813 lines
275 KiB
XML
813 lines
275 KiB
XML
<document id="05196DB8834839C1A448E00B602EA447" ID-CLB-Dataset="2386" ID-DOI="10.11646/zootaxa.5093.3.3" ID-GBIF-Dataset="d0ebc178-bd1c-434f-95bb-7cf50121af03" ID-ISSN="1175-5326" ID-Zenodo-Dep="5909853" ID-ZooBank="0A4DE130-B44E-4526-8752-DFB6E60DDFA5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643276184439" checkinUser="plazi" docAuthor="Cepeda, Diego, Sánchez, Nuria, Sørensen, Martin V. & Landers, Stephen C." docDate="2022" docId="CA7849569774FF93FF5400AEFBCCF83A" docLanguage="en" docName="zootaxa.5093.3.3.pdf" docOrigin="Zootaxa 5093 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leiocanthus satanicus Cepeda & Sánchez & Sørensen & Landers 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="328" masterDocId="3641312E977EFF9EFFC30337FA6DFFAF" masterDocTitle="Leiocanthus quinquenudus sp. nov. and L. satanicus sp. nov., two new species of pycnophyid Kinorhyncha (Allomalorhagida: Pycnophyidae) from the Gulf of Mexico" masterLastPageNumber="336" masterPageNumber="315" pageNumber="325" updateTime="1699304508666" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="CD2E7EA2E850637A3C584A3C464D0540" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="53A41138A8F0A648DF2571D7B738B54B">
|
||
<mods:title id="4934A2F8EEC0D11BABF8C4C5BE1E8962">Leiocanthus quinquenudus sp. nov. and L. satanicus sp. nov., two new species of pycnophyid Kinorhyncha (Allomalorhagida: Pycnophyidae) from the Gulf of Mexico</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="59389B35235E2B69E1EFB3918597F2B6" type="personal">
|
||
<mods:role id="9327ADB527C84BFE20369E5F7762270D">
|
||
<mods:roleTerm id="8919C34A21E71D10995160E91766EB73">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E5CF05D1BB871B9062035681986BC3CC">Cepeda, Diego</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="4B163E2A065CF2DDB6B71454C5385124" type="personal">
|
||
<mods:role id="DF652BFE58D8D13DCC97E563BD0F57A4">
|
||
<mods:roleTerm id="101EE07A5FF434D485AA48EC2EB43C99">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B3D36E4C52337BF6299C5145316DCC04">Sánchez, Nuria</mods:namePart>
|
||
<mods:nameIdentifier id="E66B7283A87137031716C65997BABC7A" type="ORCID">0000-0003-4908-8755</mods:nameIdentifier>
|
||
<mods:affiliation id="AF171329015A15383BC55C449BAAB76F">nurisanc @ ucm. es; https: // orcid. org / 0000 - 0003 - 4908 - 8755</mods:affiliation>
|
||
<mods:nameIdentifier id="BE5E11B6DC8E491D037123D11105A324" type="email">nurisanc@ucm.es</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="58CC78AEAACA6C7B58C997C4DDBD8ACD" type="personal">
|
||
<mods:role id="D0F33D0848AF3F0444C44F355557C8BA">
|
||
<mods:roleTerm id="C9D6E54059E91617B3BD6A76383B057E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D8E6E91922D7774CE99F453584253673">Sørensen, Martin V.</mods:namePart>
|
||
<mods:nameIdentifier id="053206886EFF22A638D8D5E3F7220514" type="ORCID">0000-0002-0377-0276</mods:nameIdentifier>
|
||
<mods:affiliation id="0F6E393ACADEC4B0A1E3678D22C9F008">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, 2100 Copenhagen. (Denmark). mvsorensen @ snm. ku. dk; https: // orcid. org / 0000 - 0002 - 0377 - 0276</mods:affiliation>
|
||
<mods:nameIdentifier id="64EA5BBED7F0A3CD1B95089F31B105CD" type="email">mvsorensen@snm.ku.dk</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="1DA9BA3FC9F0DF7E66841E5C0D3A506E" type="personal">
|
||
<mods:role id="6C2C084A2CBB4381783E09F8663EE8B6">
|
||
<mods:roleTerm id="73A23C29EA4D43EFBD1BB1873F3F2EB2">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8F89CBB228BE72C2A9DA0C52FB12D48D">Landers, Stephen C.</mods:namePart>
|
||
<mods:affiliation id="B602A7A2004AA27C586B87319F354010">Department of Biological and Environmental Sciences, Troy University, Troy, Alabama (U. S. A.)</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="C4511BE339B7F4C2357C3301ED47464C">text</mods:typeOfResource>
|
||
<mods:relatedItem id="335E6C4F3DA2472516C0344D334B6C3E" type="host">
|
||
<mods:titleInfo id="E81B9ECF1D97CB846BF8D8C836AFA671">
|
||
<mods:title id="923364F427F81CA49F380C3A6A4808EC">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="51698057C808CAF422FDA3E0B3B6F7F4">
|
||
<mods:date id="287A4E6C8BB030A11EF11C7F60D83171">2022</mods:date>
|
||
<mods:detail id="1A78618F08C857682693F49B925375F5" type="pubDate">
|
||
<mods:number id="ADEB2E780400DDCEE711C1924C54AFBA">2022-01-27</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="F72FDE59ABB614A343FB7B9592F6351C" type="volume">
|
||
<mods:number id="39DC475DF1470103ECBE41B474192B5B">5093</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="A731E6480BB58A3B0EF3047EA47A1DAC" type="issue">
|
||
<mods:number id="BDC8755CF6AEDE3E6BF61AE7625B93E3">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="551665E651480286C260C276A41CD34B" unit="page">
|
||
<mods:start id="6D7812F958059AA82E46F0F0357DFB5E">315</mods:start>
|
||
<mods:end id="73C9FD45D5F128E8DD5DAAFA4A7B5F83">336</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="C5D4C116D0E97085366C48723AF45609">journal article</mods:classification>
|
||
<mods:identifier id="EB07C0E1655D2CA75DA8AC6B8E93B3EC" type="CLB-Dataset">2386</mods:identifier>
|
||
<mods:identifier id="2351AFE9BE2A0215D1DB7C1D3CA1D43C" type="DOI">10.11646/zootaxa.5093.3.3</mods:identifier>
|
||
<mods:identifier id="A40141E4554A6D23B87CCD2507C25915" type="GBIF-Dataset">d0ebc178-bd1c-434f-95bb-7cf50121af03</mods:identifier>
|
||
<mods:identifier id="190C45BAB27E7073511D98EF29BDCB8D" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="C7275AABAD39F48EF2C5802917D2DF17" type="Zenodo-Dep">5909853</mods:identifier>
|
||
<mods:identifier id="F97F4082CF691B5FE8240E890440641E" type="ZooBank">0A4DE130-B44E-4526-8752-DFB6E60DDFA5</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="CA7849569774FF93FF5400AEFBCCF83A" ID-DOI="http://doi.org/10.5281/zenodo.5910867" ID-GBIF-Taxon="192659272" ID-Zenodo-Dep="5910867" LSID="urn:lsid:plazi:treatment:CA7849569774FF93FF5400AEFBCCF83A" httpUri="http://treatment.plazi.org/id/CA7849569774FF93FF5400AEFBCCF83A" lastPageId="13" lastPageNumber="328" pageId="10" pageNumber="325">
|
||
<subSubSection id="0ACBABCB9774FF94FF5400AEFE08FC77" pageId="10" pageNumber="325" type="nomenclature">
|
||
<paragraph id="426EF8409774FF94FF5400AEF866FC1B" blockId="10.[151,1125,921,1020]" box="[151,523,921,948]" pageId="10" pageNumber="325">
|
||
<heading id="19264F2C9774FF94FF5400AEF866FC1B" bold="true" box="[151,523,921,948]" fontSize="11" level="1" pageId="10" pageNumber="325" reason="1">
|
||
<emphasis id="70A524529774FF94FF5400AEF866FC1B" bold="true" box="[151,523,921,948]" pageId="10" pageNumber="325">
|
||
<taxonomicName id="85D183C39774FF94FF5400AEFBCBFC1C" ID-CoL="8TZ5Z" authority="Cepeda & Sánchez & Sørensen & Landers, 2022" authorityName="Cepeda & Sánchez & Sørensen & Landers" authorityYear="2022" box="[151,422,921,948]" class="Allomalorhagida" family="Pycnophyidae" genus="Leiocanthus" kingdom="Animalia" pageId="10" pageNumber="325" phylum="Kinorhyncha" rank="species" species="satanicus" status="sp. nov.">
|
||
<emphasis id="70A524529774FF94FF5400AEFBCBFC1C" bold="true" box="[151,422,921,948]" italics="true" pageId="10" pageNumber="325">Leiocanthus satanicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="6B9699299774FF94FE6D00ADF866FC1B" box="[430,523,922,948]" pageId="10" pageNumber="325" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="426EF8409774FF94FF54008AFE08FC77" blockId="10.[151,1125,921,1020]" box="[151,1125,957,984]" pageId="10" pageNumber="325">
|
||
Zoobank code:
|
||
<uri id="3640F4429774FF94FE86008AFE08FC77" box="[325,1125,957,984]" pageId="10" pageNumber="325">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="3677C2959774FF94FD99008AFE08FC77" box="[602,1125,957,984]" pageId="10" pageNumber="325">9B48A215-F9C4-4BE2-8D5A-D650133B133</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0ACBABCB9774FF94FF5400D6F8EBFBF2" pageId="10" pageNumber="325" type="description">
|
||
<paragraph id="426EF8409774FF94FF5400D6FB7FFC53" blockId="10.[151,1125,921,1020]" box="[151,274,993,1020]" pageId="10" pageNumber="325">
|
||
(
|
||
<figureCitation id="DAEAE4C59774FF94FF5C00D6FB67FC53" box="[159,266,993,1020]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStart-3="FIGURE 8" captionStartId-0="14.[151,250,1696,1720]" captionStartId-1="15.[151,250,1727,1751]" captionStartId-2="16.[151,250,1656,1680]" captionStartId-3="17.[151,250,1505,1529]" captionTargetBox-0="[222,1364,191,1659]" captionTargetBox-1="[218,1369,181,1703]" captionTargetBox-2="[470,1121,187,1626]" captionTargetBox-3="[151,1436,181,1481]" captionTargetId-0="figure-31@14.[216,1370,181,1672]" captionTargetId-1="figure-18@15.[218,1369,181,1703]" captionTargetId-2="figure-31@16.[463,1125,181,1632]" captionTargetId-3="figure-18@17.[151,1436,181,1481]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionTargetPageId-2="16" captionTargetPageId-3="17" captionText-0="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." captionText-1="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." captionText-2="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." captionText-3="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi-0="http://doi.org/10.5281/zenodo.5909865" figureDoi-1="http://doi.org/10.5281/zenodo.5909867" figureDoi-2="http://doi.org/10.5281/zenodo.5909869" figureDoi-3="http://doi.org/10.5281/zenodo.5909871" httpUri-0="https://zenodo.org/record/5909865/files/figure.png" httpUri-1="https://zenodo.org/record/5909867/files/figure.png" httpUri-2="https://zenodo.org/record/5909869/files/figure.png" httpUri-3="https://zenodo.org/record/5909871/files/figure.png" pageId="10" pageNumber="325">Figs. 5–8</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="426EF8409774FF94FF540711FB68FB91" blockId="10.[151,646,1062,1147]" box="[151,261,1062,1086]" pageId="10" pageNumber="325">Synonymy</paragraph>
|
||
<paragraph id="426EF8409774FF94FF540772F8EBFBF2" blockId="10.[151,646,1062,1147]" box="[151,646,1093,1117]" pageId="10" pageNumber="325">
|
||
<taxonomicName id="85D183C39774FF94FF540772FB27FBF2" box="[151,330,1093,1117]" class="Allomalorhagida" family="Pycnophyidae" genus="Pycnophyes" higherTaxonomySource="CoL" kingdom="Animalia" order="Anomoirhaga" pageId="10" pageNumber="325" phylum="Kinorhyncha" rank="species" species="undefined-a">
|
||
<emphasis id="70A524529774FF94FF540772FB7DFBF2" box="[151,272,1093,1117]" italics="true" pageId="10" pageNumber="325">Pycnophyes</emphasis>
|
||
sp. A
|
||
</taxonomicName>
|
||
—Landers
|
||
<emphasis id="70A524529774FF94FE7F0772FB82FBF2" box="[444,495,1093,1117]" italics="true" pageId="10" pageNumber="325">et al.</emphasis>
|
||
2018:
|
||
<tableCitation id="0F53CDFB9774FF94FDF70772F8EFFBF2" box="[564,642,1093,1117]" captionStart="TABLE 2" captionStartId="5.[151,239,656,680]" captionTargetBox="[166,1245,804,1553]" captionText="TABLE 2. Measurements (in μm) of six adult specimens of Leiocanthus quinquenudus sp. nov. (three females and three males) from the Gulf of Mexico. Abbreviations: LTS, lateral terminal spine length; MSW-5, maximum sternal width (measured at segment 5); n, number of measured specimens; S, segment length (followed by number of corresponding segment); SD, standard deviation; SSW, standard sternal width (measured at segment 10); TL, trunk length. Character Holotype Range Mean (SD; n) TL 475.9 475.9–661.7 556.7 (70.0; 6) MSW-5 134.0 127.4–143.5 135.9 (6.4; 6) MSW-5/TL (%) 28.2 20.6–28.2 24.7 (2.9; 6) SSW 113.8 108.5–150.4 121.6 (15.1; 6) SSW/TL (%) 23.9 17.6–23.9 22.0 (2.2; 6) S1 85.5 80.7–96.5 88.0 (6.5; 6) S2 49.5 49.5– 62.4 56.7 (5.3; 6) S3 51.5 51.5–61.2 55.8 (4.5; 6) S4 52.1 52.1–71.0 61.6 (8.2; 6) S5 52.9 52.9–72.1 62.0 (7.4; 6) S6 56.0 56.0–71.2 64.1 (5.7; 6) S7 59.7 57.9–78.9 65.4 (9.2; 6) S8 58.9 58.9–78.6 66.7 (8.4; 6) S9 54.4 54.4–80.4 64.6 (9.5; 6) S10 54.1 48.0–75.6 62.7 (12.4; 6) S11 42.1 34.8–51.5 40.0 (6.2; 6) LTS 157.1 130.1–157.1 145.4 (10.4; 6) LTS/TL (%) 33.0 19.7–33.0 27.2 (5.0; 6)" pageId="10" pageNumber="325">Table 2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0ACBABCB9774FF94FF540754F80CFBD4" box="[151,609,1123,1147]" pageId="10" pageNumber="325" type="reference_group">
|
||
<paragraph id="426EF8409774FF94FF540754F80CFBD4" blockId="10.[151,646,1062,1147]" box="[151,609,1123,1147]" pageId="10" pageNumber="325">
|
||
<treatmentCitationGroup id="62C1DF6E9774FF94FF540754F80CFBD4" box="[151,609,1123,1147]" pageId="10" pageNumber="325">
|
||
<taxonomicName id="85D183C39774FF94FF540754FB5CFBD4" ID-CoL="7Z5PF" box="[151,305,1123,1147]" class="Allomalorhagida" family="Pycnophyidae" genus="Leiocanthus" kingdom="Animalia" pageId="10" pageNumber="325" phylum="Kinorhyncha" rank="genus">
|
||
<emphasis id="70A524529774FF94FF540754FB7EFBD4" box="[151,275,1123,1147]" italics="true" pageId="10" pageNumber="325">Leiocanthus</emphasis>
|
||
W
|
||
</taxonomicName>
|
||
—in
|
||
