151 lines
15 KiB
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151 lines
15 KiB
XML
<document id="84A195F2D95AD04EB2A358EAD119B316" ID-CLB-Dataset="31983" ID-DOI="http://dx.doi.org/10.3897/zookeys.702.14728" ID-GBIF-Dataset="fb87e82a-bc10-4e27-8c9c-b0cbc89d35fb" ID-PMC="PMC5673967" ID-Pensoft-Pub="1313-2970-702-113" ID-PubMed="29118601" ID-ZooBank="CB6B3D9E27494ACCA78CD0A58C41A348" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-702-113" ModsDocOrigin="ZooKeys 702" ModsDocTitle="Two new species of the Phanaeusendymion species group (Coleoptera, Scarabaeidae, Scarabaeinae)" checkinTime="1506490258390" checkinUser="pensoft" docAuthor="Moctezuma, Victor, Sanchez-Huerta, Jose Luis & Halffter, Gonzalo" docDate="2017" docId="DD6BB5FB8EDC36DE0A5A792CDAD4EFE1" docLanguage="en" docName="ZooKeys 702: 113-135" docOrigin="ZooKeys 702" docSource="http://dx.doi.org/10.3897/zookeys.702.14728" docTitle="Phanaeus halffterorum Edmonds 1979" docType="treatment" docVersion="6" lastPageNumber="113" masterDocId="C52EFFB05F74FF992A2DFFD3572CFFD8" masterDocTitle="Two new species of the Phanaeusendymion species group (Coleoptera, Scarabaeidae, Scarabaeinae)" masterLastPageNumber="135" masterPageNumber="113" pageNumber="113" updateTime="1701382113965" updateUser="ExternalLinkService">
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<mods:titleInfo id="189083EA4CAB36C34CE6FA027B095AB2">
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<mods:title id="2EE67DCE91E8642D57E9814F9CEC7BA0">Two new species of the Phanaeusendymion species group (Coleoptera, Scarabaeidae, Scarabaeinae)</mods:title>
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<mods:name id="8FBAEE08FF0C77F27504003E6A1E2A11" type="personal">
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<mods:namePart id="B9AF694652AE6AF7E70059D70E358464">Moctezuma, Victor</mods:namePart>
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<mods:namePart id="6FDCEA4B066002CDCFC8CB3A593E25E1">Sanchez-Huerta, Jose Luis</mods:namePart>
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<mods:namePart id="3E213823877530B8098A68A43411FFF3">Halffter, Gonzalo</mods:namePart>
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<mods:typeOfResource id="0F3B64333D4AEB0575E42BE564CD782F">text</mods:typeOfResource>
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<mods:title id="9913EB4DBDA58E6D094A1D0BAA2B2A0D">ZooKeys</mods:title>
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<mods:date id="20F3C01A816A55429F8E0F64432FF9A8">2017</mods:date>
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<mods:detail id="645475C2582A9836BE9D0A949C9366D1" type="volume">
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<mods:number id="855D66C8EE6D5DAEDB189F1EB321E22F">702</mods:number>
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<mods:url id="AB9D3785C6CC21A92BEEDD6A8B8854D4">http://dx.doi.org/10.3897/zookeys.702.14728</mods:url>
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<mods:classification id="5481F1C74B60B39323B59F819F899C5C">journal article</mods:classification>
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<mods:identifier id="BA402C1A8ABDF8D9B52528CC7F6A86A2" type="DOI">http://dx.doi.org/10.3897/zookeys.702.14728</mods:identifier>
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<treatment id="DD6BB5FB8EDC36DE0A5A792CDAD4EFE1" ID-GBIF-Taxon="134842823" LSID="urn:lsid:plazi:treatment:DD6BB5FB8EDC36DE0A5A792CDAD4EFE1" httpUri="http://treatment.plazi.org/id/DD6BB5FB8EDC36DE0A5A792CDAD4EFE1" lastPageNumber="113" pageId="0" pageNumber="113">
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<subSubSection id="8F3BB7D1A15C6D001AD672E9C765B13B" pageId="0" pageNumber="113" type="nomenclature">
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<paragraph id="293C40E03D3888BF9D59C4C5F0DC7646" pageId="0" pageNumber="113">
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<taxonomicName id="264BEF70DEC42E29082AAEB6C9FFBE5B" ID-CoL="4FSJM" authority="Edmonds, 1979" authorityName="Edmonds" authorityYear="1979" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus halffterorum" order="Coleoptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="halffterorum">Phanaeus halffterorum Edmonds, 1979</taxonomicName>
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Figs 1, 2, 3, 4
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</paragraph>
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</subSubSection>
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<subSubSection id="054DE85A3E6278005EFAE1A0ABCBA844" pageId="0" pageNumber="113" type="reference_group">
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<paragraph id="9FC325FCFC5078CDF430CE38797B99D9" pageId="0" pageNumber="113">
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<taxonomicName id="D615F88CE6A4F679B7D501FF0C87895B" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus halffterorum" order="Coleoptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="halffterorum">Phanaeus halffterorum</taxonomicName>
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:
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<bibRefCitation id="0AE894129C358E804251B5E8315E32BE" author="Edmonds, WD" journalOrPublisher="Pan-Pacific Entomologist" pageId="9" pageNumber="122" pagination="99 - 105" title="A new species of Phanaeus from Mexico (Coleoptera: Scarabaeidae)." volume="55" year="1979">Edmonds (1979</bibRefCitation>
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: 99; partim),
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<bibRefCitation id="10C561A185610A6ADC79D44E1FB2C957" author="Halffter, G" journalOrPublisher="Instituto de Ecologia, Mexico City" pageId="10" pageNumber="123" title="The nesting behavior of dung beetles (Scarabaeinae). An ecological and evolutive approach." year="1982">Halffter and Edmonds (1982</bibRefCitation>
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: 88-89),
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<bibRefCitation id="6272F69280C346C2C3804918E4E6FED7" pageId="0" pageNumber="113">Anduaga and Halffter (1991</bibRefCitation>
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: 157),
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<bibRefCitation id="009D400FD2812310CC98D1EC86A8BFE5" author="Delgado-Castillo, L" journalOrPublisher="The Coleopterists Bulletin" pageId="9" pageNumber="122" pagination="121 - 126" title="A new Mexican species of Onthophagus with mycophagous habits (Coleoptera: Scarabaeidae: Scarabaeinae)." volume="47" year="1993">Delgado-Castillo et al. (1993</bibRefCitation>
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: 125),
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<bibRefCitation id="A55183FEE367284A7ADC4D85DB578A04" author="Edmonds, WD" journalOrPublisher="Natural History Museum of Los Angeles County, Contributions in Science" pageId="10" pageNumber="123" pagination="1 - 105" title="Revision of Phanaeus MacLeay, a New World genus of Scarabaeinae dung beetles (Coleoptera: Scarabaeidae, Scarabaeinae)." volume="443" year="1994">Edmonds (1994</bibRefCitation>
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: 39-43, 101),
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<bibRefCitation id="C7564060764B47E0DE4F04857327792E" author="Anduaga, S" journalOrPublisher=")" pageId="9" pageNumber="122" pagination="119 - 130" title="Escarabajos coprofagos (Coleoptera: Scarabaeoidea) asociados a hongos en la Sierra Madre Occidental, Durango, Mexico: con una compilacion de las especies micetofagas. Acta Zoologica Mexicana (n. s." volume="80" year="2000">Anduaga (2000</bibRefCitation>
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: 125, 130),
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<bibRefCitation id="6706D9CF3B36B83268DA01E3538A3B59" author="Arnaud, P" journalOrPublisher="Hillside Books, Canterbury, Kent" pageId="9" pageNumber="122" title="Phanaeini. Les Coleopteres du Monde, vol. 28." year="2002">Arnaud (2002</bibRefCitation>
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: 95-96),
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<bibRefCitation id="0DB52A24B9470F634251AC069DD92ADD" pageId="0" pageNumber="113">Edmonds (2003</bibRefCitation>
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: 61, 65),
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<bibRefCitation id="A896403BC86DFD8FE47A0F7A7A4A10BE" author="Edmonds, WD" journalOrPublisher="Zootaxa" pageId="10" pageNumber="123" pagination="31 - 37" title="A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini) from Oaxaca, Mexico." volume="1171" year="2006">Edmonds (2006</bibRefCitation>
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: 31, 34, 36),
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<bibRefCitation id="281FE1E513CD476A08178C51E0204FDF" author="Ceballos, G" journalOrPublisher="Gobierno del Estado de Mexico, Toluca de Lerdo" pageId="9" pageNumber="122" title="La diversidad biologica del Estado de Mexico. Estudio de estado." url="http://www.biodiversidad.gob.mx/region/EEB/estudios.html" year="2009">Ceballos et al. (2009</bibRefCitation>
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: 397),
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<bibRefCitation id="0D00A0F25E02E2452BF8D72614277A11" author="Edmonds, WD" journalOrPublisher="Insecta Mundi" pageId="10" pageNumber="123" pagination="1 - 108" title="Taxonomy of Phanaeus revisited: revised keys to and comments on species of the New World dung beetle genus Phanaeus MacLeay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini)." url="http://digitalcommons.unl.edu/insectamundi/784/" volume="274" year="2012">
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Edmonds and
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<normalizedToken id="6847C3A079C960B076234753449115C6" originalValue="Zídek">Zidek</normalizedToken>
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(2012
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</bibRefCitation>
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: 3, 5, 12, 52, 54),
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<bibRefCitation id="4B8A946B8013526D3F0E7EBEF66F900C" pageId="0" pageNumber="113">Deloya et al. (2014</bibRefCitation>
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: 77),
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<bibRefCitation id="2EAE06E35C35ACF736816759784A6D49" author="Moctezuma, V" journalOrPublisher="The Coleopterists Bulletin" pageId="10" pageNumber="123" pagination="47 - 56" title="A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae) from Los Chimalapas, Oaxaca, Mexico." url="https://doi.org/10.1649/0010-065X-71.1.47" volume="71" year="2017">Moctezuma and Halffter (2017</bibRefCitation>
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: 52, 54-55),
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<bibRefCitation id="236FAAFC3D48DDEC98CCEC9AD39720C3" pageId="0" pageNumber="113">Lizardo et al. (in press)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="95E526FD5C2313DC46E09B1E5C433397" pageId="0" pageNumber="113" type="type material">
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<paragraph id="48BB089F839C71D20C41B2AE051C1E90" pageId="0" pageNumber="113">Type material examined</paragraph>
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<paragraph id="D03871EF0C5768ACBF34E21AF0DD6F8B" pageId="0" pageNumber="113">(5 ♂♂, 2 ♀♀). Paratypes (TAMU): 2 ♂♂, 2 ♀♀ labeled "MEXICO: Mexico, 5 km E Temascaltepec, Real de Arriba, 2200 m, 10-VII-1976, Fungus, oak-pine forest, W.D. Edmonds, P. Reyes, B. Kohlmann cols."; 2 ♂♂ labeled "MEXICO: Mexico, 8 km W Temascaltepec, 2360 m, 11-VII-76, Fungus in pine-oak forest, W.D. Edmonds, P. Reyes and B. Kohlmann cols."; 1 ♂ labeled "Real de Arriba, Dist. Temascaltepec, Edo. Mex., VII-1932, 6300 ft, Mexico D.F., Hinton Coll. B.M.1939-583".</paragraph>
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<caption id="19C4105EA95958D9F9E7EDBBA8865FCC" pageId="0" pageNumber="113">
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<paragraph id="28FB2EEDBE5D5D7AB5A2318FCFF2DC30" pageId="0" pageNumber="113">
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Figure 1.
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<taxonomicName id="733AF48738F061E850F8300371B34041" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus halffterorum" order="Coleoptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="halffterorum">Phanaeus halffterorum</taxonomicName>
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major male, dorsal view (paratype).
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</paragraph>
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</caption>
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<caption id="D1912C02C57A9D84BC9B20022C183809" pageId="0" pageNumber="113">
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<paragraph id="268F023CDFB821F6A81C7EC3D1687DF2" pageId="0" pageNumber="113">
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Figure 2.
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<taxonomicName id="B6CD99E594CE78B145C8FCA3B47FEE02" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus halffterorum" order="Coleoptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="halffterorum">Phanaeus halffterorum</taxonomicName>
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major male, lateral view (paratype).
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</paragraph>
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</caption>
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<caption id="FA14B0627474987676B6270AA10EA3B1" pageId="0" pageNumber="113">
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<paragraph id="B68A8057EBDAA11FAB0B8F9BCA08EFF8" pageId="0" pageNumber="113">
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Figure 3. Lamella copulatrix of
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<taxonomicName id="8243C5BAFAA62C830ECD7EEE57387B96" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus halffterorum" order="Coleoptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="halffterorum">Phanaeus halffterorum</taxonomicName>
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(paratype). Abbreviations: LL: left lobe, CR: central ridge, RL: right lobe.
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</paragraph>
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</caption>
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<caption id="4399EEB9A12F9699223F4C5343994BA6" pageId="0" pageNumber="113">
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<paragraph id="C08F5AE4A240E7D78CA1B5002E9B9B85" pageId="0" pageNumber="113">
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Figure 4.
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<taxonomicName id="F4BF235D57D8E5A14E2D82A57FC3C0DD" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus halffterorum" order="Coleoptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="halffterorum">Phanaeus halffterorum</taxonomicName>
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female, dorsal view (paratype).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="C23633AE7883D82F52386E6277206B24" pageId="0" pageNumber="113" type="distribution">
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<paragraph id="F75C9B0009DD71BE625C64BE15C15691" pageId="0" pageNumber="113">Distribution and ecology.</paragraph>
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<paragraph id="0C0C61839343E09C0C6470A4A9876763" pageId="0" pageNumber="113">
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This species is known from the environs of Temascaltepec, State of Mexico (Fig. 5), and inhabits pine-oak forests from 2200-2360 m. a.s.l.
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<taxonomicName id="D82174EDCB5D7385FD6AE4CDC39AF09E" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus halffterorum" order="Coleoptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="halffterorum">Phanaeus halffterorum</taxonomicName>
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is considered a mycetophagous species (
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<bibRefCitation id="A93CEFBD9C6F5E58C005FA9368765B23" author="Edmonds, WD" journalOrPublisher="Pan-Pacific Entomologist" pageId="9" pageNumber="122" pagination="99 - 105" title="A new species of Phanaeus from Mexico (Coleoptera: Scarabaeidae)." volume="55" year="1979">Edmonds 1979</bibRefCitation>
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,
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<bibRefCitation id="C96D5F367E2F3528FA25C8EF3A07B5FA" author="Edmonds, WD" journalOrPublisher="Natural History Museum of Los Angeles County, Contributions in Science" pageId="10" pageNumber="123" pagination="1 - 105" title="Revision of Phanaeus MacLeay, a New World genus of Scarabaeinae dung beetles (Coleoptera: Scarabaeidae, Scarabaeinae)." volume="443" year="1994">1994</bibRefCitation>
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,
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<bibRefCitation id="776D500CCCB0BEC3C975DDD21A419B0E" author="Halffter, G" journalOrPublisher="Instituto de Ecologia, Mexico City" pageId="10" pageNumber="123" title="The nesting behavior of dung beetles (Scarabaeinae). An ecological and evolutive approach." year="1982">Halffter and Edmonds 1982</bibRefCitation>
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).
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</paragraph>
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<caption id="022AC96D6E6EDF2293E2438C0673C1C7" pageId="0" pageNumber="113">
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<paragraph id="0F965CB831793B9B86A2309F5099985A" pageId="0" pageNumber="113">
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Figure 5. Distribution of
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<taxonomicName id="183106E34FBD90B846DC081648F475B4" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus halffterorum" order="Coleoptera" pageId="0" pageNumber="113" phylum="Arthropoda" rank="species" species="halffterorum">Phanaeus halffterorum</taxonomicName>
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(red triangle),
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<taxonomicName id="A8AF9FAD9B32DB4B152B80A9D3C777B2" lsidName="P. bravoensis" pageId="0" pageNumber="113" rank="species" species="bravoensis">P. bravoensis</taxonomicName>
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sp. n. (blue circle) and
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<taxonomicName id="EB254895BE75D029F01D5BF1FBFDEB6E" lsidName="P. huichol" pageId="0" pageNumber="113" rank="species" species="huichol">P. huichol</taxonomicName>
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sp. n. (purple diamonds).
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |