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<mods:title id="03FA89C4BF5C59C27626655E2C3C59FC">Review of the millipede genus Levizonus Attems, 1898, with description of a new species from the Far East of Russia (Diplopoda, Polydesmida, Xystodesmidae)</mods:title>
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<heading id="D0A1AE12FFD4FFF4555CFA6985B94895" box="[571,1017,1483,1510]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="163" reason="2">
<taxonomicName id="4C5662FDFFD4FFF4555CFA6985B94895" ID-CoL="727B6" authority="Takakuwa, 1942" authorityName="Takakuwa" authorityYear="1942" box="[571,1017,1483,1510]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="163" phylum="Arthropoda" rank="species" species="circularis">
<emphasis id="B922C56CFFD4FFF4555CFA6985704895" bold="true" box="[571,816,1483,1510]" italics="true" pageId="4" pageNumber="163">Levizonus circularis</emphasis>
<bibRefCitation id="EFC7648FFFD4FFF45450FA6985B94895" author="Takakuwa Y." box="[823,1017,1483,1510]" pageId="4" pageNumber="163" pagination="39 - 47" refId="ref10993" refString="Takakuwa Y. 1942. Zur Kenntnis der Japanischen Diplopoden. Annotationes Zoologicae Japonenses 21: 39 - 47." type="journal article" year="1942">Takakuwa, 1942</bibRefCitation>
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<paragraph id="8BE9197EFFD4FFF455B6FA4C85224B7B" blockId="4.[571,1017,1483,1544]" box="[721,866,1518,1544]" pageId="4" pageNumber="163">
<figureCitation id="136D05FBFFD4FFF455B6FA4C85784B7B" box="[721,824,1518,1544]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="2.[189,232,1824,1850]" captionStartId-1="5.[189,232,1773,1799]" captionStartId-2="6.[189,232,1789,1815]" captionTargetBox-0="[189,1398,267,1785]" captionTargetBox-1="[189,1397,265,1735]" captionTargetBox-2="[203,1359,1160,1746]" captionTargetId-0="figure-47@2.[186,1401,264,1789]" captionTargetId-1="figure-93@5.[186,1401,262,1738]" captionTargetId-2="figure-434@6.[186,1401,1152,1752]" captionTargetPageId-0="2" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="Fig. 1. Levizonus circularis Takakuwa, 1942, ♂, from Russia (FSCB 142005). A. Habitus, dorsal view. B. Midbody rings, dorsal view. Scale bars: A = 2 mm; B = 1 mm." captionText-1="Fig. 2. Levizonus circularis Takakuwa, 1942, ♂♂, ♀, from China (FSCB 12019Ch) (AB) and Russia (FSCB 232008, 212008) (CG). A. Gonopod, lateral view, longitudinal striae are not visible. B. Gonopod apex, lateral view. C, F. Gonopod apex, ventral view. D. Gonopod, lateral view, coxite partially destroyed. E. Middle part of gonopod, mesal view. G. Vulva, mesal view. Scale bars: A, E = 200 μm; D, G = 100 μm; BC, F = 20 μm." captionText-2="Fig. 3. Levizonus circularis Takakuwa, 1942, ♂♂, from Russia (AI) and North Korea(J). AD. Variations in shape of distal part of gonopod telopodite, ventral view. EJ. Variations in shape of distal part of gonopod telopodite, lateral view. Scale bars: AD, J = 0.25 mm; EG = 0.2 mm; HI = 0.1 mm. AH: after Mikhaljova 1981a; I: after Lokschina &amp; Golovatch 1977; J: after Mikhaljova et al. 2000." figureDoi-0="http://doi.org/10.5281/zenodo.4923496" figureDoi-1="http://doi.org/10.5281/zenodo.4923498" figureDoi-2="http://doi.org/10.5281/zenodo.4923500" httpUri-0="https://zenodo.org/record/4923496/files/figure.png" httpUri-1="https://zenodo.org/record/4923498/files/figure.png" httpUri-2="https://zenodo.org/record/4923500/files/figure.png" pageId="4" pageNumber="163">Figs 13</figureCitation>
,
<figureCitation id="136D05FBFFD4FFF45422FA4C85224B7B" box="[837,866,1518,1544]" captionStart="Fig" captionStartId="21.[189,232,1635,1661]" captionTargetBox="[190,1396,263,1593]" captionTargetId="figure-12@21.[186,1402,263,1599]" captionTargetPageId="21" captionText="Fig. 13. Distribution of species of the genus Levizonus Attems, 1898. Filled square: L. circularis Takakuwa, 1942. Filled circle: L. distinctus Mikhaljova, 1990. Open square: L. laqueatus Mikhaljova, 1981. Open triangle: L. nakhodka Mikhaljova sp. nov. Filled triangle: L. malewitschi Lokschina &amp; Golovatch, 1977. Filled diamond: L. thaumasius Attems, 1898. Heart: L. takakuwai (Verhoeff, 1941). Oval: L. montanus (Takakuwa, 1941)." figureDoi="http://doi.org/10.5281/zenodo.4923524" httpUri="https://zenodo.org/record/4923524/files/figure.png" pageId="4" pageNumber="163">13</figureCitation>
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<treatmentCitation id="0AF73F6FFFD4FFF457DAF99284934B39" author="Takakuwa Y." box="[189,723,1583,1610]" page="44" pageId="4" pageNumber="163" year="1942">
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<emphasis id="B922C56CFFD4FFF457DAF99287EC4B3A" box="[189,428,1583,1609]" italics="true" pageId="4" pageNumber="163">Levizonus circularis</emphasis>
<bibRefCitation id="EFC7648FFFD4FFF456D3F98D84934B39" author="Takakuwa Y." box="[436,723,1583,1610]" pageId="4" pageNumber="163" pagination="39 - 47" refId="ref10993" refString="Takakuwa Y. 1942. Zur Kenntnis der Japanischen Diplopoden. Annotationes Zoologicae Japonenses 21: 39 - 47." type="journal article" year="1942">Takakuwa, 1942: 4344</bibRefCitation>
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, 44, fig. 8 (
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or
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not designated; number of specimens in the type series unclear; location of types is unknown; type locality: Sangnam-ri,
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).
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<treatmentCitation id="0AF73F6FFFD4FFF457DAF93B85214BC1" author="Lokschina I. E. &amp; Golovatch S. I." box="[189,865,1688,1714]" page="77" pageId="4" pageNumber="163" year="1977">
<taxonomicName id="4C5662FDFFD4FFF457DAF93B85214BC1" ID-CoL="8S8WV" authority="Lokschina &amp; Golovatch, 1977: 76 - 77" authorityName="Lokschina &amp; Golovatch" authorityPageNumber="76 - 77" authorityYear="1977" box="[189,865,1688,1714]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="163" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B922C56CFFD4FFF457DAF93B87E74BC1" box="[189,423,1688,1714]" italics="true" pageId="4" pageNumber="163">Levizonus variabilis</emphasis>
<bibRefCitation id="EFC7648FFFD4FFF456CBF93A85214BC1" author="Lokschina I. E. &amp; Golovatch S. I." box="[428,865,1688,1714]" pageId="4" pageNumber="163" pagination="73 - 78" refId="ref9709" refString="Lokschina I. E. &amp; Golovatch S. I. 1977. The new myriapods (Diplopoda) from the USSR. Byulleten Moskovskogo obshchestva ispytateley prirodi, Biology 82 (1): 73 - 78. [In Russian.]" type="journal article" year="1977">Lokschina &amp; Golovatch, 1977: 7677</bibRefCitation>
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, fig. 3 (
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, from Olenevod, Lazovskiy District,
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,
<collectingCountry id="F34159EEFFD4FFF45570F91984284BA6" box="[535,616,1723,1749]" name="Russia" pageId="4" pageNumber="163">Russia</collectingCountry>
, in ZMUM).
<emphasis id="B922C56CFFD4FFF45461F91985374BA6" bold="true" box="[774,887,1723,1749]" pageId="4" pageNumber="163">Syn. nov.</emphasis>
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<taxonomicName id="4C5662FDFFD4FFF457DAF95F87EC4A65" ID-CoL="727B6" authorityName="Takakuwa" authorityYear="1942" box="[189,428,1788,1814]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="163" phylum="Arthropoda" rank="species" species="circularis">
<emphasis id="B922C56CFFD4FFF457DAF95F87EC4A65" box="[189,428,1788,1814]" italics="true" pageId="4" pageNumber="163">Levizonus circularis</emphasis>
</taxonomicName>
<treatmentCitation id="0AF73F6FFFD4FFF456ADF95E84A24A64" author="Takakuwa Y." box="[458,738,1788,1815]" page="74" pageId="4" pageNumber="163" year="1954">
<bibRefCitation id="EFC7648FFFD4FFF456ADF95E84A24A64" author="Takakuwa Y." box="[458,738,1788,1815]" pageId="4" pageNumber="163" refId="ref11013" refString="Takakuwa Y. 1954. Diplopoden aus Japan und ihn angrenzenden Gebieten. Japan Society for the Promotion of Science, Tokyo. [In Japanese.]" type="book" year="1954">Takakuwa 1954: 7475</bibRefCitation>
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, 74, fig. 79. —
<treatmentCitation id="0AF73F6FFFD4FFF454FBF95E82134A65" author="Paik K. - Y." box="[924,1107,1788,1815]" page="362" pageId="4" pageNumber="163" year="1958">
<bibRefCitation id="EFC7648FFFD4FFF454FBF95E82134A65" author="Paik K. - Y." box="[924,1107,1788,1815]" pageId="4" pageNumber="163" pagination="351 - 369" refId="ref10809" refString="Paik K. - Y. 1958. A list of Chilopoda, Symphila and Diplopoda from Korea. Kyungpook University, Theses Collection 2: 351 - 369." type="journal article" year="1958">Paik 1958: 362</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AF73F6FFFD4FFF453E3F95E83324A65" author="Golovatch S. I." box="[1156,1394,1788,1814]" page="17" pageId="4" pageNumber="163" year="1979">
<bibRefCitation id="EFC7648FFFD4FFF453E3F95E83324A65" author="Golovatch S. I." box="[1156,1394,1788,1814]" pageId="4" pageNumber="163" pagination="17 - 20" refId="ref9350" refString="Golovatch S. I. 1979. Notes on Levizonus thaumasius Att. (Diplopoda). In: Ivliev L. A. (ed.) Ekologiya i biologiya chlenistonogikh yuga Dalnego Vostoka: 17 - 20. DVNC AN SSSR, Vladivostok. [In Russian.]" type="book chapter" year="1979">Golovatch 1979: 17</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AF73F6FFFD4FFF45668F8BD84184A4A" author="Mikhaljova E. V. &amp; Golovatch S. I. &amp; Wytwer J." box="[271,600,1823,1849]" page="117" pageId="4" pageNumber="163" year="2000">
<bibRefCitation id="EFC7648FFFD4FFF45668F8BD84184A4A" author="Mikhaljova E. V. &amp; Golovatch S. I. &amp; Wytwer J." box="[271,600,1823,1849]" pageId="4" pageNumber="163" pagination="109 - 122" refId="ref10667" refString="Mikhaljova E. V., Golovatch S. I. &amp; Wytwer J. 2000. On some new or poorly-known millipedes (Diplopoda) from North Korea. Fragmenta Faunistica 43 (10): 109 - 122. https: // doi. org / 10.3161 / 00159301 FF 2000.43.10.109" type="journal article" year="2000">
Mikhaljova
<emphasis id="B922C56CFFD4FFF456F9F88287984A4A" box="[414,472,1823,1849]" italics="true" pageId="4" pageNumber="163">et al.</emphasis>
2000: 117
</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AF73F6FFFD4FFF455EFF8BD85C94A4A" author="Marek P. &amp; Tanabe T. &amp; Sierwald P." box="[648,905,1823,1850]" page="72" pageId="4" pageNumber="163" year="2014">
<bibRefCitation id="EFC7648FFFD4FFF455EFF8BD85C94A4A" author="Marek P. &amp; Tanabe T. &amp; Sierwald P." box="[648,905,1823,1850]" pageId="4" pageNumber="163" refId="ref9782" refString="Marek P., Tanabe T. &amp; Sierwald P. 2014. A Species Catalog of the Millipede Family Xystodesmidae (Diplopoda: Polydesmida). Virginia Museum of Natural History, Martinsville, Virginia." type="book" year="2014">
Marek
<emphasis id="B922C56CFFD4FFF455BBF882854F4A4A" box="[732,783,1823,1849]" italics="true" pageId="4" pageNumber="163">et al</emphasis>
. 2014: 72
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.