<treatmentCitation id="C370DE519774FF94FEA70754F830FBD4" author="Landers, S. & Sorensen, M. V. & Sanchez, N. & Beaton, K. R. & Miller, J. M. & Ingels, J." box="[356,605,1123,1147]" page="5" pageId="10" pageNumber="325" year="2019">
|
||
<bibRefCitation id="264085B19774FF94FEA70754F841FBD4" author="Landers, S. & Sorensen, M. V. & Sanchez, N. & Beaton, K. R. & Miller, J. M. & Ingels, J." box="[356,556,1123,1147]" pageId="10" pageNumber="325" pagination="1 - 14" refId="ref12905" refString="Landers, S., Sorensen, M. V., Sanchez, N., Beaton, K. R., Miller, J. M. & Ingels, J. (2019) Kinorhynch communities on the Louisiana continental shelf. Proceedings of the Biological Society of Washington, 132, 1 - 14. https: // doi. org / 10.2988 / 18 - 00008" type="journal article" year="2019">
|
||
Landers
|
||
<emphasis id="70A524529774FF94FE7F0753FB82FBD4" box="[444,495,1123,1147]" italics="true" pageId="10" pageNumber="325">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
: p. 5
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0ACBABCB9774FF94FF54079EFB8EFA33" pageId="10" pageNumber="325" type="materials_examined">
|
||
<paragraph id="426EF8409774FF94FF54079EFB8EFA33" blockId="10.[151,1437,1193,1976]" pageId="10" pageNumber="325">
|
||
<emphasis id="70A524529774FF94FF54079EFBEDFB6C" bold="true" box="[151,384,1193,1219]" pageId="10" pageNumber="325">Material examined.</emphasis>
|
||
<materialsCitation id="F2B9F21D9774FF94FE46079EF974FAA3" ID-GBIF-Occurrence="3458066306" collectingDate="2013-11-12" collectionCode="NHMD" collectorName="Fluoromount G" elevation="59" latitude="29.2998" location="northern Gulf" longLatPrecision="6" longitude="-88.72" pageId="10" pageNumber="325" specimenCount="1" stateProvince="Mexico" typeStatus="holotype">
|
||
<typeStatus id="9D6A46E29774FF94FE46079EFB9DFB6C" box="[389,496,1193,1219]" pageId="10" pageNumber="325" type="holotype">Holotype</typeStatus>
|
||
, adult female, collected on
|
||
<date id="366FDE809774FF94FCDD079EF996FB6B" box="[798,1019,1193,1220]" pageId="10" pageNumber="325" value="2013-11-12">
|
||
<collectingDate id="262B27689774FF94FCDD079EF996FB6B" box="[798,1019,1193,1220]" pageId="10" pageNumber="325" value="2013-11-12">November 12, 2013</collectingDate>
|
||
</date>
|
||
at St.
|
||
<quantity id="852955A59774FF94FB83079EFED4FB6C" box="[1088,1209,1193,1219]" metricMagnitude="1" metricUnit="m" metricValue="2.64287" metricValueMax="5.11302" metricValueMin="0.17271999999999998" pageId="10" pageNumber="325" unit="in" value="1040.5" valueMax="2013.0" valueMin="68.0">68-2013 in</quantity>
|
||
the
|
||
<location id="470EAE9B9774FF94FB24079EFF12FB6C" LSID="urn:lsid:plazi:treatment:CA7849569774FF93FF5400AEFBCCF83A:470EAE9B9774FF94FB24079EFF12FB6C" box="[1255,1407,1193,1219]" latitude="29.2998" longLatPrecision="6" longitude="-88.72" name="northern Gulf" pageId="10" pageNumber="325" stateProvince="Mexico">northern Gulf</location>
|
||
of
|
||
<collectingRegion id="801536A29774FF94FF5407FAFA80FB48" box="[151,237,1229,1255]" country="Mexico" name="Mexico" pageId="10" pageNumber="325">Mexico</collectingRegion>
|
||
continental shelf:
|
||
<geoCoordinate id="27E59E879774FF94FE7F07F9F82DFB47" box="[444,576,1230,1256]" degrees="29.2998" direction="north" orientation="latitude" pageId="10" pageNumber="325" precision="5" value="29.2998">29.2998° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="27E59E879774FF94FD8907F9F8A7FB47" box="[586,714,1230,1256]" degrees="88.7200" direction="west" orientation="longitude" pageId="10" pageNumber="325" precision="5" value="-88.72">88.7200°W</geoCoordinate>
|
||
(
|
||
<tableCitation id="0F53CDFB9774FF94FD1A07FAF946FB47" box="[729,811,1229,1256]" captionStart="TABLE 1" captionStartId="1.[151,239,657,681]" captionTargetBox="[166,1409,732,1986]" captionText="TABLE 1. Summary of data on stations for specimens of Leiocanthus quinquenudus sp. nov. and Leiocanthus satanicus sp. nov. Associated kinorhynch fauna at each station is also provided." httpUri="http://table.plazi.org/id/16AEA8C8977FFF9FFF5401A6F912FD62" pageId="10" pageNumber="325" tableUuid="16AEA8C8977FFF9FFF5401A6F912FD62">Table 1</tableCitation>
|
||
) at
|
||
<elevation id="C9FC1F739774FF94FC9B07FAF9B7FB48" box="[856,986,1229,1256]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="10" pageNumber="325" unit="m" value="59.0">
|
||
<quantity id="852955A59774FF94FC9B07FAF9FCFB47" box="[856,913,1229,1256]" metricMagnitude="1" metricUnit="m" metricValue="5.9" pageId="10" pageNumber="325" unit="m" value="59.0">59 m</quantity>
|
||
depth
|
||
</elevation>
|
||
; mounted in
|
||
<collectorName id="EF249D969774FF94FBA907FAFF7AFB47" box="[1130,1303,1229,1256]" pageId="10" pageNumber="325">Fluoromount G</collectorName>
|
||
<superScript id="B5A455089774FF94FAD407FAFF4EFB73" attach="left" box="[1303,1315,1229,1244]" fontSize="6" pageId="10" pageNumber="325">®</superScript>
|
||
, deposited at the
|
||
<collectionCode id="24C060859774FF94FF2307C5FB5BFAA3" box="[224,310,1266,1292]" pageId="10" pageNumber="325">NHMD</collectionCode>
|
||
under catalogue number: NHMD-915202
|
||
</materialsCitation>
|
||
.
|
||
<typeStatus id="9D6A46E29774FF94FCE507C5F9FBFAA3" box="[806,918,1266,1292]" pageId="10" pageNumber="325" type="paratype">Paratypes</typeStatus>
|
||
,
|
||
<specimenCount id="54D733C99774FF94FC6007C6FE71FAA4" box="[931,1052,1265,1291]" pageId="10" pageNumber="325" type="adult">three adult</specimenCount>
|
||
males and
|
||
<materialsCitation id="F2B9F21D9774FF94FB6207C6F935FAFB" ID-GBIF-Occurrence="3458066302" collectionCode="NHMD" pageId="10" pageNumber="325" specimenCount="1" specimenCount-adult="1" stateProvince="Mexico" typeStatus="holotype">
|
||
<specimenCount id="54D733C99774FF94FB6207C6FF6EFAA4" box="[1185,1283,1265,1291]" pageId="10" pageNumber="325" type="adult">six adult</specimenCount>
|
||
females, one of the females collected at the same station as the
|
||
<typeStatus id="9D6A46E29774FF94FD200622F925FA80" box="[739,840,1301,1327]" pageId="10" pageNumber="325" type="holotype">holotype</typeStatus>
|
||
, remaining specimens collected at different stations (
|
||
<tableCitation id="0F53CDFB9774FF94FF5C060EFA9CFAFB" box="[159,241,1337,1364]" captionStart="TABLE 1" captionStartId="1.[151,239,657,681]" captionTargetBox="[166,1409,732,1986]" captionText="TABLE 1. Summary of data on stations for specimens of Leiocanthus quinquenudus sp. nov. and Leiocanthus satanicus sp. nov. Associated kinorhynch fauna at each station is also provided." httpUri="http://table.plazi.org/id/16AEA8C8977FFF9FFF5401A6F912FD62" pageId="10" pageNumber="325" tableUuid="16AEA8C8977FFF9FFF5401A6F912FD62">Table 1</tableCitation>
|
||
), mounted in Fluoromount G
|
||
<superScript id="B5A455089774FF94FDF4060EF82EFAE7" attach="left" box="[567,579,1337,1352]" fontSize="6" pageId="10" pageNumber="325">®</superScript>
|
||
and deposited at
|
||
<collectionCode id="24C060859774FF94FCC1060DF935FAFB" box="[770,856,1338,1364]" pageId="10" pageNumber="325">NHMD</collectionCode>
|
||
</materialsCitation>
|
||
under catalogue numbers: NHMD-915203–915211. Two additional specimens,
|
||
<materialsCitation id="F2B9F21D9774FF94FE130669FBB2FA33" ID-GBIF-Occurrence="3458066304" collectionCode="SCL, MVS" pageId="10" pageNumber="325" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Mexico" typeStatus="paratype">
|
||
<specimenCount id="54D733C99774FF94FE130669F855FAD8" box="[464,568,1373,1400]" pageId="10" pageNumber="325" type="male">one male</specimenCount>
|
||
and
|
||
<specimenCount id="54D733C99774FF94FDB70669F898FAD8" box="[628,757,1373,1400]" pageId="10" pageNumber="325" type="female">one female</specimenCount>
|
||
, mounted for SEM and deposited in the personal reference collections of
|
||
<collectionCode id="24C060859774FF94FEF506B5FB05FA33" box="[310,360,1410,1436]" country="USA" lsid="urn:lsid:biocol.org:col:14865" name="St. Cloud State University" pageId="10" pageNumber="325" type="Herbarium">SCL</collectionCode>
|
||
and
|
||
<collectionCode id="24C060859774FF94FE5C06B5FBB2FA33" box="[415,479,1410,1436]" pageId="10" pageNumber="325">MVS</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0ACBABCB9774FF94FF040692FB0EF970" pageId="10" pageNumber="325" type="diagnosis">
|
||
<paragraph id="426EF8409774FF94FF040692FB0EF970" blockId="10.[151,1437,1193,1976]" pageId="10" pageNumber="325">
|
||
<emphasis id="70A524529774FF94FF040692FB2FFA10" bold="true" box="[199,322,1445,1471]" pageId="10" pageNumber="325">Diagnosis.</emphasis>
|
||
<taxonomicName id="85D183C39774FF94FE880692FBB9FA10" box="[331,468,1445,1471]" class="Allomalorhagida" family="Pycnophyidae" genus="Leiocanthus" kingdom="Animalia" pageId="10" pageNumber="325" phylum="Kinorhyncha" rank="genus">
|
||
<emphasis id="70A524529774FF94FE880692FBB9FA10" box="[331,468,1445,1471]" italics="true" pageId="10" pageNumber="325">Leiocanthus</emphasis>
|
||
</taxonomicName>
|
||
with middorsal cuticular elevations on segments 1–6. Unpaired paradorsal setae on segments 4 and 6; paired setae in subdorsal position on segments 2, 4, 6 and 8, in laterodorsal position on segments 3, 5, 7 and 9, paralateral position on segment 1, lateroventral position on segments 2, 4, 6, 8 and 10 (two pairs in the latter), ventrolateral position on segments 1 and 5, ventromedial position on segments 4–7 and 9, and paraventral position on segments 3 and 8. Males with sexually dimorphic ventromedial tubes on segment 2, females with setae instead. Lateral terminal spines absent, minute and bulbous protuberances instead. Anterolateral margins of the tergal plate of segment 1 extended into horn-like, medially recurved, distally pointed projections; anterior margin of tergal plate strongly denticulated, posteriorly followed by a crenulated area; cuticular wrinkles at the central region of the tergal plate.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0ACBABCB9774FF94FF0405DDFE5AF8C0" pageId="10" pageNumber="325" type="etymology">
|
||
<paragraph id="426EF8409774FF94FF0405DDFE5AF8C0" blockId="10.[151,1437,1193,1976]" pageId="10" pageNumber="325">
|
||
<emphasis id="70A524529774FF94FF0405DDFB23F8AB" bold="true" box="[199,334,1770,1796]" pageId="10" pageNumber="325">Etymology.</emphasis>
|
||
The specific epithet of the species refers to the Latin name Satâna, derived from the Hebraic הַשָּׂטָן (ha-shatán), which is a negative entity of the Abrahamic religions that seduces humanity to commit sin. Satan has been traditionally represented in graphic and literary arts as a fallen angel with conspicuous horns, which resembles the recurved anterolateral margins of the segment 1 tergal plate of the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0ACBABCB9774FF93FF04044EFBCCF83A" lastPageId="13" lastPageNumber="328" pageId="10" pageNumber="325" type="description">
|
||
<paragraph id="426EF8409774FF94FF04044EF82EF818" blockId="10.[151,1437,1193,1976]" pageId="10" pageNumber="325">
|
||
<emphasis id="70A524529774FF94FF04044EFB34F83C" bold="true" box="[199,345,1913,1939]" pageId="10" pageNumber="325">Description.</emphasis>
|
||
See
|
||
<tableCitation id="0F53CDFB9774FF94FE52044EFB89F83B" box="[401,484,1913,1940]" captionStart="TABLE 4" captionStartId="11.[151,239,151,175]" captionTargetPageId="11" captionText="TABLE 4. Measurements (in μm) of nine adult specimens of Leiocanthus satanicus sp. nov. (six females and three males) from the Gulf of Mexico. Abbreviations: MSW-3, maximum sternal width (measured at segment 3); n, number of measured specimens; S, segment length (followed by number of corresponding segment); SD, standard deviation; SSW, standard sternal width (measured at segment 10); TL, trunk length." httpUri="http://table.plazi.org/id/16AEA8C89775FF95FF5403A0F90CFEB4" pageId="10" pageNumber="325" tableUuid="16AEA8C89775FF95FF5403A0F90CFEB4">Table 4</tableCitation>
|
||
for measurements and dimensions and
|
||
<tableCitation id="0F53CDFB9774FF94FC66044EF994F83B" box="[933,1017,1913,1940]" captionStart="TABLE 5" captionStartId="11.[151,239,1017,1041]" captionTargetPageId="11" captionText="TABLE 5. Summary of nature and arrangement of cuticular elevations, tubes, setae, sensory spots, nephridiopores and glandular cell outlets in Leiocanthus satanicus sp. nov. Abbreviations: ce, cuticular elevation; gco, glandular cell outlet; LD, laterodorsal; LV, lateroventral; MD, middorsal; ne, nephridiopore; PD, paradorsal; PL, paralateral; ps, penile spine; PV, paraventral; SD, subdorsal; se, seta; ss, sensory spot; ss3, type 3 sensory spot; tu, tube; VL, ventrolateral; VM, ventromedial; * indicates unpaired structures; ♂/♀ indicate the presence of sexually dimorphic structures; brackets indicate deviations from the bilateral pattern in some specimens." httpUri="http://table.plazi.org/id/16AEA8C89775FF95FF5400CEF91CFB6A" pageId="10" pageNumber="325" tableUuid="16AEA8C89775FF95FF5400CEF91CFB6A">Table 5</tableCitation>
|
||
for summary of seta, tube, glandular cell outlet, and sensory spot positions.
