British Museum of Natural History, London. Vol. 4, 334 pp., 74 figs." type="book" year="1933">Collin (1933</bibRefCitation>
, fig. 52b) with
<figureCitation id="91F7318EFFC2FF84D48AFEACFD5AD77B" box="[683,765,295,319]" captionStart="FIGURES 7077" captionStartId="14.[151,263,1276,1299]" captionTargetBox="[267,1300,216,1255]" captionTargetId="figure-345@14.[266,1320,193,1255]" captionTargetPageId="14" captionText="FIGURES 7077. Empis spinifera Bezzi, lectotype ♂. 70, head, lateral view; 71; thorax, dorsal view; 72, left foreleg, posterior view; 73, right midleg, anterior view; 74, right hindleg, anterior view; 75, terminalia, lateral view; 76, epandrium, dorsal view; 77, schematic of ejaculatory apodeme, posterior view. Abbreviations: EJAP = ejaculatory apodeme; CE = cercus; EP = epandrium; Ph = phallus; Hyp = hypandrium; St = sternite; Scales: Figures 70, 7274 = 1.0 mm; 71 = 0.5 mm; 7576 = 0.2 mm." pageId="16" pageNumber="79">Fig. 75</figureCitation>
).
</paragraph>
<paragraph id="09732D0BFFC2FF84D6B6FEE4FB2BD7C3" blockId="16.[151,1164,367,392]" box="[151,1164,367,392]" pageId="16" pageNumber="79">
The following four Neotropical
<taxonomicName id="CECC5688FFC2FF84D7DFFEFBFDE3D7C3" authorityName="Linnaeus" authorityYear="1758" box="[510,580,368,391]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="3BB8F119FFC2FF84D7DFFEFBFDE3D7C3" box="[510,580,368,391]" italics="true" pageId="16" pageNumber="79">Empis</emphasis>
</taxonomicName>
<typeStatus id="D67793A9FFC2FF84D46AFEFBFDDCD7CC" box="[587,635,368,392]" pageId="16" pageNumber="79">type</typeStatus>
specimens described by Bezzi were destroyed:
</paragraph>
<paragraph id="09732D0BFFC2FF84D6B6FE32FCB4D78B" blockId="16.[151,811,439,572]" box="[151,787,439,464]" pageId="16" pageNumber="79">
<taxonomicName id="CECC5688FFC2FF84D6B6FE32FE00D78B" authority="Bezzi, 1905" authorityName="Bezzi" authorityYear="1905" box="[151,423,439,464]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="79" phylum="Arthropoda" rank="species" species="ambigua">
<emphasis id="3BB8F119FFC2FF84D6B6FE32FEBFD794" box="[151,280,440,464]" italics="true" pageId="16" pageNumber="79">E. ambigua</emphasis>
<materialsCitation id="B9A42756FFC2FF84D73FFE3CFE00D78B" box="[286,423,439,463]" country="Peru" location="Vilcanota" pageId="16" pageNumber="79" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="6D5D50FAFFC2FF84D73FFE3CFE00D78B" author="Bezzi, M." box="[286,423,439,463]" pageId="16" pageNumber="79" pagination="424 - 460" refId="ref10956" refString="Bezzi, M. (1905) Empididae neotropicae Musei Nationalis Hungarici. Annales Historia Naturalis Musei Nationalis Hungarici, 33, 424 - 460." type="journal article" year="1905">Bezzi, 1905</bibRefCitation>
</materialsCitation>
</taxonomicName>
.
<typeStatus id="D67793A9FFC2FF84D793FE3CFDBDD78B" box="[434,538,439,463]" pageId="16" pageNumber="79" type="holotype">Holotype</typeStatus>
from
<location id="0C137BD0FFC2FF84D47EFE3CFD6AD78B" LSID="urn:lsid:plazi:treatment:81659C1DFFDFFF84D6B6F92DFCA8D478:0C137BD0FFC2FF84D47EFE3CFD6AD78B" box="[607,717,439,463]" country="Peru" name="Vilcanota" pageId="16" pageNumber="79">Vilcanota</location>
,
<collectingCountry id="71DB6D9BFFC2FF84D4F9FE33FCA8D78B" box="[728,783,440,463]" name="Peru" pageId="16" pageNumber="79">Peru</collectingCountry>
.
</paragraph>
<paragraph id="09732D0BFFC2FF84D6B6FE56FC8CD7B7" blockId="16.[151,811,439,572]" box="[151,811,476,500]" pageId="16" pageNumber="79">
<taxonomicName id="CECC5688FFC2FF84D6B6FE56FE60D7B0" authority="Bezzi, 1905" authorityName="Bezzi" authorityYear="1905" box="[151,455,476,500]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="79" phylum="Arthropoda" rank="species" species="lucidilabris">
<emphasis id="3BB8F119FFC2FF84D6B6FE56FE90D7B0" box="[151,311,476,500]" italics="true" pageId="16" pageNumber="79">E. lucidilabris</emphasis>
<materialsCitation id="B9A42756FFC2FF84D71FFE57FE60D7B0" box="[318,455,476,500]" country="Peru" location="Callanga" pageId="16" pageNumber="79" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="6D5D50FAFFC2FF84D71FFE57FE60D7B0" author="Bezzi, M." box="[318,455,476,500]" pageId="16" pageNumber="79" pagination="424 - 460" refId="ref10956" refString="Bezzi, M. (1905) Empididae neotropicae Musei Nationalis Hungarici. Annales Historia Naturalis Musei Nationalis Hungarici, 33, 424 - 460." type="journal article" year="1905">Bezzi, 1905</bibRefCitation>
</materialsCitation>
</taxonomicName>
.
<typeStatus id="D67793A9FFC2FF84D7F3FE57FD9DD7B0" box="[466,570,476,500]" pageId="16" pageNumber="79" type="holotype">Holotype</typeStatus>
from
<location id="0C137BD0FFC2FF84D45FFE57FD43D7B0" LSID="urn:lsid:plazi:treatment:81659C1DFFDFFF84D6B6F92DFCA8D478:0C137BD0FFC2FF84D45FFE57FD43D7B0" box="[638,740,476,500]" country="Peru" name="Callanga" pageId="16" pageNumber="79">Callanga</location>
,
<collectingCountry id="71DB6D9BFFC2FF84D4D1FE57FC80D7B7" box="[752,807,476,499]" name="Peru" pageId="16" pageNumber="79">Peru</collectingCountry>
.
</paragraph>
<paragraph id="09732D0BFFC2FF84D6B6FD8AFC87D453" blockId="16.[151,811,439,572]" box="[151,800,511,536]" pageId="16" pageNumber="79">
<taxonomicName id="CECC5688FFC2FF84D6B6FD8AFE1BD453" authority="Bezzi, 1905" authorityName="Bezzi" authorityYear="1905" box="[151,444,511,536]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="79" phylum="Arthropoda" rank="species" species="micropyga">
<emphasis id="3BB8F119FFC2FF84D6B6FD8AFE8AD453" box="[151,301,512,536]" italics="true" pageId="16" pageNumber="79">E. micropyga</emphasis>
<materialsCitation id="B9A42756FFC2FF84D715FE74FE1BD453" box="[308,444,511,535]" country="Peru" location="Callanga" pageId="16" pageNumber="79" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="6D5D50FAFFC2FF84D715FE74FE1BD453" author="Bezzi, M." box="[308,444,511,535]" pageId="16" pageNumber="79" pagination="424 - 460" refId="ref10956" refString="Bezzi, M. (1905) Empididae neotropicae Musei Nationalis Hungarici. Annales Historia Naturalis Musei Nationalis Hungarici, 33, 424 - 460." type="journal article" year="1905">Bezzi, 1905</bibRefCitation>
</materialsCitation>
</taxonomicName>
.
<typeStatus id="D67793A9FFC2FF84D7E6FE74FD88D453" box="[455,559,511,535]" pageId="16" pageNumber="79" type="holotype">Holotype</typeStatus>
from
<location id="0C137BD0FFC2FF84D455FE74FD7DD453" LSID="urn:lsid:plazi:treatment:81659C1DFFDFFF84D6B6F92DFCA8D478:0C137BD0FFC2FF84D455FE74FD7DD453" box="[628,730,511,535]" country="Peru" name="Callanga" pageId="16" pageNumber="79">Callanga</location>
,
<collectingCountry id="71DB6D9BFFC2FF84D4C4FD8BFCBBD453" box="[741,796,512,535]" name="Peru" pageId="16" pageNumber="79">Peru</collectingCountry>
.
</paragraph>
<paragraph id="09732D0BFFC2FF84D6B6FDAEFCA8D478" blockId="16.[151,811,439,572]" box="[151,783,548,572]" pageId="16" pageNumber="79">
<taxonomicName id="CECC5688FFC2FF84D6B6FDAEFE32D478" authority="Bezzi, 1905" authorityName="Bezzi" authorityYear="1905" box="[151,405,548,572]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="79" phylum="Arthropoda" rank="species" species="bullata">
<emphasis id="3BB8F119FFC2FF84D6B6FDAEFEA1D478" box="[151,262,548,572]" italics="true" pageId="16" pageNumber="79">E. bullata</emphasis>
<materialsCitation id="B9A42756FFC2FF84D72DFDAFFE32D478" box="[268,405,548,572]" country="Bolivia" location="Coroico" pageId="16" pageNumber="79" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="6D5D50FAFFC2FF84D72DFDAFFE32D478" author="Bezzi, M." box="[268,405,548,572]" pageId="16" pageNumber="79" pagination="424 - 460" refId="ref10956" refString="Bezzi, M. (1905) Empididae neotropicae Musei Nationalis Hungarici. Annales Historia Naturalis Musei Nationalis Hungarici, 33, 424 - 460." type="journal article" year="1905">Bezzi, 1905</bibRefCitation>
</materialsCitation>
</taxonomicName>
.
<typeStatus id="D67793A9FFC2FF84D781FDAFFDAFD478" box="[416,520,548,572]" pageId="16" pageNumber="79" type="holotype">Holotype</typeStatus>
from
<location id="0C137BD0FFC2FF84D46DFDAFFD0ED478" LSID="urn:lsid:plazi:treatment:81659C1DFFDFFF84D6B6F92DFCA8D478:0C137BD0FFC2FF84D46DFDAFFD0ED478" box="[588,681,548,572]" country="Bolivia" name="Coroico" pageId="16" pageNumber="79">Coroico</location>
,
<collectingCountry id="71DB6D9BFFC2FF84D494FDAFFCACD478" box="[693,779,548,572]" name="Bolivia" pageId="16" pageNumber="79">Bolivia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>