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</paragraph>
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<taxonomicName id="4C5662FDFFD4FFF457DAF8E187F14A2F" ID-CoL="8S8WV" authorityName="Lokschina &amp; Golovatch" authorityYear="1977" box="[189,433,1858,1884]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="163" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B922C56CFFD4FFF457DAF8E187F14A2F" box="[189,433,1858,1884]" italics="true" pageId="4" pageNumber="163">Levizonus variabilis</emphasis>
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<treatmentCitation id="0AF73F6FFFD4FFF456BAF8E085394A2E" author="Loksina I. E. &amp; Golovatch S. I." box="[477,889,1858,1885]" page="385" pageId="4" pageNumber="163" year="1979">
<bibRefCitation id="EFC7648FFFD4FFF456BAF8E085394A2E" author="Loksina I. E. &amp; Golovatch S. I." box="[477,889,1858,1885]" pageId="4" pageNumber="163" pagination="381 - 389" refId="ref9753" refString="Loksina I. E. &amp; Golovatch S. I. 1979. Diplopoda of the USSR fauna. Pedobiologia 19 (6): 381 - 389." type="journal article" year="1979">Lokšina &amp; Golovatch 1979: 385</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AF73F6FFFD4FFF454D1F8E083324A2F" author="Kurcheva G. F. &amp; Mikhaljova E. V." box="[950,1394,1858,1884]" page="120" pageId="4" pageNumber="163" year="1980">
<bibRefCitation id="EFC7648FFFD4FFF454D1F8E083324A2F" author="Kurcheva G. F. &amp; Mikhaljova E. V." box="[950,1394,1858,1884]" pageId="4" pageNumber="163" pagination="117 - 127" refId="ref9651" refString="Kurcheva G. F. &amp; Mikhaljova E. V. 1980. Millipedes (Diplopoda) of some forest types of the Verkhne-Ussuriyskii Research Station. In: Rozenberg V. A. (ed.) Kompleksnye issledovaniya lesnykh biogeotsenozov: 117 - 127. DVNC AN SSSR, Vladivostok. [In Russian.]" type="book chapter" year="1980">Kurcheva &amp; Mikhaljova 1980: 120</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AF73F6FFFD4FFF4567FF8C7845F4A0C" author="Mikhaljova E. V." box="[280,543,1893,1919]" page="64" pageId="4" pageNumber="163" year="1981">
<bibRefCitation id="EFC7648FFFD4FFF4567FF8C7845F4A0C" author="Mikhaljova E. V." box="[280,543,1893,1919]" pageId="4" pageNumber="163" pagination="62 - 67" refId="ref9873" refString="Mikhaljova E. V. 1981 a. The millipede genus Levizonus (Diplopoda, Xystodesmidae) from the USSR Far East. Byulleten Moskovskogo obshchestva ispytateley prirodi, Biology 86 (3): 62 - 67. [In Russian, English summary.]" type="journal article" year="1981">Mikhaljova 1981a: 64</bibRefCitation>
</treatmentCitation>
, figs 2, 4, map (fig. 5);
<treatmentCitation id="0AF73F6FFFD4FFF45452F8C785EF4A0C" author="Mikhaljova" box="[821,943,1893,1919]" page="87" pageId="4" pageNumber="163" year="1981">1981b: 87</treatmentCitation>
;
<treatmentCitation id="0AF73F6FFFD4FFF454DBF8C782674A0C" author="Mikhaljova" box="[956,1063,1893,1919]" page="81" pageId="4" pageNumber="163" year="1983">1983: 81</treatmentCitation>
;
<treatmentCitation id="0AF73F6FFFD4FFF45353F8C782EE4A0C" author="Mikhaljova" box="[1076,1198,1893,1919]" page="136" pageId="4" pageNumber="163" year="1990">1990: 136</treatmentCitation>
;
<treatmentCitation id="0AF73F6FFFD4FFF453DCF8C783674A0C" author="Mikhaljova" box="[1211,1319,1893,1919]" page="34" pageId="4" pageNumber="163" year="1993">1993: 34</treatmentCitation>
;
<treatmentCitation id="0AF73F6FFFD4FFF45253F8C784654AD1" author="Mikhaljova" page="55" pageId="4" pageNumber="163" year="1998">1998: 55, figs 218223, map 12</treatmentCitation>
;
<treatmentCitation id="0AF73F6FFFD4FFF45551F82A858E4AD1" author="Mikhaljova" box="[566,974,1928,1955]" page="251" pageId="4" pageNumber="163" year="2004">2004: 251, figs 638650, map 32</treatmentCitation>
;
<treatmentCitation id="0AF73F6FFFD4FFF454B9F82A82094AD0" author="Mikhaljova" box="[990,1097,1928,1955]" page="5" pageId="4" pageNumber="163" year="2009">2009a: 5</treatmentCitation>
;
<treatmentCitation id="0AF73F6FFFD4FFF4533DF82A82A64AD1" author="Mikhaljova" box="[1114,1254,1928,1954]" page="394" pageId="4" pageNumber="163" year="2009">2009b: 394</treatmentCitation>
;
<treatmentCitation id="0AF73F6FFFD4FFF45392F82A87AA4AB6" author="Mikhaljova" page="301" pageId="4" pageNumber="163" year="2017">2017: 301, figs 685697, map 43</treatmentCitation>
. —
<treatmentCitation id="0AF73F6FFFD4FFF4557FF80985F04AB5" author="Mikhaljova E. V. &amp; Petukhova E. L." box="[536,944,1963,1990]" page="53" pageId="4" pageNumber="163" year="1983">
<bibRefCitation id="EFC7648FFFD4FFF4557FF80985F04AB5" author="Mikhaljova E. V. &amp; Petukhova E. L." box="[536,944,1963,1990]" pageId="4" pageNumber="163" pagination="48 - 66" refId="ref10594" refString="Mikhaljova E. V. &amp; Petukhova E. L. 1983. A comparative analysis of the millipede faunas (Diplopoda) of Primorskiy Province's forests with the help of inclusion and similarity indices. In: Semkin B. I. &amp; Sukhanov V. V. (eds) Teoretiko-grafovye metody v biogeograficheskikh issledovaniyakh: 48 - 66. DVNC AN SSSR, Vladivostok. [In Russian.]" type="book chapter" year="1983">Mikhaljova &amp; Petukhova 1983: 53</bibRefCitation>
</treatmentCitation>
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<bibRefCitation id="EFC7648FFFD4FFF454B9F80982E54AB6" author="Ganin G. N." box="[990,1189,1963,1989]" pageId="4" pageNumber="163" refId="ref9237" refString="Ganin G. N. 1997. Soil Animals of the Ussuri Region. Dalnauka, Vladivostok / Khabarovsk. [In Russian, English summary.]" type="book" year="1997">Ganin 1997: 124</bibRefCitation>
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<treatmentCitation id="0AF73F6FFFD4FFF453D6F80983694AB5" author="Ganin G. N." box="[1201,1321,1963,1990]" page="153" pageId="4" pageNumber="163" year="2009">
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<treatmentCitation id="0AF73F6FFFD4FFF45252F80987594A9B" author="Ganin G. N." page="341" pageId="4" pageNumber="163" year="2011">
<bibRefCitation id="EFC7648FFFD4FFF45252F80987594A9B" author="Ganin G. N." pageId="4" pageNumber="163" refId="ref9320" refString="Ganin G. N. 2011. Structural and Functional Organization of Mezopedobiont Communities of the Southern Russian Far East. Dalnauka, Vladivostok. [In Russian.]" type="book" year="2011">2011: 341</bibRefCitation>
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<treatmentCitation id="0AF73F6FFFD4FFF4562EF86C845E4A9B" author="Tanabe T." box="[329,542,1998,2024]" page="108" pageId="4" pageNumber="163" year="1994">
<bibRefCitation id="EFC7648FFFD4FFF4562EF86C845E4A9B" author="Tanabe T." box="[329,542,1998,2024]" pageId="4" pageNumber="163" pagination="101 - 113" refId="ref11077" refString="Tanabe T. 1994. The millipede genus Levizonus (Polydesmida, Xystodesmidae) in Japan. Japanese Journal of Entomology 62 (1): 101 - 113." type="journal article" year="1994">Tanabe 1994: 108</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF73F6FFFD4FFF4554DF86C84F34A9B" author="Tanabe T." box="[554,691,1998,2024]" page="2178" pageId="4" pageNumber="163" year="2002">
<bibRefCitation id="EFC7648FFFD4FFF4554DF86C84F34A9B" author="Tanabe T." box="[554,691,1998,2024]" pageId="4" pageNumber="163" pagination="2139 - 2183" refId="ref11107" refString="Tanabe T. 2002. Revision of the millipede genus Parafontaria Verhoeff, 1936 (Diplopoda, Xystodesmidae). Journal of Natural History 36: 2139 - 2183. https: // doi. org / 10.1080 / 00222930110085610" type="journal article" year="2002">2002: 2178</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="DF2949F6FFD5FFF557DAF94F84054AE7" ID-DOI="http://doi.org/10.5281/zenodo.4923498" ID-Zenodo-Dep="4923498" httpUri="https://zenodo.org/record/4923498/files/figure.png" pageId="5" pageNumber="164" startId="5.[189,232,1773,1799]" targetBox="[189,1397,265,1735]" targetPageId="5">
<paragraph id="8BE9197EFFD5FFF557DAF94F84054AE7" blockId="5.[189,1399,1773,1940]" pageId="5" pageNumber="164">
<emphasis id="B922C56CFFD5FFF557DAF94F874B4A74" bold="true" box="[189,267,1773,1799]" pageId="5" pageNumber="164">Fig. 2.</emphasis>
<taxonomicName id="4C5662FDFFD5FFF55674F94C848E4A74" authority="Takakuwa, 1942" authorityName="Takakuwa" authorityYear="1942" box="[275,718,1773,1799]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="164" phylum="Arthropoda" rank="species" species="circularis">
<emphasis id="B922C56CFFD5FFF55674F94C84424A74" box="[275,514,1773,1799]" italics="true" pageId="5" pageNumber="164">Levizonus circularis</emphasis>
<bibRefCitation id="EFC7648FFFD5FFF5556DF94F848E4A74" author="Takakuwa Y." box="[522,718,1773,1799]" pageId="5" pageNumber="164" pagination="39 - 47" refId="ref10993" refString="Takakuwa Y. 1942. Zur Kenntnis der Japanischen Diplopoden. Annotationes Zoologicae Japonenses 21: 39 - 47." type="journal article" year="1942">Takakuwa, 1942</bibRefCitation>
</taxonomicName>
, ♂♂, ♀, from China (FSCB 12019Ch) (AB) and Russia (FSCB 232008, 212008) (CG).
<emphasis id="B922C56CFFD5FFF55551F8B2840B4A59" bold="true" box="[566,587,1808,1834]" pageId="5" pageNumber="164">A</emphasis>
. Gonopod, lateral view, longitudinal striae are not visible.
<emphasis id="B922C56CFFD5FFF5538EF8B282BD4A59" bold="true" box="[1257,1277,1808,1834]" pageId="5" pageNumber="164">B</emphasis>
. Gonopod apex, lateral view.
<emphasis id="B922C56CFFD5FFF556CBF89187A94A3D" bold="true" box="[428,489,1843,1870]" pageId="5" pageNumber="164">C, F</emphasis>
. Gonopod apex, ventral view.
<emphasis id="B922C56CFFD5FFF55417F89685C54A3D" bold="true" box="[880,901,1844,1870]" pageId="5" pageNumber="164">D</emphasis>
. Gonopod, lateral view, coxite partially destroyed.
<emphasis id="B922C56CFFD5FFF55658F8F587134A02" bold="true" box="[319,339,1879,1905]" pageId="5" pageNumber="164">E</emphasis>
. Middle part of gonopod, mesal view.
<emphasis id="B922C56CFFD5FFF55445F8F485794A03" bold="true" box="[802,825,1878,1904]" pageId="5" pageNumber="164">G</emphasis>
. Vulva, mesal view. Scale bars: A, E = 200 μm; D, G = 100 μm; BC, F = 20 μm.
</paragraph>
</caption>
<subSubSection id="C34C4AF5FFD6FFF657DAFEAA82F44F71" pageId="6" pageNumber="165" type="diagnosis">
<paragraph id="8BE9197EFFD6FFF657DAFEAA87764C51" blockId="6.[189,310,264,290]" box="[189,310,264,290]" pageId="6" pageNumber="165">
<heading id="D0A1AE12FFD6FFF657DAFEAA87764C51" bold="true" box="[189,310,264,290]" fontSize="11" level="3" pageId="6" pageNumber="165" reason="3">
<emphasis id="B922C56CFFD6FFF657DAFEAA87764C51" bold="true" box="[189,310,264,290]" pageId="6" pageNumber="165">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BE9197EFFD6FFF657DAFE9582F44F71" blockId="6.[189,1399,311,514]" pageId="6" pageNumber="165">
The main distinguishing characters of the species are as follows: apex of gonopod telopodite (
<figureCitation id="136D05FBFFD6FFF65274FE95832D4C22" box="[1299,1389,311,337]" captionStart="Fig" captionStartId="5.[189,232,1773,1799]" captionTargetBox="[189,1397,265,1735]" captionTargetId="figure-93@5.[186,1401,262,1738]" captionTargetPageId="5" captionText="Fig. 2. Levizonus circularis Takakuwa, 1942, ♂♂, ♀, from China (FSCB 12019Ch) (AB) and Russia (FSCB 232008, 212008) (CG). A. Gonopod, lateral view, longitudinal striae are not visible. B. Gonopod apex, lateral view. C, F. Gonopod apex, ventral view. D. Gonopod, lateral view, coxite partially destroyed. E. Middle part of gonopod, mesal view. G. Vulva, mesal view. Scale bars: A, E = 200 μm; D, G = 100 μm; BC, F = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.4923498" httpUri="https://zenodo.org/record/4923498/files/figure.png" pageId="6" pageNumber="165">Fig. 2C</figureCitation>
) not like two plates placed perpendicular to each other as in
<taxonomicName id="4C5662FDFFD6FFF654E2FEF982714C07" authorityName="Lokschina &amp; Golovatch" authorityYear="1977" box="[901,1073,346,372]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="165" phylum="Arthropoda" rank="species" species="malewitschi">
<emphasis id="B922C56CFFD6FFF654E2FEF982714C07" box="[901,1073,346,372]" italics="true" pageId="6" pageNumber="165">L. malewitschi</emphasis>
</taxonomicName>
; gonopod SL, more or less strongly curved mesad (vs not strongly curved mesad in other congeners); the central edge of the gonopod telopodite apex with outgrowths and spinules (
<figureCitation id="136D05FBFFD6FFF65421FE0385844CC8" box="[838,964,417,443]" captionStart="Fig" captionStartId="6.[189,232,1789,1815]" captionTargetBox="[203,1359,1160,1746]" captionTargetId="figure-434@6.[186,1401,1152,1752]" captionTargetPageId="6" captionText="Fig. 3. Levizonus circularis Takakuwa, 1942, ♂♂, from Russia (AI) and North Korea(J). AD. Variations in shape of distal part of gonopod telopodite, ventral view. EJ. Variations in shape of distal part of gonopod telopodite, lateral view. Scale bars: AD, J = 0.25 mm; EG = 0.2 mm; HI = 0.1 mm. AH: after Mikhaljova 1981a; I: after Lokschina &amp; Golovatch 1977; J: after Mikhaljova et al. 2000." figureDoi="http://doi.org/10.5281/zenodo.4923500" httpUri="https://zenodo.org/record/4923500/files/figure.png" pageId="6" pageNumber="165">Fig. 3AD</figureCitation>
) (vs without outgrowths and spinules in other congeners excluding
<taxonomicName id="4C5662FDFFD6FFF6557BFE6784824CAD" authorityName="Attems" authorityYear="1898" box="[540,706,452,478]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="165" phylum="Arthropoda" rank="species" species="thaumasius">
<emphasis id="B922C56CFFD6FFF6557BFE6784824CAD" box="[540,706,452,478]" italics="true" pageId="6" pageNumber="165">L. thaumasius</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5662FDFFD6FFF655B6FE6785234CAD" authority="Mikhaljova, 2021" authorityName="Mikhaljova" authorityYear="2021" box="[721,867,452,478]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="165" phylum="Arthropoda" rank="species" species="nakhodka" status="sp. nov.">
<emphasis id="B922C56CFFD6FFF655B6FE6785234CAD" box="[721,867,452,478]" italics="true" pageId="6" pageNumber="165">L. nakhodka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2117817FFD6FFF6540CFE6685864CAD" box="[875,966,452,478]" pageId="6" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C5662FDFFD6FFF65366FE6782ED4CAD" authorityName="Lokschina &amp; Golovatch" authorityYear="1977" box="[1025,1197,452,478]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="165" phylum="Arthropoda" rank="species" species="malewitschi">
<emphasis id="B922C56CFFD6FFF65366FE6782ED4CAD" box="[1025,1197,452,478]" italics="true" pageId="6" pageNumber="165">L. malewitschi</emphasis>
</taxonomicName>
); metaterga with low bosses either everywhere (
<figureCitation id="136D05FBFFD6FFF65540FE45842B4F72" box="[551,619,487,513]" captionStart="Fig" captionStartId="2.[189,232,1824,1850]" captionTargetBox="[189,1398,267,1785]" captionTargetId="figure-47@2.[186,1401,264,1789]" captionTargetPageId="2" captionText="Fig. 1. Levizonus circularis Takakuwa, 1942, ♂, from Russia (FSCB 142005). A. Habitus, dorsal view. B. Midbody rings, dorsal view. Scale bars: A = 2 mm; B = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4923496" httpUri="https://zenodo.org/record/4923496/files/figure.png" pageId="6" pageNumber="165">Fig. 1</figureCitation>
) or only laterally (vs smooth in other congeners).
