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<document id="40EFC161B438706D7F0945792E191227" ID-CLB-Dataset="22140" ID-DOI="10.5852/ejt.2017.318" ID-GBIF-Dataset="3a1f57d6-6cad-4bb9-8fdd-6b2b471bce0c" ID-ISSN="2118-9773" ID-Zenodo-Dep="3827451" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589469308205" checkinUser="carolina" docAuthor="Alwin-Kownacka, Alicja, Szadziewski, Ryszard &amp; Szwedo, Jacek" docDate="2017" docId="37097034F871FFB3FEBF3BFEFDFBFE50" docLanguage="en" docName="ejt-2017-318.pdf.imf" docOrigin="European Journal of Taxonomy 318" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Bezzia (Homobezzia) sharjahi Alwin-Kownacka &amp; Szadziewski &amp; Szwedo 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="10" masterDocId="CB30084CF877FFBAFFD33E19FF86FFED" masterDocTitle="Predatory midges of the tribes Palpomyiini and Sphaeromiini (Diptera: Ceratopogonidae) from the Middle East, with keys and descriptions of new species" masterLastPageNumber="30" masterPageNumber="1" pageNumber="7" updateTime="1698839735362" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="62ADD091F22D4CCA480E6A362B8B9C52" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5305FF174B8519F3C8019D5E7FF76610">
<mods:title id="D29F0B72098D7D6D6F06DC1D02452D49">Predatory midges of the tribes Palpomyiini and Sphaeromiini (Diptera: Ceratopogonidae) from the Middle East, with keys and descriptions of new species</mods:title>
</mods:titleInfo>
<mods:name id="D268FAF0237A59E80003473B87E83C3A" type="personal">
<mods:role id="1101C0555337F6DEAA85364C7A10846A">
<mods:roleTerm id="B5197C4595BBA8F45EA3F7FC0948A3EA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F3A2BAACA5F8453F108854D526316E0C">Alwin-Kownacka, Alicja</mods:namePart>
<mods:nameIdentifier id="E84A76A70AC319B5EC8B7420C35AC508" type="LSID">0995CC31-5CC8-46F3-BB72-A4D76919750A</mods:nameIdentifier>
<mods:affiliation id="859B9CB7414C5D53F00054C5A9730E4B">urn: lsid: zoobank. org: author: 0995 CC 31 - 5 CC 8 - 46 F 3 - BB 72 - A 4 D 76919750 A &amp; Corresponding author: alicja. alwin @ biol. ug. edu. pl</mods:affiliation>
<mods:nameIdentifier id="E9BA393267070AE2F62FA644544B4F49" type="email">alicja.alwin@biol.ug.edu.pl</mods:nameIdentifier>
</mods:name>
<mods:name id="73A17F468712E1570C19703BE45B0D20" type="personal">
<mods:role id="AC445B02587ED526C8F0A1052DD35D6A">
<mods:roleTerm id="760F4F28CC1496267C794E66536A32FA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FC9DF5D4416AEDC23FB12D562860D570">Szadziewski, Ryszard</mods:namePart>
<mods:nameIdentifier id="8D9F99329C66A19D57708861CA789EDC" type="LSID">083FF55D-C4C0-4C7D-AE23-562619624664</mods:nameIdentifier>
<mods:affiliation id="DB4EFFC3BD8836F3E3E5C62981DB8EE0">Email: ryszard. szadziewski @ biol. ug. edu. pl &amp; urn: lsid: zoobank. org: author: 083 FF 55 D-C 4 C 0 - 4 C 7 D-AE 23 - 562619624664</mods:affiliation>
<mods:nameIdentifier id="27E1B5F910966B448B2063F8FA22FEFD" type="email">ryszard.szadziewski@biol.ug.edu.pl</mods:nameIdentifier>
</mods:name>
<mods:name id="80E0D52CFE857722A80E22885502C807" type="personal">
<mods:role id="8057862110B138378A93054F6C4D74EE">
<mods:roleTerm id="013443B67CCE2FCA80ADAF0ED652B343">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D355305DA21ADB5CB3C9986AE4BB0E45">Szwedo, Jacek</mods:namePart>
<mods:nameIdentifier id="C31CA782B9B54D3260AF97C3E3862549" type="LSID">36BAE161-ECCD-4D89-B383-C7CC84FF0A89</mods:nameIdentifier>
<mods:affiliation id="378792099D3912D493BB33A052B2FACA">Email: jacek. szwedo @ biol. ug. edu. pl &amp; urn: lsid: zoobank. org: author: 36 BAE 161 - ECCD- 4 D 89 - B 383 - C 7 CC 84 FF 0 A 89</mods:affiliation>
<mods:nameIdentifier id="411DEB740FE567E05E42435DB2AFAD7B" type="email">jacek.szwedo@biol.ug.edu.pl</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="3336536C67FB6BB56D4F73EEC0C0E927">text</mods:typeOfResource>
<mods:relatedItem id="C86C7DAECF2D90D4052A60E8ADD4A137" type="host">
<mods:titleInfo id="6239D0A39227C504A842142AEF853320">
<mods:title id="2B19892320A2E1871B7FE8C9C838E32F">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="C1A048247087CAFA0D05002288A1C24B">
<mods:date id="9A750556700D48AD5DEBB231DA206FC7">2017</mods:date>
<mods:detail id="760EFFF113E20385F28CA49916A8D64D" type="pubDate">
<mods:number id="1EA66D3957B84CBCF84E7DECD03DF774">2017-05-15</mods:number>
</mods:detail>
<mods:detail id="21CB526A38D44511C3197D2C2F31159E" type="issue">