|
||
</paragraph>
|
||
<caption id="16AEA8C89775FF95FF5403A0F90CFEB4" ID-Table-UUID="16AEA8C89775FF95FF5403A0F90CFEB4" httpUri="http://table.plazi.org/id/16AEA8C89775FF95FF5403A0F90CFEB4" pageId="11" pageNumber="326" startId="11.[151,239,151,175]" targetBox="[166,1245,299,968]" targetIsTable="true" targetPageId="11">
|
||
<paragraph id="426EF8409775FF95FF5403A0F90CFEB4" blockId="11.[151,1436,151,284]" pageId="11" pageNumber="326">
|
||
<emphasis id="70A524529775FF95FF5403A0FB60FF00" bold="true" box="[151,269,151,175]" pageId="11" pageNumber="326">TABLE 4.</emphasis>
|
||
Measurements (in μm) of nine adult specimens of
|
||
<taxonomicName id="85D183C39775FF95FC8903A0FE51FF00" authority="Cepeda & Sánchez & Sørensen & Landers, 2022" authorityName="Cepeda & Sánchez & Sørensen & Landers" authorityYear="2022" box="[842,1084,151,175]" class="Allomalorhagida" family="Pycnophyidae" genus="Leiocanthus" kingdom="Animalia" pageId="11" pageNumber="326" phylum="Kinorhyncha" rank="species" species="satanicus" status="sp. nov.">
|
||
<emphasis id="70A524529775FF95FC8903A0FE51FF00" box="[842,1084,151,175]" italics="true" pageId="11" pageNumber="326">Leiocanthus satanicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="70A524529775FF95FB8603AFFEF3FF1F" bold="true" box="[1093,1182,152,176]" pageId="11" pageNumber="326">
|
||
<taxonomicNameLabel id="6B9699299775FF95FB8603AFFEF3FF1F" box="[1093,1182,152,176]" pageId="11" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(six females and three males) from the Gulf of Mexico. Abbreviations: MSW-3, maximum sternal width (measured at segment 3);
|
||
<emphasis id="70A524529775FF95FAD7038BFF4CFF7C" box="[1300,1313,188,211]" italics="true" pageId="11" pageNumber="326">n</emphasis>
|
||
, number of measured specimens; S, segment length (followed by number of corresponding segment); SD, standard deviation; SSW, standard sternal width (measured at segment 10); TL, trunk length.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="426EF8409775FF95FF65021CFEA9FC67" pageId="11" pageNumber="326">
|
||
<table id="30D10AE097750061FF65021CFEB0FC67" box="[166,1245,299,968]" gridcols="4" gridrows="17" pageId="11" pageNumber="326">
|
||
<tr id="FCE1FA0297750061FF65021CFEB0FEEB" box="[166,1245,299,324]" gridrow="0" pageId="11" pageNumber="326">
|
||
<th id="BF30937E97750061FF65021CFB2AFEEB" box="[166,327,299,324]" gridcol="0" gridrow="0" pageId="11" pageNumber="326">
|
||
<emphasis id="70A524529775FF95FF65021CFB78FEEC" bold="true" box="[166,277,299,323]" pageId="11" pageNumber="326">Character</emphasis>
|
||
</th>
|
||
<th id="BF30937E97750061FEBA021CFBB1FEEB" box="[377,476,299,324]" gridcol="1" gridrow="0" pageId="11" pageNumber="326">
|
||
<emphasis id="70A524529775FF95FEBA021BFBB1FEEB" bold="true" box="[377,476,300,324]" pageId="11" pageNumber="326">Holotype</emphasis>
|
||
</th>
|
||
<th id="BF30937E97750061FD23021CF932FEEB" box="[736,863,299,324]" gridcol="2" gridrow="0" pageId="11" pageNumber="326">
|
||
<emphasis id="70A524529775FF95FD23021BF948FEEB" bold="true" box="[736,805,300,324]" pageId="11" pageNumber="326">Range</emphasis>
|
||
</th>
|
||
<th id="BF30937E97750061FB85021CFEB0FEEB" box="[1094,1245,299,324]" gridcol="3" gridrow="0" pageId="11" pageNumber="326">
|
||
<emphasis id="70A524529775FF95FB85021BFEB0FEEC" bold="true" box="[1094,1245,299,324]" pageId="11" pageNumber="326">
|
||
Mean (SD;
|
||
<emphasis id="70A524529775FF95FB06021BFEB9FEEC" bold="true" box="[1221,1236,300,323]" italics="true" pageId="11" pageNumber="326">n</emphasis>
|
||
)
|
||
</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF650263FEB0FEC3" box="[166,1245,340,364]" gridrow="1" pageId="11" pageNumber="326">
|
||
<th id="BF30937E97750061FF650263FB2AFEC3" box="[166,327,340,364]" gridcol="0" gridrow="1" pageId="11" pageNumber="326">TL</th>
|
||
<td id="BF30937E97750061FEBA0263FBB1FEC3" box="[377,476,340,364]" gridcol="1" gridrow="1" pageId="11" pageNumber="326">585.7</td>
|
||
<td id="BF30937E97750061FD230263F932FEC3" box="[736,863,340,364]" gridcol="2" gridrow="1" pageId="11" pageNumber="326">573.0–635.5</td>
|
||
<td id="BF30937E97750061FB850263FEB0FEC3" box="[1094,1245,340,364]" gridcol="3" gridrow="1" pageId="11" pageNumber="326">610.7 (21.2; 9)</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF65024BFEB0FE3B" box="[166,1245,380,404]" gridrow="2" pageId="11" pageNumber="326">
|
||
<th id="BF30937E97750061FF65024BFB2AFE3B" box="[166,327,380,404]" gridcol="0" gridrow="2" pageId="11" pageNumber="326">MSW-3</th>
|
||
<td id="BF30937E97750061FEBA024BFBB1FE3B" box="[377,476,380,404]" gridcol="1" gridrow="2" pageId="11" pageNumber="326">156.9</td>
|
||
<td id="BF30937E97750061FD23024BF932FE3B" box="[736,863,380,404]" gridcol="2" gridrow="2" pageId="11" pageNumber="326">133.5–156.9</td>
|
||
<td id="BF30937E97750061FB85024BFEB0FE3B" box="[1094,1245,380,404]" gridcol="3" gridrow="2" pageId="11" pageNumber="326">143.9 (7.9; 9)</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF650293FEB0FE12" box="[166,1245,420,445]" gridrow="3" pageId="11" pageNumber="326">
|
||
<th id="BF30937E97750061FF650293FB2AFE12" box="[166,327,420,445]" gridcol="0" gridrow="3" pageId="11" pageNumber="326">MSW-3/TL (%)</th>
|
||
<td id="BF30937E97750061FEBA0293FBB1FE12" box="[377,476,420,445]" gridcol="1" gridrow="3" pageId="11" pageNumber="326">26.8</td>
|
||
<td id="BF30937E97750061FD230293F932FE12" box="[736,863,420,445]" gridcol="2" gridrow="3" pageId="11" pageNumber="326">21.8–26.8</td>
|
||
<td id="BF30937E97750061FB850293FEB0FE12" box="[1094,1245,420,445]" gridcol="3" gridrow="3" pageId="11" pageNumber="326">23.6 (1.7; 9)</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF6502FAFEB0FE4A" box="[166,1245,461,485]" gridrow="4" pageId="11" pageNumber="326">
|
||
<th id="BF30937E97750061FF6502FAFB2AFE4A" box="[166,327,461,485]" gridcol="0" gridrow="4" pageId="11" pageNumber="326">SSW</th>
|
||
<td id="BF30937E97750061FEBA02FAFBB1FE4A" box="[377,476,461,485]" gridcol="1" gridrow="4" pageId="11" pageNumber="326">118.3</td>
|
||
<td id="BF30937E97750061FD2302FAF932FE4A" box="[736,863,461,485]" gridcol="2" gridrow="4" pageId="11" pageNumber="326">108.2–132.7</td>
|
||
<td id="BF30937E97750061FB8502FAFEB0FE4A" box="[1094,1245,461,485]" gridcol="3" gridrow="4" pageId="11" pageNumber="326">120.1 (6.7; 9)</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF6502C2FEB0FDA2" box="[166,1245,501,525]" gridrow="5" pageId="11" pageNumber="326">
|
||
<th id="BF30937E97750061FF6502C2FB2AFDA2" box="[166,327,501,525]" gridcol="0" gridrow="5" pageId="11" pageNumber="326">SSW/TL (%)</th>
|
||
<td id="BF30937E97750061FEBA02C2FBB1FDA2" box="[377,476,501,525]" gridcol="1" gridrow="5" pageId="11" pageNumber="326">20.2</td>
|
||
<td id="BF30937E97750061FD2302C2F932FDA2" box="[736,863,501,525]" gridcol="2" gridrow="5" pageId="11" pageNumber="326">18.6–21.8</td>
|
||
<td id="BF30937E97750061FB8502C2FEB0FDA2" box="[1094,1245,501,525]" gridcol="3" gridrow="5" pageId="11" pageNumber="326">19.7 (1.0; 9)</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF65012AFEB0FD9A" box="[166,1245,541,565]" gridrow="6" pageId="11" pageNumber="326">
|
||
<th id="BF30937E97750061FF65012AFB2AFD9A" box="[166,327,541,565]" gridcol="0" gridrow="6" pageId="11" pageNumber="326">S1</th>
|
||
<td id="BF30937E97750061FEBA012AFBB1FD9A" box="[377,476,541,565]" gridcol="1" gridrow="6" pageId="11" pageNumber="326">103.5</td>
|
||
<td id="BF30937E97750061FD23012AF932FD9A" box="[736,863,541,565]" gridcol="2" gridrow="6" pageId="11" pageNumber="326">99.8–122.5</td>
|
||
<td id="BF30937E97750061FB85012AFEB0FD9A" box="[1094,1245,541,565]" gridcol="3" gridrow="6" pageId="11" pageNumber="326">106.4 (8.2; 9)</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF650172FEB0FDF1" box="[166,1245,581,606]" gridrow="7" pageId="11" pageNumber="326">
|
||
<th id="BF30937E97750061FF650172FB2AFDF1" box="[166,327,581,606]" gridcol="0" gridrow="7" pageId="11" pageNumber="326">S2</th>
|
||
<td id="BF30937E97750061FEBA0172FBB1FDF1" box="[377,476,581,606]" gridcol="1" gridrow="7" pageId="11" pageNumber="326">57.9</td>
|
||
<td id="BF30937E97750061FD230172F932FDF1" box="[736,863,581,606]" gridcol="2" gridrow="7" pageId="11" pageNumber="326">54.5– 67.5</td>
|
||
<td id="BF30937E97750061FB850172FEB0FDF1" box="[1094,1245,581,606]" gridcol="3" gridrow="7" pageId="11" pageNumber="326">59.6 (4.7; 9)</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF65015AFEB0FD29" box="[166,1245,621,646]" gridrow="8" pageId="11" pageNumber="326">
|
||
<th id="BF30937E97750061FF65015AFB2AFD29" box="[166,327,621,646]" gridcol="0" gridrow="8" pageId="11" pageNumber="326">S3</th>
|
||
<td id="BF30937E97750061FEBA015AFBB1FD29" box="[377,476,621,646]" gridcol="1" gridrow="8" pageId="11" pageNumber="326">64.8</td>
|
||
<td id="BF30937E97750061FD23015AF932FD29" box="[736,863,621,646]" gridcol="2" gridrow="8" pageId="11" pageNumber="326">57.7–69.6</td>
|
||
<td id="BF30937E97750061FB85015AFEB0FD29" box="[1094,1245,621,646]" gridcol="3" gridrow="8" pageId="11" pageNumber="326">64.0 (4.0; 9)</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF6501A1FEB0FD01" box="[166,1245,662,686]" gridrow="9" pageId="11" pageNumber="326">
|
||
<th id="BF30937E97750061FF6501A1FB2AFD01" box="[166,327,662,686]" gridcol="0" gridrow="9" pageId="11" pageNumber="326">S4</th>
|
||
<td id="BF30937E97750061FEBA01A1FBB1FD01" box="[377,476,662,686]" gridcol="1" gridrow="9" pageId="11" pageNumber="326">69.6</td>
|
||
<td id="BF30937E97750061FD2301A1F932FD01" box="[736,863,662,686]" gridcol="2" gridrow="9" pageId="11" pageNumber="326">65.7–80.3</td>
|
||
<td id="BF30937E97750061FB8501A1FEB0FD01" box="[1094,1245,662,686]" gridcol="3" gridrow="9" pageId="11" pageNumber="326">71.3 (5.1; 9)</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF650189FEB0FD79" box="[166,1245,702,726]" gridrow="10" pageId="11" pageNumber="326">
|
||
<th id="BF30937E97750061FF650189FB2AFD79" box="[166,327,702,726]" gridcol="0" gridrow="10" pageId="11" pageNumber="326">S5</th>
|
||
<td id="BF30937E97750061FEBA0189FBB1FD79" box="[377,476,702,726]" gridcol="1" gridrow="10" pageId="11" pageNumber="326">70.1</td>
|
||
<td id="BF30937E97750061FD230189F932FD79" box="[736,863,702,726]" gridcol="2" gridrow="10" pageId="11" pageNumber="326">66.1–75.4</td>
|
||
<td id="BF30937E97750061FB850189FEB0FD79" box="[1094,1245,702,726]" gridcol="3" gridrow="10" pageId="11" pageNumber="326">70.8 (3.3; 9)</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF6501D1FEB0FD51" box="[166,1245,742,766]" gridrow="11" pageId="11" pageNumber="326">
|
||
<th id="BF30937E97750061FF6501D1FB2AFD51" box="[166,327,742,766]" gridcol="0" gridrow="11" pageId="11" pageNumber="326">S6</th>
|
||
<td id="BF30937E97750061FEBA01D1FBB1FD51" box="[377,476,742,766]" gridcol="1" gridrow="11" pageId="11" pageNumber="326">70.3</td>
|
||
<td id="BF30937E97750061FD2301D1F932FD51" box="[736,863,742,766]" gridcol="2" gridrow="11" pageId="11" pageNumber="326">70.3–76.6</td>
|
||
<td id="BF30937E97750061FB8501D1FEB0FD51" box="[1094,1245,742,766]" gridcol="3" gridrow="11" pageId="11" pageNumber="326">72.8 (2.4; 9)</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF650039FEB0FC88" box="[166,1245,782,807]" gridrow="12" pageId="11" pageNumber="326">
|
||
<th id="BF30937E97750061FF650039FB2AFC88" box="[166,327,782,807]" gridcol="0" gridrow="12" pageId="11" pageNumber="326">S7</th>
|
||
<td id="BF30937E97750061FEBA0039FBB1FC88" box="[377,476,782,807]" gridcol="1" gridrow="12" pageId="11" pageNumber="326">75.3</td>
|
||
<td id="BF30937E97750061FD230039F932FC88" box="[736,863,782,807]" gridcol="2" gridrow="12" pageId="11" pageNumber="326">68.1–77.2</td>