</paragraph>
</subSubSection>
<subSubSection id="C34C4AF5FFD6FFF757DAFD8A84144FFC" lastPageId="7" lastPageNumber="166" pageId="6" pageNumber="165" type="materials_examined">
<paragraph id="8BE9197EFFD6FFF657DAFD8A87EA4F31" blockId="6.[189,426,552,578]" box="[189,426,552,578]" pageId="6" pageNumber="165">
<heading id="D0A1AE12FFD6FFF657DAFD8A87EA4F31" bold="true" box="[189,426,552,578]" fontSize="11" level="3" pageId="6" pageNumber="165" reason="3">
<emphasis id="B922C56CFFD6FFF657DAFD8A87EA4F31" bold="true" box="[189,426,552,578]" pageId="6" pageNumber="165">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BE9197EFFD6FFF757DAFDF5872C4CE3" blockId="6.[189,1400,599,1083]" lastBlockId="7.[189,1398,265,400]" lastPageId="7" lastPageNumber="166" pageId="6" pageNumber="165">
<materialsCitation id="3B3E1323FFD6FFF657DAFDF582314FE7" ID-GBIF-Occurrence="3128564331" collectingDate="1968-09-14" collectionCode="ZMUM" collectorName="L. S. Shvetsova" country="Russia" location="Olenevod" municipality="Lazovskiy District" pageId="6" pageNumber="165" specimenCode="ZMUM 1912" specimenCount="1" specimenCount-male="1" stateProvince="Primorskiy krai" typeStatus="holotype">
<collectingCountry id="F34159EEFFD6FFF657DAFDF587614F02" box="[189,289,599,625]" name="Russia" pageId="6" pageNumber="165">RUSSIA</collectingCountry>
<specimenCount id="9D50D2F7FFD6FFF65656FDF587184F01" box="[305,344,599,626]" pageId="6" pageNumber="165" type="male">1♂</specimenCount>
(holotype of
<taxonomicName id="4C5662FDFFD6FFF65682FDFA84834F02" authorityName="Lokschina &amp; Golovatch" authorityYear="1977" box="[485,707,599,625]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="165" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B922C56CFFD6FFF65682FDFA84834F02" box="[485,707,599,625]" italics="true" pageId="6" pageNumber="165">Levizonus variabilis</emphasis>
</taxonomicName>
;unfortunately,the holotype is devoid of gonopods);
<collectingRegion id="4992D79CFFD6FFF6539EFDF586AB4FE7" country="Russia" name="Primorskiy" pageId="6" pageNumber="165">Primorskiy krai</collectingRegion>
,
<collectingMunicipality id="6B8D8304FFD6FFF65791FDD887894FE7" box="[246,457,634,660]" pageId="6" pageNumber="165">Lazovskiy District</collectingMunicipality>
,
<location id="8E894FA5FFD6FFF656B4FDD884044FE7" LSID="urn:lsid:plazi:treatment:03FFA868FFD4FFF8555CFA6984B94953:8E894FA5FFD6FFF656B4FDD884044FE7" box="[467,580,634,660]" country="Russia" municipality="Lazovskiy District" name="Olenevod" pageId="6" pageNumber="165" stateProvince="Primorskiy krai">Olenevod</location>
;
<date id="FFE83FBEFFD6FFF65528FDD884A64FE7" box="[591,742,634,660]" pageId="6" pageNumber="165" value="1968-09-14">
<collectingDate id="EFACC656FFD6FFF65528FDD884A64FE7" box="[591,742,634,660]" pageId="6" pageNumber="165" value="1968-09-14">14 Sep. 1968</collectingDate>
</date>
;
<collectorName id="26A37CA8FFD6FFF65596FDD885DA4FE7" box="[753,922,634,660]" pageId="6" pageNumber="165">L.S. Shvetsova</collectorName>
leg.;
<specimenCode id="DBF0B105FFD6FFF654BFFDD982314FE7" box="[984,1137,634,661]" collectionCode="ZMUM" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="6" pageNumber="165">ZMUM 1912</specimenCode>
</materialsCitation>
<materialsCitation id="3B3E1323FFD6FFF653E0FDD887D04FA8" ID-GBIF-Occurrence="3128564329" collectedFrom="valley of stream, litter" collectingDate="1979-06-08" collectionCode="V" collectorName="E. V. Mikhaljova" country="Russia" location="Zvyozdochka Cordon" municipality="Lazovskiy State Nature Reserve" pageId="6" pageNumber="165" specimenCode="FSCB 11979" specimenCount="1" specimenCount-female="1" stateProvince="Primorskiy krai" typeStatus="holotype">
<specimenCount id="9D50D2F7FFD6FFF653E0FDD882F24FE6" box="[1159,1202,634,661]" pageId="6" pageNumber="165" type="female">1 ♀</specimenCount>
;
<collectingRegion id="4992D79CFFD6FFF653A7FDD883334FE7" box="[1216,1395,634,660]" country="Russia" name="Primorskiy" pageId="6" pageNumber="165">Primorskiy krai</collectingRegion>
,
<collectingMunicipality id="6B8D8304FFD6FFF657DAFD3F84684FCB" box="[189,552,669,696]" pageId="6" pageNumber="165">Lazovskiy State Nature Reserve</collectingMunicipality>
,
<location id="8E894FA5FFD6FFF65555FD3F856A4FC4" LSID="urn:lsid:plazi:treatment:03FFA868FFD4FFF8555CFA6984B94953:8E894FA5FFD6FFF65555FD3F856A4FC4" box="[562,810,669,695]" country="Russia" municipality="Lazovskiy State Nature Reserve" name="Zvyozdochka Cordon" pageId="6" pageNumber="165" stateProvince="Primorskiy krai">Zvyozdochka Cordon</location>
;
<date id="FFE83FBEFFD6FFF65452FD3C85FB4FCB" box="[821,955,670,696]" pageId="6" pageNumber="165" value="1979-06-08">
<collectingDate id="EFACC656FFD6FFF65452FD3C85FB4FCB" box="[821,955,670,696]" pageId="6" pageNumber="165" value="1979-06-08">8 Jun. 1979</collectingDate>
</date>
;
<collectorName id="26A37CA8FFD6FFF654A2FD3C823E4FC4" box="[965,1150,669,696]" pageId="6" pageNumber="165">E.V. Mikhaljova</collectorName>
leg.;
<collectedFrom id="384A5A83FFD6FFF653DCFD3F86B54FA8" pageId="6" pageNumber="165">valley of stream, litter</collectedFrom>
;
<specimenCode id="DBF0B105FFD6FFF65799FD6387D04FA8" box="[254,400,705,731]" collectionCode="FSCB" pageId="6" pageNumber="165">FSCB 11979</specimenCode>
</materialsCitation>
<materialsCitation id="3B3E1323FFD6FFF656C2FD6385234F8D" ID-GBIF-Occurrence="3128564328" collectedFrom="litter" collectingDate="1979-06-09" collectionCode="V" collectorName="E. V. Mikhaljova" country="Russia" location="Petrova Island" municipality="Lazovskiy State Nature Reserve" pageId="6" pageNumber="165" specimenCode="FSCB 21979" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Primorskiy" typeStatus="holotype">
<specimenCount id="9D50D2F7FFD6FFF656C2FD6387A64FA8" box="[421,486,705,731]" pageId="6" pageNumber="165" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D50D2F7FFD6FFF65697FD6384704FA8" box="[496,560,705,731]" pageId="6" pageNumber="165" type="female">3 ♀♀</specimenCount>
; same locality as for preceding but
<locationDeviation id="9A5B005CFFD6FFF654D0FD6387454F8D" pageId="6" pageNumber="165">
<taxonomicName id="4C5662FDFFD6FFF654D0FD6382564FA8" box="[951,1046,705,731]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="6" pageNumber="165" phylum="Tracheophyta" rank="genus">
<emphasis id="B922C56CFFD6FFF654D0FD6382564FA8" box="[951,1046,705,731]" italics="true" pageId="6" pageNumber="165">Quercus</emphasis>
</taxonomicName>
forest on the coast near Petrova Island
</locationDeviation>
;
<date id="FFE83FBEFFD6FFF65668FD4687D24F8D" box="[271,402,740,766]" pageId="6" pageNumber="165" value="1979-06-09">
<collectingDate id="EFACC656FFD6FFF65668FD4687D24F8D" box="[271,402,740,766]" pageId="6" pageNumber="165" value="1979-06-09">9 Jun. 1979</collectingDate>
</date>
;
<collectorName id="26A37CA8FFD6FFF656FBFD4684144F8D" box="[412,596,740,766]" pageId="6" pageNumber="165">E.V. Mikhaljova</collectorName>
leg.;
<collectedFrom id="384A5A83FFD6FFF655F7FD4684884F8D" box="[656,712,740,766]" pageId="6" pageNumber="165">litter</collectedFrom>
;
<specimenCode id="DBF0B105FFD6FFF655B6FD4685234F8D" box="[721,867,740,766]" collectionCode="FSCB" pageId="6" pageNumber="165">FSCB 21979</specimenCode>
</materialsCitation>
<materialsCitation id="3B3E1323FFD6FFF6541FFD46820B4E52" ID-GBIF-Occurrence="3128564303" collectedFrom="forest, litter" collectingDate="1979-06-11" collectionCode="V" collectorName="E. V. Mikhaljova" country="Russia" location="Sokolovka Cordon" municipality="Lazovskiy State Nature Reserve" pageId="6" pageNumber="165" specimenCode="FSCB 41979" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Primorskiy" typeStatus="holotype">
<specimenCount id="9D50D2F7FFD6FFF6541FFD4685FA4F8D" box="[888,954,740,766]" pageId="6" pageNumber="165" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D50D2F7FFD6FFF654A3FD4685AE4F8D" box="[964,1006,740,766]" pageId="6" pageNumber="165" type="female">1 ♀</specimenCount>
; same locality as for preceding but
<location id="8E894FA5FFD6FFF657DAFCA587D34E52" LSID="urn:lsid:plazi:treatment:03FFA868FFD4FFF8555CFA6984B94953:8E894FA5FFD6FFF657DAFCA587D34E52" box="[189,403,775,801]" country="Russia" municipality="Lazovskiy State Nature Reserve" name="Sokolovka Cordon" pageId="6" pageNumber="165" stateProvince="Primorskiy">Sokolovka Cordon</location>
;
<date id="FFE83FBEFFD6FFF656FAFCA5846E4E52" box="[413,558,775,802]" pageId="6" pageNumber="165" value="1979-06-11">
<collectingDate id="EFACC656FFD6FFF656FAFCA5846E4E52" box="[413,558,775,802]" pageId="6" pageNumber="165" value="1979-06-11">11 Jun. 1979</collectingDate>
</date>
;
<collectorName id="26A37CA8FFD6FFF6555FFCAA84B04E52" box="[568,752,775,802]" pageId="6" pageNumber="165">E.V. Mikhaljova</collectorName>
leg.;
<collectedFrom id="384A5A83FFD6FFF6544CFCA585EF4E52" box="[811,943,775,801]" pageId="6" pageNumber="165">forest, litter</collectedFrom>
;
<specimenCode id="DBF0B105FFD6FFF654DEFCA5820B4E52" box="[953,1099,775,801]" collectionCode="FSCB" pageId="6" pageNumber="165">FSCB 41979</specimenCode>
</materialsCitation>
<materialsCitation id="3B3E1323FFD6FFF65307FCA583384E37" ID-GBIF-Occurrence="3128564326" collectedFrom="valley forest, litter" collectingDate="1979-06-17" collectionCode="V" collectorName="E. V. Mikhaljova" country="Russia" location="Amerika Cordon" municipality="Lazovskiy State Nature Reserve" pageId="6" pageNumber="165" specimenCode="FSCB 71979" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Primorskiy" typeStatus="holotype">
<specimenCount id="9D50D2F7FFD6FFF65307FCA582CB4E51" box="[1120,1163,775,802]" pageId="6" pageNumber="165" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D50D2F7FFD6FFF653F1FCA582964E51" box="[1174,1238,775,802]" pageId="6" pageNumber="165" type="female">3 ♀♀</specimenCount>
; same locality as for preceding but
<location id="8E894FA5FFD6FFF656C4FC8884254E37" LSID="urn:lsid:plazi:treatment:03FFA868FFD4FFF8555CFA6984B94953:8E894FA5FFD6FFF656C4FC8884254E37" box="[419,613,810,836]" country="Russia" municipality="Lazovskiy State Nature Reserve" name="Amerika Cordon" pageId="6" pageNumber="165" stateProvince="Primorskiy">Amerika Cordon</location>
;
<date id="FFE83FBEFFD6FFF65517FC8885444E37" box="[624,772,810,837]" pageId="6" pageNumber="165" value="1979-06-17">
<collectingDate id="EFACC656FFD6FFF65517FC8885444E37" box="[624,772,810,837]" pageId="6" pageNumber="165" value="1979-06-17">17 Jun. 1979</collectingDate>
</date>
;
<collectorName id="26A37CA8FFD6FFF65477FC8985894E37" box="[784,969,810,837]" pageId="6" pageNumber="165">E.V. Mikhaljova</collectorName>
leg.;
<collectedFrom id="384A5A83FFD6FFF65360FC8882984E37" box="[1031,1240,810,836]" pageId="6" pageNumber="165">valley forest, litter</collectedFrom>
;
<specimenCode id="DBF0B105FFD6FFF65384FC8883384E37" box="[1251,1400,810,836]" collectionCode="FSCB" pageId="6" pageNumber="165">FSCB 71979</specimenCode>
</materialsCitation>
<materialsCitation id="3B3E1323FFD6FFF657A8FCEC87224EF8" ID-GBIF-Occurrence="3128564337" collectedFrom="litter" collectingDate="1979-06-20" collectionCode="V" collectorName="E. V. Mikhaljova" country="Russia" location="Pinus koraiensis forest" municipality="Lazovskiy State Nature Reserve" pageId="6" pageNumber="165" specimenCode="FSCB 141979" specimenCount="1" specimenCount-male="1" stateProvince="Primorskiy" typeStatus="holotype">
<specimenCount id="9D50D2F7FFD6FFF657A8FCEC86BD4E1B" box="[207,253,846,872]" pageId="6" pageNumber="165" type="male">1 ♂</specimenCount>
; same locality as for preceding but
<location id="8E894FA5FFD6FFF655F7FCEC85D54E14" LSID="urn:lsid:plazi:treatment:03FFA868FFD4FFF8555CFA6984B94953:8E894FA5FFD6FFF655F7FCEC85D54E14" box="[656,917,845,871]" country="Russia" municipality="Lazovskiy State Nature Reserve" name="Pinus koraiensis forest" pageId="6" pageNumber="165" stateProvince="Primorskiy">
<taxonomicName id="4C5662FDFFD6FFF655F7FCEC850B4E14" box="[656,843,845,871]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="165" phylum="Tracheophyta" rank="species" species="koraiensis">
<emphasis id="B922C56CFFD6FFF655F7FCEC850B4E14" box="[656,843,845,871]" italics="true" pageId="6" pageNumber="165">Pinus koraiensis</emphasis>
</taxonomicName>
forest
</location>
;
<date id="FFE83FBEFFD6FFF654C6FCEC82764E1B" box="[929,1078,846,872]" pageId="6" pageNumber="165" value="1979-06-20">
<collectingDate id="EFACC656FFD6FFF654C6FCEC82764E1B" box="[929,1078,846,872]" pageId="6" pageNumber="165" value="1979-06-20">20 Jun. 1979</collectingDate>
</date>
,
<collectorName id="26A37CA8FFD6FFF65325FCEC82BB4E14" box="[1090,1275,845,872]" pageId="6" pageNumber="165">E.V. Mikhaljova</collectorName>
leg.;
<collectedFrom id="384A5A83FFD6FFF6525CFCEF83334E14" box="[1339,1395,845,871]" pageId="6" pageNumber="165">litter</collectedFrom>
;
<specimenCode id="DBF0B105FFD6FFF657DAFCD387224EF8" box="[189,354,881,907]" collectionCode="FSCB" pageId="6" pageNumber="165">FSCB 141979</specimenCode>
</materialsCitation>
<materialsCitation id="3B3E1323FFD6FFF656E6FCD382314EDC" ID-GBIF-Occurrence="3128564316" collectedFrom="litter" collectingDate="2005-09-04" collectingDateMax="2005-09-05" collectingDateMin="2005-09-04" collectingMethod="pitfall traps" collectionCode="V" collectorName="S. Yu. Storozhenko &amp; V. S. Sidorenko &amp; S. K. Kholin &amp; Yu. N. Sundukov" country="Russia" location="glade" municipality="Lazovskiy State Nature Reserve" pageId="6" pageNumber="165" specimenCode="FSCB 162005" specimenCount="1" specimenCount-male="1" stateProvince="Primorskiy" typeStatus="holotype">
<specimenCount id="9D50D2F7FFD6FFF656E6FCD387F24EF8" box="[385,434,881,907]" pageId="6" pageNumber="165" type="male">1 ♂</specimenCount>
; same locality as for preceding but in a
<location id="8E894FA5FFD6FFF654F7FCD385904EF8" LSID="urn:lsid:plazi:treatment:03FFA868FFD4FFF8555CFA6984B94953:8E894FA5FFD6FFF654F7FCD385904EF8" box="[912,976,881,907]" country="Russia" municipality="Lazovskiy State Nature Reserve" name="glade" pageId="6" pageNumber="165" stateProvince="Primorskiy">glade</location>
;
<date id="FFE83FBEFFD6FFF654B8FCD282CB4EF8" box="[991,1163,880,907]" pageId="6" pageNumber="165" value="2005-09-04" valueMax="2005-09-05" valueMin="2005-09-04">
<collectingDate id="EFACC656FFD6FFF654B8FCD282CB4EF8" box="[991,1163,880,907]" pageId="6" pageNumber="165" value="2005-09-04" valueMax="2005-09-05" valueMin="2005-09-04">45 Sep. 2005</collectingDate>
</date>
;
<collectorName id="26A37CA8FFD6FFF653FDFCD383334EF8" box="[1178,1395,881,907]" pageId="6" pageNumber="165">S.Yu. Storozhenko</collectorName>
,
<collectorName id="26A37CA8FFD6FFF657DAFC36872B4EDD" box="[189,363,916,942]" pageId="6" pageNumber="165">V.S. Sidorenko</collectorName>
,
<collectorName id="26A37CA8FFD6FFF6561FFC3684424EDD" box="[376,514,916,942]" pageId="6" pageNumber="165">S.K. Kholin</collectorName>
and
<collectorName id="26A37CA8FFD6FFF6555CFC3684B94EDD" box="[571,761,916,942]" pageId="6" pageNumber="165">Yu.N. Sundukov</collectorName>
leg.;
<collectingMethod id="52176169FFD6FFF6545CFC3685804EDD" box="[827,960,916,942]" pageId="6" pageNumber="165">pitfall traps</collectingMethod>
;
<specimenCode id="DBF0B105FFD6FFF654AAFC3682314EDC" box="[973,1137,916,943]" collectionCode="FSCB" pageId="6" pageNumber="165">FSCB 162005</specimenCode>
</materialsCitation>
<materialsCitation id="3B3E1323FFD6FFF653EBFC3685C24E86" ID-GBIF-Occurrence="3128564308" collectedFrom="grass meadow" collectingDate="2005-09-02" collectingDateMax="2005-09-05" collectingDateMin="2005-09-04" collectingMethod="pitfall traps" collectionCode="V" collectorName="S. Yu. Storozhenko &amp; V. S. Sidorenko &amp; S. K. Kholin &amp; Yu. N. Sundukov" country="Russia" location="Korpad Cordon" municipality="Lazovskiy State Nature Reserve" pageId="6" pageNumber="165" specimenCode="FSCB 142005" specimenCount="2" specimenCount-male="2" stateProvince="Primorskiy" typeStatus="holotype">
<specimenCount id="9D50D2F7FFD6FFF653EBFC3682924EDD" box="[1164,1234,916,942]" pageId="6" pageNumber="165" type="male">2 ♂♂</specimenCount>
; same locality as for preceding but
<location id="8E894FA5FFD6FFF656D6FC1584284EA2" LSID="urn:lsid:plazi:treatment:03FFA868FFD4FFF8555CFA6984B94953:8E894FA5FFD6FFF656D6FC1584284EA2" box="[433,616,951,977]" country="Russia" municipality="Lazovskiy State Nature Reserve" name="Korpad Cordon" pageId="6" pageNumber="165" stateProvince="Primorskiy">Korpad Cordon</location>
;
<date id="FFE83FBEFFD6FFF65510FC1585474EA1" box="[631,775,951,978]" pageId="6" pageNumber="165" value="2005-09-02">
<collectingDate id="EFACC656FFD6FFF65510FC1585474EA1" box="[631,775,951,978]" pageId="6" pageNumber="165" value="2005-09-02">2 Sep. 2005</collectingDate>
</date>
;
<collectorName id="26A37CA8FFD6FFF65472FC1585AE4EA2" box="[789,1006,951,977]" pageId="6" pageNumber="165">S.Yu. Storozhenko</collectorName>
,
<collectorName id="26A37CA8FFD6FFF6549CFC1582EB4EA2" box="[1019,1195,951,977]" pageId="6" pageNumber="165">V.S. Sidorenko</collectorName>
,
<collectorName id="26A37CA8FFD6FFF653DFFC1583044EA2" box="[1208,1348,951,977]" pageId="6" pageNumber="165">S.K. Kholin</collectorName>
and
<collectorName id="26A37CA8FFD6FFF657DAFC7987374E87" box="[189,375,986,1013]" pageId="6" pageNumber="165">Yu.N. Sundukov</collectorName>
leg.;
<collectedFrom id="384A5A83FFD6FFF656D7FC7984104E87" box="[432,592,986,1013]" pageId="6" pageNumber="165">grass meadow</collectedFrom>
,
<collectingMethod id="52176169FFD6FFF6553EFC78849A4E86" box="[601,730,986,1013]" pageId="6" pageNumber="165">pitfall traps</collectingMethod>
;
<specimenCode id="DBF0B105FFD6FFF65585FC7885C24E86" box="[738,898,986,1013]" collectionCode="FSCB" pageId="6" pageNumber="165">FSCB 142005</specimenCode>
</materialsCitation>
<materialsCitation id="3B3E1323FFD6FFF654F2FC788294496B" ID-GBIF-Occurrence="3128564336" collectedFrom="forest" collectingDate="1973-08-13" collectionCode="FSCB" collectorName="G. F. Kurcheva" country="Russia" location="Verchneussuriiskiy Research Station" municipality="Chuguevskiy District" pageId="6" pageNumber="165" specimenCode="FSCB 11973" specimenCount="1" specimenCount-male="1" stateProvince="Primorskiy krai" typeStatus="holotype">
<specimenCount id="9D50D2F7FFD6FFF654F2FC7885804E86" box="[917,960,986,1013]" pageId="6" pageNumber="165" type="male">1 ♂</specimenCount>
;
<collectingRegion id="4992D79CFFD6FFF654ADFC78823A4E87" box="[970,1146,986,1012]" country="Russia" name="Primorskiy" pageId="6" pageNumber="165">Primorskiy krai</collectingRegion>
,
<collectingMunicipality id="6B8D8304FFD6FFF653E4FC7883334E87" box="[1155,1395,986,1012]" pageId="6" pageNumber="165">Chuguevskiy District</collectingMunicipality>
,
<location id="8E894FA5FFD6FFF657DAFC5F841B4964" LSID="urn:lsid:plazi:treatment:03FFA868FFD4FFF8555CFA6984B94953:8E894FA5FFD6FFF657DAFC5F841B4964" box="[189,603,1021,1047]" country="Russia" municipality="Chuguevskiy District" name="Verchneussuriiskiy Research Station" pageId="6" pageNumber="165" stateProvince="Primorskiy krai">Verchneussuriiskiy Research Station</location>
;
<date id="FFE83FBEFFD6FFF65501FC5C8543496B" box="[614,771,1022,1048]" pageId="6" pageNumber="165" value="1973-08-13">
<collectingDate id="EFACC656FFD6FFF65501FC5C8543496B" box="[614,771,1022,1048]" pageId="6" pageNumber="165" value="1973-08-13">13 Aug. 1973</collectingDate>
</date>
;
<collectorName id="26A37CA8FFD6FFF65468FC5C85F44964" box="[783,948,1021,1048]" pageId="6" pageNumber="165">G.F. Kurcheva</collectorName>
leg.;
<collectedFrom id="384A5A83FFD6FFF65494FC5F82764964" box="[1011,1078,1021,1047]" pageId="6" pageNumber="165">forest</collectedFrom>
;
<specimenCode id="DBF0B105FFD6FFF65326FC5C8294496B" box="[1089,1236,1022,1048]" collectionCode="FSCB" pageId="6" pageNumber="165">FSCB 11973</specimenCode>
</materialsCitation>
<materialsCitation id="3B3E1323FFD6FFF6538BFC5C82C04948" ID-GBIF-Occurrence="3128564322" collectedFrom="forest" collectingDate="2008-05-21" collectingDateMax="2008-05-25" collectingDateMin="2008-05-21" collectionCode="FSCB" collectorName="S. A. Shabalin" country="Russia" location="Verchneussuriiskiy Research Station" municipality="Chuguevskiy District" pageId="6" pageNumber="165" specimenCode="FSCB 212008" specimenCount="2" specimenCount-male="2" stateProvince="Primorskiy" typeStatus="holotype">
<specimenCount id="9D50D2F7FFD6FFF6538BFC5C8370496B" box="[1260,1328,1022,1048]" pageId="6" pageNumber="165" type="male">2 ♂♂</specimenCount>
; same locality as for preceding;
<date id="FFE83FBEFFD6FFF656B8FB8284E64948" box="[479,678,1056,1083]" pageId="6" pageNumber="165" value="2008-05-21" valueMax="2008-05-25" valueMin="2008-05-21">
<collectingDate id="EFACC656FFD6FFF656B8FB8284E64948" box="[479,678,1056,1083]" pageId="6" pageNumber="165" value="2008-05-21" valueMax="2008-05-25" valueMin="2008-05-21">2125 May 2008</collectingDate>
</date>
;
<collectorName id="26A37CA8FFD6FFF655D5FB83850F4948" box="[690,847,1057,1083]" pageId="6" pageNumber="165">S.A. Shabalin</collectorName>
leg.;
<collectedFrom id="384A5A83FFD6FFF654E8FB8385924948" box="[911,978,1057,1083]" pageId="6" pageNumber="165">forest</collectedFrom>
;
<specimenCode id="DBF0B105FFD6FFF654B9FB8382C04948" box="[990,1152,1057,1083]" collectionCode="FSCB" pageId="6" pageNumber="165">FSCB 212008</specimenCode>
</materialsCitation>
<materialsCitation id="3B3E1323FFD6FFF753FCFB83827C4C50" ID-GBIF-Occurrence="3128564310" collectedFrom="forest" collectingDate="2008-06-27" collectingDateMax="2008-06-30" collectingDateMin="2008-06-27" collectionCode="FSCB" collectorName="S. A. Shabalin" country="Russia" lastPageId="7" lastPageNumber="166" location="Verchneussuriiskiy Research Station" municipality="Chuguevskiy District" pageId="6" pageNumber="165" specimenCode="FSCB 222008" specimenCount="1" specimenCount-male="1" stateProvince="Primorskiy" typeStatus="holotype">
<specimenCount id="9D50D2F7FFD6FFF653FCFB83828C4948" box="[1179,1228,1057,1083]" pageId="6" pageNumber="165" type="male">1 ♂</specimenCount>
; same locality as for preceding;
<date id="FFE83FBEFFD7FFF756EDFEAB84114C50" box="[394,593,265,291]" pageId="7" pageNumber="166" value="2008-06-27" valueMax="2008-06-30" valueMin="2008-06-27">
<collectingDate id="EFACC656FFD7FFF756EDFEAB84114C50" box="[394,593,265,291]" pageId="7" pageNumber="166" value="2008-06-27" valueMax="2008-06-30" valueMin="2008-06-27">2730 Jun. 2008</collectingDate>
</date>
;
<collectorName id="26A37CA8FFD7FFF7553AFEAB85404C50" box="[605,768,265,291]" pageId="7" pageNumber="166">S.A. Shabalin</collectorName>
leg.;
<collectedFrom id="384A5A83FFD7FFF75425FEAB85C94C50" box="[834,905,265,291]" pageId="7" pageNumber="166">forest</collectedFrom>
;
<specimenCode id="DBF0B105FFD7FFF754F3FEAB827C4C50" box="[916,1084,265,291]" collectionCode="FSCB" pageId="7" pageNumber="166">FSCB 222008</specimenCode>
</materialsCitation>
<materialsCitation id="3B3E1323FFD7FFF75332FEAB85AB4C3B" ID-GBIF-Occurrence="3128564301" collectedFrom="forest" collectingDate="2008-08-21" collectingDateMax="2008-08-24" collectingDateMin="2008-08-21" collectionCode="FSCB" collectorName="S. A. Shabalin" country="Russia" location="Verchneussuriiskiy Research Station" municipality="Chuguevskiy District" pageId="7" pageNumber="166" specimenCode="FSCB 232008" specimenCount="1" specimenCount-male="1" stateProvince="Primorskiy" typeStatus="holotype">
<specimenCount id="9D50D2F7FFD7FFF75332FEAB82C44C50" box="[1109,1156,265,291]" pageId="7" pageNumber="166" type="male">1 ♂</specimenCount>
; same locality as for preceding;
<date id="FFE83FBEFFD7FFF75658FE8C844A4C3B" box="[319,522,302,328]" pageId="7" pageNumber="166" value="2008-08-21" valueMax="2008-08-24" valueMin="2008-08-21">
<collectingDate id="EFACC656FFD7FFF75658FE8C844A4C3B" box="[319,522,302,328]" pageId="7" pageNumber="166" value="2008-08-21" valueMax="2008-08-24" valueMin="2008-08-21">2124 Aug. 2008</collectingDate>
</date>
;
<collectorName id="26A37CA8FFD7FFF75572FE8C84F64C34" box="[533,694,301,328]" pageId="7" pageNumber="166">S.A. Shabalin</collectorName>
leg.;
<collectedFrom id="384A5A83FFD7FFF75593FE8F857B4C34" box="[756,827,301,327]" pageId="7" pageNumber="166">forest</collectedFrom>
;
<specimenCode id="DBF0B105FFD7FFF75422FE8C85AB4C3B" box="[837,1003,302,328]" collectionCode="FSCB" pageId="7" pageNumber="166">FSCB 232008</specimenCode>
</materialsCitation>
<materialsCitation id="3B3E1323FFD7FFF75366FE8C87284CE3" ID-GBIF-Occurrence="3128564312" collectedFrom="dark coniferous forest" collectingDate="2012-08" collectionCode="FSCB" collectorName="G. N. Ganin" country="Russia" elevation="1000" location="Shpilka" municipality="Lazovskiy mountain range" pageId="7" pageNumber="166" specimenCode="FSCB 292012" specimenCount="1" specimenCount-male="1" stateProvince="Primorskiy krai" typeStatus="holotype">
<specimenCount id="9D50D2F7FFD7FFF75366FE8C826E4C3B" box="[1025,1070,302,328]" pageId="7" pageNumber="166" type="male">1 ♂</specimenCount>
;
<collectingRegion id="4992D79CFFD7FFF7535EFE8F82B24C34" box="[1081,1266,301,327]" country="Russia" name="Primorskiy" pageId="7" pageNumber="166">Primorskiy krai</collectingRegion>
,
<collectingMunicipality id="6B8D8304FFD7FFF7539BFE8F87384C1F" pageId="7" pageNumber="166">Lazovskiy mountain range</collectingMunicipality>
,
<location id="8E894FA5FFD7FFF756E1FEF087A04C1F" LSID="urn:lsid:plazi:treatment:03FFA868FFD4FFF8555CFA6984B94953:8E894FA5FFD7FFF756E1FEF087A04C1F" box="[390,480,338,364]" country="Russia" municipality="Lazovskiy mountain range" name="Shpilka" pageId="7" pageNumber="166" stateProvince="Primorskiy krai">Shpilka</location>
pass, near
<elevation id="007BFE4DFFD7FFF75501FEF085414C1F" box="[614,769,338,364]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="166" unit="m" value="1000.0">
<quantity id="4CAEB49BFFD7FFF75501FEF084804C1F" box="[614,704,338,364]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="166" unit="m" value="1000.0">1000 m</quantity>
a.s.l.
</elevation>
;
<date id="FFE83FBEFFD7FFF75469FEF085CF4C1F" box="[782,911,338,364]" pageId="7" pageNumber="166" value="2012-08">
<collectingDate id="EFACC656FFD7FFF75469FEF085CF4C1F" box="[782,911,338,364]" pageId="7" pageNumber="166" value="2012-08">Aug. 2012</collectingDate>
</date>
;
<collectorName id="26A37CA8FFD7FFF754FBFEF082664C1F" box="[924,1062,338,364]" pageId="7" pageNumber="166">G.N. Ganin</collectorName>
leg.;
<collectedFrom id="384A5A83FFD7FFF7530DFEF083324C1F" box="[1130,1394,338,364]" pageId="7" pageNumber="166">dark coniferous forest</collectedFrom>
;
<specimenCode id="DBF0B105FFD7FFF757DAFED487284CE3" box="[189,360,374,400]" collectionCode="FSCB" pageId="7" pageNumber="166">FSCB 292012</specimenCode>
</materialsCitation>
.
</paragraph>
<caption id="DF2949F6FFD6FFF657DAF95F834D4AF2" ID-DOI="http://doi.org/10.5281/zenodo.4923500" ID-Zenodo-Dep="4923500" httpUri="https://zenodo.org/record/4923500/files/figure.png" pageId="6" pageNumber="165" startId="6.[189,232,1789,1815]" targetBox="[203,1359,1160,1746]" targetPageId="6">
<paragraph id="8BE9197EFFD6FFF657DAF95F834D4AF2" blockId="6.[189,1399,1789,1922]" pageId="6" pageNumber="165">
<emphasis id="B922C56CFFD6FFF657DAF95F87474A64" bold="true" box="[189,263,1789,1815]" pageId="6" pageNumber="165">Fig. 3.</emphasis>
<taxonomicName id="4C5662FDFFD6FFF6566CF95C84FA4A6B" authority="Takakuwa, 1942" authorityName="Takakuwa" authorityYear="1942" box="[267,698,1789,1816]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="165" phylum="Arthropoda" rank="species" species="circularis">
<emphasis id="B922C56CFFD6FFF6566CF95C87B64A64" box="[267,502,1789,1815]" italics="true" pageId="6" pageNumber="165">Levizonus circularis</emphasis>
<bibRefCitation id="EFC7648FFFD6FFF6569DF95F84FA4A6B" author="Takakuwa Y." box="[506,698,1789,1816]" pageId="6" pageNumber="165" pagination="39 - 47" refId="ref10993" refString="Takakuwa Y. 1942. Zur Kenntnis der Japanischen Diplopoden. Annotationes Zoologicae Japonenses 21: 39 - 47." type="journal article" year="1942">Takakuwa, 1942</bibRefCitation>
</taxonomicName>
, ♂♂, from Russia (AI) and North Korea(J).
<emphasis id="B922C56CFFD6FFF653DBF95F82B54A64" bold="true" box="[1212,1269,1789,1815]" pageId="6" pageNumber="165">AD</emphasis>
. Variations in shape of distal part of gonopod telopodite, ventral view.
<emphasis id="B922C56CFFD6FFF654ECF88385FC4A48" bold="true" box="[907,956,1825,1851]" pageId="6" pageNumber="165">EJ</emphasis>
. Variations in shape of distal part of gonopod telopodite, lateral view. Scale bars: AD, J = 0.25 mm; EG = 0.2 mm; HI = 0.1 mm. AH: after
<bibRefCitation id="EFC7648FFFD6FFF6579DF8C587944AF2" author="Mikhaljova E. V." box="[250,468,1895,1921]" pageId="6" pageNumber="165" pagination="62 - 67" refId="ref9873" refString="Mikhaljova E. V. 1981 a. The millipede genus Levizonus (Diplopoda, Xystodesmidae) from the USSR Far East. Byulleten Moskovskogo obshchestva ispytateley prirodi, Biology 86 (3): 62 - 67. [In Russian, English summary.]" type="journal article" year="1981">Mikhaljova 1981a</bibRefCitation>
; I: after
<bibRefCitation id="EFC7648FFFD6FFF65551F8C585D54AF2" author="Lokschina I. E. &amp; Golovatch S. I." box="[566,917,1895,1921]" pageId="6" pageNumber="165" pagination="73 - 78" refId="ref9709" refString="Lokschina I. E. &amp; Golovatch S. I. 1977. The new myriapods (Diplopoda) from the USSR. Byulleten Moskovskogo obshchestva ispytateley prirodi, Biology 82 (1): 73 - 78. [In Russian.]" type="journal article" year="1977">Lokschina &amp; Golovatch 1977</bibRefCitation>
; J: after
<bibRefCitation id="EFC7648FFFD6FFF6549DF8C583484AF2" author="Mikhaljova E. V. &amp; Golovatch S. I. &amp; Wytwer J." box="[1018,1288,1895,1921]" pageId="6" pageNumber="165" pagination="109 - 122" refId="ref10667" refString="Mikhaljova E. V., Golovatch S. I. &amp; Wytwer J. 2000. On some new or poorly-known millipedes (Diplopoda) from North Korea. Fragmenta Faunistica 43 (10): 109 - 122. https: // doi. org / 10.3161 / 00159301 FF 2000.43.10.109" type="journal article" year="2000">
Mikhaljova
<emphasis id="B922C56CFFD6FFF653EFF8CA82834AF2" box="[1160,1219,1895,1921]" italics="true" pageId="6" pageNumber="165">et al.</emphasis>
2000
</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="8BE9197EFFD7FFF757DAFE1B84CF4C84" blockId="7.[189,1399,441,504]" pageId="7" pageNumber="166">
<materialsCitation id="3B3E1323FFD7FFF757DAFE1B84D84CA0" ID-GBIF-Occurrence="3128564313" box="[189,664,441,468]" country="China" location="Jilin Province" pageId="7" pageNumber="166" specimenCount="1" stateProvince="Jilin Province">
NORTH-EAST
<collectingCountry id="F34159EEFFD7FFF7561FFE1B87934CA0" box="[376,467,441,467]" name="China" pageId="7" pageNumber="166">CHINA</collectingCountry>
<emphasis id="B922C56CFFD7FFF7568AFE1B84D84CA0" bold="true" box="[493,664,441,467]" pageId="7" pageNumber="166">
<collectingRegion id="4992D79CFFD7FFF7568AFE1B84D84CA0" box="[493,664,441,467]" country="China" name="Jilin" pageId="7" pageNumber="166">Jilin Province</collectingRegion>
</emphasis>
</materialsCitation>
<materialsCitation id="3B3E1323FFD7FFF755CAFE1B84CB4C84" ID-GBIF-Occurrence="3128564315" collectingDate="2019-08-30" collectionCode="FSCB" collectorName="L. A. Prozorova &amp; Guoyi Zhang" country="China" location="Baishan town" pageId="7" pageNumber="166" specimenCode="FSCB 12019Ch" specimenCount="1" specimenCount-male="1" stateProvince="Jilin">
<specimenCount id="9D50D2F7FFD7FFF755CAFE1B849B4CA7" box="[685,731,441,468]" pageId="7" pageNumber="166" type="male">1 ♂</specimenCount>
;
<locationDeviation id="9A5B005CFFD7FFF75582FE1B82504CA7" box="[741,1040,441,468]" pageId="7" pageNumber="166">environs of Baishan town</locationDeviation>
;
<date id="FFE83FBEFFD7FFF7537DFE1B82F84CA0" box="[1050,1208,441,467]" pageId="7" pageNumber="166" value="2019-08-30">
<collectingDate id="EFACC656FFD7FFF7537DFE1B82F84CA0" box="[1050,1208,441,467]" pageId="7" pageNumber="166" value="2019-08-30">30 Aug. 2019</collectingDate>
</date>
;
<collectorName id="26A37CA8FFD7FFF753A4FE1B83374CA7" box="[1219,1399,441,468]" pageId="7" pageNumber="166">L.A. Prozorova</collectorName>
and
<collectorName id="26A37CA8FFD7FFF75788FE7F87CA4C84" box="[239,394,477,503]" pageId="7" pageNumber="166">Guoyi Zhang</collectorName>
leg.;
<specimenCode id="DBF0B105FFD7FFF756ABFE7C84CB4C84" box="[460,651,477,504]" pageId="7" pageNumber="166">FSCB 12019Ch</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE9197EFFD7FFF757DAFD8282484F49" blockId="7.[189,1032,544,570]" box="[189,1032,544,570]" pageId="7" pageNumber="166">
<emphasis id="B922C56CFFD7FFF757DAFD82878D4F49" bold="true" box="[189,461,544,570]" pageId="7" pageNumber="166">Material re-examined</emphasis>
(specimen published by
<bibRefCitation id="EFC7648FFFD7FFF75594FD8285BF4F49" author="Mikhaljova E. V. &amp; Golovatch S. I. &amp; Wytwer J." box="[755,1023,544,570]" pageId="7" pageNumber="166" pagination="109 - 122" refId="ref10667" refString="Mikhaljova E. V., Golovatch S. I. &amp; Wytwer J. 2000. On some new or poorly-known millipedes (Diplopoda) from North Korea. Fragmenta Faunistica 43 (10): 109 - 122. https: // doi. org / 10.3161 / 00159301 FF 2000.43.10.109" type="journal article" year="2000">
Mikhaljova
<emphasis id="B922C56CFFD7FFF754E6FD8385FC4F49" box="[897,956,544,570]" italics="true" pageId="7" pageNumber="166">et al.</emphasis>
2000
</bibRefCitation>
)
</paragraph>
<paragraph id="8BE9197EFFD7FFF757DAFDF284144FFC" blockId="7.[189,1398,592,655]" pageId="7" pageNumber="166">
<materialsCitation id="3B3E1323FFD7FFF757DAFDF2840F4FFC" ID-GBIF-Occurrence="3128564307" collectingDate="1965-06-09" collectionCode="FSCB" collectorName="M. Mroczkowski &amp; A. Riedel" country="North Korea" location="Lake Cangdzin-ho" pageId="7" pageNumber="166" specimenCode="FSCB 11965K" specimenCount="1" specimenCount-male="1" stateProvince="Hamgjong-namdo Province">
<collectingCountry id="F34159EEFFD7FFF757DAFDF287C74F19" box="[189,391,592,618]" name="North Korea" pageId="7" pageNumber="166">NORTH KOREA</collectingCountry>
<emphasis id="B922C56CFFD7FFF756F8FDF284B64F19" bold="true" box="[415,758,592,618]" pageId="7" pageNumber="166">
<collectingRegion id="4992D79CFFD7FFF756F8FDF284B64F19" box="[415,758,592,618]" pageId="7" pageNumber="166">Hamgjŏng-namdo Province</collectingRegion>
</emphasis>
<specimenCount id="9D50D2F7FFD7FFF7546EFDF285754F18" box="[777,821,592,619]" pageId="7" pageNumber="166" type="male">1 ♂</specimenCount>
;
<location id="8E894FA5FFD7FFF75459FDF282584F1E" LSID="urn:lsid:plazi:treatment:03FFA868FFD4FFF8555CFA6984B94953:8E894FA5FFD7FFF75459FDF282584F1E" box="[830,1048,592,621]" country="North Korea" name="Lake Cangdzin-ho" pageId="7" pageNumber="166" stateProvince="Hamgjong-namdo Province">Lake Čangdžin-ho</location>
;
<date id="FFE83FBEFFD7FFF75346FDF282E64F18" box="[1057,1190,592,619]" pageId="7" pageNumber="166" value="1965-06-09">
<collectingDate id="EFACC656FFD7FFF75346FDF282E64F18" box="[1057,1190,592,619]" pageId="7" pageNumber="166" value="1965-06-09">9 Jun. 1965</collectingDate>
</date>
;
<collectorName id="26A37CA8FFD7FFF753C8FDF383364F19" box="[1199,1398,592,619]" pageId="7" pageNumber="166">M. Mroczkowski</collectorName>
and
<collectorName id="26A37CA8FFD7FFF7578AFDD7871E4FFD" box="[237,350,628,655]" pageId="7" pageNumber="166">A. Riedel</collectorName>
leg.;
<specimenCode id="DBF0B105FFD7FFF756F8FDD7840F4FFC" box="[415,591,628,655]" pageId="7" pageNumber="166">FSCB 11965K</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C4AF5FFD7FFF757DAFD1587874A05" pageId="7" pageNumber="166" type="description">
<paragraph id="8BE9197EFFD7FFF757DAFD15870E4FA2" blockId="7.[189,334,695,721]" box="[189,334,695,721]" pageId="7" pageNumber="166">
<heading id="D0A1AE12FFD7FFF757DAFD15870E4FA2" bold="true" box="[189,334,695,721]" fontSize="11" level="3" pageId="7" pageNumber="166" reason="3">
<emphasis id="B922C56CFFD7FFF757DAFD15870E4FA2" bold="true" box="[189,334,695,721]" pageId="7" pageNumber="166">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BE9197EFFD7FFF7578DFD45876A4E72" blockId="7.[189,709,743,806]" box="[234,298,743,769]" pageId="7" pageNumber="166">
<emphasis id="B922C56CFFD7FFF7578DFD45876A4E72" bold="true" box="[234,298,743,769]" pageId="7" pageNumber="166">Male</emphasis>
</paragraph>
<paragraph id="8BE9197EFFD7FFF757DAFCA984854E56" blockId="7.[189,709,743,806]" box="[189,709,779,806]" pageId="7" pageNumber="166">
SIZE. Body
<quantity id="4CAEB49BFFD7FFF75625FCA987FA4E55" box="[322,442,779,806]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="7" pageNumber="166" unit="mm" value="27.5" valueMax="30.0" valueMin="25.0">2530 mm</quantity>
long,
<quantity id="4CAEB49BFFD7FFF7569AFCA984C24E55" box="[509,642,779,806]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.8" metricValueMin="3.3" pageId="7" pageNumber="166" unit="mm" value="3.55" valueMax="3.8" valueMin="3.3">3.33.8 mm</quantity>
wide.
</paragraph>
<paragraph id="8BE9197EFFD7FFF757DAFCEC83374E1B" blockId="7.[189,1399,846,873]" box="[189,1399,846,873]" pageId="7" pageNumber="166">COLOUR. Shining whitish.Tergites often patterned with lateral marbled brown spots and transverse brown stripes.</paragraph>
<paragraph id="8BE9197EFFD7FFF757DAFC33848B4E87" blockId="7.[189,1400,913,1012]" pageId="7" pageNumber="166">HEAD. Epicranial suture not quite reaching the level of antennal sockets. Lateral excavation of head relatively deep for accommodation of antennae. Antennae rather slender, varying from short (reaching to front edge of ring 2) to long (reaching to front edge of ring 4).</paragraph>
<paragraph id="8BE9197EFFD7FFF757DBFBBF833849D7" blockId="7.[188,1400,1052,1188]" pageId="7" pageNumber="166">
BODY. Metaterga with bosses (
<figureCitation id="136D05FBFFD7FFF75573FBBE84164945" box="[532,598,1052,1078]" captionStart="Fig" captionStartId="2.[189,232,1824,1850]" captionTargetBox="[189,1398,267,1785]" captionTargetId="figure-47@2.[186,1401,264,1789]" captionTargetPageId="2" captionText="Fig. 1. Levizonus circularis Takakuwa, 1942, ♂, from Russia (FSCB 142005). A. Habitus, dorsal view. B. Midbody rings, dorsal view. Scale bars: A = 2 mm; B = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4923496" httpUri="https://zenodo.org/record/4923496/files/figure.png" pageId="7" pageNumber="166">Fig. 1</figureCitation>
) increasingly evident from ring 5 posteriorly, especially so in males in contrast to females and juveniles. The number and location of the bosses varies from a few (or one) only in lateral parts above peritremata to relatively numerous as transverse rows throughout each metatergite (
<figureCitation id="136D05FBFFD7FFF75273FBC78328490C" box="[1300,1384,1125,1151]" captionStart="Fig" captionStartId="2.[189,232,1824,1850]" captionTargetBox="[189,1398,267,1785]" captionTargetId="figure-47@2.[186,1401,264,1789]" captionTargetPageId="2" captionText="Fig. 1. Levizonus circularis Takakuwa, 1942, ♂, from Russia (FSCB 142005). A. Habitus, dorsal view. B. Midbody rings, dorsal view. Scale bars: A = 2 mm; B = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4923496" httpUri="https://zenodo.org/record/4923496/files/figure.png" pageId="7" pageNumber="166">Fig. 1B</figureCitation>
). Paraterga rounded laterally; starting from ring 1213, they have rounded, though never clear-cut, caudal corners.
</paragraph>
<paragraph id="8BE9197EFFD7FFF757DBFB6F85814995" blockId="7.[188,961,1228,1255]" box="[188,961,1228,1255]" pageId="7" pageNumber="166">TELSON. Caudal dorsal projection with long sparse setae, blunt apically.</paragraph>
<paragraph id="8BE9197EFFD7FFF757DBFAB2876E4800" blockId="7.[188,1399,1295,1395]" pageId="7" pageNumber="166">LEGS. Each coxa and prefemur of postgonopodal legs distoventrally with a short conical knob becoming spineshaped in coxa toward telson. Coxa of leg 2 with a large setigerous process curved forward and terminating with gonopore.</paragraph>
<paragraph id="8BE9197EFFD7FFF757DAFA3984374B34" blockId="7.[189,1400,1435,1607]" pageId="7" pageNumber="166">
GONOPODS. Distal part of coxite with three strong setae laterally (two) (
<figureCitation id="136D05FBFFD7FFF754B4FA39826B48C6" box="[979,1067,1435,1461]" captionStart="Fig" captionStartId="5.[189,232,1773,1799]" captionTargetBox="[189,1397,265,1735]" captionTargetId="figure-93@5.[186,1401,262,1738]" captionTargetPageId="5" captionText="Fig. 2. Levizonus circularis Takakuwa, 1942, ♂♂, ♀, from China (FSCB 12019Ch) (AB) and Russia (FSCB 232008, 212008) (CG). A. Gonopod, lateral view, longitudinal striae are not visible. B. Gonopod apex, lateral view. C, F. Gonopod apex, ventral view. D. Gonopod, lateral view, coxite partially destroyed. E. Middle part of gonopod, mesal view. G. Vulva, mesal view. Scale bars: A, E = 200 μm; D, G = 100 μm; BC, F = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.4923498" httpUri="https://zenodo.org/record/4923498/files/figure.png" pageId="7" pageNumber="166">Fig. 2A</figureCitation>
) and mesally (one) (
<figureCitation id="136D05FBFFD7FFF75274FA39832B48C6" box="[1299,1387,1435,1461]" captionStart="Fig" captionStartId="5.[189,232,1773,1799]" captionTargetBox="[189,1397,265,1735]" captionTargetId="figure-93@5.[186,1401,262,1738]" captionTargetPageId="5" captionText="Fig. 2. Levizonus circularis Takakuwa, 1942, ♂♂, ♀, from China (FSCB 12019Ch) (AB) and Russia (FSCB 232008, 212008) (CG). A. Gonopod, lateral view, longitudinal striae are not visible. B. Gonopod apex, lateral view. C, F. Gonopod apex, ventral view. D. Gonopod, lateral view, coxite partially destroyed. E. Middle part of gonopod, mesal view. G. Vulva, mesal view. Scale bars: A, E = 200 μm; D, G = 100 μm; BC, F = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.4923498" httpUri="https://zenodo.org/record/4923498/files/figure.png" pageId="7" pageNumber="166">Fig. 2E</figureCitation>
). Telopodite laterally with longitudinal striae along almost its entire length. Distal part of telopodite with an external dentiform process (= SL) of different length. SL more (
<figureCitation id="136D05FBFFD7FFF754E1FA46859D488D" box="[902,989,1508,1534]" captionStart="Fig" captionStartId="5.[189,232,1773,1799]" captionTargetBox="[189,1397,265,1735]" captionTargetId="figure-93@5.[186,1401,262,1738]" captionTargetPageId="5" captionText="Fig. 2. Levizonus circularis Takakuwa, 1942, ♂♂, ♀, from China (FSCB 12019Ch) (AB) and Russia (FSCB 232008, 212008) (CG). A. Gonopod, lateral view, longitudinal striae are not visible. B. Gonopod apex, lateral view. C, F. Gonopod apex, ventral view. D. Gonopod, lateral view, coxite partially destroyed. E. Middle part of gonopod, mesal view. G. Vulva, mesal view. Scale bars: A, E = 200 μm; D, G = 100 μm; BC, F = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.4923498" httpUri="https://zenodo.org/record/4923498/files/figure.png" pageId="7" pageNumber="166">Figs 2F</figureCitation>
,
<figureCitation id="136D05FBFFD7FFF7548EFA468263488D" box="[1001,1059,1508,1534]" captionStart="Fig" captionStartId="6.[189,232,1789,1815]" captionTargetBox="[203,1359,1160,1746]" captionTargetId="figure-434@6.[186,1401,1152,1752]" captionTargetPageId="6" captionText="Fig. 3. Levizonus circularis Takakuwa, 1942, ♂♂, from Russia (AI) and North Korea(J). AD. Variations in shape of distal part of gonopod telopodite, ventral view. EJ. Variations in shape of distal part of gonopod telopodite, lateral view. Scale bars: AD, J = 0.25 mm; EG = 0.2 mm; HI = 0.1 mm. AH: after Mikhaljova 1981a; I: after Lokschina &amp; Golovatch 1977; J: after Mikhaljova et al. 2000." figureDoi="http://doi.org/10.5281/zenodo.4923500" httpUri="https://zenodo.org/record/4923500/files/figure.png" pageId="7" pageNumber="166">3AI</figureCitation>
) or less (
<figureCitation id="136D05FBFFD7FFF753ECFA4682A7488D" box="[1163,1255,1508,1534]" captionStart="Fig" captionStartId="5.[189,232,1773,1799]" captionTargetBox="[189,1397,265,1735]" captionTargetId="figure-93@5.[186,1401,262,1738]" captionTargetPageId="5" captionText="Fig. 2. Levizonus circularis Takakuwa, 1942, ♂♂, ♀, from China (FSCB 12019Ch) (AB) and Russia (FSCB 232008, 212008) (CG). A. Gonopod, lateral view, longitudinal striae are not visible. B. Gonopod apex, lateral view. C, F. Gonopod apex, ventral view. D. Gonopod, lateral view, coxite partially destroyed. E. Middle part of gonopod, mesal view. G. Vulva, mesal view. Scale bars: A, E = 200 μm; D, G = 100 μm; BC, F = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.4923498" httpUri="https://zenodo.org/record/4923498/files/figure.png" pageId="7" pageNumber="166">Figs 2B</figureCitation>
,
<figureCitation id="136D05FBFFD7FFF75394FA46834C488D" box="[1267,1292,1508,1534]" captionStart="Fig" captionStartId="6.[189,232,1789,1815]" captionTargetBox="[203,1359,1160,1746]" captionTargetId="figure-434@6.[186,1401,1152,1752]" captionTargetPageId="6" captionText="Fig. 3. Levizonus circularis Takakuwa, 1942, ♂♂, from Russia (AI) and North Korea(J). AD. Variations in shape of distal part of gonopod telopodite, ventral view. EJ. Variations in shape of distal part of gonopod telopodite, lateral view. Scale bars: AD, J = 0.25 mm; EG = 0.2 mm; HI = 0.1 mm. AH: after Mikhaljova 1981a; I: after Lokschina &amp; Golovatch 1977; J: after Mikhaljova et al. 2000." figureDoi="http://doi.org/10.5281/zenodo.4923500" httpUri="https://zenodo.org/record/4923500/files/figure.png" pageId="7" pageNumber="166">3J</figureCitation>
) strongly curved mesad and located lower or higher. Central edge of telopodite apex with outgrowths and spinules of varying shape and size (
<figureCitation id="136D05FBFFD7FFF756A5F98F845C4B34" box="[450,540,1581,1607]" captionStart="Fig" captionStartId="5.[189,232,1773,1799]" captionTargetBox="[189,1397,265,1735]" captionTargetId="figure-93@5.[186,1401,262,1738]" captionTargetPageId="5" captionText="Fig. 2. Levizonus circularis Takakuwa, 1942, ♂♂, ♀, from China (FSCB 12019Ch) (AB) and Russia (FSCB 232008, 212008) (CG). A. Gonopod, lateral view, longitudinal striae are not visible. B. Gonopod apex, lateral view. C, F. Gonopod apex, ventral view. D. Gonopod, lateral view, coxite partially destroyed. E. Middle part of gonopod, mesal view. G. Vulva, mesal view. Scale bars: A, E = 200 μm; D, G = 100 μm; BC, F = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.4923498" httpUri="https://zenodo.org/record/4923498/files/figure.png" pageId="7" pageNumber="166">Figs 2C</figureCitation>
,
<figureCitation id="136D05FBFFD7FFF75541F98F842B4B34" box="[550,619,1581,1607]" captionStart="Fig" captionStartId="6.[189,232,1789,1815]" captionTargetBox="[203,1359,1160,1746]" captionTargetId="figure-434@6.[186,1401,1152,1752]" captionTargetPageId="6" captionText="Fig. 3. Levizonus circularis Takakuwa, 1942, ♂♂, from Russia (AI) and North Korea(J). AD. Variations in shape of distal part of gonopod telopodite, ventral view. EJ. Variations in shape of distal part of gonopod telopodite, lateral view. Scale bars: AD, J = 0.25 mm; EG = 0.2 mm; HI = 0.1 mm. AH: after Mikhaljova 1981a; I: after Lokschina &amp; Golovatch 1977; J: after Mikhaljova et al. 2000." figureDoi="http://doi.org/10.5281/zenodo.4923500" httpUri="https://zenodo.org/record/4923500/files/figure.png" pageId="7" pageNumber="166">3 AJ</figureCitation>
).
</paragraph>
<paragraph id="8BE9197EFFD7FFF7578DF9CD87054BFA" blockId="7.[189,1051,1647,1711]" box="[234,325,1647,1673]" pageId="7" pageNumber="166">
<emphasis id="B922C56CFFD7FFF7578DF9CD87054BFA" bold="true" box="[234,325,1647,1673]" pageId="7" pageNumber="166">Female</emphasis>
</paragraph>
<paragraph id="8BE9197EFFD7FFF757DAF936825B4BDD" blockId="7.[189,1051,1647,1711]" box="[189,1051,1684,1711]" pageId="7" pageNumber="166">
SIZE. Body
<quantity id="4CAEB49BFFD7FFF75624F93687814BDD" box="[323,449,1684,1711]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" metricValueMax="3.2" metricValueMin="2.5" pageId="7" pageNumber="166" unit="mm" value="28.5" valueMax="32.0" valueMin="25.0">2532 mm</quantity>
long, 3.05.0 mm wide, often stouter than in male.
</paragraph>
<paragraph id="8BE9197EFFD7FFF757DAF97582A04B83" blockId="7.[189,1248,1750,1777]" box="[189,1248,1750,1777]" pageId="7" pageNumber="166">BODY. Metatergal bosses either relatively well developed or traceable, same as in juvenile.</paragraph>
<paragraph id="8BE9197EFFD7FFF757DAF8B885024A47" blockId="7.[189,834,1817,1844]" box="[189,834,1817,1844]" pageId="7" pageNumber="166">LEGS. Coxa 2 with a small, pointed, setigerous process.</paragraph>
<paragraph id="8BE9197EFFD7FFF757DAF8FE87874A05" blockId="7.[189,455,1884,1910]" box="[189,455,1884,1910]" pageId="7" pageNumber="166">
VULVAE. As in
<figureCitation id="136D05FBFFD7FFF75601F8FE87844A05" box="[358,452,1884,1910]" captionStart="Fig" captionStartId="5.[189,232,1773,1799]" captionTargetBox="[189,1397,265,1735]" captionTargetId="figure-93@5.[186,1401,262,1738]" captionTargetPageId="5" captionText="Fig. 2. Levizonus circularis Takakuwa, 1942, ♂♂, ♀, from China (FSCB 12019Ch) (AB) and Russia (FSCB 232008, 212008) (CG). A. Gonopod, lateral view, longitudinal striae are not visible. B. Gonopod apex, lateral view. C, F. Gonopod apex, ventral view. D. Gonopod, lateral view, coxite partially destroyed. E. Middle part of gonopod, mesal view. G. Vulva, mesal view. Scale bars: A, E = 200 μm; D, G = 100 μm; BC, F = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.4923498" httpUri="https://zenodo.org/record/4923498/files/figure.png" pageId="7" pageNumber="166">Fig. 2G</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C4AF5FFD7FFF757DAF83C85964A9B" box="[189,982,1950,2025]" pageId="7" pageNumber="166" type="distribution">
<paragraph id="8BE9197EFFD7FFF757DAF83C87164ACB" blockId="7.[189,342,1950,1976]" box="[189,342,1950,1976]" pageId="7" pageNumber="166">
<heading id="D0A1AE12FFD7FFF757DAF83C87164ACB" bold="true" box="[189,342,1950,1976]" fontSize="11" level="3" pageId="7" pageNumber="166" reason="3">
<emphasis id="B922C56CFFD7FFF757DAF83C87164ACB" bold="true" box="[189,342,1950,1976]" pageId="7" pageNumber="166">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BE9197EFFD7FFF757DAF86C85964A9B" blockId="7.[189,982,1998,2025]" box="[189,982,1998,2025]" pageId="7" pageNumber="166">
<collectingCountry id="F34159EEFFD7FFF757DAF86C87504A9B" box="[189,272,1998,2024]" name="Russia" pageId="7" pageNumber="166">Russia</collectingCountry>
: Far East (
<collectingRegion id="4992D79CFFD7FFF756ECF86C84094A9A" box="[395,585,1998,2025]" country="Russia" name="Primorskiy" pageId="7" pageNumber="166">Primorskiy krai</collectingRegion>
);
<collectingCountry id="F34159EEFFD7FFF7553AF86C84B44A9A" box="[605,756,1998,2025]" name="North Korea" pageId="7" pageNumber="166">North Korea</collectingCountry>
; North-East
<collectingCountry id="F34159EEFFD7FFF754EFF86C85924A9B" box="[904,978,1998,2024]" name="China" pageId="7" pageNumber="166">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C4AF5FFD8FFF857DAFEAB84B94953" pageId="8" pageNumber="167" type="discussion">
<paragraph id="8BE9197EFFD8FFF857DAFEAB876F4C50" blockId="8.[189,303,265,291]" box="[189,303,265,291]" pageId="8" pageNumber="167">
<heading id="D0A1AE12FFD8FFF857DAFEAB876F4C50" bold="true" box="[189,303,265,291]" fontSize="11" level="3" pageId="8" pageNumber="167" reason="3">
<emphasis id="B922C56CFFD8FFF857DAFEAB876F4C50" bold="true" box="[189,303,265,291]" pageId="8" pageNumber="167">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BE9197EFFD8FFF857DAFE9A83374E5E" blockId="8.[189,1399,312,814]" pageId="8" pageNumber="167">
This species was originally described by
<bibRefCitation id="EFC7648FFFD8FFF855DFFE9A85C94C21" author="Takakuwa Y." box="[696,905,312,338]" pageId="8" pageNumber="167" pagination="39 - 47" refId="ref10993" refString="Takakuwa Y. 1942. Zur Kenntnis der Japanischen Diplopoden. Annotationes Zoologicae Japonenses 21: 39 - 47." type="journal article" year="1942">Takakuwa (1942)</bibRefCitation>
from Zyônanri (modern names: village: Sangnam-ri; county: Yeonweon-gun; province: Pyeongannam-do;
<collectingCountry id="F34159EEFFD8FFF854AAFEFF82274C04" box="[973,1127,349,375]" name="North Korea" pageId="8" pageNumber="167">North Korea</collectingCountry>
). However the original description is far too fragmentary. That
<taxonomicName id="4C5662FDFFD8FFF855ECFE20855A4CE8" authorityName="Takakuwa" authorityYear="1942" box="[651,794,385,411]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="167" phylum="Arthropoda" rank="species" species="circularis">
<emphasis id="B922C56CFFD8FFF855ECFE20855A4CE8" box="[651,794,385,411]" italics="true" pageId="8" pageNumber="167">L. circularis</emphasis>
</taxonomicName>
might prove to be a senior synonym of
<taxonomicName id="4C5662FDFFD8FFF8538FFE2083364CE8" authorityName="Lokschina &amp; Golovatch" authorityYear="1977" box="[1256,1398,385,411]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="167" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B922C56CFFD8FFF8538FFE2083364CE8" box="[1256,1398,385,411]" italics="true" pageId="8" pageNumber="167">L. variabilis</emphasis>
</taxonomicName>
has long been suspected (
<bibRefCitation id="EFC7648FFFD8FFF85693FE0484F94CB3" author="Golovatch S. I." box="[500,697,422,448]" pageId="8" pageNumber="167" pagination="17 - 20" refId="ref9350" refString="Golovatch S. I. 1979. Notes on Levizonus thaumasius Att. (Diplopoda). In: Ivliev L. A. (ed.) Ekologiya i biologiya chlenistonogikh yuga Dalnego Vostoka: 17 - 20. DVNC AN SSSR, Vladivostok. [In Russian.]" type="book chapter" year="1979">Golovatch 1979</bibRefCitation>
;
<bibRefCitation id="EFC7648FFFD8FFF855AEFE0485A44CB3" author="Mikhaljova E. V. &amp; Golovatch S. I. &amp; Wytwer J." box="[713,996,422,448]" pageId="8" pageNumber="167" pagination="109 - 122" refId="ref10667" refString="Mikhaljova E. V., Golovatch S. I. &amp; Wytwer J. 2000. On some new or poorly-known millipedes (Diplopoda) from North Korea. Fragmenta Faunistica 43 (10): 109 - 122. https: // doi. org / 10.3161 / 00159301 FF 2000.43.10.109" type="journal article" year="2000">
Mikhaljova
<emphasis id="B922C56CFFD8FFF8543CFE0585D94CB3" box="[859,921,422,448]" italics="true" pageId="8" pageNumber="167">et al.</emphasis>
2000
</bibRefCitation>
). So, the above male from
<collectingCountry id="F34159EEFFD8FFF85255FE0487484C96" name="North Korea" pageId="8" pageNumber="167">North Korea</collectingCountry>
, kept at FSCB, can be considered as near topotypic, because both localities are only a few airkm away from one another. Unfortunately, information about the
<typeStatus id="54EDA7DCFFD8FFF854D8FE4D82664F7A" box="[959,1062,495,521]" pageId="8" pageNumber="167" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C5662FDFFD8FFF8532AFE52829D4F7A" authorityName="Takakuwa" authorityYear="1942" box="[1101,1245,495,521]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="167" phylum="Arthropoda" rank="species" species="circularis">
<emphasis id="B922C56CFFD8FFF8532AFE52829D4F7A" box="[1101,1245,495,521]" italics="true" pageId="8" pageNumber="167">L. circularis</emphasis>
</taxonomicName>
could not be found; the
<typeStatus id="54EDA7DCFFD8FFF85658FDB187E64F5E" box="[319,422,531,557]" pageId="8" pageNumber="167" type="holotype">holotype</typeStatus>
is probably lost. Maybe it was deposited in the private collection of Y. Miyosi as Takakuwas other xystodesmid samples and was destroyed (see
<bibRefCitation id="EFC7648FFFD8FFF854F9FD9A82774F21" author="Tanabe T." box="[926,1079,568,594]" pageId="8" pageNumber="167" pagination="101 - 113" refId="ref11077" refString="Tanabe T. 1994. The millipede genus Levizonus (Polydesmida, Xystodesmidae) in Japan. Japanese Journal of Entomology 62 (1): 101 - 113." type="journal article" year="1994">Tanabe 1994</bibRefCitation>
). Originally described from Lazovskiy (old name: Sudzukhinskiy) and Kavalerovskiy districts,
<collectingRegion id="4992D79CFFD8FFF854A5FDFE823B4F05" box="[962,1147,604,630]" country="Russia" name="Primorskiy" pageId="8" pageNumber="167">Primorskiy krai</collectingRegion>
,
<collectingCountry id="F34159EEFFD8FFF853E3FDFE82924F05" box="[1156,1234,604,630]" name="Russia" pageId="8" pageNumber="167">Russia</collectingCountry>
(
<bibRefCitation id="EFC7648FFFD8FFF853B9FDFE873C4FE8" author="Lokschina I. E. &amp; Golovatch S. I." pageId="8" pageNumber="167" pagination="73 - 78" refId="ref9709" refString="Lokschina I. E. &amp; Golovatch S. I. 1977. The new myriapods (Diplopoda) from the USSR. Byulleten Moskovskogo obshchestva ispytateley prirodi, Biology 82 (1): 73 - 78. [In Russian.]" type="journal article" year="1977">Lokschina &amp; Golovatch 1977</bibRefCitation>
),
<taxonomicName id="4C5662FDFFD8FFF856F7FD20845E4FE8" authorityName="Lokschina &amp; Golovatch" authorityYear="1977" box="[400,542,641,667]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="167" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B922C56CFFD8FFF856F7FD20845E4FE8" box="[400,542,641,667]" italics="true" pageId="8" pageNumber="167">L. variabilis</emphasis>
</taxonomicName>
appears to be widespread in and confined to the south-eastern and eastern parts of the
<collectingRegion id="4992D79CFFD8FFF8562FFD0787BF4FCC" box="[328,511,677,703]" country="Russia" name="Primorskiy" pageId="8" pageNumber="167">Primorskiy krai</collectingRegion>
(
<bibRefCitation id="EFC7648FFFD8FFF85569FD07849E4FB3" author="Mikhaljova E. V." box="[526,734,677,704]" pageId="8" pageNumber="167" refId="ref10459" refString="Mikhaljova E. V. 2017. The Millipede Fauna (Diplopoda) of the Asian Part of Russia. Dalnauka, Vladivostok. [In Russian.]" type="book" year="2017">Mikhaljova 2017</bibRefCitation>
). The gonopods of
<taxonomicName id="4C5662FDFFD8FFF854D9FD04820D4FCC" authorityName="Lokschina &amp; Golovatch" authorityYear="1977" box="[958,1101,677,703]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="167" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B922C56CFFD8FFF854D9FD04820D4FCC" box="[958,1101,677,703]" italics="true" pageId="8" pageNumber="167">L. variabilis</emphasis>
</taxonomicName>
are variable (
<bibRefCitation id="EFC7648FFFD8FFF8538AFD0787484F97" author="Mikhaljova E. V." pageId="8" pageNumber="167" pagination="62 - 67" refId="ref9873" refString="Mikhaljova E. V. 1981 a. The millipede genus Levizonus (Diplopoda, Xystodesmidae) from the USSR Far East. Byulleten Moskovskogo obshchestva ispytateley prirodi, Biology 86 (3): 62 - 67. [In Russian, English summary.]" type="journal article" year="1981">Mikhaljova 1981a</bibRefCitation>
;
<bibRefCitation id="EFC7648FFFD8FFF85672FD68846A4F97" author="Mikhaljova E. V. &amp; Golovatch S. I. &amp; Wytwer J." box="[277,554,714,740]" pageId="8" pageNumber="167" pagination="109 - 122" refId="ref10667" refString="Mikhaljova E. V., Golovatch S. I. &amp; Wytwer J. 2000. On some new or poorly-known millipedes (Diplopoda) from North Korea. Fragmenta Faunistica 43 (10): 109 - 122. https: // doi. org / 10.3161 / 00159301 FF 2000.43.10.109" type="journal article" year="2000">
Mikhaljova
<emphasis id="B922C56CFFD8FFF856C2FD6987A14F97" box="[421,481,714,740]" italics="true" pageId="8" pageNumber="167">et al.</emphasis>
2000
</bibRefCitation>
). The telopodite apex carries outgrowths and spinules of varying shape and size (
<figureCitation id="136D05FBFFD8FFF8564AFD4C87F14E7A" box="[301,433,750,777]" captionStart="Fig" captionStartId="6.[189,232,1789,1815]" captionTargetBox="[203,1359,1160,1746]" captionTargetId="figure-434@6.[186,1401,1152,1752]" captionTargetPageId="6" captionText="Fig. 3. Levizonus circularis Takakuwa, 1942, ♂♂, from Russia (AI) and North Korea(J). AD. Variations in shape of distal part of gonopod telopodite, ventral view. EJ. Variations in shape of distal part of gonopod telopodite, lateral view. Scale bars: AD, J = 0.25 mm; EG = 0.2 mm; HI = 0.1 mm. AH: after Mikhaljova 1981a; I: after Lokschina &amp; Golovatch 1977; J: after Mikhaljova et al. 2000." figureDoi="http://doi.org/10.5281/zenodo.4923500" httpUri="https://zenodo.org/record/4923500/files/figure.png" pageId="8" pageNumber="167">Fig. 3AD</figureCitation>
). The SL is located somewhat lower and less strongly curved mesad in specimens from
<collectingCountry id="F34159EEFFD8FFF8579AFCB187D14E5D" box="[253,401,787,814]" name="North Korea" pageId="8" pageNumber="167">North Korea</collectingCountry>
(
<figureCitation id="136D05FBFFD8FFF856C6FCB187B24E5E" box="[417,498,787,813]" captionStart="Fig" captionStartId="6.[189,232,1789,1815]" captionTargetBox="[203,1359,1160,1746]" captionTargetId="figure-434@6.[186,1401,1152,1752]" captionTargetPageId="6" captionText="Fig. 3. Levizonus circularis Takakuwa, 1942, ♂♂, from Russia (AI) and North Korea(J). AD. Variations in shape of distal part of gonopod telopodite, ventral view. EJ. Variations in shape of distal part of gonopod telopodite, lateral view. Scale bars: AD, J = 0.25 mm; EG = 0.2 mm; HI = 0.1 mm. AH: after Mikhaljova 1981a; I: after Lokschina &amp; Golovatch 1977; J: after Mikhaljova et al. 2000." figureDoi="http://doi.org/10.5281/zenodo.4923500" httpUri="https://zenodo.org/record/4923500/files/figure.png" pageId="8" pageNumber="167">Fig. 3J</figureCitation>
) and North-East
<collectingCountry id="F34159EEFFD8FFF855DBFCB185424E5E" box="[700,770,787,813]" name="China" pageId="8" pageNumber="167">China</collectingCountry>
(
<figureCitation id="136D05FBFFD8FFF85475FCB185D14E5E" box="[786,913,787,813]" captionStart="Fig" captionStartId="5.[189,232,1773,1799]" captionTargetBox="[189,1397,265,1735]" captionTargetId="figure-93@5.[186,1401,262,1738]" captionTargetPageId="5" captionText="Fig. 2. Levizonus circularis Takakuwa, 1942, ♂♂, ♀, from China (FSCB 12019Ch) (AB) and Russia (FSCB 232008, 212008) (CG). A. Gonopod, lateral view, longitudinal striae are not visible. B. Gonopod apex, lateral view. C, F. Gonopod apex, ventral view. D. Gonopod, lateral view, coxite partially destroyed. E. Middle part of gonopod, mesal view. G. Vulva, mesal view. Scale bars: A, E = 200 μm; D, G = 100 μm; BC, F = 20 μm." figureDoi="http://doi.org/10.5281/zenodo.4923498" httpUri="https://zenodo.org/record/4923498/files/figure.png" pageId="8" pageNumber="167">Fig. 2AB</figureCitation>
) as compared to that of Russian samples.
</paragraph>
<paragraph id="8BE9197EFFD8FFF857DAFCF4822D4ECA" blockId="8.[189,1398,854,953]" pageId="8" pageNumber="167">
Supported by the geographical evidence and the morphological variability of the gonopods of
<taxonomicName id="4C5662FDFFD8FFF85384FCF5832F4E03" authorityName="Lokschina &amp; Golovatch" authorityYear="1977" box="[1251,1391,854,880]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="167" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B922C56CFFD8FFF85384FCF5832F4E03" box="[1251,1391,854,880]" italics="true" pageId="8" pageNumber="167">L. variabilis</emphasis>
</taxonomicName>
, the following new formal synonym is proposed:
<taxonomicName id="4C5662FDFFD8FFF85471FCD9829A4EE7" authority="Takakuwa, 1942" authorityName="Takakuwa" authorityYear="1942" box="[790,1242,890,916]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="167" phylum="Arthropoda" rank="species" species="circularis">
<emphasis id="B922C56CFFD8FFF85471FCD982484EE7" box="[790,1032,890,916]" italics="true" pageId="8" pageNumber="167">Levizonus circularis</emphasis>
<bibRefCitation id="EFC7648FFFD8FFF85374FCD8829A4EE7" author="Takakuwa Y." box="[1043,1242,890,916]" pageId="8" pageNumber="167" pagination="39 - 47" refId="ref10993" refString="Takakuwa Y. 1942. Zur Kenntnis der Japanischen Diplopoden. Annotationes Zoologicae Japonenses 21: 39 - 47." type="journal article" year="1942">Takakuwa, 1942</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="4C5662FDFFD8FFF85266FCD984D14ECA" authority="Lokschina &amp; Golovatch 1977" authorityName="Lokschina &amp; Golovatch" authorityYear="1977" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="167" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B922C56CFFD8FFF85266FCD9876D4ECA" italics="true" pageId="8" pageNumber="167">Levizonus variabilis</emphasis>
<bibRefCitation id="EFC7648FFFD8FFF85652FC3D84D14ECA" author="Lokschina I. E. &amp; Golovatch S. I." box="[309,657,927,953]" pageId="8" pageNumber="167" pagination="73 - 78" refId="ref9709" refString="Lokschina I. E. &amp; Golovatch S. I. 1977. The new myriapods (Diplopoda) from the USSR. Byulleten Moskovskogo obshchestva ispytateley prirodi, Biology 82 (1): 73 - 78. [In Russian.]" type="journal article" year="1977">Lokschina &amp; Golovatch 1977</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel id="A2117817FFD8FFF855FFFC3D85444ECA" box="[664,772,927,953]" pageId="8" pageNumber="167">syn. nov.</taxonomicNameLabel>
, the valid name being the first.
</paragraph>
<paragraph id="8BE9197EFFD8FFF857DAFC4084B94953" blockId="8.[189,1398,994,1057]" pageId="8" pageNumber="167">
<taxonomicName id="4C5662FDFFD8FFF857DAFC4087ED4E8F" authorityName="Takakuwa" authorityYear="1942" box="[189,429,994,1020]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="167" phylum="Arthropoda" rank="species" species="circularis">
<emphasis id="B922C56CFFD8FFF857DAFC4087ED4E8F" box="[189,429,994,1020]" italics="true" pageId="8" pageNumber="167">Levizonus circularis</emphasis>
</taxonomicName>
and accordingly its junior synonym
<taxonomicName id="4C5662FDFFD8FFF85400FC4085B84E8F" authorityName="Lokschina &amp; Golovatch" authorityYear="1977" box="[871,1016,994,1020]" class="Diplopoda" family="Xystodesmidae" genus="Levizonus" kingdom="Animalia" order="Polydesmida" pageId="8" pageNumber="167" phylum="Arthropoda" rank="species" species="variabilis">
<emphasis id="B922C56CFFD8FFF85400FC4085B84E8F" box="[871,1016,994,1020]" italics="true" pageId="8" pageNumber="167">L. variabilis</emphasis>
</taxonomicName>
have never been recorded from
<collectingCountry id="F34159EEFFD8FFF857DAFBA487474953" box="[189,263,1030,1056]" name="China" pageId="8" pageNumber="167">China</collectingCountry>
. The species is new for the Chinese fauna.
</paragraph>
</subSubSection>
</treatment>
</document>