<mods:number id="60732E304B80BD36BFE29794BBA2101D">318</mods:number>
</mods:detail>
<mods:extent id="1F87D3C00D69B0967CEF21177BF991CB" unit="page">
<mods:start id="BA40203D804E8BBA73CC6E2E0F50C685">1</mods:start>
<mods:end id="36D30111E99E87E914054FE19018A844">30</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="C5429847CCD345D28927C523985698B7">journal article</mods:classification>
<mods:identifier id="9F4F2A2557AB794ED9A1BEEBB64DE8A4" type="CLB-Dataset">22140</mods:identifier>
<mods:identifier id="D7E6E58C809FE88E47795EA5FD832271" type="DOI">10.5852/ejt.2017.318</mods:identifier>
<mods:identifier id="678686D4092239514EF4A8E325576989" type="GBIF-Dataset">3a1f57d6-6cad-4bb9-8fdd-6b2b471bce0c</mods:identifier>
<mods:identifier id="2A694B386E9006DC3AA94070D3B0049E" type="ISSN">2118-9773</mods:identifier>
<mods:identifier id="9487A90595E483E5079E601C476B34E5" type="Zenodo-Dep">3827451</mods:identifier>
<mods:identifier id="96A8C717A78C7BA73F5DF3F6939BD0CB" type="ZooBank">541EA4D0-3883-4190-8E10-FF7CAC1BB035</mods:identifier>
</mods:mods>
<treatment id="37097034F871FFB3FEBF3BFEFDFBFE50" ID-DOI="http://doi.org/10.5281/zenodo.3847266" ID-GBIF-Taxon="164151097" ID-Zenodo-Dep="3847266" LSID="urn:lsid:plazi:treatment:37097034F871FFB3FEBF3BFEFDFBFE50" httpUri="http://treatment.plazi.org/id/37097034F871FFB3FEBF3BFEFDFBFE50" lastPageId="9" lastPageNumber="10" pageId="6" pageNumber="7">
<subSubSection id="F7BA92A9F871FFBCFEBF3BFEFB4EF9EF" box="[364,1224,1511,1538]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="BF1FC122F871FFBCFEBF3BFEFB4EF9EF" blockId="6.[364,1224,1511,1608]" box="[364,1224,1511,1538]" pageId="6" pageNumber="7">
<heading id="E457764EF871FFBCFEBF3BFEFB4EF9EF" box="[364,1224,1511,1538]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="7" reason="2">
<taxonomicName id="78A0BAA1F871FFBCFEBF3BFEFBE0F9EC" ID-CoL="BCJW4" authority="Alwin-Kownacka &amp; Szadziewski" authorityName="Alwin-Kownacka &amp; Szadziewski &amp; Szwedo" authorityYear="2017" box="[364,1126,1511,1538]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="sharjahi" status="sp. nov." subGenus="Homobezzia">
<emphasis id="8DD41D30F871FFBCFEBF3BFEFD50F9EF" bold="true" box="[364,726,1511,1538]" pageId="6" pageNumber="7">
<emphasis id="8DD41D30F871FFBCFEBF3BFEFE3CF9EC" bold="true" box="[364,442,1511,1537]" italics="true" pageId="6" pageNumber="7">Bezzia</emphasis>
(
<emphasis id="8DD41D30F871FFBCFE183BFEFDD9F9EF" bold="true" box="[459,607,1511,1538]" italics="true" pageId="6" pageNumber="7">Homobezzia</emphasis>
)
<emphasis id="8DD41D30F871FFBCFDA23BFEFD50F9EF" bold="true" box="[625,726,1511,1538]" italics="true" pageId="6" pageNumber="7">sharjahi</emphasis>
</emphasis>
Alwin-Kownacka &amp; Szadziewski
</taxonomicName>
<taxonomicNameLabel id="96E7A04BF871FFBCFBBE3BF1FB4EF9EF" box="[1133,1224,1512,1538]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F7BA92A9F871FFBCFEBC3813FCBAF9A5" pageId="6" pageNumber="7" type="description">
<paragraph id="BF1FC122F871FFBCFEBC3813FB43F9C8" blockId="6.[364,1224,1511,1608]" box="[367,1221,1546,1573]" pageId="6" pageNumber="7">
<uri id="CB31CD20F871FFBCFEBC3813FB43F9C8" box="[367,1221,1546,1573]" pageId="6" pageNumber="7">
urn:lsid:zoobank.org:act:
<uuid id="CB06FBF7F871FFBCFD5E3813FB43F9C8" box="[653,1221,1546,1573]" pageId="6" pageNumber="7">07610F8D-C955-4FD8-851A-395BA06AFB87</uuid>
</uri>
</paragraph>
<paragraph id="BF1FC122F871FFBCFD243837FCBAF9A5" blockId="6.[364,1224,1511,1608]" box="[759,828,1582,1608]" pageId="6" pageNumber="7">
<figureCitation id="279BDDA7F871FFBCFD243837FCBAF9A5" box="[759,828,1582,1608]" captionStart="Fig" captionStartId="8.[189,232,1964,1990]" captionTargetBox="[233,1354,263,1929]" captionTargetId="figure@8.[233,1354,263,1929]" captionTargetPageId="8" captionText="Fig. 2. Bezzia sharjahi Szadziewski &amp; Alwin sp. nov. A. Wing, ♀. B. Antennal flagellum, ♂. C. Antennal flagellum, ♀. D. Genitalia, ♂. E. Parameres. F. Aedeagus. G. Seminal capsules, ♀." figureDoi="http://doi.org/10.5281/zenodo.3827455" httpUri="https://zenodo.org/record/3827455/files/figure.png" pageId="6" pageNumber="7">Fig. 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F7BA92A9F871FFBCFF6E3876FB09F8CC" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="BF1FC122F871FFBCFF6E3876FEB0F964" blockId="6.[189,310,1647,1673]" box="[189,310,1647,1673]" pageId="6" pageNumber="7">
<heading id="E457764EF871FFBCFF6E3876FEB0F964" bold="true" box="[189,310,1647,1673]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="8DD41D30F871FFBCFF6E3876FEB0F964" bold="true" box="[189,310,1647,1673]" pageId="6" pageNumber="7">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="BF1FC122F871FFBCFF6E3887FB09F8CC" blockId="6.[189,1398,1694,1826]" pageId="6" pageNumber="7">This new species differs from all other Middle East congeners in having the following combination of characters: thorax orange-brown with distinctly paler scutellum; legs uniformly brownish; fore femur with 6–9 ventral spines; male gonostylus long, with pointed apex; parameres long, slender, slightly curved dorsally; female with 2 subspherical seminal capsules with very long necks.</paragraph>
</subSubSection>
<subSubSection id="F7BA92A9F871FFBCFF6E3950FB68F87C" box="[189,1262,1865,1938]" pageId="6" pageNumber="7" type="etymology">
<paragraph id="BF1FC122F871FFBCFF6E3950FEC2F88E" blockId="6.[189,324,1865,1891]" box="[189,324,1865,1891]" pageId="6" pageNumber="7">
<heading id="E457764EF871FFBCFF6E3950FEC2F88E" bold="true" box="[189,324,1865,1891]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="8DD41D30F871FFBCFF6E3950FEC2F88E" bold="true" box="[189,324,1865,1891]" pageId="6" pageNumber="7">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="BF1FC122F871FFBCFF6E396EFB68F87C" blockId="6.[189,1262,1911,1938]" box="[189,1262,1911,1938]" pageId="6" pageNumber="7">
The specific name refers to the
<collectingRegion id="7D640FC0F871FFBCFDFE396EFD01F87C" box="[557,647,1911,1937]" country="United Arab Emirates" name="Ash Shariqah" pageId="6" pageNumber="7">Sharjah</collectingRegion>
Desert, where most of the
<typeStatus id="601B7F80F871FFBCFC173961FC71F87F" box="[964,1015,1912,1938]" pageId="6" pageNumber="7">type</typeStatus>
series was collected.
</paragraph>
</subSubSection>
<subSubSection id="F7BA92A9F870FFBDFF6E3F12FB00FD6B" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="BF1FC122F870FFBDFF6E3F12FE2CFEC8" blockId="7.[189,426,267,293]" box="[189,426,267,293]" pageId="7" pageNumber="8">
<heading id="E457764EF870FFBDFF6E3F12FE2CFEC8" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="8DD41D30F870FFBDFF6E3F12FE2CFEC8" bold="true" box="[189,426,267,293]" pageId="7" pageNumber="8">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="0FC8CB7FF870FFBDFF393F23FAF1FE9A" ID-GBIF-Occurrence="2618817302" collectingDate="2005-01-18" collectionCode="CEIUG" collectorName="A. van Harten" country="United Arab Emirates" location="Desert Park" pageId="7" pageNumber="8" specimenCount="1" specimenCount-male="1" stateProvince="Sharjah" typeStatus="holotype">
<paragraph id="BF1FC122F870FFBDFF393F23FEDAFEB9" blockId="7.[189,1399,314,375]" box="[234,348,314,340]" pageId="7" pageNumber="8">
<emphasis id="8DD41D30F870FFBDFF393F23FEDAFEB9" bold="true" box="[234,348,314,340]" pageId="7" pageNumber="8">
<typeStatus id="601B7F80F870FFBDFF393F23FEDAFEB9" box="[234,348,314,340]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="BF1FC122F870FFBDFF6E3F44FAF1FE9A" blockId="7.[189,1399,314,375]" box="[189,1399,348,375]" pageId="7" pageNumber="8">
<collectingCountry id="C7B781B2F870FFBDFF6E3F44FD9DFE9A" box="[189,539,349,375]" name="United Arab Emirates" pageId="7" pageNumber="8">UNITED ARAB EMIRATES</collectingCountry>
:
<specimenCount id="A9A60AABF870FFBDFDF63F44FDBBFE9A" box="[549,573,349,375]" pageId="7" pageNumber="8" type="male">♂</specimenCount>
, adult,
<collectingRegion id="7D640FC0F870FFBDFD423F45FD6DFE9B" box="[657,747,348,374]" country="United Arab Emirates" name="Ash Shariqah" pageId="7" pageNumber="8">Sharjah</collectingRegion>
<location id="BA7F97F9F870FFBDFD223F44FCFBFE9B" LSID="urn:lsid:plazi:treatment:37097034F871FFB3FEBF3BFEFDFBFE50:BA7F97F9F870FFBDFD223F44FCFBFE9B" box="[753,893,348,375]" country="United Arab Emirates" name="Desert Park" pageId="7" pageNumber="8" stateProvince="Sharjah">Desert Park</location>
,
<date id="CB1EE7E2F870FFBDFC543F44FB98FE9A" box="[903,1054,348,375]" pageId="7" pageNumber="8" value="2005-01-18">
<collectingDate id="DB5A1E0AF870FFBDFC543F44FB98FE9A" box="[903,1054,348,375]" pageId="7" pageNumber="8" value="2005-01-18">18 Jan. 2005</collectingDate>
</date>
,
<collectorName id="1255A4F4F870FFBDFBF43F44FB4CFE9A" box="[1063,1226,349,375]" pageId="7" pageNumber="8">A. van Harten</collectorName>
leg. (
<collectionCode id="D9B159E7F870FFBDFAD93F44FAECFE9A" box="[1290,1386,349,375]" pageId="7" pageNumber="8">CEIUG</collectionCode>
).
</paragraph>
</materialsCitation>
<paragraph id="BF1FC122F870FFBDFF393F87FEE1FE55" blockId="7.[189,1399,414,582]" box="[234,359,414,440]" pageId="7" pageNumber="8">
<emphasis id="8DD41D30F870FFBDFF393F87FEE1FE55" bold="true" box="[234,359,414,440]" pageId="7" pageNumber="8">
<typeStatus id="601B7F80F870FFBDFF393F87FEE1FE55" box="[234,359,414,440]" pageId="7" pageNumber="8" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="BF1FC122F870FFBDFF6E3FDBFB0CFDA8" blockId="7.[189,1399,414,582]" pageId="7" pageNumber="8">
<collectingCountry id="C7B781B2F870FFBDFF6E3FDBFD9FFE36" box="[189,537,449,476]" name="United Arab Emirates" pageId="7" pageNumber="8">UNITED ARAB EMIRATES</collectingCountry>
:
<specimenCount id="A9A60AABF870FFBDFDF93FD8FDE8FE31" box="[554,622,449,476]" pageId="7" pageNumber="8" type="female">3 ♀♀</specimenCount>
,
<specimenCount id="A9A60AABF870FFBDFDAF3FD8FD47FE31" box="[636,705,449,476]" pageId="7" pageNumber="8" type="male">3 ♂♂</specimenCount>
, same collection data as
<typeStatus id="601B7F80F870FFBDFC3A3FD8FBD6FE36" box="[1001,1104,449,475]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
except
<date id="CB1EE7E2F870FFBDFB7F3FD8FAF4FE31" box="[1196,1394,449,476]" pageId="7" pageNumber="8" value="2005-01-18" valueMax="2005-01-25" valueMin="2005-01-18">18–25 Jan. 2005</date>
, light trap;
<specimenCount id="A9A60AABF870FFBDFEE53FFCFEFEFE12" box="[310,376,485,511]" pageId="7" pageNumber="8" type="female">3 ♀♀</specimenCount>
,
<specimenCount id="A9A60AABF870FFBDFE563FFCFE34FE12" box="[389,434,485,511]" pageId="7" pageNumber="8" type="male">1 ♂</specimenCount>
, same collection data except
<date id="CB1EE7E2F870FFBDFCD63FFCFC14FE12" box="[773,914,484,511]" pageId="7" pageNumber="8" value="2005-03-09">9 Mar. 2005</date>
;
<specimenCount id="A9A60AABF870FFBDFC733FFCFC4AFE12" box="[928,972,485,511]" pageId="7" pageNumber="8" type="female">1 ♀</specimenCount>
, same collection data except
<date id="CB1EE7E2F870FFBDFAF23FFCFF7EFDCF" pageId="7" pageNumber="8" value="2005-04-30">30 Apr. 2005</date>
;
<specimenCount id="A9A60AABF870FFBDFED53C11FEB7FDCF" box="[262,305,520,546]" pageId="7" pageNumber="8" type="female">1 ♀</specimenCount>
, same collection data except
<date id="CB1EE7E2F870FFBDFD563C11FC9CFDCF" box="[645,794,519,546]" pageId="7" pageNumber="8" value="2005-06-30">30 Jun. 2005</date>
;
<specimenCount id="A9A60AABF870FFBDFCFB3C11FCD5FDCF" box="[808,851,520,546]" pageId="7" pageNumber="8" type="female">1 ♀</specimenCount>
,
<specimenCount id="A9A60AABF870FFBDFCB23C11FC08FDCF" box="[865,910,520,546]" pageId="7" pageNumber="8" type="male">1 ♂</specimenCount>
, same collection data except
<date id="CB1EE7E2F870FFBDFB333C11FAF4FDCF" box="[1248,1394,519,546]" pageId="7" pageNumber="8" value="2005-07-21">21 Jul. 2005</date>
;
<specimenCount id="A9A60AABF870FFBDFF6E3C32FF6FFDA8" box="[189,233,555,581]" pageId="7" pageNumber="8" type="female">1 ♀</specimenCount>
, Wadi Wurayah, light trap,
<date id="CB1EE7E2F870FFBDFDFE3C32FCABFDA8" box="[557,813,555,582]" pageId="7" pageNumber="8" value="2009-01-15" valueMax="2009-02-22" valueMin="2009-01-15">15 Jan.–22 Feb. 2009</date>
, A. van Harten leg. (
<collectionCode id="D9B159E7F870FFBDFBCD3C32FBF8FDA8" box="[1054,1150,555,581]" pageId="7" pageNumber="8">CEIUG</collectionCode>
).
</paragraph>
<paragraph id="BF1FC122F870FFBDFF6E3C74FB00FD6B" blockId="7.[189,1158,620,647]" box="[189,1158,620,647]" pageId="7" pageNumber="8">
<collectingCountry id="C7B781B2F870FFBDFF6E3C74FEACFD6A" box="[189,298,621,647]" name="Yemen" pageId="7" pageNumber="8">YEMEN</collectingCountry>
:
<specimenCount id="A9A60AABF870FFBDFEE63C75FEFEFD6A" box="[309,376,620,647]" pageId="7" pageNumber="8" type="female">2 ♀♀</specimenCount>
, Al Mukalla, light trap,
<date id="CB1EE7E2F870FFBDFD413C75FCA1FD6B" box="[658,807,620,646]" pageId="7" pageNumber="8" value="2003-02-01">1. Feb. 2003</date>
, A. van Harten leg. (
<collectionCode id="D9B159E7F870FFBDFBCA3C75FBFFFD6B" box="[1049,1145,620,646]" pageId="7" pageNumber="8">CEIUG</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="F7BA92A9F870FFBDFF6E3CB4FD61F8F1" pageId="7" pageNumber="8" type="description">
<paragraph id="BF1FC122F870FFBDFF6E3CB4FEC8FD2A" blockId="7.[189,334,685,711]" box="[189,334,685,711]" pageId="7" pageNumber="8">
<heading id="E457764EF870FFBDFF6E3CB4FEC8FD2A" bold="true" box="[189,334,685,711]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="8DD41D30F870FFBDFF6E3CB4FEC8FD2A" bold="true" box="[189,334,685,711]" pageId="7" pageNumber="8">Description</emphasis>
</heading>
</paragraph>
<paragraph id="BF1FC122F870FFBDFF393CC5FEACFD1B" blockId="7.[189,1398,732,899]" box="[234,298,732,758]" pageId="7" pageNumber="8">
<emphasis id="8DD41D30F870FFBDFF393CC5FEACFD1B" bold="true" box="[234,298,732,758]" pageId="7" pageNumber="8">Male</emphasis>
</paragraph>
<paragraph id="BF1FC122F870FFBDFF6E3D19FDC8FC6E" blockId="7.[189,1398,732,899]" pageId="7" pageNumber="8">
HEAD. Uniformly pale brown. Antenna (
<figureCitation id="279BDDA7F870FFBDFD6B3CE6FC9CFCF7" box="[696,794,767,794]" captionStart="Fig" captionStartId="8.[189,232,1964,1990]" captionTargetBox="[233,1354,263,1929]" captionTargetId="figure@8.[233,1354,263,1929]" captionTargetPageId="8" captionText="Fig. 2. Bezzia sharjahi Szadziewski &amp; Alwin sp. nov. A. Wing, ♀. B. Antennal flagellum, ♂. C. Antennal flagellum, ♀. D. Genitalia, ♂. E. Parameres. F. Aedeagus. G. Seminal capsules, ♀." figureDoi="http://doi.org/10.5281/zenodo.3827455" httpUri="https://zenodo.org/record/3827455/files/figure.png" pageId="7" pageNumber="8">Fig. 2B</figureCitation>
) pale brown; flagellar plume poorly developed; flagellomeres 2(3)–13 with paler basal halves; flagellomeres 10–13 short; total flagellum length
<quantity id="78586CC7F870FFBDFAE73D3BFEAAFC8D" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="7" pageNumber="8" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.50– 0.70 mm</quantity>
; antennal ratio 0.50–0.70. Palpus yellow, slender; third palpal segment short, with distinct capitate sensilla; palpal ratio 2.00.
</paragraph>
<paragraph id="BF1FC122F870FFBDFF6E3DB2FB21FBC3" blockId="7.[189,1398,938,1070]" pageId="7" pageNumber="8">
THORAX. Orange-brown; scutellum paler. Wing pale, with barely visible veins; 1 poorly marked long radial cell; wing length
<quantity id="78586CC7F870FFBDFE013DD7FD06FC05" box="[466,640,974,1000]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="7" pageNumber="8" unit="mm" value="1.1" valueMax="1.3" valueMin="0.9">0.90–1.30 mm</quantity>
; costal ratio 0.70–0.80. Legs uniformly pale brown, tarsi slightly paler, coxae dark brown; claws small and equal on all legs; fore femur armed with 4–8 short ventral spines; tarsal ratio of fore leg 1.60–1.90, of mid leg 1.80–2.30, of hind leg 2.10–2.20.
</paragraph>
<paragraph id="BF1FC122F870FFBDFF6E3A4FFE6CFB82" blockId="7.[189,490,1109,1136]" box="[189,490,1109,1136]" pageId="7" pageNumber="8">ABDOMEN. Orange brown.</paragraph>
<paragraph id="BF1FC122F870FFBDFF6E3A8EFDBDFAF6" blockId="7.[189,1399,1175,1307]" pageId="7" pageNumber="8">
GENITALIA (
<figureCitation id="279BDDA7F870FFBDFE903A8EFE0AFB5C" box="[323,396,1175,1201]" captionStart="Fig" captionStartId="8.[189,232,1964,1990]" captionTargetBox="[233,1354,263,1929]" captionTargetId="figure@8.[233,1354,263,1929]" captionTargetPageId="8" captionText="Fig. 2. Bezzia sharjahi Szadziewski &amp; Alwin sp. nov. A. Wing, ♀. B. Antennal flagellum, ♂. C. Antennal flagellum, ♀. D. Genitalia, ♂. E. Parameres. F. Aedeagus. G. Seminal capsules, ♀." figureDoi="http://doi.org/10.5281/zenodo.3827455" httpUri="https://zenodo.org/record/3827455/files/figure.png" pageId="7" pageNumber="8">Fig. 2</figureCitation>
D–F). Tergite IX short, reaching apex of gonocoxite; sternite IX short. Gonocoxite short, stout; gonostylus long, narrow, distal half curved, with sharp apex. Parameres (
<figureCitation id="279BDDA7F870FFBDFB113AA3FA9BFB39" box="[1218,1309,1210,1236]" captionStart="Fig" captionStartId="8.[189,232,1964,1990]" captionTargetBox="[233,1354,263,1929]" captionTargetId="figure@8.[233,1354,263,1929]" captionTargetPageId="8" captionText="Fig. 2. Bezzia sharjahi Szadziewski &amp; Alwin sp. nov. A. Wing, ♀. B. Antennal flagellum, ♂. C. Antennal flagellum, ♀. D. Genitalia, ♂. E. Parameres. F. Aedeagus. G. Seminal capsules, ♀." figureDoi="http://doi.org/10.5281/zenodo.3827455" httpUri="https://zenodo.org/record/3827455/files/figure.png" pageId="7" pageNumber="8">Fig. 2E</figureCitation>
) fused, tongue-like, slightly curved dorsally, with blunt apex. Aedeagus (
<figureCitation id="279BDDA7F870FFBDFC613AC4FB8EFB1A" box="[946,1032,1245,1271]" captionStart="Fig" captionStartId="8.[189,232,1964,1990]" captionTargetBox="[233,1354,263,1929]" captionTargetId="figure@8.[233,1354,263,1929]" captionTargetPageId="8" captionText="Fig. 2. Bezzia sharjahi Szadziewski &amp; Alwin sp. nov. A. Wing, ♀. B. Antennal flagellum, ♂. C. Antennal flagellum, ♀. D. Genitalia, ♂. E. Parameres. F. Aedeagus. G. Seminal capsules, ♀." figureDoi="http://doi.org/10.5281/zenodo.3827455" httpUri="https://zenodo.org/record/3827455/files/figure.png" pageId="7" pageNumber="8">Fig. 2F</figureCitation>
) triangular, with low basal arch and barely visible rounded apex.
</paragraph>
<paragraph id="BF1FC122F870FFBDFF393B5BFEC3FAB1" blockId="7.[189,1399,1346,1513]" box="[234,325,1346,1372]" pageId="7" pageNumber="8">
<emphasis id="8DD41D30F870FFBDFF393B5BFEC3FAB1" bold="true" box="[234,325,1346,1372]" pageId="7" pageNumber="8">Female</emphasis>
</paragraph>
<paragraph id="BF1FC122F870FFBDFF6E3B7FFC87FA04" blockId="7.[189,1399,1346,1513]" pageId="7" pageNumber="8">
HEAD. Brown. Antennal flagellum (
<figureCitation id="279BDDA7F870FFBDFD8A3B7CFD35FA92" box="[601,691,1381,1407]" captionStart="Fig" captionStartId="8.[189,232,1964,1990]" captionTargetBox="[233,1354,263,1929]" captionTargetId="figure@8.[233,1354,263,1929]" captionTargetPageId="8" captionText="Fig. 2. Bezzia sharjahi Szadziewski &amp; Alwin sp. nov. A. Wing, ♀. B. Antennal flagellum, ♂. C. Antennal flagellum, ♀. D. Genitalia, ♂. E. Parameres. F. Aedeagus. G. Seminal capsules, ♀." figureDoi="http://doi.org/10.5281/zenodo.3827455" httpUri="https://zenodo.org/record/3827455/files/figure.png" pageId="7" pageNumber="8">Fig. 2C</figureCitation>
) with short flagellomeres, brown, proximal portions yellow; total flagellum length
<quantity id="78586CC7F870FFBDFE1F3B91FDF8FA4E" box="[460,638,1416,1443]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="7" pageNumber="8" unit="mm" value="0.6" valueMax="0.7" valueMin="0.5">0.50–0.70 mm</quantity>
; antennal ratio 0.80–1.10. Palpus slender, yellow; third palpal segment short, rather stout, without sensory pit; palpal ratio 2.00–2.50. Mandible with 6–8 massive distal teeth and numerous smaller proximal teeth.
</paragraph>
<paragraph id="BF1FC122F870FFBDFF6E3808FE47F955" blockId="7.[189,1398,1552,1720]" pageId="7" pageNumber="8">
THORAX. Orange brown; scutellum paler. Wing (
<figureCitation id="279BDDA7F870FFBDFD253809FCD2F9C7" box="[758,852,1552,1578]" captionStart="Fig" captionStartId="8.[189,232,1964,1990]" captionTargetBox="[233,1354,263,1929]" captionTargetId="figure@8.[233,1354,263,1929]" captionTargetPageId="8" captionText="Fig. 2. Bezzia sharjahi Szadziewski &amp; Alwin sp. nov. A. Wing, ♀. B. Antennal flagellum, ♂. C. Antennal flagellum, ♀. D. Genitalia, ♂. E. Parameres. F. Aedeagus. G. Seminal capsules, ♀." figureDoi="http://doi.org/10.5281/zenodo.3827455" httpUri="https://zenodo.org/record/3827455/files/figure.png" pageId="7" pageNumber="8">Fig. 2A</figureCitation>
) venation similar to that of male; wing length
<quantity id="78586CC7F870FFBDFF6E382DFEE8F9A3" box="[189,366,1588,1614]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="7" pageNumber="8" unit="mm" value="1.75" valueMax="1.8" valueMin="1.7">1.70–1.80 mm</quantity>
; costal ratio 0.80. Femoral coloration as in male; fore femur armed with 6–9 ventral spines; fore and mid tibiae with slightly dark bases, occasionally with slightly darker central areas, and dark apices; hind tibia with slightly darker base; tarsal ratio of fore leg 1.80–2.10, of mid leg 1.90–2.50, of hind leg 2.00–2.50.
</paragraph>
<paragraph id="BF1FC122F870FFBDFF6E38C6FD61F8F1" blockId="7.[189,1399,1758,1820]" pageId="7" pageNumber="8">
ABDOMEN. Two subspherical seminal capsules (
<figureCitation id="279BDDA7F870FFBDFD3438C7FCC4F914" box="[743,834,1758,1785]" captionStart="Fig" captionStartId="8.[189,232,1964,1990]" captionTargetBox="[233,1354,263,1929]" captionTargetId="figure@8.[233,1354,263,1929]" captionTargetPageId="8" captionText="Fig. 2. Bezzia sharjahi Szadziewski &amp; Alwin sp. nov. A. Wing, ♀. B. Antennal flagellum, ♂. C. Antennal flagellum, ♀. D. Genitalia, ♂. E. Parameres. F. Aedeagus. G. Seminal capsules, ♀." figureDoi="http://doi.org/10.5281/zenodo.3827455" httpUri="https://zenodo.org/record/3827455/files/figure.png" pageId="7" pageNumber="8">Fig. 2G</figureCitation>
) with long, thick necks; measuring 0.04–0.05 ×
<quantity id="78586CC7F870FFBDFF6E391BFEEEF8F1" box="[189,360,1794,1820]" metricMagnitude="-5" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="7" pageNumber="8" unit="mm" value="0.035" valueMax="0.04" valueMin="0.03">0.03–0.04 mm</quantity>
and 0.05–0.06 ×
<quantity id="78586CC7F870FFBDFDE63918FD65F8F1" box="[565,739,1793,1820]" metricMagnitude="-5" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="7" pageNumber="8" unit="mm" value="0.045" valueMax="0.05" valueMin="0.04">0.04–0.05 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="F7BA92A9F870FFBDFF6E395AFDC8F861" box="[189,590,1859,1932]" pageId="7" pageNumber="8" type="distribution">
<paragraph id="BF1FC122F870FFBDFF6E395AFED0F8B0" blockId="7.[189,342,1859,1885]" box="[189,342,1859,1885]" pageId="7" pageNumber="8">
<heading id="E457764EF870FFBDFF6E395AFED0F8B0" bold="true" box="[189,342,1859,1885]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="8DD41D30F870FFBDFF6E395AFED0F8B0" bold="true" box="[189,342,1859,1885]" pageId="7" pageNumber="8">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="BF1FC122F870FFBDFF6E396BFDC8F861" blockId="7.[189,590,1906,1932]" box="[189,590,1906,1932]" pageId="7" pageNumber="8">
<collectingCountry id="C7B781B2F870FFBDFF6E396BFE3AF861" box="[189,444,1906,1932]" name="United Arab Emirates" pageId="7" pageNumber="8">United Arab Emirates</collectingCountry>
and
<collectingCountry id="C7B781B2F870FFBDFE26396BFDCDF861" box="[501,587,1906,1932]" name="Yemen" pageId="7" pageNumber="8">Yemen</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="EBDF91AAF87FFFB2FF6E39B5FB0CF807" ID-DOI="http://doi.org/10.5281/zenodo.3827455" ID-Zenodo-Dep="3827455" httpUri="https://zenodo.org/record/3827455/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1964,1990]" targetBox="[233,1354,263,1929]" targetPageId="8">
<paragraph id="BF1FC122F87FFFB2FF6E39B5FB0CF807" blockId="8.[189,1399,1964,2026]" pageId="8" pageNumber="9">
<emphasis id="8DD41D30F87FFFB2FF6E39B5FE8FF82B" bold="true" box="[189,265,1964,1990]" pageId="8" pageNumber="9">Fig. 2.</emphasis>
<taxonomicName id="78A0BAA1F87FFFB2FEDC39B4FD43F82B" authority="Szadziewski &amp; Alwin" authorityName="Szadziewski &amp; Alwin" authorityYear="2017" box="[271,709,1964,1990]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="sharjahi" status="sp. nov.">
<emphasis id="8DD41D30F87FFFB2FEDC39B4FE45F82B" box="[271,451,1964,1990]" italics="true" pageId="8" pageNumber="9">Bezzia sharjahi</emphasis>
Szadziewski &amp; Alwin
</taxonomicName>
<taxonomicNameLabel id="96E7A04BF87FFFB2FD1839B4FCA2F82A" box="[715,804,1965,1991]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="8DD41D30F87FFFB2FCFA39B5FCB8F82B" bold="true" box="[809,830,1964,1990]" pageId="8" pageNumber="9">A</emphasis>
. Wing, ♀.
<emphasis id="8DD41D30F87FFFB2FC6839B4FC49F82A" bold="true" box="[955,975,1965,1991]" pageId="8" pageNumber="9">B</emphasis>
. Antennal flagellum, ♂.
<emphasis id="8DD41D30F87FFFB2FB3939B5FB79F82B" bold="true" box="[1258,1279,1964,1990]" pageId="8" pageNumber="9">C</emphasis>
. Antennal flagellum, ♀.
<emphasis id="8DD41D30F87FFFB2FE8C39C9FEF2F807" bold="true" box="[351,372,2000,2026]" pageId="8" pageNumber="9">D</emphasis>
. Genitalia, ♂.
<emphasis id="8DD41D30F87FFFB2FDF139C9FDB0F807" bold="true" box="[546,566,2000,2026]" pageId="8" pageNumber="9">E</emphasis>
. Parameres.
<emphasis id="8DD41D30F87FFFB2FD1E39C9FD59F807" bold="true" box="[717,735,2000,2026]" pageId="8" pageNumber="9">F</emphasis>
. Aedeagus.
<emphasis id="8DD41D30F87FFFB2FCBD39D6FC03F804" bold="true" box="[878,901,1999,2025]" pageId="8" pageNumber="9">G</emphasis>
. Seminal capsules, ♀.
</paragraph>
</caption>
<subSubSection id="F7BA92A9F87EFFB3FF6E3F12FDFBFE50" pageId="9" pageNumber="10" type="discussion">
<paragraph id="BF1FC122F87EFFB3FF6E3F12FEA9FEC8" blockId="9.[189,303,267,293]" box="[189,303,267,293]" pageId="9" pageNumber="10">
<heading id="E457764EF87EFFB3FF6E3F12FEA9FEC8" bold="true" box="[189,303,267,293]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="8DD41D30F87EFFB3FF6E3F12FEA9FEC8" bold="true" box="[189,303,267,293]" pageId="9" pageNumber="10">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="BF1FC122F87EFFB3FF6E3F20FDFBFE50" blockId="9.[189,1398,313,446]" pageId="9" pageNumber="10">
This new species is a typical member of the subgenus
<taxonomicName id="78A0BAA1F87EFFB3FC5F3F20FB9BFEBE" authorityName="Macfie" authorityYear="1932" box="[908,1053,313,339]" class="Insecta" family="Ceratopogonidae" genus="Homobezzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="8DD41D30F87EFFB3FC5F3F20FB9BFEBE" box="[908,1053,313,339]" italics="true" pageId="9" pageNumber="10">Homobezzia</emphasis>
</taxonomicName>
. It most closely resembles
<taxonomicName id="78A0BAA1F87EFFB3FF6E3F44FEE7FE9A" authorityName="Remm" authorityYear="1967" box="[189,353,349,375]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="kuhetiensis">
<emphasis id="8DD41D30F87EFFB3FF6E3F44FEE7FE9A" box="[189,353,349,375]" italics="true" pageId="9" pageNumber="10">B. kuhetiensis</emphasis>
</taxonomicName>
, which has similar male genitalia and seminal capsules. However,
<taxonomicName id="78A0BAA1F87EFFB3FBAD3F44FAA4FE9A" authorityName="Remm" authorityYear="1967" box="[1150,1314,349,375]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="kuhetiensis">
<emphasis id="8DD41D30F87EFFB3FBAD3F44FAA4FE9A" box="[1150,1314,349,375]" italics="true" pageId="9" pageNumber="10">B. kuhetiensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="78A0BAA1F87EFFB3FF2E3F98FE06FE77" authority="Alwin-Kownacka &amp; Szadziewski &amp; Szwedo, 2017" authorityName="Alwin-Kownacka &amp; Szadziewski &amp; Szwedo" authorityYear="2017" box="[253,384,384,410]" class="Insecta" family="Ceratopogonidae" genus="Bezzia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sharjahi" status="sp. nov.">
<emphasis id="8DD41D30F87EFFB3FF2E3F98FE89FE77" box="[253,271,385,410]" italics="true" pageId="9" pageNumber="10">B</emphasis>
.
<emphasis id="8DD41D30F87EFFB3FECD3F99FE06FE77" box="[286,384,384,410]" italics="true" pageId="9" pageNumber="10">sharjahi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="96E7A04BF87EFFB3FE543F99FE64FE77" box="[391,482,384,410]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
by its whitish legs with distinct, dark bands, the fore femur has only 2 ventral spines, and the parameres are straight.
</paragraph>
</subSubSection>
</treatment>
</document>
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