|
||
<td id="BF30937E97750061FB850039FEB0FC88" box="[1094,1245,782,807]" gridcol="3" gridrow="12" pageId="11" pageNumber="326">73.5 (3.1; 9)</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF650000FEB0FCE0" box="[166,1245,823,847]" gridrow="13" pageId="11" pageNumber="326">
|
||
<th id="BF30937E97750061FF650000FB2AFCE0" box="[166,327,823,847]" gridcol="0" gridrow="13" pageId="11" pageNumber="326">S8</th>
|
||
<td id="BF30937E97750061FEBA0000FBB1FCE0" box="[377,476,823,847]" gridcol="1" gridrow="13" pageId="11" pageNumber="326">86.5</td>
|
||
<td id="BF30937E97750061FD230000F932FCE0" box="[736,863,823,847]" gridcol="2" gridrow="13" pageId="11" pageNumber="326">71.4–86.5</td>
|
||
<td id="BF30937E97750061FB850000FEB0FCE0" box="[1094,1245,823,847]" gridcol="3" gridrow="13" pageId="11" pageNumber="326">77.6 (4.8; 9)</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF650068FEB0FCD8" box="[166,1245,863,887]" gridrow="14" pageId="11" pageNumber="326">
|
||
<th id="BF30937E97750061FF650068FB2AFCD8" box="[166,327,863,887]" gridcol="0" gridrow="14" pageId="11" pageNumber="326">S9</th>
|
||
<td id="BF30937E97750061FEBA0068FBB1FCD8" box="[377,476,863,887]" gridcol="1" gridrow="14" pageId="11" pageNumber="326">87.5</td>
|
||
<td id="BF30937E97750061FD230068F932FCD8" box="[736,863,863,887]" gridcol="2" gridrow="14" pageId="11" pageNumber="326">67.1–87.5</td>
|
||
<td id="BF30937E97750061FB850068FEB0FCD8" box="[1094,1245,863,887]" gridcol="3" gridrow="14" pageId="11" pageNumber="326">78.1 (7.0; 9)</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF6500B0FEB0FC30" box="[166,1245,903,927]" gridrow="15" pageId="11" pageNumber="326">
|
||
<th id="BF30937E97750061FF6500B0FB2AFC30" box="[166,327,903,927]" gridcol="0" gridrow="15" pageId="11" pageNumber="326">S10</th>
|
||
<td id="BF30937E97750061FEBA00B0FBB1FC30" box="[377,476,903,927]" gridcol="1" gridrow="15" pageId="11" pageNumber="326">111.5</td>
|
||
<td id="BF30937E97750061FD2300B0F932FC30" box="[736,863,903,927]" gridcol="2" gridrow="15" pageId="11" pageNumber="326">91.9–111.5</td>
|
||
<td id="BF30937E97750061FB8500B0FEB0FC30" box="[1094,1245,903,927]" gridcol="3" gridrow="15" pageId="11" pageNumber="326">100.8 (6.9; 9)</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF650098FEB0FC67" box="[166,1245,943,968]" gridrow="16" pageId="11" pageNumber="326">
|
||
<th id="BF30937E97750061FF650098FB2AFC67" box="[166,327,943,968]" gridcol="0" gridrow="16" pageId="11" pageNumber="326">S11</th>
|
||
<td id="BF30937E97750061FEBA0098FBB1FC67" box="[377,476,943,968]" gridcol="1" gridrow="16" pageId="11" pageNumber="326">32.0</td>
|
||
<td id="BF30937E97750061FD230098F932FC67" box="[736,863,943,968]" gridcol="2" gridrow="16" pageId="11" pageNumber="326">25.9–38.6</td>
|
||
<td id="BF30937E97750061FB850098FEB0FC67" box="[1094,1245,943,968]" gridcol="3" gridrow="16" pageId="11" pageNumber="326">31.8 (5.0; 9)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="16AEA8C89775FF95FF5400CEF91CFB6A" ID-Table-UUID="16AEA8C89775FF95FF5400CEF91CFB6A" httpUri="http://table.plazi.org/id/16AEA8C89775FF95FF5400CEF91CFB6A" pageId="11" pageNumber="326" startId="11.[151,239,1017,1041]" targetBox="[166,1407,1236,1704]" targetIsTable="true" targetPageId="11">
|
||
<paragraph id="426EF8409775FF95FF5400CEF91CFB6A" blockId="11.[151,1437,1017,1221]" pageId="11" pageNumber="326">
|
||
<emphasis id="70A524529775FF95FF5400CEFB66FBBE" bold="true" box="[151,267,1017,1041]" pageId="11" pageNumber="326">TABLE 5.</emphasis>
|
||
Summary of nature and arrangement of cuticular elevations, tubes, setae, sensory spots, nephridiopores and glandular cell outlets in
|
||
<taxonomicName id="85D183C39775FF95FE58072AF8E4FB9A" authority="Cepeda & Sánchez & Sørensen & Landers, 2022" authorityName="Cepeda & Sánchez & Sørensen & Landers" authorityYear="2022" box="[411,649,1053,1077]" class="Allomalorhagida" family="Pycnophyidae" genus="Leiocanthus" kingdom="Animalia" pageId="11" pageNumber="326" phylum="Kinorhyncha" rank="species" species="satanicus" status="sp. nov.">
|
||
<emphasis id="70A524529775FF95FE58072AF8E4FB9A" box="[411,649,1053,1077]" italics="true" pageId="11" pageNumber="326">Leiocanthus satanicus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="70A524529775FF95FD53072AF88BFB9A" bold="true" box="[656,742,1053,1077]" pageId="11" pageNumber="326">
|
||
<taxonomicNameLabel id="6B9699299775FF95FD53072AF88BFB9A" box="[656,742,1053,1077]" pageId="11" pageNumber="326" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Abbreviations: ce, cuticular elevation; gco, glandular cell outlet; LD, laterodorsal; LV, lateroventral; MD, middorsal; ne, nephridiopore; PD, paradorsal; PL, paralateral; ps, penile spine; PV, paraventral; SD, subdorsal; se, seta; ss, sensory spot; ss3, type 3 sensory spot; tu, tube; VL, ventrolateral; VM, ventromedial;
|
||
<superScript id="B5A455089775FF95FEF007B0FB56FB3A" attach="right" box="[307,315,1159,1173]" fontSize="6" pageId="11" pageNumber="326">*</superScript>
|
||
indicates unpaired structures; ♂/♀ indicate the presence of sexually dimorphic structures; brackets indicate deviations from the bilateral pattern in some specimens.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="426EF8409775FF95FF6507E3FE25F908" pageId="11" pageNumber="326">
|
||
<table id="30D10AE097750061FF6507E3FF12F907" box="[166,1407,1236,1704]" gridcols="10" gridrows="12" pageId="11" pageNumber="326">
|
||
<tr id="FCE1FA0297750061FF6507E3FF12FB42" box="[166,1407,1236,1261]" gridrow="0" pageId="11" pageNumber="326">
|
||
<th id="BF30937E97750061FF6507E3FB6EFB42" box="[166,259,1236,1261]" gridcol="0" gridrow="0" pageId="11" pageNumber="326">
|
||
<emphasis id="70A524529775FF95FF6507E3FB6EFB43" bold="true" box="[166,259,1236,1260]" pageId="11" pageNumber="326">Segment</emphasis>
|
||
</th>
|
||
<th id="BF30937E97750061FEF007E3FB30FB42" box="[307,349,1236,1261]" gridcol="1" gridrow="0" pageId="11" pageNumber="326">
|
||
<emphasis id="70A524529775FF95FEF007E2FB30FB42" bold="true" box="[307,349,1237,1261]" pageId="11" pageNumber="326">MD</emphasis>
|
||
</th>
|
||
<th id="BF30937E97750061FE4707E3FBADFB42" box="[388,448,1236,1261]" gridcol="2" gridrow="0" pageId="11" pageNumber="326">
|
||
<emphasis id="70A524529775FF95FE4707E2FBCBFB42" bold="true" box="[388,422,1237,1261]" pageId="11" pageNumber="326">PD</emphasis>
|
||
</th>
|
||
<th id="BF30937E97750061FE2A07E3F8FDFB42" box="[489,656,1236,1261]" gridcol="3" gridrow="0" pageId="11" pageNumber="326">
|
||
<emphasis id="70A524529775FF95FE2A07E3F864FB43" bold="true" box="[489,521,1236,1260]" pageId="11" pageNumber="326">SD</emphasis>
|
||
</th>
|
||
<th id="BF30937E97750061FD0507E3F960FB42" box="[710,781,1236,1261]" gridcol="4" gridrow="0" pageId="11" pageNumber="326">
|
||
<emphasis id="70A524529775FF95FD0507E2F884FB42" bold="true" box="[710,745,1237,1261]" pageId="11" pageNumber="326">LD</emphasis>
|
||
</th>
|
||
<th id="BF30937E97750061FCF607E3F907FB42" box="[821,874,1236,1261]" gridcol="5" gridrow="0" pageId="11" pageNumber="326">
|
||
<emphasis id="70A524529775FF95FCF607E2F938FB42" bold="true" box="[821,853,1237,1261]" pageId="11" pageNumber="326">PL</emphasis>
|
||
</th>
|
||
<th id="BF30937E97750061FC5707E3F9A5FB42" box="[916,968,1236,1261]" gridcol="6" gridrow="0" pageId="11" pageNumber="326">
|
||
<emphasis id="70A524529775FF95FC5707E2F9D8FB42" bold="true" box="[916,949,1237,1261]" pageId="11" pageNumber="326">LV</emphasis>
|
||
</th>
|
||
<th id="BF30937E97750061FC3A07E3FE25FB42" box="[1017,1096,1236,1261]" gridcol="7" gridrow="0" pageId="11" pageNumber="326">
|
||
<emphasis id="70A524529775FF95FC3A07E2FE71FB42" bold="true" box="[1017,1052,1237,1261]" pageId="11" pageNumber="326">VL</emphasis>
|
||
</th>
|
||
<th id="BF30937E97750061FBAE07E3FE86FB42" box="[1133,1259,1236,1261]" gridcol="8" gridrow="0" pageId="11" pageNumber="326">
|
||
<emphasis id="70A524529775FF95FBAE07E2FEFAFB42" bold="true" box="[1133,1175,1237,1261]" pageId="11" pageNumber="326">VM</emphasis>
|
||
</th>
|
||
<th id="BF30937E97750061FAE407E3FF12FB42" box="[1319,1407,1236,1261]" gridcol="9" gridrow="0" pageId="11" pageNumber="326">
|
||
<emphasis id="70A524529775FF95FAE407E2FF24FB42" bold="true" box="[1319,1353,1237,1261]" pageId="11" pageNumber="326">PV</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF6507CAFF12FABA" box="[166,1407,1277,1301]" gridrow="1" pageId="11" pageNumber="326" rowspan-6="1" rowspan-9="1">
|
||
<th id="BF30937E97750061FF6507CAFB6EFABA" box="[166,259,1277,1301]" gridcol="0" gridrow="1" pageId="11" pageNumber="326">1</th>
|
||
<td id="BF30937E97750061FEF007CAFB30FABA" box="[307,349,1277,1301]" gridcol="1" gridrow="1" pageId="11" pageNumber="326">ce</td>
|
||
<td id="BF30937E97750061FE4707CAFBADFABA" box="[388,448,1277,1301]" gridcol="2" gridrow="1" pageId="11" pageNumber="326">ss</td>
|
||
<td id="BF30937E97750061FE2A07CAF8FDFABA" box="[489,656,1277,1301]" gridcol="3" gridrow="1" pageId="11" pageNumber="326">gco, ss x2</td>
|
||
<td id="BF30937E97750061FD0507CAF960FABA" box="[710,781,1277,1301]" gridcol="4" gridrow="1" pageId="11" pageNumber="326">ss</td>
|
||
<td id="BF30937E97750061FCF607CAF907FABA" box="[821,874,1277,1301]" gridcol="5" gridrow="1" pageId="11" pageNumber="326">se, ss</td>
|
||
<td id="BF30937E97750061FC3A07CAFE25FABA" box="[1017,1096,1277,1301]" gridcol="7" gridrow="1" pageId="11" pageNumber="326">se, ss</td>
|
||
<td id="BF30937E97750061FBAE07CAFE86FABA" box="[1133,1259,1277,1301]" gridcol="8" gridrow="1" pageId="11" pageNumber="326">gco, ss</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF650612FF12FA91" box="[166,1407,1317,1342]" gridrow="2" pageId="11" pageNumber="326" rowspan-5="1" rowspan-7="1">
|
||
<th id="BF30937E97750061FF650612FB6EFA91" box="[166,259,1317,1342]" gridcol="0" gridrow="2" pageId="11" pageNumber="326">2</th>
|
||
<td id="BF30937E97750061FEF00612FB30FA91" box="[307,349,1317,1342]" gridcol="1" gridrow="2" pageId="11" pageNumber="326">ce</td>
|
||
<td id="BF30937E97750061FE470612FBADFA91" box="[388,448,1317,1342]" gridcol="2" gridrow="2" pageId="11" pageNumber="326">ss</td>
|
||
<td id="BF30937E97750061FE2A0612F8FDFA91" box="[489,656,1317,1342]" gridcol="3" gridrow="2" pageId="11" pageNumber="326">se, ss, gco, ss</td>
|
||
<td id="BF30937E97750061FD050612F960FA91" box="[710,781,1317,1342]" gridcol="4" gridrow="2" pageId="11" pageNumber="326">ss</td>
|
||
<td id="BF30937E97750061FC570612F9A5FA91" box="[916,968,1317,1342]" gridcol="6" gridrow="2" pageId="11" pageNumber="326">se</td>
|
||
<td id="BF30937E97750061FBAE0612FE86FA91" box="[1133,1259,1317,1342]" gridcol="8" gridrow="2" pageId="11" pageNumber="326">tu♂/se♀, ss</td>
|
||
<td id="BF30937E97750061FAE40612FF12FA91" box="[1319,1407,1317,1342]" gridcol="9" gridrow="2" pageId="11" pageNumber="326">gco</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF650679FF12FAC9" box="[166,1407,1358,1382]" gridrow="3" pageId="11" pageNumber="326" rowspan-5="1" rowspan-6="1" rowspan-7="1">
|
||
<th id="BF30937E97750061FF650679FB6EFAC9" box="[166,259,1358,1382]" gridcol="0" gridrow="3" pageId="11" pageNumber="326">3</th>
|
||
<td id="BF30937E97750061FEF00679FB30FAC9" box="[307,349,1358,1382]" gridcol="1" gridrow="3" pageId="11" pageNumber="326">ce</td>
|
||
<td id="BF30937E97750061FE470679FBADFAC9" box="[388,448,1358,1382]" gridcol="2" gridrow="3" pageId="11" pageNumber="326">ss</td>
|
||
<td id="BF30937E97750061FE2A0679F8FDFAC9" box="[489,656,1358,1382]" gridcol="3" gridrow="3" pageId="11" pageNumber="326">ss x2, gco,</td>
|
||
<td id="BF30937E97750061FD050679F960FAC9" box="[710,781,1358,1382]" gridcol="4" gridrow="3" pageId="11" pageNumber="326">(se), ss</td>
|
||
<td id="BF30937E97750061FBAE0679FE86FAC9" box="[1133,1259,1358,1382]" gridcol="8" gridrow="3" pageId="11" pageNumber="326">ss</td>
|
||
<td id="BF30937E97750061FAE40679FF12FAC9" box="[1319,1407,1358,1382]" gridcol="9" gridrow="3" pageId="11" pageNumber="326">gco, se</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF650642FF12FA21" box="[166,1407,1397,1422]" gridrow="4" pageId="11" pageNumber="326" rowspan-5="1" rowspan-7="1">
|
||
<th id="BF30937E97750061FF650642FB6EFA21" box="[166,259,1397,1422]" gridcol="0" gridrow="4" pageId="11" pageNumber="326">4</th>
|
||
<td id="BF30937E97750061FEF00642FB30FA21" box="[307,349,1397,1422]" gridcol="1" gridrow="4" pageId="11" pageNumber="326">ce</td>
|
||
<td id="BF30937E97750061FE470642FBADFA21" box="[388,448,1397,1422]" gridcol="2" gridrow="4" pageId="11" pageNumber="326">ss, se*</td>
|
||
<td id="BF30937E97750061FE2A0642F8FDFA21" box="[489,656,1397,1422]" gridcol="3" gridrow="4" pageId="11" pageNumber="326">ss, se, gco, ss</td>
|
||
<td id="BF30937E97750061FD050642F960FA21" box="[710,781,1397,1422]" gridcol="4" gridrow="4" pageId="11" pageNumber="326">ss</td>
|
||
<td id="BF30937E97750061FC570642F9A5FA21" box="[916,968,1397,1422]" gridcol="6" gridrow="4" pageId="11" pageNumber="326">se</td>
|
||
<td id="BF30937E97750061FBAE0642FE86FA21" box="[1133,1259,1397,1422]" gridcol="8" gridrow="4" pageId="11" pageNumber="326">ss, se</td>
|
||
<td id="BF30937E97750061FAE40642FF12FA21" box="[1319,1407,1397,1422]" gridcol="9" gridrow="4" pageId="11" pageNumber="326">gco</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF6506A9FF12FA19" box="[166,1407,1438,1462]" gridrow="5" pageId="11" pageNumber="326" rowspan-5="1" rowspan-6="1">
|
||
<th id="BF30937E97750061FF6506A9FB6EFA19" box="[166,259,1438,1462]" gridcol="0" gridrow="5" pageId="11" pageNumber="326">5</th>
|
||
<td id="BF30937E97750061FEF006A9FB30FA19" box="[307,349,1438,1462]" gridcol="1" gridrow="5" pageId="11" pageNumber="326">ce</td>
|
||
<td id="BF30937E97750061FE4706A9FBADFA19" box="[388,448,1438,1462]" gridcol="2" gridrow="5" pageId="11" pageNumber="326">ss</td>
|
||
<td id="BF30937E97750061FE2A06A9F8FDFA19" box="[489,656,1438,1462]" gridcol="3" gridrow="5" pageId="11" pageNumber="326">ss, gco, ss</td>
|
||
<td id="BF30937E97750061FD0506A9F960FA19" box="[710,781,1438,1462]" gridcol="4" gridrow="5" pageId="11" pageNumber="326">(se), ss</td>
|
||
<td id="BF30937E97750061FC3A06A9FE25FA19" box="[1017,1096,1438,1462]" gridcol="7" gridrow="5" pageId="11" pageNumber="326">se</td>
|
||
<td id="BF30937E97750061FBAE06A9FE86FA19" box="[1133,1259,1438,1462]" gridcol="8" gridrow="5" pageId="11" pageNumber="326">se, ss</td>
|
||
<td id="BF30937E97750061FAE406A9FF12FA19" box="[1319,1407,1438,1462]" gridcol="9" gridrow="5" pageId="11" pageNumber="326">gco</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF6506F2FF12FA70" box="[166,1407,1477,1503]" gridrow="6" pageId="11" pageNumber="326" rowspan-5="1" rowspan-7="1">
|
||
<th id="BF30937E97750061FF6506F2FB6EFA70" box="[166,259,1477,1503]" gridcol="0" gridrow="6" pageId="11" pageNumber="326">6</th>
|
||
<td id="BF30937E97750061FEF006F2FB30FA70" box="[307,349,1477,1503]" gridcol="1" gridrow="6" pageId="11" pageNumber="326">ce</td>
|
||
<td id="BF30937E97750061FE4706F2FBADFA70" box="[388,448,1477,1503]" gridcol="2" gridrow="6" pageId="11" pageNumber="326">ss, se*</td>
|
||
<td id="BF30937E97750061FE2A06F2F8FDFA70" box="[489,656,1477,1503]" gridcol="3" gridrow="6" pageId="11" pageNumber="326">ss, se, gco, ss</td>
|
||
<td id="BF30937E97750061FD0506F2F960FA70" box="[710,781,1477,1503]" gridcol="4" gridrow="6" pageId="11" pageNumber="326">ss</td>
|
||
<td id="BF30937E97750061FC5706F2F9A5FA70" box="[916,968,1477,1503]" gridcol="6" gridrow="6" pageId="11" pageNumber="326">se</td>
|
||
<td id="BF30937E97750061FBAE06F2FE86FA70" box="[1133,1259,1477,1503]" gridcol="8" gridrow="6" pageId="11" pageNumber="326">ss,(se)</td>
|
||
<td id="BF30937E97750061FAE406F2FF12FA70" box="[1319,1407,1477,1503]" gridcol="9" gridrow="6" pageId="11" pageNumber="326">gco</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF6506D8FF12F9A8" box="[166,1407,1519,1543]" gridrow="7" pageId="11" pageNumber="326" rowspan-1="1" rowspan-2="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
|
||
<th id="BF30937E97750061FF6506D8FB6EF9A8" box="[166,259,1519,1543]" gridcol="0" gridrow="7" pageId="11" pageNumber="326">7</th>
|
||
<td id="BF30937E97750061FE2A06D8F8FDF9A8" box="[489,656,1519,1543]" gridcol="3" gridrow="7" pageId="11" pageNumber="326">ss, gco, ss</td>
|
||
<td id="BF30937E97750061FD0506D8F960F9A8" box="[710,781,1519,1543]" gridcol="4" gridrow="7" pageId="11" pageNumber="326">ss, (se)</td>
|
||
<td id="BF30937E97750061FBAE06D8FE86F9A8" box="[1133,1259,1519,1543]" gridcol="8" gridrow="7" pageId="11" pageNumber="326">se, ss</td>
|
||
<td id="BF30937E97750061FAE406D8FF12F9A8" box="[1319,1407,1519,1543]" gridcol="9" gridrow="7" pageId="11" pageNumber="326">gco</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF650520FF12F980" box="[166,1407,1559,1583]" gridrow="8" pageId="11" pageNumber="326" rowspan-1="1" rowspan-2="1" rowspan-5="1" rowspan-7="1">
|
||
<th id="BF30937E97750061FF650520FB6EF980" box="[166,259,1559,1583]" gridcol="0" gridrow="8" pageId="11" pageNumber="326">8</th>
|
||
<td id="BF30937E97750061FE2A0520F8FDF980" box="[489,656,1559,1583]" gridcol="3" gridrow="8" pageId="11" pageNumber="326">ss x2, gco, se, ss</td>
|
||
<td id="BF30937E97750061FD050520F960F980" box="[710,781,1559,1583]" gridcol="4" gridrow="8" pageId="11" pageNumber="326">ss</td>
|
||
<td id="BF30937E97750061FC570520F9A5F980" box="[916,968,1559,1583]" gridcol="6" gridrow="8" pageId="11" pageNumber="326">se</td>
|
||
<td id="BF30937E97750061FBAE0520FE86F980" box="[1133,1259,1559,1583]" gridcol="8" gridrow="8" pageId="11" pageNumber="326">ss</td>
|
||
<td id="BF30937E97750061FAE40520FF12F980" box="[1319,1407,1559,1583]" gridcol="9" gridrow="8" pageId="11" pageNumber="326">gco, (se)</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF650508FF12F9F8" box="[166,1407,1599,1623]" gridrow="9" pageId="11" pageNumber="326" rowspan-1="1" rowspan-2="1" rowspan-5="1" rowspan-9="1">
|
||
<th id="BF30937E97750061FF650508FB6EF9F8" box="[166,259,1599,1623]" gridcol="0" gridrow="9" pageId="11" pageNumber="326">9</th>
|
||
<td id="BF30937E97750061FE2A0508F8FDF9F8" box="[489,656,1599,1623]" gridcol="3" gridrow="9" pageId="11" pageNumber="326">ss x2, gco, ss</td>
|
||
<td id="BF30937E97750061FD050508F960F9F8" box="[710,781,1599,1623]" gridcol="4" gridrow="9" pageId="11" pageNumber="326">se, ss</td>
|
||
<td id="BF30937E97750061FC570508F9A5F9F8" box="[916,968,1599,1623]" gridcol="6" gridrow="9" pageId="11" pageNumber="326">ne</td>
|
||
<td id="BF30937E97750061FC3A0508FE25F9F8" box="[1017,1096,1599,1623]" gridcol="7" gridrow="9" pageId="11" pageNumber="326">ss</td>
|
||
<td id="BF30937E97750061FBAE0508FE86F9F8" box="[1133,1259,1599,1623]" gridcol="8" gridrow="9" pageId="11" pageNumber="326">se, gco, ss</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF650550FF12F92F" box="[166,1407,1639,1664]" gridrow="10" pageId="11" pageNumber="326" rowspan-1="1" rowspan-2="1" rowspan-5="1" rowspan-9="1">
|
||
<th id="BF30937E97750061FF650550FB6EF92F" box="[166,259,1639,1664]" gridcol="0" gridrow="10" pageId="11" pageNumber="326">10</th>
|
||
<td id="BF30937E97750061FE2A0550F8FDF92F" box="[489,656,1639,1664]" gridcol="3" gridrow="10" pageId="11" pageNumber="326">ss, gco</td>
|
||
<td id="BF30937E97750061FD050550F960F92F" box="[710,781,1639,1664]" gridcol="4" gridrow="10" pageId="11" pageNumber="326">ss</td>
|
||
<td id="BF30937E97750061FC570550F9A5F92F" box="[916,968,1639,1664]" gridcol="6" gridrow="10" pageId="11" pageNumber="326">se x2</td>
|
||
<td id="BF30937E97750061FC3A0550FE25F92F" box="[1017,1096,1639,1664]" gridcol="7" gridrow="10" pageId="11" pageNumber="326">ss</td>
|
||
<td id="BF30937E97750061FBAE0550FE86F92F" box="[1133,1259,1639,1664]" gridcol="8" gridrow="10" pageId="11" pageNumber="326">gco, ss</td>
|
||
</tr>
|
||
<tr id="FCE1FA0297750061FF6505A7FF12F907" box="[166,1407,1680,1704]" gridrow="11" pageId="11" pageNumber="326" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-8="1" rowspan-9="1">
|
||
<th id="BF30937E97750061FF6505A7FB6EF907" box="[166,259,1680,1704]" gridcol="0" gridrow="11" pageId="11" pageNumber="326">11</th>
|
||
<td id="BF30937E97750061FE2A05A7F8FDF907" box="[489,656,1680,1704]" gridcol="3" gridrow="11" pageId="11" pageNumber="326">gco, ss3</td>
|
||
<td id="BF30937E97750061FC3A05A7FE25F907" box="[1017,1096,1680,1704]" gridcol="7" gridrow="11" pageId="11" pageNumber="326">ps x2♂</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="426EF8409775FF95FF0405E0F8ABF80A" blockId="11.[151,1436,1751,2029]" pageId="11" pageNumber="326">
|
||
<emphasis id="70A524529775FF95FF0405E0FB66F95E" box="[199,267,1751,1777]" italics="true" pageId="11" pageNumber="326">Head.</emphasis>
|
||
Only
|
||
<specimenCount id="54D733C99775FF95FE8C05E0FB99F95E" box="[335,500,1751,1777]" pageId="11" pageNumber="326" type="generic">two specimens</specimenCount>
|
||
had the head everted, hence exact details on the morphology and arrangement of the mouth cone and introvert structures cannot be completely provided. Internal rings of mouth cone not observed. Ring 00 of mouth cone with nine, equally sized outer oral styles, each one composed of a single, flexible unit, wider at the base, bearing a fringed basal sheath, with a lateral incision about one third from the proximal end, progressively tapering towards a distally rounded tip (
|
||
<figureCitation id="DAEAE4C59775FF95FD990450F8D9F82D" box="[602,692,1895,1922]" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="11" pageNumber="326">Fig. 6C</figureCitation>
|
||
). Outer oral styles located anterior to each introvert sector, except in the middorsal sector 6 where a style is missing.
|
||
</paragraph>
|
||
<paragraph id="426EF8409775FF92FF040498FE13FE7E" blockId="11.[151,1436,1751,2029]" lastBlockId="12.[151,1437,151,2014]" lastPageId="12" lastPageNumber="327" pageId="11" pageNumber="326">
|
||
Introvert with six transverse rings of scalids and 10 longitudinal sectors defined by the position of the primary spinoscalids. Ring 01 with 10 primary spinoscalids, larger than the remaining ones, each one composed of a basal, rectangular, wide sheath and a distal, elongated, flexible, laterally compressed, distally rounded end-piece (
|
||
<figureCitation id="DAEAE4C59772FF92FAAC03A0FAB7FF7A" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="12" pageNumber="327">Fig. 6D, F</figureCitation>
|
||
). Basal sheath of primary spinoscalids superficially fringed, bearing a rhomboid cuticular piece superficially fringed near the articulation point with the end-piece (
|
||
<figureCitation id="DAEAE4C59772FF92FCC903E8F90EFF56" box="[778,867,223,249]" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="12" pageNumber="327">Fig. 6D</figureCitation>
|
||
inset, F). Rings 02–06 with several regular-sized scalids, conspicuously smaller than the primary spinoscalids, also composed of a basal, rectangular, superficially fringed, wide sheath and a distal, elongated, flexible, distally rounded end–piece (
|
||
<figureCitation id="DAEAE4C59772FF92FBFF0210FEFEFEED" box="[1084,1171,295,322]" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="12" pageNumber="327">Fig. 6F</figureCitation>
|
||
). Exact arrangement of regular-sized scalids cannot be provided due to the collapsed condition of the only available introverts. Despite this, the odd-numbered sectors seem to possess seven regular-sized scalids arranged as a double diamond (
|
||
<figureCitation id="DAEAE4C59772FF92FADC0258FF1BFE25" box="[1311,1398,367,394]" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="12" pageNumber="327">Fig. 6F</figureCitation>
|
||
). A ring of 14 trichoscalids posterior to the scalid rings (
|
||
<figureCitation id="DAEAE4C59772FF92FD2102A4F955FE01" box="[738,824,403,430]" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="12" pageNumber="327">Fig. 6E</figureCitation>
|
||
), arranged as two in the odd-numbered sectors (except sector 1 with a single trichoscalid) and one in the even-numbered sectors of the introvert.
|
||
</paragraph>
|
||
<paragraph id="426EF8409772FF92FF0402ECF9CAFD2A" blockId="12.[151,1437,151,2014]" pageId="12" pageNumber="327">
|
||
<emphasis id="70A524529772FF92FF0402ECFB6AFE5A" box="[199,263,475,501]" italics="true" pageId="12" pageNumber="327">Neck.</emphasis>
|
||
Neck with four dorsal and two ventral sclerotized placids (
|
||
<figureCitation id="DAEAE4C59772FF92FC7502ECFE50FE5A" box="[950,1085,474,501]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FB8802ECFEF9FE59" box="[1099,1172,475,502]" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="12" pageNumber="327">6G–H</figureCitation>
|
||
). Mesial dorsal placids sub-quadrangular, conspicuously higher than lateral ones (
|
||
<figureCitation id="DAEAE4C59772FF92FCF702C8F9F5FDB6" box="[820,920,510,537]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FC6402C8F9A1FDB5" box="[935,972,511,538]" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="12" pageNumber="327">6G</figureCitation>
|
||
), ca. 33–41 μm wide at the base; lateral dorsal placids rectangular (
|
||
<figureCitation id="DAEAE4C59772FF92FE0F0114F842FD92" box="[460,559,546,573]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FDFE0114F80EFD91" box="[573,611,547,574]" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="12" pageNumber="327">6G</figureCitation>
|
||
), ca. 23–26 μm wide at the base. Ventral placids even more rectangular, longitudinally compressed, with the posterolateral margins straight to slightly curved towards the sternal plates of the first trunk segment (
|
||
<figureCitation id="DAEAE4C59772FF92FE60015CF86AFD2A" box="[419,519,618,645]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5A</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FDD0015CF855FD29" box="[531,568,619,646]" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="12" pageNumber="327">6H</figureCitation>
|
||
), ca. 21–30 μm wide at the base.
|
||
</paragraph>
|
||
<paragraph id="426EF8409772FF92FF0401B8FB68FA32" blockId="12.[151,1437,151,2014]" pageId="12" pageNumber="327">
|
||
<emphasis id="70A524529772FF92FF0401B8FB7DFD06" box="[199,272,655,681]" italics="true" pageId="12" pageNumber="327">Trunk.</emphasis>
|
||
Trunk rectangular, stout, triangular in cross–section, composed of 11 segments (
|
||
<figureCitation id="DAEAE4C59772FF92FB4801B8FF62FD06" box="[1163,1295,654,681]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FAD901B8FF32FD06" box="[1306,1375,655,681]" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="12" pageNumber="327">6A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FAA901B8FFE2FD06" box="[1386,1423,655,681]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="12" pageNumber="327">8A</figureCitation>
|
||
). Segment 1 with one tergal, two episternal, and one midsternal plates; remaining segments with one tergal and two sternal plates (
|
||
<figureCitation id="DAEAE4C59772FF92FEF801E0FBAFFD5E" box="[315,450,726,753]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5A–D</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FE0D01E0F87EFD5E" box="[462,531,727,753]" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="12" pageNumber="327">6A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FDE301E0F80AFD5E" box="[544,615,727,753]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="12" pageNumber="327">7A–K</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FDB001E0F8DAFD5E" box="[627,695,727,753]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="12" pageNumber="327">8A–F</figureCitation>
|
||
). Maximum sternal width at segment 3 (
|
||
<tableCitation id="0F53CDFB9772FF92FBBC01E0FEBBFD5E" box="[1151,1238,727,753]" captionStart="TABLE 4" captionStartId="11.[151,239,151,175]" captionTargetPageId="11" captionText="TABLE 4. Measurements (in μm) of nine adult specimens of Leiocanthus satanicus sp. nov. (six females and three males) from the Gulf of Mexico. Abbreviations: MSW-3, maximum sternal width (measured at segment 3); n, number of measured specimens; S, segment length (followed by number of corresponding segment); SD, standard deviation; SSW, standard sternal width (measured at segment 10); TL, trunk length." httpUri="http://table.plazi.org/id/16AEA8C89775FF95FF5403A0F90CFEB4" pageId="12" pageNumber="327" tableUuid="16AEA8C89775FF95FF5403A0F90CFEB4">Table 4</tableCitation>
|
||
), almost constant in width throughout the trunk until segment 3, and progressively tapering along the last trunk segments (
|
||
<figureCitation id="DAEAE4C59772FF92FAA701CCFAB1FC96" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FF2A0028FB43FC96" box="[233,302,799,825]" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="12" pageNumber="327">6A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FEF80028FB0DFC96" box="[315,352,799,825]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="12" pageNumber="327">8A</figureCitation>
|
||
). Sternal cuticular plates relatively narrow in relation to trunk length (MSW–3:TL interval ratio = 21.8–26.8) (
|
||
<tableCitation id="0F53CDFB9772FF92FEF50074FBE0FCF2" box="[310,397,835,861]" captionStart="TABLE 4" captionStartId="11.[151,239,151,175]" captionTargetPageId="11" captionText="TABLE 4. Measurements (in μm) of nine adult specimens of Leiocanthus satanicus sp. nov. (six females and three males) from the Gulf of Mexico. Abbreviations: MSW-3, maximum sternal width (measured at segment 3); n, number of measured specimens; S, segment length (followed by number of corresponding segment); SD, standard deviation; SSW, standard sternal width (measured at segment 10); TL, trunk length." httpUri="http://table.plazi.org/id/16AEA8C89775FF95FF5403A0F90CFEB4" pageId="12" pageNumber="327" tableUuid="16AEA8C89775FF95FF5403A0F90CFEB4">Table 4</tableCitation>
|
||
), giving the animal a relatively slender appearance (
|
||
<figureCitation id="DAEAE4C59772FF92FC140074FE36FCF2" box="[983,1115,834,861]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FBA40074FEC1FCF2" box="[1127,1196,835,861]" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="12" pageNumber="327">6A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FB7B0074FEB0FCF2" box="[1208,1245,835,861]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="12" pageNumber="327">8A</figureCitation>
|
||
). Segments 1–11 with glandular cell outlets with a single round to oval opening: segments 1 and 9–10 with these glandular cell outlets in subdorsal and ventromedial positions, segments
|
||
<quantity id="852955A59772FF92FD1F00BCF949FC0A" box="[732,804,907,933]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" metricValueMax="2.032" metricValueMin="0.508" pageId="12" pageNumber="327" unit="in" value="5.0" valueMax="8.0" valueMin="2.0">2–8 in</quantity>
|
||
subdorsal and paraventral positions, and segment
|
||
<quantity id="852955A59772FF92FAA000BCFFF1FC0A" box="[1379,1436,907,933]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="12" pageNumber="327" unit="in" value="11.0">11 in</quantity>
|
||
subdorsal position (
|
||
<figureCitation id="DAEAE4C59772FF92FEB00098FB94FC66" box="[371,505,942,969]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5A–D</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FDC70098F829FC66" box="[516,580,943,969]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="12" pageNumber="327">7A–J</figureCitation>
|
||
); intraspecific variation of this character was found in a couple of specimens that showed up to three round openings per glandular outlet (
|
||
<figureCitation id="DAEAE4C59772FF92FC8600E4F9CDFC42" box="[837,928,979,1005]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="12" pageNumber="327">Fig. 7D</figureCitation>
|
||
); in addition, in some specimens the bilateral pattern was lost as the glandular cell outlet was missing in one side of some tergal plates. Segments 2–10 with paired, small, not always conspicuous cuticular ridges in ventrolateral position, with adjacent, minute glandular cell outlets (
|
||
<figureCitation id="DAEAE4C59772FF92FF630708FB73FBF6" box="[160,286,1086,1113]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Fig. 5A–D</figureCitation>
|
||
). Minute, acicular cuticular hairs, widely covering the cuticular surface of segments 1–10 (except the most anterior halves of the episternal plates of segment 1 where hairs are absent), denser at the tergosternal junctions (
|
||
<figureCitation id="DAEAE4C59772FF92FF6307B0FB7BFB0E" box="[160,278,1159,1185]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="12" pageNumber="327">Fig. 8B–F</figureCitation>
|
||
). Males with sexually dimorphic longitudinal bands of cuticle densely covered by smaller acicular hairs in ventromedial position of segment 2, under the tubes (
|
||
<figureCitation id="DAEAE4C59772FF92FCC7079CF931FB6A" box="[772,860,1195,1221]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="12" pageNumber="327">Fig. 8C</figureCitation>
|
||
). Very conspicuous, rounded to oval muscular scars without hair-covering in laterodorsal and ventromedial positions (
|
||
<figureCitation id="DAEAE4C59772FF92FC5307F8FE77FB46" box="[912,1050,1230,1257]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5A–D</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FBEA07F8FE04FB46" box="[1065,1129,1231,1257]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="12" pageNumber="327">7A–J</figureCitation>
|
||
). Pachycycli and ball-andsocket joints well-developed and thick on segments 2–10 (
|
||
<figureCitation id="DAEAE4C59772FF92FCE007C4F9C4FAA2" box="[803,937,1266,1293]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5A–D</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FC7707C4F993FAA2" box="[948,1022,1267,1293]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="12" pageNumber="327">7A–H</figureCitation>
|
||
). Apodemes absent. Posterior margin of segments straight, with primary pectinate fringes showing weakly serrated free flaps (
|
||
<figureCitation id="DAEAE4C59772FF92FBA80620FE9DFA9E" box="[1131,1264,1302,1329]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5A–D</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FB380620FF2FFA9E" box="[1275,1346,1303,1329]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="12" pageNumber="327">7A–H</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FA8F0620FFFDFA9E" box="[1356,1424,1303,1329]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="12" pageNumber="327">8A–F</figureCitation>
|
||
). Secondary pectinate fringes as three (dorsal) to two (ventral) transverse rows, finely serrated, becoming wavier at the sternal plates, extending posteriorly in triangular extensions near the location of the muscular scars (
|
||
<figureCitation id="DAEAE4C59772FF92FAD00668FFF5FAD6" box="[1299,1432,1374,1401]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5A–D</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FF5406B4FA95FA32" box="[151,248,1411,1437]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="12" pageNumber="327">8C, E, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="426EF8409772FF92FF040690FBFBF8E2" blockId="12.[151,1437,151,2014]" pageId="12" pageNumber="327">
|
||
Segment 1 with middorsal elevation not projecting beyond the posterior margin of segment, with conspicuous paradorsal, butterfly-like atria of sensory spots (
|
||
<figureCitation id="DAEAE4C59772FF92FD0306FCF949FA4A" box="[704,804,1482,1509]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FCF106FCF935FA4A" box="[818,856,1483,1509]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="12" pageNumber="327">7A</figureCitation>
|
||
). Anterolateral margins of the tergal plate forming conspicuous horn-like, medially recurved, distally pointed projections (
|
||
<figureCitation id="DAEAE4C59772FF92FC1506D8FE32F9A6" box="[982,1119,1518,1545]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FBAC06D8FEA2F9A6" box="[1135,1231,1519,1545]" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="12" pageNumber="327">6A–B, I</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FB2306D8FF4AF9A6" box="[1248,1319,1519,1545]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="12" pageNumber="327">8A–B</figureCitation>
|
||
). Anterior margin of tergal plate strongly denticulated, posteriorly followed by a crenulated area (
|
||
<figureCitation id="DAEAE4C59772FF92FB4E0524FE9FF982" box="[1165,1266,1554,1581]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FAC10524FF45F981" box="[1282,1320,1555,1582]" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="12" pageNumber="327">6K</figureCitation>
|
||
). Anterior margins of sternal plates with several, small, cuticular depressions and some semi-circular ridges (
|
||
<figureCitation id="DAEAE4C59772FF92FB340500FF31F9FE" box="[1271,1372,1590,1617]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5A</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FAA90500FFE2F9FD" box="[1386,1423,1591,1618]" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="12" pageNumber="327">6H</figureCitation>
|
||
). Characteristic cuticular wrinkles at the central region of the tergal plate in subdorsal towards laterodorsal position (
|
||
<figureCitation id="DAEAE4C59772FF92FF5C0548FB6FF936" box="[159,258,1662,1689]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FED30548FB5BF935" box="[272,310,1663,1690]" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="12" pageNumber="327">6K</figureCitation>
|
||
). Midsternal plate almost rectangular, not extended at its base, with a small lateral indentation near its anterior margin, and a straight posterior margin (
|
||
<figureCitation id="DAEAE4C59772FF92FD1E0594F92CF912" box="[733,833,1698,1725]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5A</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FC8E0594F91DF912" box="[845,880,1699,1725]" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="12" pageNumber="327">6B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FCBF0594F9CFF912" box="[892,930,1699,1725]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="12" pageNumber="327">8A</figureCitation>
|
||
). Paired setae in paralateral and ventrolateral positions (
|
||
<figureCitation id="DAEAE4C59772FF92FEC905F0FBE0F94E" box="[266,397,1734,1761]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FE5405F0FBD0F94E" box="[407,445,1735,1761]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="12" pageNumber="327">7A</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FE0505F0FB87F94E" box="[454,490,1735,1761]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="12" pageNumber="327">8B</figureCitation>
|
||
). Sensory spots in paradorsal (one pair), subdorsal (two pairs, longitudinally aligned), laterodorsal (one pair), paralateral (one pair), ventrolateral (one pair) and ventromedial (one pair) positions (
|
||
<figureCitation id="DAEAE4C59772FF92FAA705DCFAB1F886" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FF2A0438FB63F886" box="[233,270,1807,1833]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="12" pageNumber="327">7A</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FED90438FB50F886" box="[282,317,1807,1833]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="12" pageNumber="327">8B</figureCitation>
|
||
). Sensory spots on this and remaining segments oval, with up to three pores surrounded by several rings of micropapillae.
|
||
</paragraph>
|
||
<paragraph id="426EF8409772FF93FF040460FBCCFF7A" blockId="12.[151,1437,151,2014]" lastBlockId="13.[151,1437,150,1941]" lastPageId="13" lastPageNumber="328" pageId="12" pageNumber="327">
|
||
Segment 2 with middorsal elevation as on the preceding segment (
|
||
<figureCitation id="DAEAE4C59772FF92FC0E0460FE5FF8DE" box="[973,1074,1878,1905]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59772FF92FB820460FE0AF8DE" box="[1089,1127,1879,1905]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="12" pageNumber="327">7D</figureCitation>
|
||
). Paired setae in subdorsal and lateroventral positions; subdorsal setae more mesial than subdorsal sensory spots; ventromedial setae only in females (sexually dimorphic) and located more lateral than the ventromedial sensory spots (
|
||
<figureCitation id="DAEAE4C59772FF92FB7404A8FF68F816" box="[1207,1285,1950,1977]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="12" pageNumber="327">Figs. 5</figureCitation>
|
||
A-C, 7B, D). Sensory spots in paradorsal (one pair), subdorsal (two pairs), laterodorsal (one pair) and ventromedial (one pair) positions (
|
||
<figureCitation id="DAEAE4C59773FF93FECC03A0FBF5FF1E" box="[271,408,150,177]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A–C</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FE6403A0FB9CFF1E" box="[423,497,151,177]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7B, D</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FDC303A0F849FF1E" box="[512,548,151,177]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="13" pageNumber="328">8C</figureCitation>
|
||
). Males with conspicuously large, sexually dimorphic tubes in ventromedial position (
|
||
<figureCitation id="DAEAE4C59773FF93FEC2038CFB08FF7A" box="[257,357,186,213]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FEB3038CFBF9FF7A" box="[368,404,187,213]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="13" pageNumber="328">8C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="426EF8409773FF93FF0403E8FEE6FE26" blockId="13.[151,1437,150,1941]" pageId="13" pageNumber="328">
|
||
Segment 3 with middorsal elevation as on the preceding segments (
|
||
<figureCitation id="DAEAE4C59773FF93FC0103E8FE49FF56" box="[962,1060,222,249]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FBF203E8FE3AFF56" box="[1073,1111,223,249]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7D</figureCitation>
|
||
). Paired setae in laterodorsal and paraventral positions (
|
||
<figureCitation id="DAEAE4C59773FF93FE060234F826FEB2" box="[453,587,258,285]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FD9A0234F8F3FEB2" box="[601,670,259,285]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7C–D</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FD680234F8A2FEB2" box="[683,719,259,285]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="13" pageNumber="328">8C</figureCitation>
|
||
). Sensory spots in paradorsal (one pair), subdorsal (two pairs), laterodorsal (one pair) and ventromedial (one pair) positions (
|
||
<figureCitation id="DAEAE4C59773FF93FC8B0210F9A1FEEE" box="[840,972,294,321]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FC1B0210FE70FEEE" box="[984,1053,295,321]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7C–D</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FBEA0210FE20FEEE" box="[1065,1101,295,321]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="13" pageNumber="328">8C</figureCitation>
|
||
). Intraspecific variation in the arrangement of the laterodorsal setae has been observed in
|
||
<specimenCount id="54D733C99773FF93FC81027CF980FECA" box="[834,1005,331,357]" pageId="13" pageNumber="328" type="generic">two specimens</specimenCount>
|
||
, where these structures were laterally displaced, still in laterodorsal position but more lateral than the laterodorsal sensory spots.
|
||
</paragraph>
|
||
<paragraph id="426EF8409773FF93FF0402A4F854FD92" blockId="13.[151,1437,150,1941]" pageId="13" pageNumber="328">
|
||
Segment 4 with middorsal elevation as on the preceding segments (
|
||
<figureCitation id="DAEAE4C59773FF93FC2502A4FE2EFE02" box="[998,1091,402,429]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Fig. 5B</figureCitation>
|
||
). Unpaired seta in paradorsal position, and paired setae in subdorsal, lateroventral and ventromedial positions; subdorsal setae flanked by the two pairs of subdorsal sensory spots, ventromedial setae more mesial than those of female segment 2 (
|
||
<figureCitation id="DAEAE4C59773FF93FB2702ECFF05FE5A" box="[1252,1384,474,501]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FAB602ECFFFAFE5A" box="[1397,1431,475,501]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7C</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FF5402C8FAD0FDB6" box="[151,189,511,537]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="13" pageNumber="328">8D</figureCitation>
|
||
). Sensory spots in paradorsal (one pair), subdorsal (two pairs), laterodorsal (one pair) and ventromedial (one pair) positions (
|
||
<figureCitation id="DAEAE4C59773FF93FE840114FBA1FD92" box="[327,460,546,573]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FE1B0114FB97FD92" box="[472,506,547,573]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7C</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FDC50114F841FD92" box="[518,556,547,573]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="13" pageNumber="328">8D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="426EF8409773FF93FF040170FAB5FC96" blockId="13.[151,1437,150,1941]" pageId="13" pageNumber="328">
|
||
Segment 5 with middorsal elevation as on the preceding segments (
|
||
<figureCitation id="DAEAE4C59773FF93FC030170FE4FFDCE" box="[960,1058,582,609]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FBEC0170FE3DFDCE" box="[1071,1104,583,609]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7F</figureCitation>
|
||
). Paired setae in laterodorsal, ventrolateral and ventromedial positions; ventromedial setae more lateral than those of segment 4 (not aligned longitudinally) (
|
||
<figureCitation id="DAEAE4C59773FF93FE8D01B8FBBFFD06" box="[334,466,654,681]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FE1C01B8F871FD06" box="[479,540,655,681]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7E–F</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FDEB01B8F823FD06" box="[552,590,655,681]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="13" pageNumber="328">8D</figureCitation>
|
||
). Sensory spots in paradorsal (one pair), subdorsal (two pairs), laterodorsal (one pair) and ventromedial (one pair) positions (
|
||
<figureCitation id="DAEAE4C59773FF93FD7B0184F951FD62" box="[696,828,690,717]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FC840184F9E9FD62" box="[839,900,691,717]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7E–F</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FC4D0184F9D9FD62" box="[910,948,691,717]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="13" pageNumber="328">8D</figureCitation>
|
||
); the latter pair aligned with that of segment 3. Intraspecific variation in the arrangement of the laterodorsal setae has been observed in a single specimen, where these structures were laterally displaced, still in laterodorsal position but more lateral than the laterodorsal sensory spots.
|
||
</paragraph>
|
||
<paragraph id="426EF8409773FF93FF040074FB0DFB9A" blockId="13.[151,1437,150,1941]" pageId="13" pageNumber="328">
|
||
Segment 6 with middorsal elevation as on the preceding segments (
|
||
<figureCitation id="DAEAE4C59773FF93FC7C0074FE4CFCF2" box="[959,1057,834,861]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FBED0074FE3EFCF2" box="[1070,1107,835,861]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7H</figureCitation>
|
||
). Unpaired seta in paradorsal position, and paired setae in subdorsal, lateroventral and ventromedial positions (the former flanked by the subdorsal sensory spots, the latter longitudinally aligned with those of segment 4) (
|
||
<figureCitation id="DAEAE4C59773FF93FC2A00BCFE1DFC0A" box="[1001,1136,906,933]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FBBC00BCFEC8FC0A" box="[1151,1189,907,933]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7H</figureCitation>
|
||
); deviations from the bilateral symmetry in the position of the ventromedial setae has been observed in a single specimen that had one seta mesially displaced, though still in ventromedial position. Sensory spots in paradorsal (one pair), subdorsal (two pairs), laterodorsal (one pair) and ventromedial (one pair) positions (
|
||
<figureCitation id="DAEAE4C59773FF93FC6100C0FE45FBBE" box="[930,1064,1014,1041]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FBF600C0FE37FBBE" box="[1077,1114,1015,1041]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7H</figureCitation>
|
||
); the latter pair aligned with that of segment 4.
|
||
</paragraph>
|
||
<paragraph id="426EF8409773FF93FF040708FF5CFAA2" blockId="13.[151,1437,150,1941]" pageId="13" pageNumber="328">
|
||
Segment 7 without middorsal process or elevation. Paired setae in laterodorsal and ventromedial positions, the laterodorsal ones more lateral than the laterodorsal sensory spots, and the ventromedial ones aligned with those of segment 5 and female segment 2 (
|
||
<figureCitation id="DAEAE4C59773FF93FDDA07B0F8F3FB0E" box="[537,670,1158,1185]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FD6807B0F8BDFB0E" box="[683,720,1159,1185]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7H</figureCitation>
|
||
). Sensory spots in subdorsal (two pairs), laterodorsal (one pair) and ventromedial (one pair) positions (
|
||
<figureCitation id="DAEAE4C59773FF93FD89079CF8A3FB6A" box="[586,718,1194,1221]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FD1A079CF892FB6A" box="[729,767,1195,1221]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7H</figureCitation>
|
||
); the latter pair aligned with that of segment 3. Intraspecific variation in the arrangement of the laterodorsal setae has been observed in a single specimen, where these structures were mesially displaced, still in laterodorsal position but more mesial than the laterodorsal sensory spots.
|
||
</paragraph>
|
||
<paragraph id="426EF8409773FF93FF040620FBD4FA4A" blockId="13.[151,1437,150,1941]" pageId="13" pageNumber="328">
|
||
Segment 8 without middorsal process or elevation. Paired setae in subdorsal, lateroventral and paraventral positions; the former flanked by the subdorsal sensory spots (
|
||
<figureCitation id="DAEAE4C59773FF93FC95060CF9B1FAFA" box="[854,988,1338,1365]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FC29060CFE63FAFA" box="[1002,1038,1339,1365]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7G</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FBDF060CFE52FAFA" box="[1052,1087,1339,1365]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="13" pageNumber="328">8E</figureCitation>
|
||
). Deviations from the bilateral pattern in the arrangement of the paraventral setae has been observed in a single specimen, which had the left sternal plate with the seta in ventromedial instead of paraventral position. Sensory spots in subdorsal (three pairs), laterodorsal (one pair) and ventromedial (one pair) positions (
|
||
<figureCitation id="DAEAE4C59773FF93FC880690F9A2FA6E" box="[843,975,1446,1473]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FC180690FE6DFA6E" box="[987,1024,1447,1473]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7G</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FBCF0690FE42FA6E" box="[1036,1071,1447,1473]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="13" pageNumber="328">8E</figureCitation>
|
||
); the latter pair aligned with that of the preceding segment.
|
||
</paragraph>
|
||
<paragraph id="426EF8409773FF93FF0406D8FE73F9DA" blockId="13.[151,1437,150,1941]" pageId="13" pageNumber="328">
|
||
Segment 9 without middorsal process or elevation. Paired setae in laterodorsal and ventromedial position; the latter longitudinally aligned with those of segments 5 and 7 (
|
||
<figureCitation id="DAEAE4C59773FF93FC980524F98FF982" box="[859,994,1554,1581]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FC330524FE4EF982" box="[1008,1059,1555,1581]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7I–J</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FBF20524FE39F982" box="[1073,1108,1555,1581]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="13" pageNumber="328">8E</figureCitation>
|
||
). Sensory spots in subdorsal (three pairs), laterodorsal (one pair), ventrolateral (one pair) and ventromedial positions (
|
||
<figureCitation id="DAEAE4C59773FF93FBBF0500FF6DF9FE" box="[1148,1280,1590,1617]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A–B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FACF0500FF52F9FE" box="[1292,1343,1591,1617]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7I–J</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FA890500FFFDF9FE" box="[1354,1424,1591,1617]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="13" pageNumber="328">8E, H</figureCitation>
|
||
). Nephridia externally opening as short cuticular tubes with fringed tips (
|
||
<figureCitation id="DAEAE4C59773FF93FC74056CFE7FF9DA" box="[951,1042,1626,1653]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Fig. 5A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="426EF8409773FF93FF040548FB55F8AA" blockId="13.[151,1437,150,1941]" pageId="13" pageNumber="328">
|
||
Segment 10 without middorsal process or elevation. Tergal plates with elongated, dagger-like projections at midlateral posterior margins (
|
||
<figureCitation id="DAEAE4C59773FF93FE2A0594F8F9F912" box="[489,660,1698,1725]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A–B, D</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FD620594F96CF912" box="[673,769,1699,1725]" captionStart="FIGURE 6" captionStartId="15.[151,250,1727,1751]" captionTargetBox="[218,1369,181,1703]" captionTargetId="figure-18@15.[218,1369,181,1703]" captionTargetPageId="15" captionText="FIGURE 6. Light micrographs showing trunk overviews and details in the head, neck and main cuticular ornamentations of female holotype NHMD-915202 (A–B, I–J) and male paratype NHMD-915207 (C–H, K) of Leiocanthus satanicus sp. nov. A, dorsal overview; B, ventral overview; C, outer oral styles of mouth cone (arrows indicate the lateral incisions); D, detail of a spinoscalid (arrow indicates the fringes of the basal sheath), inset shows the rhomboid cuticular piece; E, trichoscalids; F, regular-sized scalids of sector 03 (black head arrows indicate the arrangement of the scalids); G, dorsal placids; H, ventral placids (arrows indicate the posterolateral margins); I, horn-shaped anterolateral projections of segment 1 tergal plate; J, daggerlike extensions of segment 10 tergal plate; K, anterior margin of segment 1 tergal plate and conspicuous cuticular wrinkles (indicated by arrow and circle). Scales: A–B, 100 μm; C–K, 20 μm; inset of D, 10 μm. Abbreviations: bs, basal sheath; ep, endpiece; s03, introvert sector 3." figureDoi="http://doi.org/10.5281/zenodo.5909867" httpUri="https://zenodo.org/record/5909867/files/figure.png" pageId="13" pageNumber="328">6A–B, J</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FCCD0594F939F912" box="[782,852,1699,1725]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="13" pageNumber="328">8A, F</figureCitation>
|
||
). Two pairs of setae in lateroventral position (
|
||
<figureCitation id="DAEAE4C59773FF93FAA70594FA92F94E" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A–B, D</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FEC805F0FB41F94E" box="[267,300,1735,1761]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="13" pageNumber="328">8F</figureCitation>
|
||
). One pair of sensory spots in subdorsal, laterodorsal, ventrolateral and ventromedial positions (
|
||
<figureCitation id="DAEAE4C59773FF93FAA705F0FA92F8AA" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A–B, D</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FEC905DCFB46F8AA" box="[266,299,1771,1797]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="13" pageNumber="328">8F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="426EF8409773FF93FF040438FBCCF83A" blockId="13.[151,1437,150,1941]" pageId="13" pageNumber="328">
|
||
Segment 11 with one pair of
|
||
<typeStatus id="9D6A46E29773FF93FDC90438F857F886" box="[522,570,1807,1833]" pageId="13" pageNumber="328">type</typeStatus>
|
||
3 sensory spots in subdorsal position (
|
||
<figureCitation id="DAEAE4C59773FF93FC2F0438FE20F886" box="[1004,1101,1806,1833]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5B</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FB9A0438FE12F886" box="[1113,1151,1807,1833]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7K</figureCitation>
|
||
). Males with two pairs of sexually dimorphic penile spines and genital pores (
|
||
<figureCitation id="DAEAE4C59773FF93FD0E0404F943F8E2" box="[717,814,1842,1869]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FCFB0404F935F8E2" box="[824,856,1843,1869]" captionStart="FIGURE 8" captionStartId="17.[151,250,1505,1529]" captionTargetBox="[151,1436,181,1481]" captionTargetId="figure-18@17.[151,1436,181,1481]" captionTargetPageId="17" captionText="FIGURE 8. Scanning electron micrographs showing general overview and details of the cuticular morphology of a male (A–F) and a female (G–J) non-type specimens of Leiocanthus satanicus sp. nov. A, ventral overview; B, ventral view of the left episternal plate of segment 1; C, ventral view of the left sternal plates of segments 2–3; D, laterodorsal to ventromedial view on right half of cuticular plates of segments 4–5; E, ventral view of the left sternal plates of segments 8–9; F, ventral view of the right sternal plate of segment 10. Scales: A, 100 μm; B–J, 10 μm. Abbreviations: cr, cuticular ridge; gco, glandular cell outlet; lvse, lateroventral seta; ps, penile spine; pvse, paraventral seta; vlse, ventrolateral seta; vmse, ventromedial seta; vmt, ventromedial tube; sensory spots are marked as dashed circles; ♂ indicates the presence of male sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909871" httpUri="https://zenodo.org/record/5909871/files/figure.png" pageId="13" pageNumber="328">8F</figureCitation>
|
||
). Lateral terminal spines absent; minute, rectangular, distally rounded, bulbous protuberances emerge between tergal and sternal plates, in males superficially covered by hairs (
|
||
<figureCitation id="DAEAE4C59773FF93FF1E044CFB0EF83A" box="[221,355,1914,1941]" captionStart="FIGURE 5" captionStartId="14.[151,250,1696,1720]" captionTargetBox="[222,1364,191,1659]" captionTargetId="figure-31@14.[216,1370,181,1672]" captionTargetPageId="14" captionText="FIGURE 5. Line art illustrations of adult Leiocanthus satanicus sp. nov. A, male in ventral view; B, male in dorsal view; C, female segment 2 in ventral view; D, female segments 10–11 in ventral view. Scale: 100 μm. Abbreviations: bla, butterflylike atrium; bp, bulbous protuberance; bsj, ball-and-socket joint; cr, cuticular ridge; dle, dagger-like extension; dpl, dorsal placid; gco, glandular cell outlet; ldse, laterodorsal seta; ldss, laterodorsal sensory spot; lvse, lateroventral seta; mde, middorsal elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; pdss, paradorsal sensory spot; plse, paralateral seta; plss, paralateral sensory spot; ppf, primary pectinate fringe; ps, penile spine; pvse, paraventral seta; S, segment (followed by number of corresponding segment); sdgco, subdorsal glandular cell outlet; sdse, subdorsal seta; sdss, subdorsal sensory spot; ss3, subdorsal type 3 sensory spot; spf, secondary pectinate fringes; vlse, ventrolateral seta; vlss, ventrolateral sensory spot; vmgco, ventromedial glandular cell outlet; vmse, ventromedial seta; vmss, ventromedial sensory spot; vmt, ventromedial tube; vpl, ventral placid." figureDoi="http://doi.org/10.5281/zenodo.5909865" httpUri="https://zenodo.org/record/5909865/files/figure.png" pageId="13" pageNumber="328">Figs. 5A, D</figureCitation>
|
||
,
|
||
<figureCitation id="DAEAE4C59773FF93FEAC044CFBF9F83A" box="[367,404,1915,1941]" captionStart="FIGURE 7" captionStartId="16.[151,250,1656,1680]" captionTargetBox="[470,1121,187,1626]" captionTargetId="figure-31@16.[463,1125,181,1632]" captionTargetPageId="16" captionText="FIGURE 7. Light micrographs showing trunk cuticular details of female holotype NHMD-915202 (B, C, E, G, I, K) and male paratype NHMD-915207 (A, D, F, H, J) of Leiocanthus satanicus sp. nov. A, middorsal to paralateral view on left half of tergal plate of segment 1; B, left sternal plate of segment 2; C, left sternal plates of segments 3–4; D, middorsal to laterodorsal view on left half of tergal plates of segments 2–3; E, left sternal plates of segments 5–6; F, middorsal to laterodorsal view on left half of tergal plate of segment 5; G, left sternal plate of segment 8; H, middorsal to laterodorsal view on left half of tergal plates of segments 6–7; I, left sternal plate of segment 9; J, middorsal to laterodorsal view on left half of tergal plate of segment 9; K, dorsal overview of segment 11 (inset shows the bulbous protuberance of a male). Scales: A–K, 20 μm; inset of K, 5 μm. Abbreviations: bla, butterfly–like intracuticular atria; bp, bulbous protuberance; bsj, ball-and-socket joint; ldse, laterodorsal seta; mde, middorsal cuticular elevation; ms, muscular scar; pc, pachycyclus; pdse, paradorsal seta; plse, paralateral seta; pvse, paraventral seta; sdse, subdorsal seta; ss3, type 3 sensory spot; vlse, ventrolateral seta; vmse, ventromedial seta; glandular cell outlets as closed circles, and sensory spots as dashed circles; ♂/♀ indicate the presence of sexually dimorphic structures." figureDoi="http://doi.org/10.5281/zenodo.5909869" httpUri="https://zenodo.org/record/5909869/files/figure.png" pageId="13" pageNumber="328">7K</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |